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<document id="EEEA7B453CF25ABAA500829380FE6524" ID-CLB-Dataset="3953" ID-DOI="10.5281/zenodo.1196003" ID-GBIF-Dataset="673e368f-06c4-465a-b40d-68dbedd81138" ID-ISSN="0035-418" ID-Zenodo-Dep="1196003" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634783046189" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N." docDate="2018" docId="BC2887A2FFAD99205EE8F1364DBBFC38" docLanguage="en" docName="1_Anisyutkin_1.pdf" docOrigin="Revue suisse de Zoologie 125 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Morphnodes goliath" docType="treatment" docVersion="6" lastPageNumber="10" masterDocId="4011FFDAFFA599295E10F6034F12FF98" masterDocTitle="Little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Muséum d'histoire naturelle de Genève. Part 3" masterLastPageNumber="16" masterPageNumber="1" pageNumber="9" updateTime="1699460453851" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FC99BE40A52CED090E2CEBEAE1518D61">Little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Muséum d'histoire naturelle de Genève. Part 3</mods:title>
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<mods:namePart id="F7BEE3B137B1D3AE7A2BD2657281ACC3">Anisyutkin, Leonid N.</mods:namePart>
<mods:affiliation id="7D02E2547F39CC6CF35A5346E5EAF096">Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia. E-mail: leonid. dictyoptera @ gmail. com, Leonid. Anisyutkin @ zin. ru</mods:affiliation>
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<treatment id="BC2887A2FFAD99205EE8F1364DBBFC38" ID-DOI="http://doi.org/10.5281/zenodo.5589902" ID-GBIF-Taxon="189149527" ID-Zenodo-Dep="5589902" LSID="urn:lsid:plazi:treatment:BC2887A2FFAD99205EE8F1364DBBFC38" httpUri="http://treatment.plazi.org/id/BC2887A2FFAD99205EE8F1364DBBFC38" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="7C9B653FFFAD99215EE8F1364D81F8D5" box="[248,659,1845,1869]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="343E36B4FFAD99215EE8F1364D81F8D5" blockId="8.[248,659,1845,1901]" box="[248,659,1845,1869]" pageId="8" pageNumber="9">
<heading id="6F7681D8FFAD99215EE8F1364D81F8D5" bold="true" box="[248,659,1845,1869]" centered="true" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="F3814D37FFAD99215EE8F1364D81F8D5" authority="(Shelford, 1906)" baseAuthorityName="Shelford" baseAuthorityYear="1906" box="[248,659,1845,1869]" class="Insecta" family="Blaberidae" genus="Morphnodes" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="goliath">
<emphasis id="06F5EAA6FFAD99215EE8F1364D81F8D5" bold="true" box="[248,659,1845,1869]" pageId="8" pageNumber="9">
<emphasis id="06F5EAA6FFAD99215EE8F1364EC5F8D5" bold="true" box="[248,471,1845,1869]" italics="true" pageId="8" pageNumber="9">Morphnodes goliath</emphasis>
(
<bibRefCitation id="50104B45FFAD99215FF4F1354D9EF8D5" author="Shelford R." box="[484,652,1846,1869]" pageId="8" pageNumber="9" pagination="265 - 275" refId="ref10757" refString="Shelford R. 1906. Studies of the Blattidae. III. Some new Blattidae from Sarawak, Borneo, in the Hope Department, Oxford University Museum. Transactions of the Entomological Society, London 1: 265 - 275." type="journal article" year="1906">Shelford, 1906</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7C9B653FFFAD99215F64F1554D0AF8F5" box="[372,536,1878,1901]" pageId="8" pageNumber="9" type="description">
<paragraph id="343E36B4FFAD99215F64F1554D0AF8F5" blockId="8.[248,659,1845,1901]" box="[372,536,1878,1901]" pageId="8" pageNumber="9">
<figureCitation id="ACBA2A31FFAD99215F64F1554EDCF8F5" box="[372,462,1878,1901]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="8" pageNumber="9">Figs 5-6</figureCitation>
,
<figureCitation id="ACBA2A31FFAD99215FC9F1554D0AF8F5" box="[473,536,1878,1901]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">44-65</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7C9B653FFFAD99205E87F1864D64FD06" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="343E36B4FFAD99215E87F1864DE6F785" blockId="8.[151,757,1925,2077]" pageId="8" pageNumber="9">
<emphasis id="06F5EAA6FFAD99215E87F1864E61F804" bold="true" box="[151,371,1925,1948]" pageId="8" pageNumber="9">Material examined</emphasis>
:
<materialsCitation id="84E93CE9FFAD99215F94F1864FFCF864" ID-GBIF-Occurrence="3391002303" collectingDate="1987-04-23" collectionCode="MHNG" collectorName="C. Besuchet &amp; Besuchet, I." country="Malaysia" elevation="1550" location="Mt. Kinabalu" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
<collectionCode id="5290AE71FFAD99215F94F1864ECBF804" box="[388,473,1925,1948]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="9" type="Museum">MHNG</collectionCode>
;
<specimenCount id="2287FD3DFFAD99215FF7F1864D20F804" box="[487,562,1925,1948]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(genital complex in prep. 240317/01);
<collectingCountry id="4C967624FFAD99215FB6F1A64D1FF824" box="[422,525,1957,1980]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
,
<collectingRegion id="F645F856FFAD99215C35F1A64DA7F824" box="[549,693,1957,1980]" country="Malaysia" name="Sabah" pageId="8" pageNumber="9">Sabah State</collectingRegion>
,
<location id="315E606FFFAD99215CDEF1A64FEFF844" LSID="urn:lsid:plazi:treatment:BC2887A2FFAD99205EE8F1364DBBFC38:315E606FFFAD99215CDEF1A64FEFF844" country="Malaysia" name="Mt. Kinabalu" pageId="8" pageNumber="9" stateProvince="Sabah">Mt. Kinabalu</location>
,
<quantity id="F3799B51FFAD99215F05F1C64E64F844" box="[277,374,1989,2012]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="9" unit="m" value="1550.0">
<elevation id="BFACD187FFAD99215F05F1C64E64F844" box="[277,374,1989,2012]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="9" unit="m" value="1550.0">1550 m</elevation>
</quantity>
;
<date id="403F1074FFAD99215F9EF1C64D1EF844" box="[398,524,1989,2012]" pageId="8" pageNumber="9" value="1987-04-23">
<collectingDate id="507BE99CFFAD99215F9EF1C64D1EF844" box="[398,524,1989,2012]" pageId="8" pageNumber="9" value="1987-04-23">23.04.1987</collectingDate>
</date>
; leg.
<collectorName id="99745362FFAD99215C4EF1C64DE2F844" box="[606,752,1989,2012]" pageId="8" pageNumber="9">C. Besuchet</collectorName>
, I. Löbl.
</materialsCitation>
<materialsCitation id="84E93CE9FFAD99215F05F1E64DE2F785" ID-GBIF-Occurrence="3391002305" collectionCode="ZIN" country="Malaysia" location="Sabah State" pageId="8" pageNumber="9" specimenCount="2" specimenCount-male="2" stateProvince="Sabah">
<collectionCode id="5290AE71FFAD99215F05F1E64E57F864" box="[277,325,2021,2044]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="9" type="Museum">ZIN</collectionCode>
;
<specimenCount id="2287FD3DFFAD99215F46F1E64EBDF864" box="[342,431,2021,2044]" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
(genital complexes in prep. 240317/02 and 240317/03);
<collectingCountry id="4C967624FFAD99215FF9FE054D5DF785" box="[489,591,2054,2077]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
,
<collectingRegion id="F645F856FFAD99215C74FE054DE2F785" box="[612,752,2054,2077]" country="Malaysia" name="Sabah" pageId="8" pageNumber="9">Sabah State</collectingRegion>
</materialsCitation>
,
</paragraph>
<paragraph id="343E36B4FFAD99215D2FF7014B12FEA1" blockId="8.[831,1436,258,313]" pageId="8" pageNumber="9">
Gunung Alab Resort,
<quantity id="F3799B51FFAD99215A52F7014BCFFE81" box="[1090,1245,258,281]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="8" pageNumber="9" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500-1600 m</quantity>
; 25.-
<date id="403F1074FFAD99215B06F7014A8AFE81" box="[1302,1432,258,281]" pageId="8" pageNumber="9" value="2008-09-28">28.09.2008</date>
; leg. A. Michailov.
</paragraph>
<paragraph id="343E36B4FFAD99215D2FF7524A99F875" blockId="8.[831,1437,337,2029]" pageId="8" pageNumber="9">
<emphasis id="06F5EAA6FFAD99215D2FF7524B90FEF0" bold="true" box="[831,1154,337,360]" pageId="8" pageNumber="9">Somatic characters of male:</emphasis>
General colour brownish, with scattered light dots (
<figureCitation id="ACBA2A31FFAD99215A70F7724BB7FE10" box="[1120,1189,369,392]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
). Colouration of head as in
<figureCitation id="ACBA2A31FFAD99215D91F7924CD7FE30" box="[897,965,401,424]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
, eyes brown or black, facial part of head with large longitudinal dark (brown or black) spot. Scapi and proximal parts of antennae yellowish, distal parts of antennae greyish. Impressions on pronotum blackish. Mouthparts, thorax from below, legs and abdomen yellowish; 3rd-7th abdominal tergites and sternites with pairs of black lateral spots. Surfaces smooth and lustrous, head with several very weak transverse wrinkles between eyes, weak punctuation present in proximal part of tegmina, mostly in costal field. Head slightly longer than wide, with indistinct transverse impression between antennal sockets (
<figureCitation id="ACBA2A31FFAD99215D57F5104C83FCB2" box="[839,913,787,810]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="8" pageNumber="9">Figs 6</figureCitation>
,
<figureCitation id="ACBA2A31FFAD99215DB1F5104CD2FCB2" box="[929,960,787,810]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">44</figureCitation>
); ocellar spots small but distinct; distance between eyes 0.6 times eye length; distance between antennal sockets about 1.5-1.6 times scape length (~
<quantity id="F3799B51FFAD99215B61F5504C8AFC12" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="8" pageNumber="9" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6- 1.7 mm</quantity>
); approximate length ratio of 3rd-5th segments of maxillary palps 1.0: 1.0: 1.2-1.3. Pronotum as in
<figureCitation id="ACBA2A31FFAD99215D2FF5B04C94FC52" box="[831,902,947,970]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="8" pageNumber="9">Figs 5</figureCitation>
,
<figureCitation id="ACBA2A31FFAD99215D85F5B04CA0FC52" box="[917,946,947,970]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">45</figureCitation>
, with a pair of impressions; anterior margin widely rounded. Tegmina and wings completely developed (
<figureCitation id="ACBA2A31FFAD99215DACF5F04CEFFB92" box="[956,1021,1011,1034]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
), surpassing abdominal apex. Tegmina with rounded apex, sclerotized in costal field; venation slightly obliterated when seen from above, well visible on ventral side of tegmen; costal field wide;
<emphasis id="06F5EAA6FFAD99215B93F2574A8EFBF3" box="[1411,1436,1108,1131]" italics="true" pageId="8" pageNumber="9">Sc</emphasis>
thickened (well visible on ventral side of tegmen);
<emphasis id="06F5EAA6FFAD99215B95F2774A87FB13" box="[1413,1429,1140,1163]" italics="true" pageId="8" pageNumber="9">
<collectionCode id="5290AE71FFAD99215B95F2774A87FB13" box="[1413,1429,1140,1163]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="9">R</collectionCode>
</emphasis>
,
<emphasis id="06F5EAA6FFAD99215D2FF2974C47FB33" box="[831,853,1172,1195]" italics="true" pageId="8" pageNumber="9">M</emphasis>
and
<emphasis id="06F5EAA6FFAD99215D87F2974CD5FB33" box="[919,967,1172,1195]" italics="true" pageId="8" pageNumber="9">CuP</emphasis>
basally fused. Wings membranous, only with weakly sclerotized area of
<emphasis id="06F5EAA6FFAD99215A8FF2B74BAAFB53" box="[1183,1208,1204,1227]" italics="true" pageId="8" pageNumber="9">Sc</emphasis>
and anterior rami of
<emphasis id="06F5EAA6FFAD99215D2FF2D74C5DFB73" box="[831,847,1236,1259]" italics="true" pageId="8" pageNumber="9">
<collectionCode id="5290AE71FFAD99215D2FF2D74C5DFB73" box="[831,847,1236,1259]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="9">R</collectionCode>
</emphasis>
. Fore tibiae not thickened distally. Anterior margin of fore femora of armed
<typeStatus id="EB3A8816FFAD99215A67F2F74BB7FA93" box="[1143,1189,1268,1291]" pageId="8" pageNumber="9">type</typeStatus>
B, with 4-7 spines, including 2 apical ones. Tibial spines well developed. Structure of hind tarsus (
<figureCitation id="ACBA2A31FFAD99215A90F3374A1AFAD3" box="[1152,1288,1332,1355]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Figs 48-49</figureCitation>
): metatarsus slightly shorter than other tarsal segments combined, with small euplantula and 2 more or less equal rows of spines along lower margin; 2nd-4th segments with large euplantulae, spines along lower margins absent; 1st-3rd segments bordered with 0-3 “additional spines” on inside and outside (
<figureCitation id="ACBA2A31FFAD99215A2AF3F64BA1F994" box="[1082,1203,1525,1548]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Figs 48-49</figureCitation>
,
<emphasis id="06F5EAA6FFAD99215AD1F3F64BF4F994" box="[1217,1254,1525,1548]" italics="true" pageId="8" pageNumber="9">a.s.</emphasis>
), in some males “additional spines” absent; claws symmetrical, simple; arolium distinct, about half as long as claw (
<figureCitation id="ACBA2A31FFAD99215B2BF0364A83F9D4" box="[1339,1425,1589,1612]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 49</figureCitation>
). Fore (
<figureCitation id="ACBA2A31FFAD99215D91F0564CC2F9F4" box="[897,976,1621,1644]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 46</figureCitation>
) and mid (
<figureCitation id="ACBA2A31FFAD99215A59F0564B8AF9F4" box="[1097,1176,1621,1644]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 47</figureCitation>
) tarsi dissimilar to hind tarsi: metatarsal euplantulae large, spines along lower margin absent; one pair of “additional spines” present only on 1st-3rd segment of mid tarsi of
<specimenCount id="2287FD3DFFAD99215B11F0B64A8BF954" box="[1281,1433,1717,1740]" pageId="8" pageNumber="9" type="generic">one specimen</specimenCount>
. Abdomen without visible glandular specializations; spiracle-bearing outgrowths of tergite VIII without attenuate posterolateral angles (
<figureCitation id="ACBA2A31FFAD99215AA0F1154A18F8B5" box="[1200,1290,1814,1837]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 50</figureCitation>
). Anal plate (tergite X) with widely rounded hind margin, with a triangular medial incision (
<figureCitation id="ACBA2A31FFAD99215A77F1554BABF8F5" box="[1127,1209,1878,1901]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 50</figureCitation>
). Cerci slender, with distinct segments (
<figureCitation id="ACBA2A31FFAD99215A0CF1754B64F815" box="[1052,1142,1910,1933]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 50</figureCitation>
). Paraprocts of blaberidtype (
<figureCitation id="ACBA2A31FFAD99215D92F1954CCAF835" box="[898,984,1942,1965]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 51</figureCitation>
). Hypandrium nearly symmetrical, with hind margin between styli projected and membranous, median incision weak (
<figureCitation id="ACBA2A31FFAD99215A27F1D54B95F875" box="[1079,1159,2006,2029]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Fig. 52</figureCitation>
); styli short, cylindrical.
</paragraph>
<paragraph id="343E36B4FFAD99215D2FFE054A8EF785" blockId="8.[831,1436,2054,2077]" box="[831,1436,2054,2077]" pageId="8" pageNumber="9">
<heading id="6F7681D8FFAD99215D2FFE054A8EF785" box="[831,1436,2054,2077]" centered="true" fontSize="10" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="06F5EAA6FFAD99215D2FFE054CEAF785" bold="true" box="[831,1016,2054,2077]" pageId="8" pageNumber="9">Male genitalia</emphasis>
(
<figureCitation id="ACBA2A31FFAD99215A08FE054BB4F785" box="[1048,1190,2054,2077]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="8" pageNumber="9">Figs 53-65</figureCitation>
): Right phallomere
</heading>
</paragraph>
<paragraph id="343E36B4FFAC99205E87F7014D64FD06" blockId="9.[151,756,258,670]" pageId="9" pageNumber="10">
(
<collectionCode id="5290AE71FFAC99205E8FF7014FA1FE81" box="[159,179,258,281]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
+N): caudal part of sclerite
<collectionCode id="5290AE71FFAC99205FEFF7014D01FE81" box="[511,531,258,281]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
1
<collectionCode id="5290AE71FFAC99205C0CF7014D3CFE81" box="[540,558,258,281]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="10">T</collectionCode>
well sclerotized, subrectangular in shape, with more or less attenuate medial angle (
<figureCitation id="ACBA2A31FFAC99205F2DF7404EAEFEC2" box="[317,444,323,346]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="9" pageNumber="10">Figs 53-54</figureCitation>
,
<emphasis id="06F5EAA6FFAC99205FDEF7404D30FEC2" box="[462,546,323,346]" italics="true" pageId="9" pageNumber="10">
c.p.
<collectionCode id="5290AE71FFAC99205FE2F7404D17FEC2" box="[498,517,323,346]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
<figureCitation id="ACBA2A31FFAC99205C15F7404D30FEC2" box="[517,546,323,346]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="9" pageNumber="10">1T</figureCitation>
</emphasis>
), densely covered with bristles;
<collectionCode id="5290AE71FFAC99205F3AF7604E2EFEE2" box="[298,316,355,378]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
2 angulate;
<collectionCode id="5290AE71FFAC99205FA9F7604EDEFEE2" box="[441,460,355,378]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
3 robust, widened caudally;
<collectionCode id="5290AE71FFAC99205E87F7874FBBFE03" box="[151,169,388,411]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
4 and
<collectionCode id="5290AE71FFAC99205EFFF7874E10FE03" box="[239,258,388,411]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
5 large, plate-like;
<collectionCode id="5290AE71FFAC99205FC0F7874EF1FE03" box="[464,483,388,411]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
5 fused with
<collectionCode id="5290AE71FFAC99205C64F7874D9BFE03" box="[628,649,388,411]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="10">R</collectionCode>
3. Sclerite L2D (L1) divided into basal and apical parts (
<figureCitation id="ACBA2A31FFAC99205C84F7A74DF5FE23" box="[660,743,420,443]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="9" pageNumber="10">Fig. 55</figureCitation>
); basal part rod-like, weakly widened cranially; “apical sclerite” variable in shape (
<figureCitation id="ACBA2A31FFAC99205FDFF7E64D5EFE64" box="[463,588,485,508]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="9" pageNumber="10">Figs 56-61</figureCitation>
), covered with recumbent bristles; “dorsal outgrowth” large (
<figureCitation id="ACBA2A31FFAC99205CD4F4054FCAFDA5" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="9" pageNumber="10">Figs 56-61</figureCitation>
,
<emphasis id="06F5EAA6FFAC99205EF9F4254E00FDA5" box="[233,274,550,573]" italics="true" pageId="9" pageNumber="10">d.o.</emphasis>
). Sclerite L3 (L2d) with basal subsclerite (
<figureCitation id="ACBA2A31FFAC99205E8FF4454FE1FDC5" box="[159,243,582,605]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="9" pageNumber="10">Fig. 62</figureCitation>
,
<emphasis id="06F5EAA6FFAC99205F13F4454E26FDC5" box="[259,308,582,605]" italics="true" pageId="9" pageNumber="10">
b.L
<figureCitation id="ACBA2A31FFAC99205F36F4454E26FDC5" box="[294,308,582,605]" captionStart="Figs 1-9" captionStartId="2.[151,192,1970,1991]" captionTargetBox="[163,1418,269,1921]" captionTargetId="figure-8@2.[151,1436,257,1935]" captionTargetPageId="2" captionText="Figs 1-9. Photos of Morphna maculata (Brunner von Wattenwyl, 1865) (1-2: female, syntype, 3-4: male), Morphnodes goliath (Shelford, 1906) (5-6: male), Pseudocalolampra inexpectata Roth &amp; Princis, 1971 (7: male), Colapteroblatta compsa Hebard, 1919 (8: male, 9: female). (1, 3, 5) Dorsal view of entire specimen. (2, 4, 6-9) Facial part of head. Abdominal apex removed in Fig. 3. Scale bars 10 mm: a = 1, b = 3, c = 5. Other photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5589893" httpUri="https://zenodo.org/record/5589893/files/figure.png" pageId="9" pageNumber="10">3</figureCitation>
</emphasis>
); “folded structure” distinct, with short bristles; apex of L3 with “small tooth” (
<figureCitation id="ACBA2A31FFAC99205C52F4644DABFDE6" box="[578,697,615,638]" captionStart="Figs 44-65" captionStartId="7.[163,204,1830,1851]" captionTargetBox="[165,1423,257,1781]" captionTargetId="figure-5@7.[155,1427,246,1790]" captionTargetPageId="7" captionText="Figs 44-65. Morphnodes goliath (Shelford, 1906), specimen in MHNG (44-53, 55, 59-65), specimen No. 200709a (54, 56-57) and specimen No. 200709b (58). (44) Facial part of head. (45) Pronotum, dorsal view. (46-47, 49) Fore (46), mid (47) and hind (49) tarsus, prolateral view. (48) Hind metatarsus, ventral view. (50) Abdominal apex, dorsal view. (51) Paraprocts, ventral view. (52) Hypandrium, ventral view. (53-54) Right phallomere, dorsal view. (55) Sclerite L2D, dorsal view, bristles not shown. (56, 59) Posterior part of sclerite L2D, lateral view. (57-58, 60) Same, dorsal view. (61) Same, contralateral view, bristles not shown. (62) Sclerite L3. (63-65) Apex of sclerite L3. Dotted areas show membranous parts.Abbreviations: a.s., b.L3, c.p.R1T, d.o., R2, R3, R4, R5, s.t. see paragraph “abbreviation used in figures”; for details see text. Scale bars 1 mm: a = 44, b = 45, c = 46-49, d = 50, e = 51, f = 52, g = 53-54, h = 55, i = 56-61, j = 62, k = 63-65." figureDoi="http://doi.org/10.5281/zenodo.5589908" httpUri="https://zenodo.org/record/5589908/files/figure.png" pageId="9" pageNumber="10">Figs 63-65</figureCitation>
,
<emphasis id="06F5EAA6FFAC99205CD4F4644DF1FDE6" box="[708,739,615,638]" italics="true" pageId="9" pageNumber="10">s.t.</emphasis>
); groove
<emphasis id="06F5EAA6FFAC99205EF8F4844E1DFD06" box="[232,271,647,670]" italics="true" pageId="9" pageNumber="10">hge</emphasis>
absent. Sclerite L4U (L3d) large.
</paragraph>
</subSubSection>
<subSubSection id="7C9B653FFFAC99205E87F4B44DFEFC97" pageId="9" pageNumber="10" type="description">
<paragraph id="343E36B4FFAC99205E87F4B44DFEFC97" blockId="9.[151,756,695,783]" pageId="9" pageNumber="10">
<emphasis id="06F5EAA6FFAC99205E87F4B44E09FD56" bold="true" box="[151,283,695,718]" pageId="9" pageNumber="10">Dimensions</emphasis>
(in mm): Head length 5.8-6.0, head width 5.5-5.7; pronotum length 9.1-10.1, pronotum width 13.6-15.1; tegmen length 32.5-45.0, tegmen width 15.0.
</paragraph>
</subSubSection>
<subSubSection id="7C9B653FFFAC99205E87F5244DBBFC38" pageId="9" pageNumber="10" type="discussion">
<paragraph id="343E36B4FFAC99205E87F5244DBBFC38" blockId="9.[151,756,807,928]" pageId="9" pageNumber="10">
<emphasis id="06F5EAA6FFAC99205E87F5244FECFCA6" bold="true" box="[151,254,807,830]" pageId="9" pageNumber="10">Remark:</emphasis>
This species was described on the basis of
<specimenCount id="2287FD3DFFAC99205CDDF5244FD9FCC7" pageId="9" pageNumber="10" type="male">one male</specimenCount>
from Mt Matang in northern Borneo (
<bibRefCitation id="50104B45FFAC99205C9FF54B4FC3FCE7" author="Shelford R." pageId="9" pageNumber="10" pagination="265 - 275" refId="ref10757" refString="Shelford R. 1906. Studies of the Blattidae. III. Some new Blattidae from Sarawak, Borneo, in the Hope Department, Oxford University Museum. Transactions of the Entomological Society, London 1: 265 - 275." type="journal article" year="1906">Shelford, 1906</bibRefCitation>
). The
<typeStatus id="EB3A8816FFAC99205F35F56B4E96FCE7" box="[293,388,872,895]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
(not examined) is kept in the University Museum of Oxford (
<bibRefCitation id="50104B45FFAC99205FFAF58A4D8CFC38" author="Beccaloni G. W." box="[490,670,905,928]" pageId="9" pageNumber="10" refId="ref10070" refString="Beccaloni G. W. 2014. Cockroach Species File Online. Version 5.0 / 5.0. World Wide Web electronic publication. Available at http: // Cockroach. SpeciesFile. org (accessed 22 March 2017)." type="url" year="2014">Beccaloni, 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>