241 lines
27 KiB
XML
241 lines
27 KiB
XML
<document id="4A8EC600B427980617BA39883C439CEC" ID-DOI="10.11646/zootaxa.3734.4.1" ID-GBIF-Dataset="f372aec6-39ab-4836-a1f3-d0eefa9efefd" ID-ISSN="1175-5326" ID-Zenodo-Dep="223309" ID-ZooBank="00DC3ED7-62FD-4D99-ABCC-0DC57D1A51E7" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460696324322" checkinUser="plazi" docAuthor="Bamber, Roger N." docDate="2013" docId="03DC9341FF90FFB0C7F31F3CFC78F859" docLanguage="en" docName="zt03734p441.pdf" docOrigin="Zootaxa 3734 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Actenos Bamber, 2013, gen. nov." docType="treatment" docVersion="8" lastPageNumber="408" masterDocId="FFE5EB39FF96FFB7C7641849FFAFFFE0" masterDocTitle="Tanaidaceans from Brunei, IV. The Families Kalliapseudidae, Pagurapseudopsidae, Parapseudidae and Apseudidae (Crustacea: Peracarida: Tanaidacea: Apseudomorpha), with descriptions of a new genus and six new species" masterLastPageNumber="441" masterPageNumber="401" pageNumber="407" updateTime="1698353876566" updateUser="plazi">
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<mods:titleInfo id="F3CA740106B6C9EE595339AE0556D6EC">
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<mods:title id="246DAC4FA781E7585FB6D24F14AE7AA8">Tanaidaceans from Brunei, IV. The Families Kalliapseudidae, Pagurapseudopsidae, Parapseudidae and Apseudidae (Crustacea: Peracarida: Tanaidacea: Apseudomorpha), with descriptions of a new genus and six new species</mods:title>
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<mods:name id="A6D8D1F31C5D59E0FAB9251EEE63494C" type="personal">
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<mods:roleTerm id="FCAE63CF6253774AA7FBA2D8A125944E">Author</mods:roleTerm>
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<mods:namePart id="F11D2718DDB6EBC28F315379BA2AC10C">Bamber, Roger N.</mods:namePart>
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<mods:typeOfResource id="A58042940B6A4F51DEBBF1EEE16F88C9">text</mods:typeOfResource>
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<mods:title id="BD366A88114E42E8AF0F2C7F3BD4486A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7C38BD53422C587F1B1EC9D5BCD4AA59">
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<mods:date id="38B2D271DA75334259A9DD2EEFEFDD9F">2013</mods:date>
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<mods:detail id="898490F635CEDCF72C4EEC21CA867EE5" type="volume">
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<mods:number id="6261F1558CAA2947546AEB6F982E60C3">3734</mods:number>
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<mods:detail id="DDB10F5D3516B5B4254D124F982CF345" type="issue">
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<mods:number id="6158CF12AE346B2DE2A4DFC533706217">4</mods:number>
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<mods:classification id="65BB45A4BF818099094ABFD26884AF09">journal article</mods:classification>
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<mods:identifier id="501041D55738C9FF14A05AFD91715806" type="DOI">10.11646/zootaxa.3734.4.1</mods:identifier>
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<mods:identifier id="ABF9070CA7F13A055829FDCAF27500C5" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DC9341FF90FFB0C7F31F3CFC78F859" ID-DOI="http://doi.org/10.5281/zenodo.6148930" ID-GBIF-Taxon="119582086" ID-Zenodo-Dep="6148930" LSID="urn:lsid:plazi:treatment:03DC9341FF90FFB0C7F31F3CFC78F859" httpUri="http://treatment.plazi.org/id/03DC9341FF90FFB0C7F31F3CFC78F859" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
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<subSubSection id="C36F71DCFF90FFB1C7F31F3CFE10F86F" box="[151,447,1909,1935]" pageId="6" pageNumber="407" type="nomenclature">
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<paragraph id="8BCA2257FF90FFB1C7F31F3CFE10F86F" blockId="6.[151,447,1909,1935]" box="[151,447,1909,1935]" pageId="6" pageNumber="407">
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<heading id="D082953BFF90FFB1C7F31F3CFE10F86F" bold="true" box="[151,447,1909,1935]" fontSize="11" level="1" pageId="6" pageNumber="407" reason="1">
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<emphasis id="B901FE45FF90FFB1C7F31F3CFE10F86F" bold="true" box="[151,447,1909,1935]" pageId="6" pageNumber="407">
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Genus
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<taxonomicName id="4C7559D4FF90FFB1C78A1F3CFEE1F86F" ID-CoL="84L5X" box="[238,334,1909,1935]" class="Malacostraca" family="Parapseudidae" genus="Actenos" kingdom="Animalia" order="Tanaidacea" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B901FE45FF90FFB1C78A1F3CFEE1F86F" bold="true" box="[238,334,1909,1935]" italics="true" pageId="6" pageNumber="407">Actenos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A232433EFF90FFB1C6311F3CFE10F86F" box="[341,447,1909,1935]" pageId="6" pageNumber="407" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F71DCFF90FFB0C7F31FF5FD34FE4D" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" type="diagnosis">
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<paragraph id="8BCA2257FF90FFB0C7F31FF5FD34FE4D" blockId="6.[151,1435,1980,2005]" lastBlockId="7.[151,1436,151,1977]" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
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<emphasis id="B901FE45FF90FFB1C7F31FF5FEA8F835" bold="true" box="[151,263,1980,2005]" pageId="6" pageNumber="407">Diagnosis</emphasis>
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: Pakistanapseudin without ventral-distal row of fine or lanceolate spines on propodus of pereopod 6; dactylus of pereopods 1 to 3 distally bifurcate, without distinct unguis; carpus of pereopod 1 dorsal margin with conspicuous array of long setae. Mandibular molar with or without rugose or denticulate distal flange. Eyes present, pigmented, rostrum centrally extended to a point; pereonite 4 (and either 3 or 5) as long as or longer than wide; all pleonites together nearly twice as long as carapace; epistome with up-curved spine-like apophysis; maxillule inner endite with proximal inner apophysis. Pereonites mostly wider at attachment of pereopod coxae. Pereopod 1 with spine(s) on coxal apophysis, without spines on dorsal margin of propodus. Pleonites expanded posterolaterally to accommodate ventrolateral attachment of pleopods; pleopod bases with both inner and outer setae. Peduncle articles 2 and 3 of mature male antenna fused.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F71DCFF91FFB0C7A319FEFC42FE15" pageId="7" pageNumber="408" type="etymology">
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<paragraph id="8BCA2257FF91FFB0C7A319FEFC42FE15" blockId="7.[151,1436,151,1977]" pageId="7" pageNumber="408">
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<emphasis id="B901FE45FF91FFB0C7A319FEFEE7FE30" bold="true" box="[199,328,439,464]" pageId="7" pageNumber="408">Etymology</emphasis>
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: From the Greek—
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<emphasis id="B901FE45FF91FFB0C54E19F1FD94FE2F" box="[554,571,440,463]" italics="true" pageId="7" pageNumber="408">A</emphasis>
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(not) and
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<emphasis id="B901FE45FF91FFB0C5D119FEFD53FE30" box="[693,764,439,464]" italics="true" pageId="7" pageNumber="408">ktenos</emphasis>
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(a comb), referring to the lack of a row of spines/spinules along the ventral to distal margin of the propodus of pereopod 6. Masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F71DCFF91FFB0C7A319B6FB18FDDD" pageId="7" pageNumber="408" type="materials_examined">
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<paragraph id="8BCA2257FF91FFB0C7A319B6FC09FDF8" blockId="7.[151,1436,151,1977]" box="[199,934,511,536]" pageId="7" pageNumber="408">
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<emphasis id="B901FE45FF91FFB0C7A319B6FEF4FDF8" bold="true" box="[199,347,511,536]" pageId="7" pageNumber="408">
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<typeStatus id="54CE9CF5FF91FFB0C7A319B6FEADFDF8" box="[199,258,511,536]" pageId="7" pageNumber="408">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C7559D4FF91FFB0C60E1A49FD9BFDF8" box="[362,564,511,536]" class="Malacostraca" family="Parapseudidae" genus="Actenos" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="marshalli" status="sp. nov.">
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<emphasis id="B901FE45FF91FFB0C60E1A49FD9BFDF8" box="[362,564,511,536]" italics="true" pageId="7" pageNumber="408">Actenos marshalli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A232433EFF91FFB0C55819B6FD3AFDF8" box="[572,661,511,536]" pageId="7" pageNumber="408" rank="species">
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<emphasis id="B901FE45FF91FFB0C55819B6FD3AFDF8" bold="true" box="[572,661,511,536]" pageId="7" pageNumber="408">sp. nov.</emphasis>
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</taxonomicNameLabel>
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by original designation.
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</paragraph>
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<paragraph id="8BCA2257FF91FFB0C7A31A6DFB18FDDD" blockId="7.[151,1436,151,1977]" box="[199,1207,548,573]" pageId="7" pageNumber="408">
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<emphasis id="B901FE45FF91FFB0C7A31A6DFEC8FDDD" bold="true" box="[199,359,548,573]" pageId="7" pageNumber="408">Other species</emphasis>
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:
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<taxonomicName id="4C7559D4FF91FFB0C6121A6DFDFEFDDD" box="[374,593,548,573]" class="Malacostraca" family="Parapseudidae" genus="Actenos" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="kalimantus" status="sp. nov.">
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<emphasis id="B901FE45FF91FFB0C6121A6DFDFEFDDD" box="[374,593,548,573]" italics="true" pageId="7" pageNumber="408">Actenos kalimantus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A232433EFF91FFB0C53C1A6DFD1DFDDD" box="[600,690,548,573]" pageId="7" pageNumber="408" rank="species">
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<emphasis id="B901FE45FF91FFB0C53C1A6DFD1DFDDD" bold="true" box="[600,690,548,573]" pageId="7" pageNumber="408">sp. nov.</emphasis>
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</taxonomicNameLabel>
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,
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<taxonomicName id="4C7559D4FF91FFB0C5DB1A6DFB9FFDDD" authority="Shiino, 1963" authorityName="Shiino" authorityYear="1963" box="[703,1072,548,573]" class="Malacostraca" family="Apseudidae" genus="Apseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tenuicorporeus" status="comb. nov.">
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<emphasis id="B901FE45FF91FFB0C5DB1A6DFC28FDDC" box="[703,903,548,572]" italics="true" pageId="7" pageNumber="408">A. tenuicorporeus</emphasis>
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(Shiino, 1963)
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</taxonomicName>
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<taxonomicNameLabel id="A232433EFF91FFB0C3531A6DFB18FDDD" box="[1079,1207,548,573]" pageId="7" pageNumber="408">
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<emphasis id="B901FE45FF91FFB0C3531A6DFB18FDDD" bold="true" box="[1079,1207,548,573]" pageId="7" pageNumber="408">comb. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F71DCFF91FFB0C7A31A0EFC78F859" pageId="7" pageNumber="408" type="discussion">
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<paragraph id="8BCA2257FF91FFB0C7A31A0EFADAFD48" blockId="7.[151,1436,151,1977]" pageId="7" pageNumber="408">
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<emphasis id="B901FE45FF91FFB0C7A31A0EFE9BFD80" bold="true" box="[199,308,583,608]" pageId="7" pageNumber="408">Remarks</emphasis>
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: With the relatively elongate pereonites and pleonites, a labial palp with an inner apophysis and two distal spines, and the secondary segmentation of the antennal and antennular flagella, the two new species described below are clearly members of the
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<taxonomicName id="4C7559D4FF91FFB0C5E21AC6FCCBFD48" box="[646,868,655,680]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subFamily" subFamily="Pakistanapseudinae">Pakistanapseudinae</taxonomicName>
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, and constitute the new genus established here.
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</paragraph>
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<paragraph id="8BCA2257FF91FFB0C7A31AFCFCB5FC45" blockId="7.[151,1436,151,1977]" pageId="7" pageNumber="408">
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However, only one other species of the subfamily,
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<taxonomicName id="4C7559D4FF91FFB0C4471AFDFA8BFD2D" authority="Shiino, 1963" authorityName="Shiino" authorityYear="1963" box="[803,1316,692,717]" class="Malacostraca" family="Parapseudidae" genus="Biropalostoma" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tenuicorporeus">
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<emphasis id="B901FE45FF91FFB0C4471AFDFBDBFD2C" box="[803,1140,692,717]" italics="true" pageId="7" pageNumber="408">Biropalostoma tenuicorporeus</emphasis>
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(Shiino, 1963)
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</taxonomicName>
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, lacks the ventral-distal row of fine spines on the propodus of pereopod 6, where that pereopod is known; it is unknown in
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<taxonomicName id="4C7559D4FF91FFB0C7F31AB5FD27FCF5" authority="Gutu, 1996" authorityName="Gutu" authorityYear="1996" box="[151,648,764,789]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B901FE45FF91FFB0C7F31AB5FE40FCF5" box="[151,495,764,789]" italics="true" pageId="7" pageNumber="408">Pakistanapseudes brasiliensis</emphasis>
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Guţu, 1996
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</taxonomicName>
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(a),
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<taxonomicName id="4C7559D4FF91FFB0C5DD1AB5FB52FCF4" authority="Bamber & Sheader, 2003" authorityName="Bamber & Sheader" authorityYear="2003" box="[697,1277,764,789]" class="Malacostraca" family="Parapseudidae" genus="Leptolicoa" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="thokozele">
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<emphasis id="B901FE45FF91FFB0C5DD1AB5FC07FCF5" box="[697,936,764,789]" italics="true" pageId="7" pageNumber="408">Leptolicoa thokozele</emphasis>
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(Bamber & Sheader, 2003)
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</taxonomicName>
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and
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<taxonomicName id="4C7559D4FF91FFB0C2271AB4FE40FCD8" authority="Bamber & Sheader, 2003" authorityName="Bamber & Sheader" authorityYear="2003" class="Malacostraca" family="Parapseudidae" genus="Biropalostoma" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="goofi">
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<emphasis id="B901FE45FF91FFB0C2271AB4FA33FCF4" box="[1347,1436,764,788]" italics="true" pageId="7" pageNumber="408">B. goofi</emphasis>
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(Bamber & Sheader, 2003)
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</taxonomicName>
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, but those species have other clear distinctions from
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<taxonomicName id="4C7559D4FF91FFB0C3D11B69FAA1FCD7" box="[1205,1294,800,823]" class="Malacostraca" family="Parapseudidae" genus="Actenos" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B901FE45FF91FFB0C3D11B69FAA1FCD7" box="[1205,1294,800,823]" italics="true" pageId="7" pageNumber="408">Actenos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A232433EFF91FFB0C2461B69FA3EFCD7" box="[1314,1425,800,824]" pageId="7" pageNumber="408" rank="genus">gen. nov</taxonomicNameLabel>
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..
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<taxonomicName id="4C7559D4FF91FFB0C7F31B0CFEF5FCBC" box="[151,346,837,860]" class="Malacostraca" family="Parapseudidae" genus="Biropalostoma" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tenuicorporeus">
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<emphasis id="B901FE45FF91FFB0C7F31B0CFEF5FCBC" box="[151,346,837,860]" italics="true" pageId="7" pageNumber="408">B. tenuicorporeus</emphasis>
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</taxonomicName>
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also has a pereopod 1 carpal setal array, no dorsal marginal spines on the carpus or propodus of pereopod 1, has very similar claw morphology on pereopods 2 to 6 to the species of
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<taxonomicName id="4C7559D4FF91FFB0C3551B21FB25FC9F" box="[1073,1162,872,895]" class="Malacostraca" family="Parapseudidae" genus="Actenos" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
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<emphasis id="B901FE45FF91FFB0C3551B21FB25FC9F" box="[1073,1162,872,895]" italics="true" pageId="7" pageNumber="408">Actenos</emphasis>
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</taxonomicName>
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, and apparently does not have a distinct unguis on the bifurcate pereopod1 dactylus.
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</paragraph>
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<paragraph id="8BCA2257FF91FFB0C7A31BE6FDF3FACF" blockId="7.[151,1436,151,1977]" pageId="7" pageNumber="408">
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Having been moved originally from
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<taxonomicName id="4C7559D4FF91FFB0C5EC1BE6FD5FFC28" box="[648,752,943,968]" class="Malacostraca" family="Apseudidae" genus="Apseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
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<emphasis id="B901FE45FF91FFB0C5EC1BE6FD5FFC28" box="[648,752,943,968]" italics="true" pageId="7" pageNumber="408">Apseudes</emphasis>
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</taxonomicName>
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to
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<taxonomicName id="4C7559D4FF91FFB0C4411BE6FB0CFC28" authority="Bacescu, 1978" authorityName="Bacescu" authorityYear="1978" box="[805,1187,943,968]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
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<emphasis id="B901FE45FF91FFB0C4411BE6FC46FC28" box="[805,1001,943,968]" italics="true" pageId="7" pageNumber="408">Pakistanapseudes</emphasis>
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Băcescu, 1978
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</taxonomicName>
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by Băcescu (1978),
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<taxonomicName id="4C7559D4FF91FFB0C7F31B9CFEF5FC0C" box="[151,346,981,1004]" class="Malacostraca" family="Parapseudidae" genus="Biropalostoma" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tenuicorporeus">
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<emphasis id="B901FE45FF91FFB0C7F31B9CFEF5FC0C" box="[151,346,981,1004]" italics="true" pageId="7" pageNumber="408">B. tenuicorporeus</emphasis>
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</taxonomicName>
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was placed in the genus
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<taxonomicName id="4C7559D4FF91FFB0C5D61B9DFBA3FC0C" authority="Bamber, 1997" authorityName="Bamber" authorityYear="1997" box="[690,1036,980,1005]" class="Malacostraca" family="Parapseudidae" genus="Swireapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C5D61B9DFCFBFC0D" box="[690,852,980,1005]" italics="true" pageId="7" pageNumber="408">Swireapseudes</emphasis>
|
||
Bamber, 1997
|
||
</taxonomicName>
|
||
by Bamber (1997), returned to
|
||
<taxonomicName id="4C7559D4FF91FFB0C7F31BBEFEF3FBF0" box="[151,348,1015,1040]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C7F31BBEFEF3FBF0" box="[151,348,1015,1040]" italics="true" pageId="7" pageNumber="408">Pakistanapseudes</emphasis>
|
||
</taxonomicName>
|
||
by Bamber & Sheader (2003), and moved to
|
||
<taxonomicName id="4C7559D4FF91FFB0C4291BBEFC40FBF0" box="[845,1007,1015,1040]" class="Malacostraca" family="Parapseudidae" genus="Biropalostoma" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C4291BBEFC40FBF0" box="[845,1007,1015,1040]" italics="true" pageId="7" pageNumber="408">Biropalostoma</emphasis>
|
||
</taxonomicName>
|
||
by Guţu & Angsupanich (2004) on the basis of having some outer ornamentation on the mandibular body, a feature not present in
|
||
<taxonomicName id="4C7559D4FF91FFB0C3181C55FB7BFBD3" box="[1148,1236,1052,1075]" class="Malacostraca" family="Parapseudidae" genus="Actenos" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C3181C55FB7BFBD3" box="[1148,1236,1052,1075]" italics="true" pageId="7" pageNumber="408">Actenos</emphasis>
|
||
</taxonomicName>
|
||
. The type-species of
|
||
<taxonomicName id="4C7559D4FF91FFB0C7D01C76FEF9FBB8" box="[180,342,1087,1112]" class="Malacostraca" family="Parapseudidae" genus="Biropalostoma" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C7D01C76FEF9FBB8" box="[180,342,1087,1112]" italics="true" pageId="7" pageNumber="408">Biropalostoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7559D4FF91FFB0C60F1C08FC98FBB8" authority="Gutu & Angsupanich, 2004" authorityName="Gutu & Angsupanich" authorityYear="2004" box="[363,823,1087,1112]" class="Malacostraca" family="Parapseudidae" genus="Biropalostoma" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="spiniferum">
|
||
<emphasis id="B901FE45FF91FFB0C60F1C08FE51FBB8" box="[363,510,1088,1112]" italics="true" pageId="7" pageNumber="408">B. spiniferum</emphasis>
|
||
Guţu & Angsupanich, 2004
|
||
</taxonomicName>
|
||
, has a distinct unguis on pereopod 1, a row of spines on the propodus of pereopod 6, and dorsal marginal spines on propodus and carpus of pereopod 1 (as well as a large tubercle on the outer margin of the mandible, a diagnostic character of that genus). Features such as the spination of the pereopod 6 propodus and carpus and the propodus of pereopod 1, as well as the presence of an unguis on the anterior pereopods, together with the ventrolateral placement of the pleopods, are considered here to be of greater phylogenetic significance than the presence or absence of some cuticular ornamentation on the mandible body, and so Shiino’s species is transferred to
|
||
<taxonomicName id="4C7559D4FF91FFB0C69B1D51FDF9FACF" box="[511,598,1304,1327]" class="Malacostraca" family="Parapseudidae" genus="Actenos" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C69B1D51FDF9FACF" box="[511,598,1304,1327]" italics="true" pageId="7" pageNumber="408">Actenos</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCA2257FF91FFB0C7A31D75FB9EF995" blockId="7.[151,1436,151,1977]" pageId="7" pageNumber="408">
|
||
Further, within the
|
||
<taxonomicName id="4C7559D4FF91FFB0C6C91D75FD24FAB5" box="[429,651,1340,1365]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subFamily" subFamily="Pakistanapseudinae">Pakistanapseudinae</taxonomicName>
|
||
, the lack of a distinct unguis on pereopod 1 is only found in the generotype of
|
||
<taxonomicName id="4C7559D4FF91FFB0C6241D16FDA6FA98" box="[320,521,1375,1400]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C6241D16FDA6FA98" box="[320,521,1375,1400]" italics="true" pageId="7" pageNumber="408">Pakistanapseudes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7559D4FF91FFB0C57F1D28FCD3FA98" authority="Bacescu, 1978" authorityName="Bacescu" authorityYear="1978" box="[539,892,1375,1400]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="leptochelatus">
|
||
<emphasis id="B901FE45FF91FFB0C57F1D28FD64FA98" box="[539,715,1375,1400]" italics="true" pageId="7" pageNumber="408">P. leptochelatus</emphasis>
|
||
Băcescu, 1978
|
||
</taxonomicName>
|
||
, based on the figure of Băcescu (1978), and no mention of this was made in his description. Unlike the two species described below,
|
||
<taxonomicName id="4C7559D4FF91FFB0C31D1DCCFA86FA7D" box="[1145,1321,1412,1437]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="leptochelatus">
|
||
<emphasis id="B901FE45FF91FFB0C31D1DCCFA86FA7D" box="[1145,1321,1412,1437]" italics="true" pageId="7" pageNumber="408">P. leptochelatus</emphasis>
|
||
</taxonomicName>
|
||
lacks the bifurcate claw on pereopods 2 and 3 and the large array of setae on the dorsal margin of the pereopod 1 carpus, as well as having far more spines on the carpus and propodus of pereopod 1, but no setae on the outer margin of the basis of the pleopods; there are other distinctions of the mouthpart proportions and setation. Băcescu (1978) makes no mention of a comb of fine spines around the ventrodistal margin of the propodus of pereopod 6, although he does describe and illustrate it for
|
||
<taxonomicName id="4C7559D4FF91FFB0C6B11E70FD4AF9B0" authority="Bacescu, 1978" authorityName="Bacescu" authorityYear="1978" box="[469,741,1591,1616]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="shiinoi">
|
||
<emphasis id="B901FE45FF91FFB0C6B11E70FD96F9B0" box="[469,569,1591,1616]" italics="true" pageId="7" pageNumber="408">P. shiinoi</emphasis>
|
||
Băcescu,
|
||
<quantity id="4C8D8FB2FF91FFB0C5CA1E71FD4AF9B0" box="[686,741,1592,1616]" metricMagnitude="1" metricUnit="m" metricValue="5.02412" pageId="7" pageNumber="408" unit="in" value="1978.0">1978</quantity>
|
||
</taxonomicName>
|
||
in the same paper. If
|
||
<taxonomicName id="4C7559D4FF91FFB0C4BA1E70FB20F9B0" box="[990,1167,1591,1616]" class="Malacostraca" family="Parapseudidae" genus="Pakistanapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="leptochelatus">
|
||
<emphasis id="B901FE45FF91FFB0C4BA1E70FB20F9B0" box="[990,1167,1591,1616]" italics="true" pageId="7" pageNumber="408">P. leptochelatus</emphasis>
|
||
</taxonomicName>
|
||
lacks the spine-row on pereopod 6, then a complete revision of the
|
||
<taxonomicName id="4C7559D4FF91FFB0C5E71E15FCF1F995" box="[643,862,1628,1653]" class="Malacostraca" family="Parapseudidae" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subFamily" subFamily="Pakistanapseudinae">Pakistanapseudinae</taxonomicName>
|
||
may be necessary.
|
||
</paragraph>
|
||
<paragraph id="8BCA2257FF91FFB0C7A31E36FC78F859" blockId="7.[151,1436,151,1977]" pageId="7" pageNumber="408">
|
||
The dorsal setal array on the carpus of pereopod 1 invites speculation as to its function. It may act as an aid in locomotion (swimming), although where such a functional modification is known within the Parapseudidae—in
|
||
<taxonomicName id="4C7559D4FF91FFB0C7F31E8EFCE2F900" authority="BlaZewicz-Paszkowycz & Bamber, 2007" authorityName="BlaZewicz-Paszkowycz & Bamber" authorityYear="2007" box="[151,845,1735,1760]" class="Malacostraca" family="Parapseudidae" genus="Remexudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="toompani">
|
||
<emphasis id="B901FE45FF91FFB0C7F31E8EFE2DF900" box="[151,386,1735,1760]" italics="true" pageId="7" pageNumber="408">Remexudes toompani</emphasis>
|
||
BłaŻewicz-Paszkowycz & Bamber, 2007
|
||
</taxonomicName>
|
||
—it has been achieved by an adaptation of the distal articles of pereopod 2. An alternative hypothesis is as some filter-feeding function: the gut contents of the species described below were found to be organic detritus rather than the sedimentary particles commonly found in guts of other parapseudids. The elaborate distal flange on the mandibular molar in both new species described below also suggests some atypical feeding strategy. Elaboration of dorsal setae on the distal articles of pereopod 1 are present in some other parapseudid genera, such as
|
||
<taxonomicName id="4C7559D4FF91FFB0C5291F35FCEFF875" authority="Gutu, 2006" authorityName="Gutu" authorityYear="2006" box="[589,832,1916,1941]" class="Malacostraca" family="Parapseudidae" genus="Platylicoa" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C5291F35FD12F875" box="[589,701,1916,1941]" italics="true" pageId="7" pageNumber="408">Platylicoa</emphasis>
|
||
Guţu, 2006
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7559D4FF91FFB0C4101F35FA38F874" authority="Bamber, Ariyananda & Silva, 1997" authorityName="Bamber, Ariyananda & Silva" authorityYear="1997" box="[884,1431,1916,1941]" class="Malacostraca" family="Parapseudidae" genus="Ctenapseudes" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901FE45FF91FFB0C4101F35FBA3F875" box="[884,1036,1916,1941]" italics="true" pageId="7" pageNumber="408">Ctenapseudes</emphasis>
|
||
Bamber, Ariyananda & Silva, 1997
|
||
</taxonomicName>
|
||
, although not in a similar conformation, but again their function is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |