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<document id="DB517D7D53A2C0D2C071C389AFE90086" ID-CLB-Dataset="3307" ID-DOI="10.11646/zootaxa.5071.1.5" ID-GBIF-Dataset="f5a054cf-41c7-40cd-bac9-63e1ea8d1b8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5723327" ID-ZooBank="C7DFE939-E932-4570-8496-7A67A51D4D7A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637742853538" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. &amp; Sangpradub, Narumon" docDate="2021" docId="03DC697BFF82FFD277CAFCA3FE23F847" docLanguage="en" docName="zootaxa.5071.1.5.pdf" docOrigin="Zootaxa 5071 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dineutus unidentatus Aube 1838" docType="treatment" docVersion="7" lastPageNumber="109" masterDocId="FFE51103FF89FFDE775DFFE0FFABFFCF" masterDocTitle="The Gyrinidae (Coleoptera) fauna of Thailand: Key to tribes and genera, with new records and keys to species of Dineutini and Gyrinini" masterLastPageNumber="117" masterPageNumber="97" pageNumber="108" updateTime="1699469841920" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="18B2E408C423A82B13A1CDA95FBB0DA6">The Gyrinidae (Coleoptera) fauna of Thailand: Key to tribes and genera, with new records and keys to species of Dineutini and Gyrinini</mods:title>
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<mods:namePart id="21A12E2930DC979C247C7ADC27B1F49F">Suksai, Benjamart</mods:namePart>
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<mods:nameIdentifier id="81AD6180DA87C2FF74B4E29F492966AE" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AF2F80B1C246B277C916CBF8CFA6F553">Gustafson, Grey T.</mods:namePart>
<mods:affiliation id="F9A2FF88C9AB89B63B72CAC53AB58A97">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. &amp; Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="085A737B832D6D0EF70817CC936E9946">Sites, Robert W.</mods:namePart>
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<mods:affiliation id="8547389752E500042A53BCA6BAA81F1E">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<treatment id="03DC697BFF82FFD277CAFCA3FE23F847" ID-DOI="http://doi.org/10.5281/zenodo.5723075" ID-GBIF-Taxon="190703580" ID-Zenodo-Dep="5723075" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847" httpUri="http://treatment.plazi.org/id/03DC697BFF82FFD277CAFCA3FE23F847" lastPageId="12" lastPageNumber="109" pageId="11" pageNumber="108">
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<taxonomicName id="4C75A3EEFF82FFD577CAFCA3FD84FC93" authority="Aube, 1838" authorityName="Aube" authorityYear="1838" box="[151,559,834,861]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="108" phylum="Arthropoda" rank="species" species="unidentatus">
<emphasis id="B901047FFF82FFD577CAFCA3FD84FC93" bold="true" box="[151,559,834,861]" pageId="11" pageNumber="108">
<emphasis id="B901047FFF82FFD577CAFCA3FE37FC92" bold="true" box="[151,412,834,861]" italics="true" pageId="11" pageNumber="108">Dineutus unidentatus</emphasis>
<bibRefCitation id="EFE4A59CFF82FFD576FEFCA2FD84FC93" author="Aube, C." box="[419,559,834,861]" pageId="11" pageNumber="108" pagination="1 - 804" refId="ref16564" refString="Aube, C. (1838) Hydrocanthares et gyriniens. In: Dejean, P. F. M. A. (Ed.), Species general des coleopteres de la collection de M. le comte Dejean. Mequignon Pere et Fils, Paris, pp. xvi + 1 - 804." type="book chapter" year="1838">Aubé, 1838</bibRefCitation>
</emphasis>
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<paragraph id="8BCAD86DFF82FFD577CAFC87FD92FC4E" blockId="11.[151,569,834,897]" box="[151,569,871,897]" pageId="11" pageNumber="108">
(
<figureCitation id="134EC4E8FF82FFD577C2FC87FEA9FC4E" box="[159,258,871,897]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (C–D) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="11" pageNumber="108">Figs. 1G</figureCitation>
,
<figureCitation id="134EC4E8FF82FFD57652FC87FE9FFC4E" box="[271,308,871,897]" captionStart="FIGURE 4" captionStartId="7.[151,250,1575,1599]" captionTargetBox="[157,1429,181,1551]" captionTargetId="figure-18@7.[157,1429,181,1551]" captionTargetPageId="7" captionText="FIGURE 4. SEM photographs of protochanters in species of Dineutus and Porrorhynchus (males): (A) D. australis, patch of thick setae (arrow), scale bar = 100 μm; (B) D. sitesi, protrochanter without setae (arrow), scale bar = 1 mm; (C) D. spinosus, setae (arrow), scale bar = 100 μm; (D) D. unidentatus, setae (arrow), scale bar = 100 μm; (E–F) P. marginatus, cluster setae (square and arrow): (E) scale bar = 500 μm; (F) scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5723335" httpUri="https://zenodo.org/record/5723335/files/figure.png" pageId="11" pageNumber="108">4D</figureCitation>
,
<figureCitation id="134EC4E8FF82FFD57662FC87FE2CFC4E" box="[319,391,871,897]" captionStart="FIGURE 6" captionStartId="12.[151,250,1219,1243]" captionTargetBox="[177,1411,192,1185]" captionTargetId="figure-22@12.[159,1428,181,1195]" captionTargetPageId="12" captionText="FIGURE 6. Elytral apex, aedeagus and gonocoxae: (A–D) D. unidentatus: (A) male elytral apex, scale bar = 2 mm; (B–C) aedeagus, scale bar = 0.5 mm: (B) dorsal view; (C) lateral view; (D) gonocoxae, ventral view, scale bar = 0.5 mm; (E–G) P. marginatus: (E–F) aedeagus, scale bars = 1 mm: (E) dorsal view; (F) lateral view; (G) gonocoxae, ventral view, scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5723339" httpUri="https://zenodo.org/record/5723339/files/figure.png" pageId="11" pageNumber="108">6A–D</figureCitation>
,
<figureCitation id="134EC4E8FF82FFD576CFFC87FDA8FC4E" box="[402,515,871,897]" captionStart="FIGURE 7" captionStartId="13.[151,250,1836,1860]" captionTargetBox="[193,1395,181,1812]" captionTargetId="figure-18@13.[193,1395,181,1812]" captionTargetPageId="13" captionText="FIGURE 7. Habitats of species of Gyrinini and Dineutini: (A–B) G. smaragdinus: (A) stream at Siriphum Waterfall, Doi Inthanon Natl. Park, Chiang Mai Province; (B) pond at Ban Buak Thoei, Chiang Mai Province; (C) D. australis, D. spinosus and D. unidentatus, pond at Sakaerat Environmental Research Station, Nakhon Ratchasima Province; (D) D. sitesi, Tum Din Stream, Thung Salaeng Luang Natl. Park, Phitsanulok Province; (E) D. spinosus, Huay Sae Stream, Ubon Ratchathani Province; (F–G) D. unidentatus, ponds at Ban Kho, Khon Kaen Province; (H) P. marginatus, Lam Takong Stream, Nakhon Ratchasima Province." figureDoi="http://doi.org/10.5281/zenodo.5723341" httpUri="https://zenodo.org/record/5723341/files/figure.png" pageId="11" pageNumber="108">7C, 7F–G</figureCitation>
,
<figureCitation id="134EC4E8FF82FFD57553FC87FD99FC4E" box="[526,562,871,897]" captionStart="FIGURE 8" captionStartId="14.[151,250,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-22@14.[151,1436,181,1155]" captionTargetPageId="14" captionText="FIGURE 8. (A–B) Distributions of species of Dineutini and Gyrinini in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5723345" httpUri="https://zenodo.org/record/5723345/files/figure.png" pageId="11" pageNumber="108">8A</figureCitation>
)
</paragraph>
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<emphasis id="B901047FFF82FFD577CAFC4EFE2FFC07" bold="true" box="[151,388,942,968]" pageId="11" pageNumber="108">Material examined.</emphasis>
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<collectingCountry id="F36298FDFF82FFD576D3FC4EFD82FC07" box="[398,553,942,968]" name="Thailand" pageId="11" pageNumber="108">THAILAND</collectingCountry>
:
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:
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, leg.
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:
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, wastewater treatment pond 8,
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<collectorName id="2680BDBBFF82FFD573F1FBFBFAB5FBFA" box="[1196,1310,1051,1077]" pageId="11" pageNumber="108">B. Suksai</collectorName>
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,
<specimenCount id="9D7313E4FF82FFD572D3FBFBFF48FB96" pageId="11" pageNumber="108" type="female">1 female</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD577A8FBDFFE93FB96" box="[245,312,1087,1113]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="11" pageNumber="108">KKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD230FF82FFD57613FBDFFBC9FBB2" ID-GBIF-Occurrence="3411799343" collectingDate="1995-02-11" collectionCode="KKU" collectorName="N. Sangpradub" country="Thailand" county="Phong River" elevation="210" latitude="16.868473" location="Si Chomphu Subdistrict" longLatPrecision="1" longitude="102.18283" municipality="Ban Huai Sai Nang" pageId="11" pageNumber="108" specimenCount="14" specimenCount-female="8" specimenCount-male="6" stateProvince="Khon Kaen">
<collectingCounty id="62ABA0E1FF82FFD57613FBDFFE4BFB96" box="[334,480,1087,1113]" pageId="11" pageNumber="108">Phong River</collectingCounty>
,
<collectingMunicipality id="6BAE4217FF82FFD576B3FBDFFD79FB96" box="[494,722,1087,1113]" pageId="11" pageNumber="108">Ban Huai Sai Nang</collectingMunicipality>
,
<location id="8EAA8EB6FF82FFD575BCFBDFFC5DFB96" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD575BCFBDFFC5DFB96" box="[737,1014,1087,1113]" country="Thailand" county="Phong River" latitude="16.868473" longLatPrecision="1" longitude="102.18283" municipality="Ban Huai Sai Nang" name="Si Chomphu Subdistrict" pageId="11" pageNumber="108" stateProvince="Khon Kaen">Si Chomphu Subdistrict</location>
,
<location id="8EAA8EB6FF82FFD57358FBDFFB5EFB96" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD57358FBDFFB5EFB96" box="[1029,1269,1087,1113]" country="Thailand" county="Phong River" latitude="16.868473" longLatPrecision="1" longitude="102.18283" municipality="Ban Huai Sai Nang" name="Si Chomphu District" pageId="11" pageNumber="108" stateProvince="Khon Kaen">Si Chomphu District</location>
,
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<geoCoordinate id="EE41BEAAFF82FFD577CAFB82FE95FBB2" box="[151,318,1122,1149]" degrees="102" direction="east" minutes="10" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="58.2" value="102.18283">102°10'58.2&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57615FB83FE61FBB2" box="[328,458,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="108" unit="m" value="210.0">
<quantity id="4C8D7588FF82FFD57615FB83FE24FBB2" box="[328,399,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="108" unit="m" value="210.0">210 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD5768BFB82FDE7FBB2" box="[470,588,1122,1149]" pageId="11" pageNumber="108" value="1995-02-11">
<collectingDate id="EF8F0745FF82FFD5768BFB82FDE7FBB2" box="[470,588,1122,1149]" pageId="11" pageNumber="108" value="1995-02-11">11.II.1995</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD575DAFB83FC9AFBB2" box="[647,817,1123,1149]" pageId="11" pageNumber="108">N. Sangpradub</collectorName>
,
<specimenCount id="9D7313E4FF82FFD57466FB83FC38FBB2" box="[827,915,1123,1149]" pageId="11" pageNumber="108" type="male">6 males</specimenCount>
,
<specimenCount id="9D7313E4FF82FFD574C3FB83FBA2FBB2" box="[926,1033,1123,1149]" pageId="11" pageNumber="108" type="female">8 females</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD57345FB83FBF0FBB2" box="[1048,1115,1123,1149]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="11" pageNumber="108">KKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD230FF82FFD57330FB83FDF1FB0A" ID-GBIF-Occurrence="3411799409" collectingDate="2016-11-26" collectionCode="KKU, THNHM" collectorName="B. Suksai" country="Thailand" county="Pond" elevation="180" latitude="16.570528" location="Ban Kho Subdistrict" longLatPrecision="1" longitude="102.78162" municipality="Ban Kho" pageId="11" pageNumber="108" specimenCount="113" specimenCount-female="65" specimenCount-male="48" stateProvince="Khon Kaen">
<collectingCounty id="62ABA0E1FF82FFD57330FB83FB0CFBB2" box="[1133,1191,1123,1149]" pageId="11" pageNumber="108">Pond</collectingCounty>
at
<collectingMunicipality id="6BAE4217FF82FFD57395FB83FA85FBB2" box="[1224,1326,1123,1149]" pageId="11" pageNumber="108">Ban Kho</collectingMunicipality>
,
<location id="8EAA8EB6FF82FFD57264FB83FEB8FB6E" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD57264FB83FEB8FB6E" country="Thailand" county="Pond" latitude="16.570528" longLatPrecision="1" longitude="102.78162" municipality="Ban Kho" name="Ban Kho Subdistrict" pageId="11" pageNumber="108" stateProvince="Khon Kaen">Ban Kho Subdistrict</location>
,
<location id="8EAA8EB6FF82FFD5767CFB67FDC2FB6E" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD5767CFB67FDC2FB6E" box="[289,617,1159,1185]" country="Thailand" county="Pond" latitude="16.570528" longLatPrecision="1" longitude="102.78162" municipality="Ban Kho" name="Mueang Khon Kaen District" pageId="11" pageNumber="108" stateProvince="Khon Kaen">Mueang Khon Kaen District</location>
,
<geoCoordinate id="EE41BEAAFF82FFD5752AFB67FCBBFB6E" box="[631,784,1159,1185]" degrees="16" direction="north" minutes="34" orientation="latitude" pageId="11" pageNumber="108" precision="1" seconds="13.9" value="16.570528">16°34'13.9&quot;N</geoCoordinate>
<geoCoordinate id="EE41BEAAFF82FFD57444FB66FC6BFB6E" box="[793,960,1158,1185]" degrees="102" direction="east" minutes="46" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="53.8" value="102.78162">102°46'53.8&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57490FB67FBFDFB6E" box="[973,1110,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="108" unit="m" value="180.0">
<quantity id="4C8D7588FF82FFD57490FB67FBBCFB6E" box="[973,1047,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="108" unit="m" value="180.0">180 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD57338FB67FB4CFB6E" box="[1125,1255,1159,1185]" pageId="11" pageNumber="108" value="2016-11-26">
<collectingDate id="EF8F0745FF82FFD57338FB67FB4CFB6E" box="[1125,1255,1159,1185]" pageId="11" pageNumber="108" value="2016-11-26">26.XI.2016</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD5727AFB67FA33FB6E" box="[1319,1432,1159,1185]" pageId="11" pageNumber="108">B. Suksai</collectorName>
,
<specimenCount id="9D7313E4FF82FFD577CAFB4BFEABFB0A" box="[151,256,1195,1221]" pageId="11" pageNumber="108" type="male">48 males</specimenCount>
,
<specimenCount id="9D7313E4FF82FFD57650FB4AFE23FB0A" box="[269,392,1194,1221]" pageId="11" pageNumber="108" type="female">65 females</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD576C4FB4BFE71FB0A" box="[409,474,1195,1221]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="11" pageNumber="108">KKU</collectionCode>
,
<collectionCode id="ED6440A8FF82FFD576BBFB4BFDF9FB0A" box="[486,594,1195,1221]" pageId="11" pageNumber="108">THNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD230FF82FFD5753BFB4AFE80FAC2" ID-GBIF-Occurrence="3411799396" collectingDate="2005-06-03" collectionCode="UMC" collectorName="R. W. Sites &amp; A. Vitheepradit &amp; T. Prommi" country="Thailand" county="Pond" elevation="27" latitude="8.033334" location="Nuea Khlong Subdistrict" longLatPrecision="1302" longitude="99.01667" municipality="Ban Paga Sai" pageId="11" pageNumber="108" specimenCount="1" specimenCount-male="1" stateProvince="Krabi">
<emphasis id="B901047FFF82FFD5753BFB4AFC83FB0B" bold="true" box="[614,808,1194,1220]" pageId="11" pageNumber="108">
<collectingRegion id="49B1168FFF82FFD5753BFB4AFC8FFB0B" box="[614,804,1194,1220]" country="Thailand" name="Krabi" pageId="11" pageNumber="108">Krabi Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62ABA0E1FF82FFD5746CFB4BFCC0FB0A" box="[817,875,1195,1221]" pageId="11" pageNumber="108">Pond</collectingCounty>
at
<collectingMunicipality id="6BAE4217FF82FFD574CDFB4BFB80FB0A" box="[912,1067,1195,1221]" pageId="11" pageNumber="108">Ban Paga Sai</collectingMunicipality>
,
<location id="8EAA8EB6FF82FFD57365FB4BFAFFFB0A" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD57365FB4BFAFFFB0A" box="[1080,1364,1195,1221]" country="Thailand" county="Pond" latitude="8.033334" longLatPrecision="1302" longitude="99.01667" municipality="Ban Paga Sai" name="Nuea Khlong Subdistrict" pageId="11" pageNumber="108" stateProvince="Krabi">Nuea Khlong Subdistrict</location>
,
<location id="8EAA8EB6FF82FFD5723CFB4BFEE0FB26" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD5723CFB4BFEE0FB26" country="Thailand" county="Pond" latitude="8.033334" longLatPrecision="1302" longitude="99.01667" municipality="Ban Paga Sai" name="Nuea Khlong District" pageId="11" pageNumber="108" stateProvince="Krabi">Nuea Khlong District</location>
,
<geoCoordinate id="EE41BEAAFF82FFD57605FB2FFE0CFB26" box="[344,423,1231,1257]" degrees="8" direction="north" minutes="02" orientation="latitude" pageId="11" pageNumber="108" precision="925" value="8.033334">8°02'N</geoCoordinate>
<geoCoordinate id="EE41BEAAFF82FFD576EDFB2FFDA0FB26" box="[432,523,1231,1257]" degrees="99" direction="east" minutes="01" orientation="longitude" pageId="11" pageNumber="108" precision="925" value="99.01667">99°01'E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57544FB2FFD39FB26" box="[537,658,1231,1257]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="108" unit="m" value="27.0">
<quantity id="4C8D7588FF82FFD57544FB2FFDFFFB26" box="[537,596,1231,1257]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="108" unit="m" value="27.0">27 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD575FCFB2EFCBEFB26" box="[673,789,1230,1257]" pageId="11" pageNumber="108" value="2005-06-03">
<collectingDate id="EF8F0745FF82FFD575FCFB2EFCBEFB26" box="[673,789,1230,1257]" pageId="11" pageNumber="108" value="2005-06-03">3.VI.2005</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD57408FB2FFC64FB26" box="[853,975,1231,1257]" pageId="11" pageNumber="108">R.W. Sites</collectorName>
,
<collectorName id="2680BDBBFF82FFD57486FB2FFB22FB26" box="[987,1161,1231,1257]" pageId="11" pageNumber="108">A. Vitheepradit</collectorName>
&amp;
<collectorName id="2680BDBBFF82FFD573ECFB2FFA86FB26" box="[1201,1325,1231,1257]" pageId="11" pageNumber="108">T. Prommi</collectorName>
, L-828,
<specimenCount id="9D7313E4FF82FFD572D3FB2FFF66FAC2" pageId="11" pageNumber="108" type="male">1 male</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD57783FB13FE8FFAC2" box="[222,292,1267,1293]" pageId="11" pageNumber="108">UMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD230FF82FFD5766AFB13FBF3FAFE" ID-GBIF-Occurrence="3411799315" collectingDate="2017-12-22" collectionCode="THNHM" collectorName="B. Suksai" country="Thailand" county="Huai Lao Waterfall" elevation="226" latitude="16.743584" location="District" longLatPrecision="1" longitude="104.3513" municipality="Ban Kho Subdistrict" pageId="11" pageNumber="108" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mukdahan">
<emphasis id="B901047FFF82FFD5766AFB13FD9DFAC3" bold="true" box="[311,566,1266,1293]" pageId="11" pageNumber="108">
<collectingRegion id="49B1168FFF82FFD5766AFB13FD9AFAC3" box="[311,561,1266,1293]" country="Thailand" name="Mukdahan" pageId="11" pageNumber="108">Mukdahan Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62ABA0E1FF82FFD57563FB13FCB1FAC2" box="[574,794,1267,1293]" pageId="11" pageNumber="108">Huai Lao Waterfall</collectingCounty>
,
<collectingMunicipality id="6BAE4217FF82FFD57478FB13FBA5FAC2" box="[805,1038,1267,1293]" pageId="11" pageNumber="108">Ban Kho Subdistrict</collectingMunicipality>
, Khamcha-I
<location id="8EAA8EB6FF82FFD573FDFB13FB5CFAC2" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD573FDFB13FB5CFAC2" box="[1184,1271,1267,1293]" country="Thailand" county="Huai Lao Waterfall" latitude="16.743584" longLatPrecision="1" longitude="104.3513" municipality="Ban Kho Subdistrict" name="District" pageId="11" pageNumber="108" stateProvince="Mukdahan">District</location>
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<geoCoordinate id="EE41BEAAFF82FFD577CAFAF7FE96FAFE" box="[151,317,1303,1329]" degrees="104" direction="east" minutes="21" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="04.7" value="104.3513">104°21'04.7&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57614FAF7FE67FAFE" box="[329,460,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="2.26" pageId="11" pageNumber="108" unit="m" value="226.0">
<quantity id="4C8D7588FF82FFD57614FAF7FE3BFAFE" box="[329,400,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="2.26" pageId="11" pageNumber="108" unit="m" value="226.0">226 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD57684FAF7FDCFFAFE" box="[473,612,1303,1329]" pageId="11" pageNumber="108" value="2017-12-22">
<collectingDate id="EF8F0745FF82FFD57684FAF7FDCFFAFE" box="[473,612,1303,1329]" pageId="11" pageNumber="108" value="2017-12-22">22.XII.2017</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD575FDFAF7FCA5FAFE" box="[672,782,1303,1329]" pageId="11" pageNumber="108">B. Suksai</collectorName>
,
<specimenCount id="9D7313E4FF82FFD57447FAF7FCC3FAFE" box="[794,872,1303,1329]" pageId="11" pageNumber="108" type="male">1 male</specimenCount>
,
<specimenCount id="9D7313E4FF82FFD5742EFAF7FC7FFAFE" box="[883,980,1303,1329]" pageId="11" pageNumber="108" type="female">1 female</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD574BEFAF7FBFBFAFE" box="[995,1104,1303,1329]" pageId="11" pageNumber="108">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD230FF82FFD5733EFAF7FEF6FAB6" ID-GBIF-Occurrence="3411799363" collectingDate="2017-12-22" collectionCode="THNHM" collectorName="B. Suksai" country="Thailand" county="Tadton Waterfall" elevation="208" latitude="16.494501" location="Nong Sung District" longLatPrecision="1" longitude="104.313225" municipality="Non Yang Subdistrict" pageId="11" pageNumber="108" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Mukdahan">
<collectingCounty id="62ABA0E1FF82FFD5733EFAF7FA89FAFE" box="[1123,1314,1303,1329]" pageId="11" pageNumber="108">Tadton Waterfall</collectingCounty>
,
<collectingMunicipality id="6BAE4217FF82FFD57271FAF7FEB8FA9A" pageId="11" pageNumber="108">Non Yang Subdistrict</collectingMunicipality>
,
<location id="8EAA8EB6FF82FFD57640FADBFE5CFA9A" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD57640FADBFE5CFA9A" box="[285,503,1339,1365]" country="Thailand" county="Tadton Waterfall" latitude="16.494501" longLatPrecision="1" longitude="104.313225" municipality="Non Yang Subdistrict" name="Nong Sung District" pageId="11" pageNumber="108" stateProvince="Mukdahan">Nong Sung District</location>
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<geoCoordinate id="EE41BEAAFF82FFD575C3FADBFCEFFA9A" box="[670,836,1339,1365]" degrees="104" direction="east" minutes="18" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="47.6" value="104.313225">104°18'47.6&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57413FADBFC64FA9A" box="[846,975,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="2.08" pageId="11" pageNumber="108" unit="m" value="208.0">
<quantity id="4C8D7588FF82FFD57413FADBFC3FFA9A" box="[846,916,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="2.08" pageId="11" pageNumber="108" unit="m" value="208.0">208 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD57484FADBFBCFFA9A" box="[985,1124,1339,1365]" pageId="11" pageNumber="108" value="2017-12-22">
<collectingDate id="EF8F0745FF82FFD57484FADBFBCFFA9A" box="[985,1124,1339,1365]" pageId="11" pageNumber="108" value="2017-12-22">22.XII.2017</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD573C0FADBFAA2FA9A" box="[1181,1289,1339,1365]" pageId="11" pageNumber="108">B. Suksai</collectorName>
,
<specimenCount id="9D7313E4FF82FFD5724EFADBFACBFA9A" box="[1299,1376,1339,1365]" pageId="11" pageNumber="108" type="male">1 male</specimenCount>
,
<specimenCount id="9D7313E4FF82FFD57237FADBFF73FAB6" pageId="11" pageNumber="108" type="female">2 females</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD577BAFABFFEFFFAB6" box="[231,340,1375,1401]" pageId="11" pageNumber="108">THNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD230FF82FFD5763AFABFFE8CFA0E" ID-GBIF-Occurrence="3411799418" collectingDate="2013-10-26" collectionCode="KKU" collectorName="B. Suksai" country="Thailand" county="Sakaerat Environmental Research Station" elevation="392" latitude="14.465805" location="Sap Subdistrict" longLatPrecision="1" longitude="101.90325" municipality="Boa" pageId="11" pageNumber="108" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima">
<emphasis id="B901047FFF82FFD5763AFABFFD7FFAB7" bold="true" box="[359,724,1374,1401]" pageId="11" pageNumber="108">
<collectingRegion id="49B1168FFF82FFD5763AFABFFD64FAB7" box="[359,719,1374,1401]" country="Thailand" name="Nakhon Ratchasima" pageId="11" pageNumber="108">Nakhon Ratchasima Province</collectingRegion>
:
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<collectingCounty id="62ABA0E1FF82FFD57586FABFFB04FAB6" box="[731,1199,1375,1401]" pageId="11" pageNumber="108">Sakaerat Environmental Research Station</collectingCounty>
,
<collectingMunicipality id="6BAE4217FF82FFD573E7FABFFB4CFAB6" box="[1210,1255,1375,1401]" pageId="11" pageNumber="108">Boa</collectingMunicipality>
yaepun, U-dom
<location id="8EAA8EB6FF82FFD577CAFA63FEEFFA52" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD577CAFA63FEEFFA52" box="[151,324,1411,1437]" country="Thailand" county="Sakaerat Environmental Research Station" latitude="14.465805" longLatPrecision="1" longitude="101.90325" municipality="Boa" name="Sap Subdistrict" pageId="11" pageNumber="108" stateProvince="Nakhon Ratchasima">Sap Subdistrict</location>
,
<location id="8EAA8EB6FF82FFD57612FA63FDDDFA52" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD57612FA63FDDDFA52" box="[335,630,1411,1437]" country="Thailand" county="Sakaerat Environmental Research Station" latitude="14.465805" longLatPrecision="1" longitude="101.90325" municipality="Boa" name="Wang Nam Khiao District" pageId="11" pageNumber="108" stateProvince="Nakhon Ratchasima">Wang Nam Khiao District</location>
,
<geoCoordinate id="EE41BEAAFF82FFD575DCFA62FCB1FA52" box="[641,794,1410,1437]" degrees="14" direction="north" minutes="27" orientation="latitude" pageId="11" pageNumber="108" precision="1" seconds="56.9" value="14.465805">14°27'56.9&quot;N</geoCoordinate>
<geoCoordinate id="EE41BEAAFF82FFD5747CFA62FC6DFA52" box="[801,966,1410,1437]" degrees="101" direction="east" minutes="54" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="11.7" value="101.90325">101°54'11.7&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD5748CFA63FBFFFA52" box="[977,1108,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="3.92" pageId="11" pageNumber="108" unit="m" value="392.0">
<quantity id="4C8D7588FF82FFD5748CFA63FBB3FA52" box="[977,1048,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="3.92" pageId="11" pageNumber="108" unit="m" value="392.0">392 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD5733CFA63FB72FA52" box="[1121,1241,1411,1437]" pageId="11" pageNumber="108" value="2013-10-26">
<collectingDate id="EF8F0745FF82FFD5733CFA63FB72FA52" box="[1121,1241,1411,1437]" pageId="11" pageNumber="108" value="2013-10-26">26.X.2013</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD57248FA63FA28FA52" box="[1301,1411,1411,1437]" pageId="11" pageNumber="108">B. Suksai</collectorName>
,
<specimenCount id="9D7313E4FF82FFD572D3FA63FF66FA0E" pageId="11" pageNumber="108" type="male">1 male</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD57780FA47FE8BFA0E" box="[221,288,1447,1473]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="11" pageNumber="108">KKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD230FF82FFD5766EFA47FB03FA2A" ID-GBIF-Occurrence="3411799335" collectingDate="2018-03-27" collectionCode="THNHM" collectorName="P. Chanteenok" country="Thailand" county="Bon" elevation="472" latitude="14.497001" location="Wang Nam Khiao District" longLatPrecision="1" longitude="101.91542" municipality="Sap Subdistrict" pageId="11" pageNumber="108" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Nakhon Ratchasima">
same locality except for,
<collectingCounty id="62ABA0E1FF82FFD57510FA47FDD7FA0E" box="[589,636,1447,1473]" pageId="11" pageNumber="108">Bon</collectingCounty>
weir, U-dom
<collectingMunicipality id="6BAE4217FF82FFD57445FA47FC6EFA0E" box="[792,965,1447,1473]" pageId="11" pageNumber="108">Sap Subdistrict</collectingMunicipality>
,
<location id="8EAA8EB6FF82FFD5748DFA47FB53FA0E" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD5748DFA47FB53FA0E" box="[976,1272,1447,1473]" country="Thailand" county="Bon" latitude="14.497001" longLatPrecision="1" longitude="101.91542" municipality="Sap Subdistrict" name="Wang Nam Khiao District" pageId="11" pageNumber="108" stateProvince="Nakhon Ratchasima">Wang Nam Khiao District</location>
,
<geoCoordinate id="EE41BEAAFF82FFD5725EFA47FA37FA0E" box="[1283,1436,1447,1473]" degrees="14" direction="north" minutes="29" orientation="latitude" pageId="11" pageNumber="108" precision="1" seconds="49.2" value="14.497001">14°29'49.2&quot;N</geoCoordinate>
<geoCoordinate id="EE41BEAAFF82FFD577CAFA2AFE96FA2A" box="[151,317,1482,1509]" degrees="101" direction="east" minutes="54" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="55.5" value="101.91542">101°54'55.5&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57617FA2BFE7AFA2A" box="[330,465,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="4.72" pageId="11" pageNumber="108" unit="m" value="472.0">
<quantity id="4C8D7588FF82FFD57617FA2BFE38FA2A" box="[330,403,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="4.72" pageId="11" pageNumber="108" unit="m" value="472.0">472 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD57683FA2BFDF4FA2A" box="[478,607,1483,1509]" pageId="11" pageNumber="108" value="2018-03-27">
<collectingDate id="EF8F0745FF82FFD57683FA2BFDF4FA2A" box="[478,607,1483,1509]" pageId="11" pageNumber="108" value="2018-03-27">27.III.2018</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD575C0FA2BFCE9FA2A" box="[669,834,1483,1509]" pageId="11" pageNumber="108">P. Chanteenok</collectorName>
,
<specimenCount id="9D7313E4FF82FFD57413FA2AFC02FA2A" box="[846,937,1482,1509]" pageId="11" pageNumber="108" type="male">5 males</specimenCount>
,
<specimenCount id="9D7313E4FF82FFD574EBFA2BFB88FA2A" box="[950,1059,1483,1509]" pageId="11" pageNumber="108" type="female">6 females</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD5736EFA2BFB0BFA2A" box="[1075,1184,1483,1509]" pageId="11" pageNumber="108">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD230FF82FFD573E8FA2BFCD6F9C6" ID-GBIF-Occurrence="3411799448" collectingDate="2018-03-29" collectionCode="THNHM" collectorName="P. Chanteenok" country="Thailand" county="Bon" elevation="472" latitude="14.497001" location="Wang Nam Khiao District" longLatPrecision="1" longitude="101.91542" municipality="Sap Subdistrict" pageId="11" pageNumber="108" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Nakhon Ratchasima">
same locality except for,
<date id="FFCBFEADFF82FFD57799FA0FFEEFF9C6" box="[196,324,1519,1545]" pageId="11" pageNumber="108" value="2018-03-29">
<collectingDate id="EF8F0745FF82FFD57799FA0FFEEFF9C6" box="[196,324,1519,1545]" pageId="11" pageNumber="108" value="2018-03-29">29.III.2018</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD57622FA0FFD8AF9C6" box="[383,545,1519,1545]" pageId="11" pageNumber="108">P. Chanteenok</collectorName>
,
<specimenCount id="9D7313E4FF82FFD57576FA0FFD28F9C6" box="[555,643,1519,1545]" pageId="11" pageNumber="108" type="male">4 males</specimenCount>
,
<specimenCount id="9D7313E4FF82FFD575D3FA0FFD52F9C6" box="[654,761,1519,1545]" pageId="11" pageNumber="108" type="female">6 females</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD5745AFA0FFCDFF9C6" box="[775,884,1519,1545]" pageId="11" pageNumber="108">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD230FF82FFD574DAFA0FFC0EF9E2" ID-GBIF-Occurrence="3411799310" collectingDate="2018-04-08" collectionCode="KKU" collectorName="B. Suksai &amp; O. Pornsophon &amp; T. Jantaranakee &amp; S. Phengsalung" country="Thailand" county="Bon" elevation="472" latitude="14.497001" location="Wang Nam Khiao District" longLatPrecision="1" longitude="101.91542" municipality="Sap Subdistrict" pageId="11" pageNumber="108" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima">
same locality except for,
<date id="FFCBFEADFF82FFD573C0FA0FFAA6F9C6" box="[1181,1293,1519,1545]" pageId="11" pageNumber="108" value="2018-04-08">
<collectingDate id="EF8F0745FF82FFD573C0FA0FFAA6F9C6" box="[1181,1293,1519,1545]" pageId="11" pageNumber="108" value="2018-04-08">8.IV.2018</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD5721AFA0FFF12F9E2" pageId="11" pageNumber="108">B. Suksai</collectorName>
,
<collectorName id="2680BDBBFF82FFD57799F9F3FEC5F9E2" box="[196,366,1555,1581]" pageId="11" pageNumber="108">O. Pornsophon</collectorName>
,
<collectorName id="2680BDBBFF82FFD57625F9F3FD8EF9E2" box="[376,549,1555,1581]" pageId="11" pageNumber="108">T. Jantaranakee</collectorName>
&amp;
<collectorName id="2680BDBBFF82FFD5751AF9F3FD5DF9E2" box="[583,758,1555,1581]" pageId="11" pageNumber="108">S. Phengsalung</collectorName>
,
<specimenCount id="9D7313E4FF82FFD5745CF9F3FCE0F9E2" box="[769,843,1555,1581]" pageId="11" pageNumber="108" type="male">1 male</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD57407F9F3FC36F9E2" box="[858,925,1555,1581]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="11" pageNumber="108">KKU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD230FF82FFD574F3F9F3FE5BF9BA" ID-GBIF-Occurrence="3411799367" collectingDate="2017-03-21" collectionCode="THNHM" collectorName="B. Suksai" country="Thailand" county="Nam Churn Stream" elevation="278" latitude="16.674166" location="Nam Nao District" longLatPrecision="1" longitude="101.73416" municipality="Kok Mon Subdistrict" pageId="11" pageNumber="108" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Phetchabun">
<emphasis id="B901047FFF82FFD574F3F9F3FB1CF9E3" bold="true" box="[942,1207,1554,1581]" pageId="11" pageNumber="108">
<collectingRegion id="49B1168FFF82FFD574F3F9F3FB19F9E3" box="[942,1202,1554,1581]" country="Thailand" name="Phetchabun" pageId="11" pageNumber="108">Phetchabun Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62ABA0E1FF82FFD573E0F9F3FA33F9E2" box="[1213,1432,1555,1581]" pageId="11" pageNumber="108">Nam Churn Stream</collectingCounty>
,
<collectingMunicipality id="6BAE4217FF82FFD577CAF9D7FE27F99E" box="[151,396,1591,1617]" pageId="11" pageNumber="108">Kok Mon Subdistrict</collectingMunicipality>
,
<location id="8EAA8EB6FF82FFD576C7F9D7FDC2F99E" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD576C7F9D7FDC2F99E" box="[410,617,1591,1617]" country="Thailand" county="Nam Churn Stream" latitude="16.674166" longLatPrecision="1" longitude="101.73416" municipality="Kok Mon Subdistrict" name="Nam Nao District" pageId="11" pageNumber="108" stateProvince="Phetchabun">Nam Nao District</location>
,
<geoCoordinate id="EE41BEAAFF82FFD5752AF9D7FCBBF99E" box="[631,784,1591,1617]" degrees="16" direction="north" minutes="40" orientation="latitude" pageId="11" pageNumber="108" precision="1" seconds="27.0" value="16.674166">16°40'27.0&quot;N</geoCoordinate>
<geoCoordinate id="EE41BEAAFF82FFD57444F9D7FC6BF99E" box="[793,960,1591,1617]" degrees="101" direction="east" minutes="44" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="03.0" value="101.73416">101°44'03.0&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57493F9D7FBFCF99E" box="[974,1111,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="2.78" pageId="11" pageNumber="108" unit="m" value="278.0">
<quantity id="4C8D7588FF82FFD57493F9D7FBBCF99E" box="[974,1047,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="2.78" pageId="11" pageNumber="108" unit="m" value="278.0">278 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD5733BF9D7FB4DF99E" box="[1126,1254,1591,1617]" pageId="11" pageNumber="108" value="2017-03-21">
<collectingDate id="EF8F0745FF82FFD5733BF9D7FB4DF99E" box="[1126,1254,1591,1617]" pageId="11" pageNumber="108" value="2017-03-21">21.III.2017</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD5727AF9D7FA33F99E" box="[1319,1432,1591,1617]" pageId="11" pageNumber="108">B. Suksai</collectorName>
,
<specimenCount id="9D7313E4FF82FFD577CAF9BBFF5AF9BA" box="[151,241,1627,1653]" pageId="11" pageNumber="108" type="male">2 males</specimenCount>
,
<specimenCount id="9D7313E4FF82FFD577A3F9BBFEC0F9BA" box="[254,363,1627,1653]" pageId="11" pageNumber="108" type="female">3 females</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD57626F9BBFE43F9BA" box="[379,488,1627,1653]" pageId="11" pageNumber="108">THNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD230FF82FFD576A6F9BBFA87F956" ID-GBIF-Occurrence="3411799354" collectingDate="2006-01-09" collectionCode="UMC" collectorName="A. Vitheepradit &amp; T. Prommi" country="Thailand" elevation="5" latitude="7.46925" location="Sikao District" longLatPrecision="1" longitude="99.335976" pageId="11" pageNumber="108" specimenCount="1" specimenCount-female="1" stateProvince="Trang">
<emphasis id="B901047FFF82FFD576A6F9BBFD14F9BB" bold="true" box="[507,703,1626,1653]" pageId="11" pageNumber="108">
<collectingRegion id="49B1168FFF82FFD576A6F9BBFD11F9BB" box="[507,698,1626,1653]" country="Thailand" name="Trang" pageId="11" pageNumber="108">Trang Province</collectingRegion>
:
</emphasis>
<location id="8EAA8EB6FF82FFD5759AF9BBFCCDF9BA" LSID="urn:lsid:plazi:treatment:03DC697BFF82FFD277CAFCA3FE23F847:8EAA8EB6FF82FFD5759AF9BBFCCDF9BA" box="[711,870,1627,1653]" country="Thailand" latitude="7.46925" longLatPrecision="1" longitude="99.335976" name="Sikao District" pageId="11" pageNumber="108" stateProvince="Trang">Sikao District</location>
, ca.
<locationDeviation id="9A78C14FFF82FFD574C7F9BBFAAEF9BA" box="[922,1285,1627,1653]" location="Chao Mai Natl. Park" pageId="11" pageNumber="108">
<quantity id="4C8D7588FF82FFD574C7F9BBFC7FF9BA" box="[922,980,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="108" unit="km" value="8.0">8 km</quantity>
N of Chao Mai Natl. Park
</locationDeviation>
,
<geoCoordinate id="EE41BEAAFF82FFD5724CF9BBFA37F9BA" box="[1297,1436,1627,1653]" degrees="7" direction="north" minutes="28" orientation="latitude" pageId="11" pageNumber="108" precision="1" seconds="09.3" value="7.46925">7°28'09.3&quot;N</geoCoordinate>
<geoCoordinate id="EE41BEAAFF82FFD577CAF99EFE9BF956" box="[151,304,1662,1689]" degrees="99" direction="east" minutes="20" orientation="longitude" pageId="11" pageNumber="108" precision="1" seconds="09.5" value="99.335976">99°20'09.5&quot;E</geoCoordinate>
,
<elevation id="00583F5EFF82FFD57666F99EFE09F956" box="[315,418,1662,1689]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="108" unit="m" value="5.0">
<quantity id="4C8D7588FF82FFD57666F99EFECDF956" box="[315,358,1662,1689]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="108" unit="m" value="5.0">5 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCBFEADFF82FFD576F2F99FFDA5F956" box="[431,526,1663,1689]" pageId="11" pageNumber="108" value="2006-01-09">
<collectingDate id="EF8F0745FF82FFD576F2F99FFDA5F956" box="[431,526,1663,1689]" pageId="11" pageNumber="108" value="2006-01-09">9.I.2006</collectingDate>
</date>
, leg.
<collectorName id="2680BDBBFF82FFD57514F99FFD5EF956" box="[585,757,1663,1689]" pageId="11" pageNumber="108">A. Vitheepradit</collectorName>
&amp;
<collectorName id="2680BDBBFF82FFD57445F99FFC39F956" box="[792,914,1663,1689]" pageId="11" pageNumber="108">T. Prommi</collectorName>
, mangrove, L-906,
<specimenCount id="9D7313E4FF82FFD57333F99FFB64F956" box="[1134,1231,1663,1689]" pageId="11" pageNumber="108" type="female">1 female</specimenCount>
(
<collectionCode id="ED6440A8FF82FFD57383F99FFA8FF956" box="[1246,1316,1663,1689]" pageId="11" pageNumber="108">UMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8BE6FF82FFD2779AF942FBB8FA7B" lastPageId="12" lastPageNumber="109" pageId="11" pageNumber="108" type="diagnosis">
<paragraph id="8BCAD86DFF82FFD5779AF942FBC8F8BE" blockId="11.[151,1437,942,1978]" pageId="11" pageNumber="108">
<emphasis id="B901047FFF82FFD5779AF942FEE9F973" bold="true" box="[199,322,1698,1724]" pageId="11" pageNumber="108">Diagnosis.</emphasis>
Male (
<figureCitation id="134EC4E8FF82FFD576CEF943FE46F972" box="[403,493,1699,1725]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (C–D) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="11" pageNumber="108">Fig. 1G</figureCitation>
): Size: 6.0–
<quantity id="4C8D7588FF82FFD5752BF943FD64F972" box="[630,719,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="11" pageNumber="108" unit="mm" value="6.7">6.7 mm</quantity>
. Body form narrowly oval; apex of flagellomere VI darkly colored; elytral apices truncate and straight without parasutural spines, with epipleural spines and serrations (
<figureCitation id="134EC4E8FF82FFD57239F927FF10F8CA" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (C–D) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="11" pageNumber="108">Figs. 1G</figureCitation>
,
<figureCitation id="134EC4E8FF82FFD57795F90BFF45F8C9" box="[200,238,1771,1798]" captionStart="FIGURE 6" captionStartId="12.[151,250,1219,1243]" captionTargetBox="[177,1411,192,1185]" captionTargetId="figure-22@12.[159,1428,181,1195]" captionTargetPageId="12" captionText="FIGURE 6. Elytral apex, aedeagus and gonocoxae: (A–D) D. unidentatus: (A) male elytral apex, scale bar = 2 mm; (B–C) aedeagus, scale bar = 0.5 mm: (B) dorsal view; (C) lateral view; (D) gonocoxae, ventral view, scale bar = 0.5 mm; (E–G) P. marginatus: (E–F) aedeagus, scale bars = 1 mm: (E) dorsal view; (F) lateral view; (G) gonocoxae, ventral view, scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5723339" httpUri="https://zenodo.org/record/5723339/files/figure.png" pageId="11" pageNumber="108">6A</figureCitation>
); lateral marginal of pronotum and elytron black; protrochanter apically with single series of few setae on ventral surface (
<figureCitation id="134EC4E8FF82FFD57616F8EFFE0DF8E6" box="[331,422,1807,1833]" captionStart="FIGURE 4" captionStartId="7.[151,250,1575,1599]" captionTargetBox="[157,1429,181,1551]" captionTargetId="figure-18@7.[157,1429,181,1551]" captionTargetPageId="7" captionText="FIGURE 4. SEM photographs of protochanters in species of Dineutus and Porrorhynchus (males): (A) D. australis, patch of thick setae (arrow), scale bar = 100 μm; (B) D. sitesi, protrochanter without setae (arrow), scale bar = 1 mm; (C) D. spinosus, setae (arrow), scale bar = 100 μm; (D) D. unidentatus, setae (arrow), scale bar = 100 μm; (E–F) P. marginatus, cluster setae (square and arrow): (E) scale bar = 500 μm; (F) scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5723335" httpUri="https://zenodo.org/record/5723335/files/figure.png" pageId="11" pageNumber="108">Fig. 4D</figureCitation>
); profemur without sub-apicoventral tooth; protibia wedge-shaped; venter, brown to black; epipleura brown; aedeagus (
<figureCitation id="134EC4E8FF82FFD5768CF8D3FDF8F881" box="[465,595,1843,1870]" captionStart="FIGURE 6" captionStartId="12.[151,250,1219,1243]" captionTargetBox="[177,1411,192,1185]" captionTargetId="figure-22@12.[159,1428,181,1195]" captionTargetPageId="12" captionText="FIGURE 6. Elytral apex, aedeagus and gonocoxae: (A–D) D. unidentatus: (A) male elytral apex, scale bar = 2 mm; (B–C) aedeagus, scale bar = 0.5 mm: (B) dorsal view; (C) lateral view; (D) gonocoxae, ventral view, scale bar = 0.5 mm; (E–G) P. marginatus: (E–F) aedeagus, scale bars = 1 mm: (E) dorsal view; (F) lateral view; (G) gonocoxae, ventral view, scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5723339" httpUri="https://zenodo.org/record/5723339/files/figure.png" pageId="11" pageNumber="108">Figs. 6B–C</figureCitation>
) with median lobe slightly longer than parameres, parallel-sided after basal fourth, strongly acuminate in apical 1/6, apex with tip rounded, straight in lateral view.
</paragraph>
<paragraph id="8BCAD86DFF82FFD5779AF89BFCAEF875" blockId="11.[151,1437,942,1978]" pageId="11" pageNumber="108">
Female: Similar to male except for size:
<quantity id="4C8D7588FF82FFD575DFF89BFCA1F85A" box="[642,778,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.9" metricValueMin="6.4" pageId="11" pageNumber="108" unit="mm" value="6.65" valueMax="6.9" valueMin="6.4">6.4–6.9 mm</quantity>
; gonocoxae elongate, lateral margin rounded, apex obliquely truncate with apicolateral corner produced (
<figureCitation id="134EC4E8FF82FFD575C3F87FFD53F875" box="[670,760,1951,1978]" captionStart="FIGURE 6" captionStartId="12.[151,250,1219,1243]" captionTargetBox="[177,1411,192,1185]" captionTargetId="figure-22@12.[159,1428,181,1195]" captionTargetPageId="12" captionText="FIGURE 6. Elytral apex, aedeagus and gonocoxae: (A–D) D. unidentatus: (A) male elytral apex, scale bar = 2 mm; (B–C) aedeagus, scale bar = 0.5 mm: (B) dorsal view; (C) lateral view; (D) gonocoxae, ventral view, scale bar = 0.5 mm; (E–G) P. marginatus: (E–F) aedeagus, scale bars = 1 mm: (E) dorsal view; (F) lateral view; (G) gonocoxae, ventral view, scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5723339" httpUri="https://zenodo.org/record/5723339/files/figure.png" pageId="11" pageNumber="108">Fig. 6D</figureCitation>
).
</paragraph>
<caption id="DF0A88E5FF85FFD277CAFB23FF6DFA88" ID-DOI="http://doi.org/10.5281/zenodo.5723339" ID-Zenodo-Dep="5723339" httpUri="https://zenodo.org/record/5723339/files/figure.png" pageId="12" pageNumber="109" startId="12.[151,250,1219,1243]" targetBox="[177,1411,192,1185]" targetPageId="12">
<paragraph id="8BCAD86DFF85FFD277CAFB23FF6DFA88" blockId="12.[151,1437,1219,1351]" pageId="12" pageNumber="109">
<emphasis id="B901047FFF85FFD277CAFB23FEBEFB14" bold="true" box="[151,277,1219,1243]" pageId="12" pageNumber="109">FIGURE 6.</emphasis>
Elytral apex, aedeagus and gonocoxae: (A–D)
<taxonomicName id="4C75A3EEFF85FFD2745EFB24FC31FB14" box="[771,922,1219,1243]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="109" phylum="Arthropoda" rank="species" species="unidentatus">
<emphasis id="B901047FFF85FFD2745EFB24FC31FB14" box="[771,922,1219,1243]" italics="true" pageId="12" pageNumber="109">D. unidentatus</emphasis>
</taxonomicName>
: (A) male elytral apex, scale bar = 2 mm; (B–C) aedeagus, scale bar = 0.5 mm: (B) dorsal view; (C) lateral view; (D) gonocoxae, ventral view, scale bar = 0.5 mm; (E–G)
<taxonomicName id="4C75A3EEFF85FFD272D7FB08FEA2FAEC" baseAuthorityName="Laporte" baseAuthorityYear="1835" class="Insecta" family="Gyrinidae" genus="Porrorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="109" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B901047FFF85FFD272D7FB08FEA2FAEC" italics="true" pageId="12" pageNumber="109">P. marginatus</emphasis>
</taxonomicName>
: (E–F) aedeagus, scale bars = 1 mm: (E) dorsal view; (F) lateral view; (G) gonocoxae, ventral view, scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BCAD86DFF85FFD2779AFA95FBB8FA7B" blockId="12.[151,1437,1397,1928]" pageId="12" pageNumber="109">
<emphasis id="B901047FFF85FFD2779AFA95FE65FA40" bold="true" box="[199,462,1397,1423]" pageId="12" pageNumber="109">Differential diagnosis.</emphasis>
<taxonomicName id="4C75A3EEFF85FFD27688FA96FD6EFA5F" box="[469,709,1398,1424]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="109" phylum="Arthropoda" rank="species" species="unidentatus">
<emphasis id="B901047FFF85FFD27688FA96FD6EFA5F" box="[469,709,1398,1424]" italics="true" pageId="12" pageNumber="109">Dineutus unidentatus</emphasis>
</taxonomicName>
is closest to
<emphasis id="B901047FFF85FFD2740AFA96FC48FA5F" box="[855,995,1398,1424]" italics="true" pageId="12" pageNumber="109">
<taxonomicName id="4C75A3EEFF85FFD2740AFA96FC74FA5F" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[855,991,1398,1424]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="109" phylum="Arthropoda" rank="species" species="australis">D. australis</taxonomicName>
,
</emphasis>
which also occurs in
<collectingCountry id="F36298FDFF85FFD27385FA96FA94FA5F" box="[1240,1343,1398,1424]" name="Thailand" pageId="12" pageNumber="109">Thailand</collectingCountry>
. See the differential diagnosis under
<taxonomicName id="4C75A3EEFF85FFD2768FFA7AFDFDFA7B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[466,598,1434,1460]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="109" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B901047FFF85FFD2768FFA7AFDFDFA7B" box="[466,598,1434,1460]" italics="true" pageId="12" pageNumber="109">D. australis</emphasis>
</taxonomicName>
to distinguish between the two species.
</paragraph>
</subSubSection>
<subSubSection id="C36F8BE6FF85FFD2779AFA5DFE78F937" pageId="12" pageNumber="109" type="distribution">
<paragraph id="8BCAD86DFF85FFD2779AFA5DFB9DF9A7" blockId="12.[151,1437,1397,1928]" pageId="12" pageNumber="109">
<emphasis id="B901047FFF85FFD2779AFA5DFECAFA18" bold="true" box="[199,353,1469,1495]" pageId="12" pageNumber="109">Distribution.</emphasis>
Widely distributed in the Indian subcontinent and Southeast Asia from western
<collectingCountry id="F36298FDFF85FFD27258FA5EFA94FA17" box="[1285,1343,1470,1496]" name="India" pageId="12" pageNumber="109">India</collectingCountry>
and
<collectingCountry id="F36298FDFF85FFD27221FA5EFF76FA33" name="Sri Lanka" pageId="12" pageNumber="109">Sri Lanka</collectingCountry>
through Assam,
<collectingCountry id="F36298FDFF85FFD276F9FA02FD81FA33" box="[420,554,1506,1532]" name="Bangladesh" pageId="12" pageNumber="109">Bangladesh</collectingCountry>
,
<collectingCountry id="F36298FDFF85FFD27565FA02FD0CFA33" box="[568,679,1506,1532]" name="Myanmar" pageId="12" pageNumber="109">Myanmar</collectingCountry>
,
<collectingCountry id="F36298FDFF85FFD275E8FA02FCB0FA33" box="[693,795,1506,1532]" name="Thailand" pageId="12" pageNumber="109">Thailand</collectingCountry>
,
<collectingCountry id="F36298FDFF85FFD27474FA02FC34FA33" box="[809,927,1506,1532]" name="Cambodia" pageId="12" pageNumber="109">Cambodia</collectingCountry>
,
<collectingCountry id="F36298FDFF85FFD274F0FA02FC4DFA33" box="[941,998,1506,1532]" name="Laos" pageId="12" pageNumber="109">Laos</collectingCountry>
, to
<collectingCountry id="F36298FDFF85FFD27349FA02FBD8FA33" box="[1044,1139,1506,1532]" name="Vietnam" pageId="12" pageNumber="109">Vietnam</collectingCountry>
and Hainan Island (
<bibRefCitation id="EFE4A59CFF85FFD2723CFA02FF79F9EF" author="Ochs" pageId="12" pageNumber="109" pagination="401 - 403" refId="ref18207" refString="Ochs, G. (1929) Papers on Malayan aquatic biology. X. Additional notes on the Gyrinidae. Journal of the Federated Malay States Museums, 14 (3 - 4), 401 - 403." type="journal article" year="1929">Ochs 1929</bibRefCitation>
,
<bibRefCitation id="EFE4A59CFF85FFD27782F9E6FEB1F9EF" author="Ochs" box="[223,282,1542,1568]" pageId="12" pageNumber="109" refId="ref18247" refString="Ochs, G. (1930) Catalogue of Indian insects. Part 19. Gyrinoidea. Government of India Central Publications Branch, Calcutta, 39 pp." type="book" year="1930">1930</bibRefCitation>
;
<bibRefCitation id="EFE4A59CFF85FFD2767AF9E6FE5EF9EF" author="Mouchamps, R." box="[295,501,1542,1568]" pageId="12" pageNumber="109" pagination="216 - 264" refId="ref18072" refString="Mouchamps, R. (1949) Contribution a la systematique des coleopteres gyrinides I. Dineutus (Spinosodineutes) Hatch. Bulletin &amp; Annales de la Societe Entomologique de Belgique, 85, 216 - 264." type="journal article" year="1949">Mouchamps 1949</bibRefCitation>
;
<bibRefCitation id="EFE4A59CFF85FFD2755FF9E6FD39F9EF" author="Brinck, P." box="[514,658,1542,1568]" pageId="12" pageNumber="109" pagination="17 - 32" refId="ref16729" refString="Brinck, P. (1981) Taxonomic status and evolutionary trends in the whirligig beetle taxon Spinosodineutes (Coleoptera: Gyrinidae). Entomologia Generalis, 7 (1), 17 - 32. https: // doi. org / 10.1127 / entom. gen / 7 / 1981 / 17" type="journal article" year="1981">Brinck 1981</bibRefCitation>
;
<bibRefCitation id="EFE4A59CFF85FFD275C2F9E6FCEBF9EF" author="Vazirani, T. G." box="[671,832,1542,1568]" pageId="12" pageNumber="109" refId="ref19005" refString="Vazirani, T. G. (1984) The fauna of India. Coleoptera. Family Gyrinidae and family Haliplidae. Zoological Survey of India, Calcutta, 138 pp." type="book" year="1984">Vazirani 1984</bibRefCitation>
;
<bibRefCitation id="EFE4A59CFF85FFD27410F9E6FC52F9EF" author="Mazzoldi, P." box="[845,1017,1541,1568]" pageId="12" pageNumber="109" refId="ref17864" refString="Mazzoldi, P. (1995) Gyrinidae: Catalogue of Chinese Gyrinidae (Coleoptera). In: J ach, M. A. &amp; Ji, L. (Eds.), Water Beetles of China. Vol. 1. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Vienna, pp. 155 - 172." type="book" year="1995">Mazzoldi 1995</bibRefCitation>
) as far south as the Malay peninsula (
<bibRefCitation id="EFE4A59CFF85FFD277C2F9CAFEC8F98B" author="Balke, M. &amp; Jach, M. A. &amp; Hendrich, L." box="[159,355,1578,1604]" pageId="12" pageNumber="109" pagination="555 - 609" refId="ref16621" refString="Balke, M., Jach, M. A. &amp; Hendrich, L. (2004) Insecta: Coleoptera. In: Yule, C. M. &amp; Yong, H. S. (Eds.), Freshwater Invertebrates of the Malaysian Region. Academy of Sciences Malaysia, Kuala Lumpur, pp. 555 - 609." type="book chapter" year="2004">
Balke
<emphasis id="B901047FFF85FFD277B4F9CAFEB0F98B" box="[233,283,1578,1604]" italics="true" pageId="12" pageNumber="109">et al</emphasis>
. 2004
</bibRefCitation>
) and Perak Island (
<bibRefCitation id="EFE4A59CFF85FFD27560F9CAFD17F98B" author="Ochs" box="[573,700,1578,1604]" pageId="12" pageNumber="109" pagination="401 - 403" refId="ref18207" refString="Ochs, G. (1929) Papers on Malayan aquatic biology. X. Additional notes on the Gyrinidae. Journal of the Federated Malay States Museums, 14 (3 - 4), 401 - 403." type="journal article" year="1929">Ochs 1929</bibRefCitation>
). The species has not been recorded from
<collectingCountry id="F36298FDFF85FFD273CBF9CAFAA3F98B" box="[1174,1288,1578,1604]" name="Singapore" pageId="12" pageNumber="109">Singapore</collectingCountry>
(
<bibRefCitation id="EFE4A59CFF85FFD2724BF9CAFF5EF9A7" author="Hendrich, L. &amp; Balke, M. &amp; Yang, C. M." pageId="12" pageNumber="109" pagination="97 - 145" refId="ref17504" refString="Hendrich, L., Balke, M. &amp; Yang, C. M. (2004) Aquatic Coleoptera of Singapore: Species richness, ecology and conservation. The Raffles Bulletin of Zoology, 52, 97 - 145." type="journal article" year="2004">
Hendrich
<emphasis id="B901047FFF85FFD272D5F9CAFF06F9A7" italics="true" pageId="12" pageNumber="109">et al</emphasis>
. 2004
</bibRefCitation>
) and historical records from Java are considered erroneous (
<bibRefCitation id="EFE4A59CFF85FFD274C5F9AEFB82F9A7" author="Brinck, P." box="[920,1065,1614,1640]" pageId="12" pageNumber="109" pagination="17 - 32" refId="ref16729" refString="Brinck, P. (1981) Taxonomic status and evolutionary trends in the whirligig beetle taxon Spinosodineutes (Coleoptera: Gyrinidae). Entomologia Generalis, 7 (1), 17 - 32. https: // doi. org / 10.1127 / entom. gen / 7 / 1981 / 17" type="journal article" year="1981">Brinck 1981</bibRefCitation>
).
</paragraph>
<paragraph id="8BCAD86DFF85FFD2779AF991FE78F937" blockId="12.[151,1437,1397,1928]" pageId="12" pageNumber="109">
<emphasis id="B901047FFF85FFD2779AF991FE81F944" bold="true" box="[199,298,1649,1675]" pageId="12" pageNumber="109">Habitat.</emphasis>
In
<collectingCountry id="F36298FDFF85FFD2760EF992FE6EF943" box="[339,453,1650,1676]" name="Sri Lanka" pageId="12" pageNumber="109">Sri Lanka</collectingCountry>
, the species has been found in rivers, streams, ponds, water tanks, paddy fields, and a slightly brackish lagoon (
<bibRefCitation id="EFE4A59CFF85FFD276F2F976FD95F97F" author="Brinck, P." box="[431,574,1686,1712]" pageId="12" pageNumber="109" pagination="17 - 32" refId="ref16729" refString="Brinck, P. (1981) Taxonomic status and evolutionary trends in the whirligig beetle taxon Spinosodineutes (Coleoptera: Gyrinidae). Entomologia Generalis, 7 (1), 17 - 32. https: // doi. org / 10.1127 / entom. gen / 7 / 1981 / 17" type="journal article" year="1981">Brinck 1981</bibRefCitation>
). On
<collectingRegion id="49B1168FFF85FFD27525F976FD13F97F" box="[632,696,1686,1712]" country="Malaysia" name="Perak" pageId="12" pageNumber="109">Perak</collectingRegion>
Island,
<collectingCountry id="F36298FDFF85FFD27453F976FCDCF97F" box="[782,887,1686,1712]" name="Malaysia" pageId="12" pageNumber="109">Malaysia</collectingCountry>
, the species was taken near a waterfall in a stream (
<bibRefCitation id="EFE4A59CFF85FFD277C2F95AFEB7F91B" author="Ochs" box="[159,284,1722,1748]" pageId="12" pageNumber="109" pagination="401 - 403" refId="ref18207" refString="Ochs, G. (1929) Papers on Malayan aquatic biology. X. Additional notes on the Gyrinidae. Journal of the Federated Malay States Museums, 14 (3 - 4), 401 - 403." type="journal article" year="1929">Ochs 1929</bibRefCitation>
). We found this species near the margins of a stagnant stream and in ponds where it formed rafts of many individuals (
<figureCitation id="134EC4E8FF85FFD2767EF93EFE63F937" box="[291,456,1758,1784]" captionStart="FIGURE 7" captionStartId="13.[151,250,1836,1860]" captionTargetBox="[193,1395,181,1812]" captionTargetId="figure-18@13.[193,1395,181,1812]" captionTargetPageId="13" captionText="FIGURE 7. Habitats of species of Gyrinini and Dineutini: (A–B) G. smaragdinus: (A) stream at Siriphum Waterfall, Doi Inthanon Natl. Park, Chiang Mai Province; (B) pond at Ban Buak Thoei, Chiang Mai Province; (C) D. australis, D. spinosus and D. unidentatus, pond at Sakaerat Environmental Research Station, Nakhon Ratchasima Province; (D) D. sitesi, Tum Din Stream, Thung Salaeng Luang Natl. Park, Phitsanulok Province; (E) D. spinosus, Huay Sae Stream, Ubon Ratchathani Province; (F–G) D. unidentatus, ponds at Ban Kho, Khon Kaen Province; (H) P. marginatus, Lam Takong Stream, Nakhon Ratchasima Province." figureDoi="http://doi.org/10.5281/zenodo.5723341" httpUri="https://zenodo.org/record/5723341/files/figure.png" pageId="12" pageNumber="109">Figs. 7C, F–G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F8BE6FF85FFD2779AF8E1FE23F847" pageId="12" pageNumber="109" type="discussion">
<paragraph id="8BCAD86DFF85FFD2779AF8E1FE23F847" blockId="12.[151,1437,1397,1928]" pageId="12" pageNumber="109">
<emphasis id="B901047FFF85FFD2779AF8E1FEE7F8D4" bold="true" box="[199,332,1793,1819]" pageId="12" pageNumber="109">Discussion.</emphasis>
We found
<taxonomicName id="4C75A3EEFF85FFD27696F8E2FDD9F8D3" box="[459,626,1794,1820]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="109" phylum="Arthropoda" rank="species" species="unidentatus">
<emphasis id="B901047FFF85FFD27696F8E2FDD9F8D3" box="[459,626,1794,1820]" italics="true" pageId="12" pageNumber="109">D. unidentatus</emphasis>
</taxonomicName>
only in the northeastern and southern regions of
<collectingCountry id="F36298FDFF85FFD273FBF8E2FAA1F8D3" box="[1190,1290,1794,1820]" name="Thailand" pageId="12" pageNumber="109">Thailand</collectingCountry>
(
<figureCitation id="134EC4E8FF85FFD27247F8E2FADDF8D3" box="[1306,1398,1794,1820]" captionStart="FIGURE 8" captionStartId="14.[151,250,1179,1203]" captionTargetBox="[151,1436,181,1155]" captionTargetId="figure-22@14.[151,1436,181,1155]" captionTargetPageId="14" captionText="FIGURE 8. (A–B) Distributions of species of Dineutini and Gyrinini in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5723345" httpUri="https://zenodo.org/record/5723345/files/figure.png" pageId="12" pageNumber="109">Fig. 8A</figureCitation>
). It was found at low to medium elevation (
<quantity id="4C8D7588FF85FFD27504F8C5FD15F88F" box="[601,702,1829,1856]" metricMagnitude="2" metricUnit="m" metricValue="2.385" metricValueMax="4.72" metricValueMin="0.05" pageId="12" pageNumber="109" unit="m" value="238.5" valueMax="472.0" valueMin="5.0">5–472 m</quantity>
a.s.l.) and co-occurred in ponds and streams with
<taxonomicName id="4C75A3EEFF85FFD273A4F8C6FAD7F8F0" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[1273,1404,1830,1855]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="109" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B901047FFF85FFD273A4F8C6FAD7F8F0" box="[1273,1404,1830,1855]" italics="true" pageId="12" pageNumber="109">D. spinosus</emphasis>
</taxonomicName>
. It has not been found at high elevation. We suspect that it might be found in other localities in
<collectingCountry id="F36298FDFF85FFD273CAF8AAFB50F8AB" box="[1175,1275,1866,1892]" name="Thailand" pageId="12" pageNumber="109">Thailand</collectingCountry>
with the same habitat and elevation.
</paragraph>
</subSubSection>
</treatment>
</document>
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