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<mods:title id="13BBBC9B37360C337163C3C88BE1BFF5">Preliminary re-examination of genus-level taxonomy of the pollen beetle subfamily Meligethinae (Coleoptera: Nitidulidae)</mods:title>
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<mods:namePart id="507D1DED70CC315FCA918EC4716C0E03">Cline, Andrew Richard</mods:namePart>
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1.
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<emphasis id="B963EAC8F66CFF82BAB2FB7AFDE9FB1C" bold="true" box="[441,604,1166,1191]" italics="true" pageId="6" pageNumber="347">Acanthogethes</emphasis>
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stat. nov.
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(
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)
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<taxonomicName id="4C174D59F66CFF82BB9CFB25FE01FB58" ID-CoL="M7N" authority="Reitter, 1871: 155" authorityName="Reitter" authorityPageNumber="155" authorityYear="1871" box="[151,436,1232,1251]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F66CFF82BB9CFB25FEA0FB58" box="[151,277,1233,1251]" italics="true" pageId="6" pageNumber="347">Acanthogethes</emphasis>
<bibRefCitation id="EF864B2BF66CFF82BA11FB24FE01FB58" author="REITTER E." box="[282,436,1232,1251]" pageId="6" pageNumber="347" pagination="39 - 169" refId="ref91274" refString="REITTER E. 1871: Revision der europaischen Meligethes-Arten. Verhandlungen des Naturforschenden Vereines in Brunn 9: 39 - 169." type="journal article" year="1871">Reitter, 1871: 155</bibRefCitation>
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(described as a subgenus of
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<emphasis id="B963EAC8F66CFF82B9A2FB25FCB2FB58" box="[681,775,1233,1251]" italics="true" pageId="6" pageNumber="347">Meligethes</emphasis>
<bibRefCitation id="EF864B2BF66CFF82B807FB24FC27FB58" author="STEPHENS J. F." box="[780,914,1232,1251]" pageId="6" pageNumber="347" pagination="16 - 19" refId="ref92184" refString="STEPHENS J. F. 1830: Illustrations of British Entomology; or, a Synopsis of Indigenous Insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, vol. III. Baldwin and Craddock, London, 379 pp, pls. 16 - 19." type="book chapter" year="1830">Stephens, 1830</bibRefCitation>
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).
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species.
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<taxonomicName id="4C174D59F66CFF82BA30FB09FC9AFAAF" authority="A. G. Olivier, 1790: 10" authorityName="A. G. Olivier" authorityPageNumber="10" authorityYear="1790" box="[315,815,1276,1300]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Fungi" order="Coleoptera" pageId="6" pageNumber="347" phylum="Ascomycota" rank="species" species="fuscum">
<emphasis id="B963EAC8F66CFF82BA30FB09FDAEFAAF" box="[315,539,1277,1300]" italics="true" pageId="6" pageNumber="347">Sphaeridium fuscum</emphasis>
A. G.
<bibRefCitation id="EF864B2BF66CFF82B964FB08FC9AFAAF" author="OLIVIER A. G." box="[623,815,1276,1300]" pageId="6" pageNumber="347" pagination="9 - 34" refId="ref91007" refString="OLIVIER A. G. 1790: Entomologie, ou Histoire Naturelle des Insectes, avec leur caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enlumiee. Coleopteres, Tome 2. Baudouin, Paris, 9 - 34 + xxii pp." type="book chapter" year="1790">Olivier, 1790: 10</bibRefCitation>
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(subsequent designation by
<bibRefCitation id="EF864B2BF66CFF82BB9CFAE8FEF2FA8F" author="PARSONS C. T." box="[151,327,1308,1332]" pageId="6" pageNumber="347" pagination="121 - 278" refId="ref91056" refString="PARSONS C. T. 1943: A revision of Nearctic Nitidulidae (Coleoptera). Bulletin of the Museum of Comparative Zoology 92: 121 - 278." type="journal article" year="1943">PARSONS (1943))</bibRefCitation>
[=
<taxonomicName id="4C174D59F66CFF82BA67FAE9FC8CFA8F" authority="(A. G. Olivier, 1790)" baseAuthorityName="A. G. Olivier" baseAuthorityYear="1790" box="[364,825,1308,1332]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="fuscus" status="comb. nov.">
<emphasis id="B963EAC8F66CFF82BA67FAE9FDE7FA8F" box="[364,594,1309,1332]" italics="true" pageId="6" pageNumber="347">Acanthogethes fuscus</emphasis>
(A. G.
<bibRefCitation id="EF864B2BF66CFF82B9A8FAE8FC84FA8F" author="OLIVIER A. G." box="[675,817,1308,1332]" pageId="6" pageNumber="347" pagination="9 - 34" refId="ref91007" refString="OLIVIER A. G. 1790: Entomologie, ou Histoire Naturelle des Insectes, avec leur caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enlumiee. Coleopteres, Tome 2. Baudouin, Paris, 9 - 34 + xxii pp." type="book chapter" year="1790">Olivier, 1790</bibRefCitation>
)
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<taxonomicNameLabel id="A25057B3F66CFF82B834FAE8FC0DFA88" box="[831,952,1308,1332]" pageId="6" pageNumber="347" rank="species">comb. nov.</taxonomicNameLabel>
]
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<emphasis id="B963EAC8F66CFF82BB9CFABEFD8CFAD9" bold="true" box="[151,569,1354,1379]" pageId="6" pageNumber="347">Generic redescription and diagnosis.</emphasis>
Inclusive species range greatly in size (
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1.2
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4.3 mm
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length), and share the following combination of characters.
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<emphasis id="B963EAC8F66CFF82BBBEFA78FE53FA18" box="[181,486,1420,1443]" italics="true" pageId="6" pageNumber="347">Body color and pubescence</emphasis>
: pubescence usually short and fine, recumbent, golden to silvery-whitish, never obscuring the variably colored (brown, blackish, reddish, or blackish with orange spots on elytra, e.g.
<figureCitation id="132C2A5FF66CFF82BAF9FA3FFD8EFA58" box="[498,571,1483,1507]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="6" pageNumber="347">Fig. 1a</figureCitation>
) dorsal body surface; pronotal and elytral sides narrowly flattened, typically same color as disc, a few species with pale reddish sides. Lateral margin of pronutum and elytra with a series of faintly distinct, small and short setae, each seta 0.3
<emphasis id="B963EAC8F66DFF83BBCAFF28FF7BFF4F" bold="true" box="[193,206,220,244]" pageId="7" pageNumber="348"></emphasis>
0.5× as long as those on elytral disc; posterior margin of pronotum with long, usually distally bifid or trifid microsetae, microsetae uniformly distributed in middle region anterior to scutellum (
<figureCitation id="132C2A5FF66DFF83BA08FEEFFEE7FE88" box="[259,338,283,307]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1d</figureCitation>
).
</paragraph>
<paragraph id="8BA836DAF66DFF83BB9BFEC9FCEEFC88" blockId="7.[113,1089,219,1620]" pageId="7" pageNumber="348">
<emphasis id="B963EAC8F66DFF83BB9BFEC9FE85FEEF" box="[144,304,317,340]" italics="true" pageId="7" pageNumber="348">Dorsal habitus</emphasis>
: body markedly convex, short, wide and oval (
<figureCitation id="132C2A5FF66DFF83B826FECFFCCCFEEF" box="[813,889,315,340]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1a</figureCitation>
); dorsal punctures on discal portion of pronotum as large as, or larger than eye facets, usually deeply impressed and densely distributed; anterior margin of clypeus always strongly arcuately emarginate, simple, i.e. without small distinct medial bulge, and distinctly bordered (
<figureCitation id="132C2A5FF66DFF83B867FE6FFC6FFE0F" box="[876,986,411,436]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Figs. 1a, c</figureCitation>
); circumocular furrows (occipital sulci) on dorsal side of head reduced, somewhat distinct anteriorly, absent posteriorly (
<figureCitation id="132C2A5FF66DFF83BA4FFE2FFE26FE4F" box="[324,403,475,500]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1c</figureCitation>
); eyes large and usually moderately projecting laterally (
<figureCitation id="132C2A5FF66DFF83BF00FE2FFF13FDAF" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Figs. 1a, c</figureCitation>
); pronotum with faintly distinct posterior angles, rounded to obtuse and never directed posteriorly (
<figureCitation id="132C2A5FF66DFF83BBF8FDEFFE8AFD8F" box="[243,319,539,564]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1a</figureCitation>
); scutellum regularly punctured on most of exposed portion; elytra with simple punctation, never transversely strigose, occasionally with faint traces of orange peellike rugosity; elytral humeral angle bluntly rounded, not protruding laterally (
<figureCitation id="132C2A5FF66DFF83B896FDAFFC5DFDCF" box="[925,1000,603,628]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1a</figureCitation>
); elytral humeral striae faint; elytral pre-sutural striae visible, originating at scutellar vertex or slightly posterior, terminating at elytral apex, and delimited on each elytron by a faintly distinct, flat sutural border, widest at posterior third; border slightly narrower than proximal width of 3
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antennomere; elytral apices faintly rounded in both sexes (
<figureCitation id="132C2A5FF66DFF83B818FD2FFCD4FD4F" box="[787,865,731,756]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1a</figureCitation>
); pygidium partially exposed, moderately convex, apically rounded in both sexes (
<figureCitation id="132C2A5FF66DFF83B9F5FD0FFCFCFCAF" box="[766,841,763,788]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1a</figureCitation>
) or distinctly produced and distally faintly explanate in females (Fig. 125
<emphasis id="B963EAC8F66DFF83B98FFCE9FD3BFC8F" box="[644,654,797,820]" italics="true" pageId="7" pageNumber="348">s</emphasis>
in
<bibRefCitation id="EF864B2BF66DFF83B9A4FCE8FCFBFC88" author="AUDISIO P." box="[687,846,795,820]" pageId="7" pageNumber="348" refId="ref82842" refString="AUDISIO P. 1993 b: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">AUDISIO 1993b</bibRefCitation>
).
</paragraph>
<paragraph id="8BA836DAF66DFF83BB9BFCC9FE15FAE8" blockId="7.[113,1089,219,1620]" pageId="7" pageNumber="348">
<emphasis id="B963EAC8F66DFF83BB9BFCC9FE80FCEF" box="[144,309,829,852]" italics="true" pageId="7" pageNumber="348">Ventral habitus</emphasis>
: antennal furrows markedly delimited, nearly parallel-sided, or slightly diverging posteriorly; mentum subpentagonal (
<figureCitation id="132C2A5FF66DFF83B963FCAFFD03FCC8" box="[616,694,859,883]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1b</figureCitation>
); prosternal antennal furrows almost completely obliterated (
<figureCitation id="132C2A5FF66DFF83BA76FC8FFE64FC28" box="[381,465,891,915]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1b</figureCitation>
); prosternal process variably shaped, usually relatively narrow, subapical portion dilated, 2.7
<emphasis id="B963EAC8F66DFF83B90AFC68FDBBFC0F" bold="true" box="[513,526,924,948]" pageId="7" pageNumber="348"></emphasis>
3.0× as wide as maximum width of 1
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antennomere, apex usually blunt, not acute (
<figureCitation id="132C2A5FF66DFF83BAB2FC4FFDB3FC6F" box="[441,518,955,980]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1e</figureCitation>
); lateral borders of prosternal process not delimiting shallowly impressed and distinct furrows, and distally terminating prior to predistal lateral expansions (
<figureCitation id="132C2A5FF66DFF83BBFDFC0FFEF4FBAF" box="[246,321,1019,1044]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1e</figureCitation>
); posterior margin of mesoventrite distinctly incised at middle (
<figureCitation id="132C2A5FF66DFF83B8EDFC0FFB84FBAF" box="[998,1073,1019,1044]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1e</figureCitation>
); usually distinct sexual dimorphism present, comprising shallow to deep impressions on metaventrite, more or less raised tubercles, or small lobes along posterior margin of metaventrite; first two visible abdominal ventrites usually simple in both sexes, without tufts of setae, but in males of the
<taxonomicName id="4C174D59F66DFF83BA12FB89FECEFB2F" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[281,379,1149,1172]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B963EAC8F66DFF83BA12FB89FECEFB2F" box="[281,379,1149,1172]" italics="true" pageId="7" pageNumber="348">A. fuscus</emphasis>
</taxonomicName>
complex with faintly distinct, minute tufts of brown setae on first four abdominal ventrites; caudal marginal lines of metacoxal cavities always simple, parallel and contiguous to posterior margin of metacoxal cavities, without deep arched impression of outer axillary line (
<figureCitation id="132C2A5FF66DFF83BA5BFB2FFE2AFB4F" box="[336,415,1243,1268]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">
Fig.
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); axillary space on first abdominal ventrite strongly reduced, axillary angle acute (
<figureCitation id="132C2A5FF66DFF83BA52FB0FFE11FAAF" box="[345,420,1275,1300]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">
Fig.
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); large shallowly impressed arched impressions on basal portion of last visible abdominal ventrite usually not covered by distal portion of penultimate visible abdominal ventrite (
<figureCitation id="132C2A5FF66DFF83BA42FACFFE26FAE8" box="[329,403,1339,1363]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1f</figureCitation>
).
</paragraph>
<paragraph id="8BA836DAF66DFF8DBB9BFAA9FE7EFE28" blockId="7.[113,1089,219,1620]" lastBlockId="9.[113,1089,219,1427]" lastPageId="9" lastPageNumber="350" pageId="7" pageNumber="348">
<emphasis id="B963EAC8F66DFF83BB9BFAA9FEA7FACF" box="[144,274,1373,1396]" italics="true" pageId="7" pageNumber="348">Appendages</emphasis>
: male 1
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antennomere 0.8
<emphasis id="B963EAC8F66DFF83B92DFAA8FD86FACF" bold="true" box="[550,563,1372,1396]" pageId="7" pageNumber="348"></emphasis>
1.0× as long as width of protibiae excluding distal teeth (
<figureCitation id="132C2A5FF66DFF83BBBFFA8FFE92FA28" box="[180,295,1403,1428]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Figs. 1a, b</figureCitation>
); 3
<superScript id="7C629B92F66DFF83BA42FA8EFEE3FA32" attach="left" box="[329,342,1402,1417]" fontSize="6" pageId="7" pageNumber="348">rd</superScript>
antennomere in both sexes 2.8
<emphasis id="B963EAC8F66DFF83B994FA88FD19FA2F" bold="true" box="[671,684,1404,1428]" pageId="7" pageNumber="348"></emphasis>
3.0× as long as wide, 1.3
<emphasis id="B963EAC8F66DFF83B8B3FA88FC70FA2F" bold="true" box="[952,965,1404,1428]" pageId="7" pageNumber="348"></emphasis>
1.4× longer and distinctly thinner than 2
<superScript id="7C629B92F66DFF83BA98FA6EFE17FA12" attach="left" box="[403,418,1434,1449]" fontSize="6" pageId="7" pageNumber="348">nd</superScript>
antennomere (
<figureCitation id="132C2A5FF66DFF83B937FA6FFD1DFA0F" box="[572,680,1435,1460]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Figs. 1a, c</figureCitation>
); 4
<superScript id="7C629B92F66DFF83B9C3FA6EFD61FA12" attach="left" box="[712,724,1434,1449]" fontSize="6" pageId="7" pageNumber="348">th</superScript>
and 5
<superScript id="7C629B92F66DFF83B81AFA6EFCA8FA12" attach="left" box="[785,797,1434,1449]" fontSize="6" pageId="7" pageNumber="348">th</superScript>
antennomeres in both sexes subequal, short, slightly longer than wide; antennal club compact, small, simple, comprising last 3 antennomeres in both sexes, never sexually dimorphic (
<figureCitation id="132C2A5FF66DFF83B80CFA2FFCE6FA4F" box="[775,851,1499,1524]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1a</figureCitation>
), much narrower than width of protibiae; labial palpi relatively short in both sexes (
<figureCitation id="132C2A5FF66DFF83B817FA0FFCD8F9A8" box="[796,877,1531,1555]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1b</figureCitation>
), terminal segment 1.7
<emphasis id="B963EAC8F66DFF83BB98F9E8FF15F98F" bold="true" box="[147,160,1564,1588]" pageId="7" pageNumber="348"></emphasis>
1.8× as long as wide; maxillary palpi moderately long and slender in both sexes (
<figureCitation id="132C2A5FF66DFF83B8ECF9EFFB87F988" box="[999,1074,1563,1587]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1b</figureCitation>
), terminal segment 2.32.4× as long as wide; mandible usually large (
<figureCitation id="132C2A5FF66DFF83B830F9CFFC30F9EF" box="[827,901,1595,1620]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="7" pageNumber="348">Fig. 1a</figureCitation>
), nearly 1.51.6× longer than wide, with peculiarly acuminate apex and denticulate distal portion of inner margin, without evident differences in shape and size among sexes; tarsal claws strongly dentate at base (as in
<figureCitation id="132C2A5FF662FF8CBA27FEEFFE25FE8F" box="[300,400,283,308]" captionStart="Fig" captionStartId="62.[151,182,1415,1435]" captionTargetBox="[152,1126,217,1360]" captionTargetId="figure-10@62.[151,1127,217,1360]" captionTargetPageId="62" captionText="Fig. 17. Aristogethes Audisio &amp; Cline, gen. nov.: a A. translatus (Grouvelle, 1913); b A. pecten (Audisio, Kirk-Spriggs &amp; Kirejtshuk, 1998); c, ek A. pubescens (Reitter, 1872); d, m A. marshalli (Grouvelle, 1914). a, b male habitus (a length 2.6 mm; b length 2.4 mm); c, d dorsal view of head; e ventral view of head and anterior portion of prosternum; f prosternal process and mesoventrite; g anterior portion of scutellum and microsetae on middle of posterior margin of pronotum; h exposed portion of last visible abdominal ventrite; k caudal marginal line of metacoxal cavity; m last tarsomere of middle leg. Scale bars: Figs. d, f, h, k, m = 100 μm; Fig. g = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.5363178" httpUri="https://zenodo.org/record/5363178/files/figure.png" pageId="8" pageNumber="349">
Fig.
<quantity id="4CEF9B3FF662FF8CBA56FEE8FE25FE8F" box="[349,400,284,308]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="349" unit="m" value="17.0">17m</quantity>
</figureCitation>
); tarsi of normal size and shape, 0.60.7× as long as corresponding tibiae (
<figureCitation id="132C2A5FF662FF8CBBE8FECFFE84FEEF" box="[227,305,315,340]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="8" pageNumber="349">Fig. 1a</figureCitation>
); protibiae with a series of usually very large, uneven, sharp or blunt teeth on lateral margin (
<figureCitation id="132C2A5FF662FF8CBA6BFEAFFE1AFECF" box="[352,431,347,372]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="8" pageNumber="349">Fig. 1a</figureCitation>
; see also Figs. 126
<emphasis id="B963EAC8F662FF8CB974FEA9FD10FECF" box="[639,677,348,372]" italics="true" pageId="8" pageNumber="349">ae</emphasis>
in
<bibRefCitation id="EF864B2BF662FF8CB9C2FEA8FCDFFEC8" author="AUDISIO P." box="[713,874,347,372]" pageId="8" pageNumber="349" refId="ref82842" refString="AUDISIO P. 1993 b: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">AUDISIO 1993b</bibRefCitation>
); meso- and metatibiae on lateral margin bearing a nearly double and usually uneven row of large and robust blackish spurs (
<figureCitation id="132C2A5FF663FF8DBBE7FF0FFED5FEA8" box="[236,352,251,276]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="9" pageNumber="350">Figs. 1a, h</figureCitation>
), without U-shaped sinuosity at distal third; meso- and metatibiae of variable width, usually moderately flat but slender (
<figureCitation id="132C2A5FF663FF8DB993FEEFFD56FE8F" box="[664,739,283,308]" captionStart="Fig" captionStartId="8.[151,182,1492,1512]" captionTargetBox="[190,1087,416,1468]" captionTargetId="figure-10@8.[189,1088,415,1468]" captionTargetPageId="8" captionText="Fig. 1. Acanthogethes Reitter, 1871: ah A. fuscus (Olivier, 1790). a male habitus (length 3.5 mm); b ventral view of head and anterior portion of prosternum; c dorsal view of head; d scutellum and microsetae on posterior margin of pronotum; e prosternal process and mesoventrite; f exposed portion of last visible abdominal ventrite; g caudal marginal lines of metacoxal cavities; h middle leg illustrating outer margin of mesotibia. Scale bars: Figs. b, c = 200 μm; Figs. d, e, f, g, h = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="9" pageNumber="350">Fig. 1a</figureCitation>
), never distinctly subtrapezoidal or axe-shaped; scarcely apparent sexual dimorphism in tibial shape; tarsal plates of prolegs slightly wider in males than females; posterior margin of metafemora simple in both sexes, without tubercles or projections.
</paragraph>
<caption id="DF686652F662FF8CBB9CFA20FD82F9E8" ID-DOI="http://doi.org/10.5281/zenodo.5319336" ID-Zenodo-Dep="5319336" httpUri="https://zenodo.org/record/5319336/files/figure.png" pageId="8" pageNumber="349" startId="8.[151,182,1492,1512]" targetBox="[190,1087,416,1468]" targetPageId="8">
<paragraph id="8BA836DAF662FF8CBB9CFA20FD82F9E8" blockId="8.[151,1128,1492,1619]" pageId="8" pageNumber="349">
<emphasis id="B963EAC8F662FF8CBB9CFA20FF7AFA53" bold="true" box="[151,207,1492,1512]" pageId="8" pageNumber="349">Fig. 1.</emphasis>
<taxonomicName id="4C174D59F662FF8CBBDEFA22FE7CFA53" ID-CoL="M7N" authority="Reitter, 1871" authorityName="Reitter" authorityYear="1871" box="[213,457,1493,1512]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F662FF8CBBDEFA22FEE6FA53" box="[213,339,1494,1512]" italics="true" pageId="8" pageNumber="349">Acanthogethes</emphasis>
<bibRefCitation id="EF864B2BF662FF8CBA5CFA21FE7CFA53" author="REITTER E." box="[343,457,1493,1512]" pageId="8" pageNumber="349" pagination="39 - 169" refId="ref91274" refString="REITTER E. 1871: Revision der europaischen Meligethes-Arten. Verhandlungen des Naturforschenden Vereines in Brunn 9: 39 - 169." type="journal article" year="1871">Reitter, 1871</bibRefCitation>
</taxonomicName>
:
<emphasis id="B963EAC8F662FF8CBADAFA21FE46FA53" bold="true" box="[465,499,1493,1512]" pageId="8" pageNumber="349">ah</emphasis>
<taxonomicName id="4C174D59F662FF8CB903FA22FD6EFA53" authority="(Olivier, 1790)" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[520,731,1493,1512]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B963EAC8F662FF8CB903FA22FDE1FA53" box="[520,596,1494,1512]" italics="true" pageId="8" pageNumber="349">A. fuscus</emphasis>
(
<bibRefCitation id="EF864B2BF662FF8CB96BFA21FD60FA53" author="OLIVIER A. G." box="[608,725,1493,1512]" pageId="8" pageNumber="349" pagination="9 - 34" refId="ref91007" refString="OLIVIER A. G. 1790: Entomologie, ou Histoire Naturelle des Insectes, avec leur caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enlumiee. Coleopteres, Tome 2. Baudouin, Paris, 9 - 34 + xxii pp." type="book chapter" year="1790">Olivier, 1790</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B963EAC8F662FF8CB9E8FA21FD5BFA53" bold="true" box="[739,750,1493,1512]" pageId="8" pageNumber="349">a</emphasis>
male habitus (length 3.5 mm);
<emphasis id="B963EAC8F662FF8CBF01FA21FBA3FA53" bold="true" box="[1034,1046,1493,1512]" pageId="8" pageNumber="349">b</emphasis>
ventral view of head and anterior portion of prosternum;
<emphasis id="B963EAC8F662FF8CB93CFA04FDF5F9B8" bold="true" box="[567,576,1520,1539]" pageId="8" pageNumber="349">c</emphasis>
dorsal view of head;
<emphasis id="B963EAC8F662FF8CB80EFA04FCA4F9B8" bold="true" box="[773,785,1520,1539]" pageId="8" pageNumber="349">d</emphasis>
scutellum and microsetae on posterior margin of pronotum;
<emphasis id="B963EAC8F662FF8CBA40F9FFFEE1F9A5" bold="true" box="[331,340,1547,1566]" pageId="8" pageNumber="349">e</emphasis>
prosternal process and mesoventrite;
<emphasis id="B963EAC8F662FF8CB9A8F9FEFD1FF9A5" bold="true" box="[675,682,1546,1566]" pageId="8" pageNumber="349">f</emphasis>
exposed portion of last visible abdominal ventrite;
<emphasis id="B963EAC8F662FF8CBB9CF9D1FF17F983" bold="true" box="[151,162,1573,1592]" pageId="8" pageNumber="349">g</emphasis>
caudal marginal lines of metacoxal cavities;
<emphasis id="B963EAC8F662FF8CB933F9D1FDF1F983" bold="true" box="[568,580,1573,1592]" pageId="8" pageNumber="349">h</emphasis>
middle leg illustrating outer margin of mesotibia. Scale bars: Figs.
<emphasis id="B963EAC8F662FF8CBBCCF9B4FF66F9E8" bold="true" box="[199,211,1600,1619]" pageId="8" pageNumber="349">b</emphasis>
,
<emphasis id="B963EAC8F662FF8CBBD6F9B4FF53F9E8" bold="true" box="[221,230,1600,1619]" pageId="8" pageNumber="349">c</emphasis>
= 200 μm; Figs.
<emphasis id="B963EAC8F662FF8CBA71F9B4FE33F9E8" bold="true" box="[378,390,1600,1619]" pageId="8" pageNumber="349">d</emphasis>
,
<emphasis id="B963EAC8F662FF8CBA9AF9B4FE2FF9E8" bold="true" box="[401,410,1600,1619]" pageId="8" pageNumber="349">e</emphasis>
,
<emphasis id="B963EAC8F662FF8CBAAEF9CBFE19F9E8" bold="true" box="[421,428,1599,1619]" pageId="8" pageNumber="349">f</emphasis>
,
<emphasis id="B963EAC8F662FF8CBABCF9B4FE77F9E8" bold="true" box="[439,450,1600,1619]" pageId="8" pageNumber="349">g</emphasis>
,
<emphasis id="B963EAC8F662FF8CBAC7F9B4FE6DF9E8" bold="true" box="[460,472,1600,1619]" pageId="8" pageNumber="349">h</emphasis>
= 100 μm.
</paragraph>
</caption>
<paragraph id="8BA836DAF663FF8DBB9BFE69FC90FD88" blockId="9.[113,1089,219,1427]" pageId="9" pageNumber="350">
<emphasis id="B963EAC8F663FF8DBB9BFE69FE9CFE0F" box="[144,297,413,436]" italics="true" pageId="9" pageNumber="350">Male genitalia</emphasis>
: processes along inner side of parameres absent (Fig.
<quantity id="4CEF9B3FF663FF8DB85FFE68FC23FE08" box="[852,918,411,436]" metricMagnitude="0" metricUnit="m" metricValue="3.3528" pageId="9" pageNumber="350" unit="in" value="132.0">132 in</quantity>
<bibRefCitation id="EF864B2BF663FF8DB892FE68FB81FE08" author="AUDISIO P." box="[921,1076,411,436]" pageId="9" pageNumber="350" refId="ref82842" refString="AUDISIO P. 1993 b: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">AUDISIO 1993b</bibRefCitation>
), usually with deep and narrow excision along distal margin, without deep median longitudinal desclerotization from proximal portion of tegmen extending to a medial distal V-shaped excision; median lobe of aedeagus variable, without emargination laterally, rounded, subtruncate to acute apically, without distal minute excision or emargination.
</paragraph>
<paragraph id="8BA836DAF663FF8DBB9BFDC9FE28FCA8" blockId="9.[113,1089,219,1427]" pageId="9" pageNumber="350">
<emphasis id="B963EAC8F663FF8DBB9BFDC9FEF9FDEF" box="[144,332,573,596]" italics="true" pageId="9" pageNumber="350">Female genitalia</emphasis>
(
<emphasis id="B963EAC8F663FF8DBA6BFDC9FE7BFDEF" box="[352,462,573,596]" italics="true" pageId="9" pageNumber="350">ovipositor</emphasis>
): variably shaped, usually large; styli usually short but distinct, simple, cylindrical, not distinctly pigmented, inserted close to apex of contiguous gonostyloids; each gonostyloid lightly sclerotized and unpigmented distally, with a simple, never indentate outer portion of basicoxites (Figs. 153
<emphasis id="B963EAC8F663FF8DB9B6FD69FD6AFD0F" box="[701,735,668,692]" italics="true" pageId="9" pageNumber="350">af</emphasis>
in
<bibRefCitation id="EF864B2BF663FF8DB808FD68FC16FD08" author="AUDISIO P." box="[771,931,667,692]" pageId="9" pageNumber="350" refId="ref82842" refString="AUDISIO P. 1993 b: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">AUDISIO 1993b</bibRefCitation>
), and a single, narrow, scarcely pigmented and sclerotized arcuate area along outer subdistal portion of gonostyloids; central point of ovipositor usually more distad than middle, with or without proximally directed spicule.
</paragraph>
</subSubSection>
<subSubSection id="C30D6551F663FF8DBB7AFCE8FCBDFCC8" pageId="9" pageNumber="350" type="etymology">
<paragraph id="8BA836DAF663FF8DBB7AFCE8FCBDFCC8" blockId="9.[113,1089,219,1427]" pageId="9" pageNumber="350">
<emphasis id="B963EAC8F663FF8DBB7AFCE8FF47FC8F" bold="true" box="[113,242,796,820]" pageId="9" pageNumber="350">Etymology.</emphasis>
The generic name is derived from a combination of the Greek ‘ακανθα’ (= spine), to emphasize the strongly toothed tarsal claws, and -
<emphasis id="B963EAC8F663FF8DB9D5FCC9FC97FCEF" box="[734,802,829,852]" italics="true" pageId="9" pageNumber="350">gethes</emphasis>
which is indicative of the phylogenetic relationship with
<taxonomicName id="4C174D59F663FF8DBAB7FCA9FD84FCCF" authorityName="Stephens" authorityYear="1830" box="[444,561,861,884]" class="Insecta" family="Nitidulidae" genus="Meligethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F663FF8DBAB7FCA9FD84FCCF" box="[444,561,861,884]" italics="true" pageId="9" pageNumber="350">Meligethes</emphasis>
</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C30D6551F663FF8DBB7AFC8FFCC3F9FA" pageId="9" pageNumber="350" type="biology_ecology">
<paragraph id="8BA836DAF663FF8DBB7AFC8FFE7DFC68" blockId="9.[113,1089,219,1427]" pageId="9" pageNumber="350">
<emphasis id="B963EAC8F663FF8DBB7AFC8FFF78FC28" bold="true" box="[113,205,891,915]" pageId="9" pageNumber="350">Biology.</emphasis>
All species are strictly associated with flowers of
<taxonomicName id="4C174D59F663FF8DB9D6FC8FFCF6FC28" box="[733,835,891,915]" class="Magnoliopsida" family="Cistaceae" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="350" phylum="Tracheophyta" rank="family">Cistaceae</taxonomicName>
for larval development, especially on small bushes of the following genera:
<taxonomicName id="4C174D59F663FF8DB98FFC69FD73FC0F" box="[644,710,925,948]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="350" phylum="Tracheophyta" rank="genus">
<emphasis id="B963EAC8F663FF8DB98FFC69FD73FC0F" box="[644,710,925,948]" italics="true" pageId="9" pageNumber="350">Cistus</emphasis>
</taxonomicName>
L.,
<taxonomicName id="4C174D59F663FF8DB9E7FC69FC7AFC08" authority="Miller" authorityName="Miller" box="[748,975,923,948]" class="Magnoliopsida" family="Cistaceae" genus="Helianthemum" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="350" phylum="Tracheophyta" rank="genus">
<emphasis id="B963EAC8F663FF8DB9E7FC69FC32FC0F" box="[748,903,925,948]" italics="true" pageId="9" pageNumber="350">Helianthemum</emphasis>
Miller
</taxonomicName>
,
<taxonomicName id="4C174D59F663FF8DB8DCFC69FE71FC68" authority="(Dunal) Spach (AUDISIO 1993 b)" authorityName="Spach (AUDISIO" authorityYear="1993" class="Magnoliopsida" family="Cistaceae" genus="Halimium" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="350" phylum="Tracheophyta" rank="genus">
<emphasis id="B963EAC8F663FF8DB8DCFC69FBF5FC0F" box="[983,1088,925,948]" italics="true" pageId="9" pageNumber="350">Halimium</emphasis>
(Dunal) Spach (
<bibRefCitation id="EF864B2BF663FF8DBA11FC48FE0EFC68" author="AUDISIO P." box="[282,443,955,980]" pageId="9" pageNumber="350" refId="ref82842" refString="AUDISIO P. 1993 b: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">AUDISIO 1993b</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BA836DAF663FF8DBB7AFC2FFDBBFB28" blockId="9.[113,1089,219,1427]" pageId="9" pageNumber="350">
<emphasis id="B963EAC8F663FF8DBB7AFC2FFEC4FC48" bold="true" box="[113,369,987,1011]" pageId="9" pageNumber="350">Phylogenetic position.</emphasis>
Available morphological datasets suggest a likely monophyletic clade that includes
<taxonomicName id="4C174D59F663FF8DBA34FC09FE69FBAF" authorityName="Reitter" authorityYear="1871" box="[319,476,1021,1044]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F663FF8DBA34FC09FE69FBAF" box="[319,476,1021,1044]" italics="true" pageId="9" pageNumber="350">Acanthogethes</emphasis>
</taxonomicName>
, members of the
<taxonomicName id="4C174D59F663FF8DB99CFC09FD49FBAF" authorityName="Kirejtshuk" authorityYear="1989" box="[663,764,1021,1044]" class="Insecta" family="Nitidulidae" genus="Lariopsis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F663FF8DB99CFC09FD49FBAF" box="[663,764,1021,1044]" italics="true" pageId="9" pageNumber="350">Lariopsis</emphasis>
</taxonomicName>
complex of genera, and likely
<taxonomicName id="4C174D59F663FF8DBB7AFBE9FE22FB8F" authority="Scholz, 1932" authorityName="Scholz" authorityYear="1932" box="[113,407,1051,1076]" class="Insecta" family="Nitidulidae" genus="Clypeogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F663FF8DBB7AFBE9FEB4FB8F" box="[113,257,1053,1076]" italics="true" pageId="9" pageNumber="350">Clypeogethes</emphasis>
<bibRefCitation id="EF864B2BF663FF8DBA03FBEFFE22FB8F" author="SCHOLZ M. F. R." box="[264,407,1051,1076]" pageId="9" pageNumber="350" pagination="97 - 100" refId="ref91881" refString="SCHOLZ M. F. R. 1932: Ein neuer Meligethes aus Sudeuropa und Bemerkungen zu einigen Arten. Entomologische Blatter 28: 97 - 100." type="journal article" year="1932">Scholz, 1932</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C174D59F663FF8DBAC4FBE9FDF3FB8F" authority="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti, 2009" authorityName="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti" authorityYear="2009" box="[463,582,1053,1076]" class="Insecta" family="Nitidulidae" genus="Xerogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B963EAC8F663FF8DBAC4FBE9FDF3FB8F" box="[463,582,1053,1076]" italics="true" pageId="9" pageNumber="350">Xerogethes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057B3F663FF8DB945FBE8FD05FB88" box="[590,688,1052,1076]" pageId="9" pageNumber="350" rank="genus">gen. nov.</taxonomicNameLabel>
Molecular data weakly support these relationships. With regards to
<taxonomicName id="4C174D59F663FF8DBAAFFBC9FD8BFBEF" authorityName="Reitter" authorityYear="1871" box="[420,574,1085,1108]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F663FF8DBAAFFBC9FD8BFBEF" box="[420,574,1085,1108]" italics="true" pageId="9" pageNumber="350">Acanthogethes</emphasis>
</taxonomicName>
, this genus possesses a combined series of several symplesiomorphic and few autapomorphic characters, and likely occupies a relatively basal phylogenetic position in
<taxonomicName id="4C174D59F663FF8DBA71FB8FFDBFFB28" authorityName="C. G. Thomson" authorityYear="1859" box="[378,522,1147,1171]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="subFamily" subFamily="Meligethinae">Meligethinae</taxonomicName>
.
</paragraph>
<paragraph id="8BA836DAF663FF8DBB7AFB68FF73FA28" blockId="9.[113,1089,219,1427]" pageId="9" pageNumber="350">
<emphasis id="B963EAC8F663FF8DBB7AFB68FD8FFB08" bold="true" box="[113,570,1179,1204]" pageId="9" pageNumber="350">Taxonomy and geographic distribution.</emphasis>
<taxonomicName id="4C174D59F663FF8DB948FB69FD50FB0F" authorityName="Reitter" authorityYear="1871" box="[579,741,1181,1204]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F663FF8DB948FB69FD50FB0F" box="[579,741,1181,1204]" italics="true" pageId="9" pageNumber="350">Acanthogethes</emphasis>
</taxonomicName>
includes six described species distributed from the
<collectingRegion id="49D3F838F663FF8DBA50FB48FDB0FB68" box="[347,517,1211,1236]" country="Spain" name="Canarias" pageId="9" pageNumber="350">Canary Islands</collectingRegion>
, western Europe and northern Africa to Caucasian areas, which were attributed to three formerly recognized species-complexes, the
<taxonomicName id="4C174D59F663FF8DBF0EFB29FEB0FAAF" baseAuthorityName="Olivier" baseAuthorityYear="1790" class="Insecta" family="Nitidulidae" genus="Meligethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B963EAC8F663FF8DBF0EFB29FEB0FAAF" italics="true" pageId="9" pageNumber="350">Meligethes fuscus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C174D59F663FF8DBA2CFB09FE3AFAAF" authorityName="Sturm" authorityYear="1845" box="[295,399,1277,1300]" class="Insecta" family="Nitidulidae" genus="Meligethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="B963EAC8F663FF8DBA2CFB09FE3AFAAF" box="[295,399,1277,1300]" italics="true" pageId="9" pageNumber="350">M. brevis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C174D59F663FF8DBAEAFB09FDE0FAAF" baseAuthorityName="Kugelann" baseAuthorityYear="1794" box="[481,597,1277,1300]" class="Insecta" family="Nitidulidae" genus="Meligethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="solidus">
<emphasis id="B963EAC8F663FF8DBAEAFB09FDE0FAAF" box="[481,597,1277,1300]" italics="true" pageId="9" pageNumber="350">M. solidus</emphasis>
</taxonomicName>
species-complexes. Two additional species described from North Africa by
<bibRefCitation id="EF864B2BF663FF8DBACFFAE8FDDBFA88" author="EASTON A. M." box="[452,622,1307,1331]" pageId="9" pageNumber="350" pagination="225 - 240" refId="ref85400" refString="EASTON A. M. 1955 a: The Palearctic species of the subgenus Acanthogethes of the genus Meligethes Stephens (Coleoptera, Nitidulidae). Annals and Magazine of Natural History, Series 12 8: 225 - 240." type="journal article" year="1955">EASTON (1955a)</bibRefCitation>
, likely dubiously attributed to differentiated populations of
<taxonomicName id="4C174D59F663FF8DBA17FAC9FD53FAE8" authority="(Rosenhauer, 1856)" authorityName="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti" authorityYear="2009" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[284,742,1339,1364]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="lamii">
<emphasis id="B963EAC8F663FF8DBA17FAC9FE4AFAEF" box="[284,511,1341,1364]" italics="true" pageId="9" pageNumber="350">Acanthogethes lamii</emphasis>
(
<bibRefCitation id="EF864B2BF663FF8DB91AFACFFD6BFAE8" author="ROSENHAUER W. G." box="[529,734,1339,1363]" pageId="9" pageNumber="350" refId="ref91743" refString="ROSENHAUER W. G. 1856: Die Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt, nebst den Beschreibungen von 249 neuen oder bis jetz noch unbeschriebenen Gattungen und Arten. T. Blaesing Verlag, Erlangen, 429 pp." type="book" year="1856">Rosenhauer, 1856</bibRefCitation>
)
</taxonomicName>
, require further morphological and molecular analyses to definitively ascertain their true taxonomic status (
<bibRefCitation id="EF864B2BF663FF8DB8ACFAA8FB89FAC8" author="AUDISIO P." box="[935,1084,1371,1396]" pageId="9" pageNumber="350" pagination="543 - 556" refId="ref82734" refString="AUDISIO P. 1986: La posizione tassonomica di Meligethes fuscus (Olivier) e specie correlate (Coleoptera, Nitidulidae). Bollettino del Museo Regionale di Scienze Naturali Torino 4: 543 - 556." type="journal article" year="1986">AUDISIO 1986</bibRefCitation>
,
<bibRefCitation id="EF864B2BF663FF8DBB7AFA8FFF0FFA28" author="AUDISIO P." box="[113,186,1403,1427]" pageId="9" pageNumber="350" refId="ref82842" refString="AUDISIO P. 1993 b: Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. Vol. 32. Calderini Edizione, Bologna, xvi + 971 pp." type="book" year="1993">1993b</bibRefCitation>
).
</paragraph>
<paragraph id="8BA836DAF663FF8DBB79FA5EFCC3FA07" blockId="9.[113,947,1449,1602]" box="[114,886,1449,1468]" pageId="9" pageNumber="350">
<taxonomicName id="4C174D59F663FF8DBB79FA5EFE10FA07" authority="(Sturm, 1845)" authorityName="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti" authorityYear="2009" baseAuthorityName="Sturm" baseAuthorityYear="1845" box="[114,421,1449,1468]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="brevis" status="comb. nov.">
<emphasis id="B963EAC8F663FF8DBB79FA5EFE9DFA07" box="[114,296,1450,1468]" italics="true" pageId="9" pageNumber="350">Acanthogethes brevis</emphasis>
(
<bibRefCitation id="EF864B2BF663FF8DBA38FA5DFE2AFA07" author="STURM J." box="[307,415,1449,1468]" pageId="9" pageNumber="350" pagination="ccciv - cccix" refId="ref92410" refString="STURM J. 1845: Deutschlands Fauna in Abbildungen nach der Natur mit Beschreibungen. V. Abtheilung. Die Insecten. Zweites Bandchen. Kafer. Nurnberg, 114 pp + pls. ccciv - cccix." type="book chapter" year="1845">Sturm, 1845</bibRefCitation>
)
</taxonomicName>
<emphasis id="B963EAC8F663FF8DBAA1FA5DFDB9FA07" bold="true" box="[426,524,1449,1468]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A25057B3F663FF8DBAA1FA5DFDB9FA07" box="[426,524,1449,1468]" pageId="9" pageNumber="350" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Europe, N
<collectingCountry id="F300764AF663FF8DB9D4FA5DFCA8FA07" box="[735,797,1449,1468]" name="Turkey" pageId="9" pageNumber="350">Turkey</collectingCountry>
, Caucasus
</paragraph>
<paragraph id="8BA836DAF663FF8DBB7AFA31FC06FA6C" blockId="9.[113,947,1449,1602]" box="[113,947,1476,1495]" pageId="9" pageNumber="350">
<taxonomicName id="4C174D59F663FF8DBB7AFA31FE57FA6C" authority="(A. G. Olivier, 1790)" baseAuthorityName="A. G. Olivier" baseAuthorityYear="1790" box="[113,482,1476,1495]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="fuscus" status="comb. nov.">
<emphasis id="B963EAC8F663FF8DBB7AFA31FE9CFA6C" box="[113,297,1477,1495]" italics="true" pageId="9" pageNumber="350">Acanthogethes fuscus</emphasis>
(A. G.
<bibRefCitation id="EF864B2BF663FF8DBA61FA30FE69FA6C" author="OLIVIER A. G." box="[362,476,1476,1495]" pageId="9" pageNumber="350" pagination="9 - 34" refId="ref91007" refString="OLIVIER A. G. 1790: Entomologie, ou Histoire Naturelle des Insectes, avec leur caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enlumiee. Coleopteres, Tome 2. Baudouin, Paris, 9 - 34 + xxii pp." type="book chapter" year="1790">Olivier, 1790</bibRefCitation>
)
</taxonomicName>
<emphasis id="B963EAC8F663FF8DBAE3FA30FDFFFA6C" bold="true" box="[488,586,1476,1495]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A25057B3F663FF8DBAE3FA30FDFFFA6C" box="[488,586,1476,1495]" pageId="9" pageNumber="350" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
W Mediterranean region, W
<collectingCountry id="F300764AF663FF8DB87CFA30FC06FA6C" box="[887,947,1476,1495]" name="Greece" pageId="9" pageNumber="350">Greece</collectingCountry>
</paragraph>
<paragraph id="8BA836DAF663FF8DBB7AFA2BFCE1FA4A" blockId="9.[113,947,1449,1602]" box="[113,852,1502,1521]" pageId="9" pageNumber="350">
<taxonomicName id="4C174D59F663FF8DBB7AFA2BFE7DFA4A" authority="(Audisio, 1986)" authorityName="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti" authorityYear="2009" baseAuthorityName="Audisio" baseAuthorityYear="1986" box="[113,456,1502,1521]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="hercules" status="comb. nov.">
<emphasis id="B963EAC8F663FF8DBB7AFA2BFE8EFA4A" box="[113,315,1503,1521]" italics="true" pageId="9" pageNumber="350">Acanthogethes hercules</emphasis>
(
<bibRefCitation id="EF864B2BF663FF8DBA4CFA2AFE77FA4A" author="AUDISIO P." box="[327,450,1502,1521]" pageId="9" pageNumber="350" pagination="543 - 556" refId="ref82734" refString="AUDISIO P. 1986: La posizione tassonomica di Meligethes fuscus (Olivier) e specie correlate (Coleoptera, Nitidulidae). Bollettino del Museo Regionale di Scienze Naturali Torino 4: 543 - 556." type="journal article" year="1986">Audisio, 1986</bibRefCitation>
)
</taxonomicName>
<emphasis id="B963EAC8F663FF8DBAC5FA2AFD85FA4A" bold="true" box="[462,560,1502,1521]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A25057B3F663FF8DBAC5FA2AFD85FA4A" box="[462,560,1502,1521]" pageId="9" pageNumber="350" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
<collectingRegion id="49D3F838F663FF8DB989FA2AFCB0FA4A" box="[642,773,1502,1521]" country="Spain" name="Canarias" pageId="9" pageNumber="350">Canary Islands</collectingRegion>
: Tenerife
</paragraph>
<paragraph id="8BA836DAF663FF8DBB7AFA0EFCE1F9B7" blockId="9.[113,947,1449,1602]" box="[113,852,1529,1548]" pageId="9" pageNumber="350">
<taxonomicName id="4C174D59F663FF8DBB7AFA0EFE78F9B7" authority="(Rosenhauer, 1856)" authorityName="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti" authorityYear="2009" baseAuthorityName="Rosenhauer" baseAuthorityYear="1856" box="[113,461,1529,1548]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="lamii" status="comb. nov.">
<emphasis id="B963EAC8F663FF8DBB7AFA0EFE95F9B7" box="[113,288,1530,1548]" italics="true" pageId="9" pageNumber="350">Acanthogethes lamii</emphasis>
(
<bibRefCitation id="EF864B2BF663FF8DBA20FA0DFE72F9B7" author="ROSENHAUER W. G." box="[299,455,1529,1548]" pageId="9" pageNumber="350" refId="ref91743" refString="ROSENHAUER W. G. 1856: Die Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt, nebst den Beschreibungen von 249 neuen oder bis jetz noch unbeschriebenen Gattungen und Arten. T. Blaesing Verlag, Erlangen, 429 pp." type="book" year="1856">Rosenhauer, 1856</bibRefCitation>
)
</taxonomicName>
<emphasis id="B963EAC8F663FF8DBAD9FA0DFD81F9B7" bold="true" box="[466,564,1529,1548]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A25057B3F663FF8DBAD9FA0DFD81F9B7" box="[466,564,1529,1548]" pageId="9" pageNumber="350" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
W Mediterranean region
</paragraph>
<paragraph id="8BA836DAF663FF8DBB7AF9E1FCACF99C" blockId="9.[113,947,1449,1602]" box="[113,793,1555,1575]" pageId="9" pageNumber="350">
<taxonomicName id="4C174D59F663FF8DBB7AF9E1FE0EF99D" authority="(Guillebeau, 1885)" authorityName="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti" authorityYear="2009" baseAuthorityName="Guillebeau" baseAuthorityYear="1885" box="[113,443,1555,1575]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="reyi" status="comb. nov.">
<emphasis id="B963EAC8F663FF8DBB7AF9E1FEA1F99C" box="[113,276,1557,1575]" italics="true" pageId="9" pageNumber="350">Acanthogethes reyi</emphasis>
(Guillebeau, 1885)
</taxonomicName>
<emphasis id="B963EAC8F663FF8DBACBF9E0FD97F99C" bold="true" box="[448,546,1556,1575]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A25057B3F663FF8DBACBF9E0FD97F99C" box="[448,546,1556,1575]" pageId="9" pageNumber="350" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
S Europe,
<collectingCountry id="F300764AF663FF8DB9D7F9E0FCACF99C" box="[732,793,1556,1575]" name="Turkey" pageId="9" pageNumber="350">Turkey</collectingCountry>
</paragraph>
<paragraph id="8BA836DAF663FF8DBB7AF9DBFCC3F9FA" blockId="9.[113,947,1449,1602]" box="[113,886,1582,1602]" pageId="9" pageNumber="350">
<taxonomicName id="4C174D59F663FF8DBB7AF9DBFE78F9FA" authority="(Kugelann, 1794)" authorityName="Audisio &amp; Cline &amp; Biase &amp; Antonini &amp; Mancini &amp; Trizzino &amp; Costantini &amp; Strika &amp; Lamanna &amp; Cerretti" authorityYear="2009" baseAuthorityName="Kugelann" baseAuthorityYear="1794" box="[113,461,1582,1601]" class="Insecta" family="Nitidulidae" genus="Acanthogethes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="solidus" status="comb. nov.">
<emphasis id="B963EAC8F663FF8DBB7AF9DBFE85F9FA" box="[113,304,1583,1601]" italics="true" pageId="9" pageNumber="350">Acanthogethes solidus</emphasis>
(Kugelann, 1794)
</taxonomicName>
<emphasis id="B963EAC8F663FF8DBAD9F9DAFD81F9FA" bold="true" box="[466,564,1582,1601]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A25057B3F663FF8DBAD9F9DAFD81F9FA" box="[466,564,1582,1601]" pageId="9" pageNumber="350" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Europe, N
<collectingCountry id="F300764AF663FF8DB9D4F9DAFCA8F9FA" box="[735,797,1582,1601]" name="Turkey" pageId="9" pageNumber="350">Turkey</collectingCountry>
, Caucasus
</paragraph>
</subSubSection>
</treatment>
</document>