243 lines
29 KiB
XML
243 lines
29 KiB
XML
<document id="98351B12ADE6C20499F019D80A2DC5D2" ID-CLB-Dataset="55857" ID-DOI="10.11646/zootaxa.5128.1.4" ID-GBIF-Dataset="d0690c80-bf6f-4ae7-9397-9e504dfad497" ID-ISSN="1175-5326" ID-Zenodo-Dep="6498040" ID-ZooBank="A5F79F57-8C14-47C8-BA3F-5AED0D6B83C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650719515444" checkinUser="plazi" docAuthor="KAZANTSEV, SERGEY V. & PERKOVSKY, EVGENY E." docDate="2022" docId="03A8241AFFB9F77DF2C8FE23EAF7D616" docLanguage="en" docName="zootaxa.5128.1.4.pdf" docOrigin="Zootaxa 5128 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5128.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helcophorus (Thibeteros) ekaterinae Kazantsev et Perkovsky, sp. nov." docType="treatment" docVersion="6" lastPageNumber="87" masterDocId="FF915C62FFBBF77EF25FFFECEE78D01C" masterDocTitle="Imprint of a Helcophorus Fairmaire, 1881: the first net-winged beetle (Coleoptera Lycidae) from Rovno amber" masterLastPageNumber="90" masterPageNumber="84" pageNumber="86" updateTime="1699329033473" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="28DED644C99334D0B4A40960E2BA3466">
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<mods:title id="AAF43C8657E03D85AD0AE06ECDB5C8AB">Imprint of a Helcophorus Fairmaire, 1881: the first net-winged beetle (Coleoptera Lycidae) from Rovno amber</mods:title>
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<mods:name id="E4D61DF549737B8DF35E84023BCA2C95" type="personal">
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<mods:namePart id="4584329747DF60CDF06014922155AFDB">KAZANTSEV, SERGEY V.</mods:namePart>
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<mods:namePart id="65BAC301293507C365C0A2B8EB2C97B1">PERKOVSKY, EVGENY E.</mods:namePart>
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<mods:title id="765AB3C5630AFBBFA3BA538D9E36719B">Zootaxa</mods:title>
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<mods:part id="B457A790999ED3D83CCEFCCCA0D2A716">
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<mods:date id="DB9D46516B9BB1BDB937BD4B37BAD5B7">2022</mods:date>
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<mods:number id="348E417E939D0BA22AA120AAD250F608">2022-04-19</mods:number>
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<mods:number id="913E75A027495EB07C07D7F180FE21CF">5128</mods:number>
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<mods:url id="251CEE63F0CCACDF8B6748916CBAA501">http://dx.doi.org/10.11646/zootaxa.5128.1.4</mods:url>
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<treatment id="03A8241AFFB9F77DF2C8FE23EAF7D616" ID-DOI="http://doi.org/10.5281/zenodo.6498155" ID-GBIF-Taxon="194847470" ID-Zenodo-Dep="6498155" LSID="urn:lsid:plazi:treatment:03A8241AFFB9F77DF2C8FE23EAF7D616" httpUri="http://treatment.plazi.org/id/03A8241AFFB9F77DF2C8FE23EAF7D616" lastPageId="3" lastPageNumber="87" pageId="2" pageNumber="86">
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<subSubSection id="C31BC687FFB9F77CF2C8FE23ED8BD1F6" box="[151,1011,463,490]" pageId="2" pageNumber="86" type="nomenclature">
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<paragraph id="8BBE950CFFB9F77CF2C8FE23ED8BD1F6" blockId="2.[151,1011,463,526]" box="[151,1011,463,490]" pageId="2" pageNumber="86">
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<heading id="D0F62260FFB9F77CF2C8FE23ED8BD1F6" bold="true" box="[151,1011,463,490]" fontSize="11" level="1" pageId="2" pageNumber="86" reason="1">
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<emphasis id="B975491EFFB9F77CF2C8FE23ED8BD1F6" bold="true" box="[151,1011,463,490]" pageId="2" pageNumber="86">
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<taxonomicName id="4C01EE8FFFB9F77CF2C8FE23EDF1D1F6" authority="Kazantsev et Perkovsky" authorityName="Kazantsev et Perkovsky" box="[151,905,463,490]" class="Insecta" family="Lycidae" genus="Helcophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="86" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov." subGenus="Thibeteros">
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<emphasis id="B975491EFFB9F77CF2C8FE23EF4BD1F6" bold="true" box="[151,307,463,490]" italics="true" pageId="2" pageNumber="86">Helcophorus</emphasis>
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(
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<emphasis id="B975491EFFB9F77CF31AFE23EFBED1F6" bold="true" box="[325,454,463,490]" italics="true" pageId="2" pageNumber="86">Thibeteros</emphasis>
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)
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<emphasis id="B975491EFFB9F77CF388FE23EC2ED1F6" bold="true" box="[471,598,463,490]" italics="true" pageId="2" pageNumber="86">ekaterinae</emphasis>
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Kazantsev et Perkovsky
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A246F465FFB9F77CF1C9FE3CED8BD1F6" box="[918,1011,464,490]" pageId="2" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31BC687FFB9F77CF2C8FE18EF74D212" box="[151,268,500,526]" pageId="2" pageNumber="86" type="description">
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<paragraph id="8BBE950CFFB9F77CF2C8FE18EF74D212" blockId="2.[151,1011,463,526]" box="[151,268,500,526]" pageId="2" pageNumber="86">
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(
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||
<figureCitation id="133A8989FFB9F77CF2C0FE18EE9ED212" box="[159,230,500,526]" captionStart="FIGURE 1" captionStartId="2.[151,250,1555,1579]" captionTargetBox="[156,1432,728,1529]" captionTargetId="figure-137@2.[156,1432,728,1529]" captionTargetPageId="2" captionText="FIGURE 1. General view of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498052" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="2" pageNumber="86">Figs 1</figureCitation>
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,
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<figureCitation id="133A8989FFB9F77CF2ABFE18EF7BD212" box="[244,259,500,526]" captionStart="FIGURE 2" captionStartId="3.[151,250,1126,1150]" captionTargetBox="[161,1426,181,1101]" captionTargetId="figure-24@3.[161,1426,181,1101]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum and the base of elytra of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498067" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="2" pageNumber="86">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31BC687FFB9F77CF2C8FDD0EA42D24B" box="[151,1082,572,599]" pageId="2" pageNumber="86" type="materials_examined">
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<paragraph id="8BBE950CFFB9F77CF2C8FDD0EA42D24B" blockId="2.[151,1436,572,706]" box="[151,1082,572,599]" pageId="2" pageNumber="86">
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||
<emphasis id="B975491EFFB9F77CF2C8FDD0EF7ED24A" bold="true" box="[151,262,572,598]" pageId="2" pageNumber="86">Material.</emphasis>
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<materialsCitation id="3B699F51FFB9F77CF352FDD0EA42D24B" ID-GBIF-Occurrence="3756432301" box="[269,1082,572,599]" collectionCode="SIZK" location="Rovno" pageId="2" pageNumber="86" specimenCode="UA-28623" specimenCount="1" specimenCount-male="1" stateProvince="Rivne" typeStatus="holotype">
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<typeStatus id="54BA2BAEFFB9F77CF352FDD0EF0DD24A" box="[269,373,572,598]" pageId="2" pageNumber="86" type="holotype">Holotype</typeStatus>
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||
<specimenCount id="9D075E85FFB9F77CF322FDD1EFECD24B" box="[381,404,573,599]" pageId="2" pageNumber="86" type="male">♂</specimenCount>
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, specimen No.
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<collectionCode id="ED100DC9FFB9F77CF01EFDD0EC07D24A" box="[577,639,572,598]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="2" pageNumber="86">SIZK</collectionCode>
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||
<specimenCode id="DBA73D77FFB9F77CF0D9FDD0ED79D24A" box="[646,769,572,598]" collectionCode="UA-" pageId="2" pageNumber="86">UA-28623</specimenCode>
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||
,
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||
<collectingRegion id="49C55BEEFFB9F77CF153FDD1ED2FD24B" box="[780,855,573,599]" country="Ukraine" name="Rivne" pageId="2" pageNumber="86">Rovno</collectingRegion>
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amber, late Eocene.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31BC687FFB9F77DF298FD8CEA93D4F6" lastPageId="3" lastPageNumber="87" pageId="2" pageNumber="86" type="description">
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||
<paragraph id="8BBE950CFFB9F77CF298FD8CECF5D2DE" blockId="2.[151,1436,572,706]" pageId="2" pageNumber="86">
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||
<emphasis id="B975491EFFB9F77CF298FD8CEF21D266" bold="true" box="[199,345,608,634]" pageId="2" pageNumber="86">Description.</emphasis>
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Sex unknown. Most of the body missing, the amber piece only carries an imprint of the anterior half of its upper part, the inclusion of antennae and the anterior leg in milky substance. Upperside light brown (quite probably red in the live specimen;
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<figureCitation id="133A8989FFB9F77CF043FD44EC19D2DE" box="[540,609,680,706]" captionStart="FIGURE 1" captionStartId="2.[151,250,1555,1579]" captionTargetBox="[156,1432,728,1529]" captionTargetId="figure-137@2.[156,1432,728,1529]" captionTargetPageId="2" captionText="FIGURE 1. General view of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498052" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="2" pageNumber="86">Figs 1</figureCitation>
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,
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<figureCitation id="133A8989FFB9F77CF030FD44ECF8D2DE" box="[623,640,680,706]" captionStart="FIGURE 2" captionStartId="3.[151,250,1126,1150]" captionTargetBox="[161,1426,181,1101]" captionTargetId="figure-24@3.[161,1426,181,1101]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum and the base of elytra of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498067" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="2" pageNumber="86">2</figureCitation>
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).
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</paragraph>
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<caption id="DF7EC584FFB9F77CF2C8F9FFEDEFD637" ID-DOI="http://doi.org/10.5281/zenodo.6498052" ID-Zenodo-Dep="6498052" box="[151,919,1555,1579]" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="2" pageNumber="86" startId="2.[151,250,1555,1579]" targetBox="[156,1432,728,1529]" targetPageId="2">
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<paragraph id="8BBE950CFFB9F77CF2C8F9FFEDEFD637" blockId="2.[151,919,1555,1579]" box="[151,919,1555,1579]" pageId="2" pageNumber="86">
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||
<emphasis id="B975491EFFB9F77CF2C8F9FFEF75D637" bold="true" box="[151,269,1555,1579]" pageId="2" pageNumber="86">FIGURE 1</emphasis>
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. General view of
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<taxonomicName id="4C01EE8FFFB9F77CF39CF9FFED42D637" box="[451,826,1555,1579]" class="Insecta" family="Lycidae" genus="Helcophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="86" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov." subGenus="Thibeteros">
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<emphasis id="B975491EFFB9F77CF39CF9FFEC3ED637" box="[451,582,1555,1579]" italics="true" pageId="2" pageNumber="86">Helcophorus</emphasis>
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||
(
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<emphasis id="B975491EFFB9F77CF00AF9FFECB8D637" box="[597,704,1555,1579]" italics="true" pageId="2" pageNumber="86">Thibeteros</emphasis>
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)
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<emphasis id="B975491EFFB9F77CF090F9FFED42D637" box="[719,826,1555,1579]" italics="true" pageId="2" pageNumber="86">ekaterinae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A246F465FFB9F77CF119F9FFEDEFD637" box="[838,919,1555,1579]" pageId="2" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</caption>
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<paragraph id="8BBE950CFFB9F77CF298F9B5ED4BD68B" blockId="2.[151,1437,1625,2012]" pageId="2" pageNumber="86">
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Head about as wide as pronotum. Eyes bulging, relatively large, their diameter ca. 1/2 length of pronotum (
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<figureCitation id="133A8989FFB9F77CF734F9B5EEDDD68B" captionStart="FIGURE 1" captionStartId="2.[151,250,1555,1579]" captionTargetBox="[156,1432,728,1529]" captionTargetId="figure-137@2.[156,1432,728,1529]" captionTargetPageId="2" captionText="FIGURE 1. General view of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498052" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="2" pageNumber="86">Figs 1</figureCitation>
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,
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<figureCitation id="133A8989FFB9F77CF2ECF991EEBCD68B" box="[179,196,1661,1687]" captionStart="FIGURE 2" captionStartId="3.[151,250,1126,1150]" captionTargetBox="[161,1426,181,1101]" captionTargetId="figure-24@3.[161,1426,181,1101]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum and the base of elytra of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498067" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="2" pageNumber="86">2</figureCitation>
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). Antennomeres 2 and 3 of approximately equal length.
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</paragraph>
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<paragraph id="8BBE950CFFB9F77CF298F94EED49D73B" blockId="2.[151,1437,1625,2012]" pageId="2" pageNumber="86">
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Pronotum transverse, ca. 1.5 times wider than long, semi-trapezoidal, with pronounced acute posterior angles; median areole rhomboidal, broad in the middle, connected with lateral sides with narrow oblique, slightly curved costae. Scutellum (postnotal plate of mesonotal scutellum) transverse, parallel-sided, medially moderately triangularly notched, with broadly rounded distal lobes (
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<figureCitation id="133A8989FFB9F77CF0E1F8E1ED7DD73B" box="[702,773,1805,1831]" captionStart="FIGURE 1" captionStartId="2.[151,250,1555,1579]" captionTargetBox="[156,1432,728,1529]" captionTargetId="figure-137@2.[156,1432,728,1529]" captionTargetPageId="2" captionText="FIGURE 1. General view of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498052" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="2" pageNumber="86">Figs 1</figureCitation>
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,
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<figureCitation id="133A8989FFB9F77CF14CF8E1ED5CD73B" box="[787,804,1805,1831]" captionStart="FIGURE 2" captionStartId="3.[151,250,1126,1150]" captionTargetBox="[161,1426,181,1101]" captionTargetId="figure-24@3.[161,1426,181,1101]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum and the base of elytra of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498067" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="2" pageNumber="86">2</figureCitation>
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||
).
|
||
</paragraph>
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<paragraph id="8BBE950CFFB9F77CF298F8DDECE7D78F" blockId="2.[151,1437,1625,2012]" pageId="2" pageNumber="86">
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Elytra elongate, flattened, noticeably widened distally; with four primary costae, of which third distinctly stouter; interstices with rectangular transverse, their width ca. 2 times exceeding their length, cells; pubescence sparse and short, almost indistinguishable (
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<figureCitation id="133A8989FFB9F77CF073F895EC0BD78F" box="[556,627,1913,1939]" captionStart="FIGURE 1" captionStartId="2.[151,250,1555,1579]" captionTargetBox="[156,1432,728,1529]" captionTargetId="figure-137@2.[156,1432,728,1529]" captionTargetPageId="2" captionText="FIGURE 1. General view of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498052" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="2" pageNumber="86">Figs 1</figureCitation>
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,
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<figureCitation id="133A8989FFB9F77CF0DEF895ECEAD78F" box="[641,658,1913,1939]" captionStart="FIGURE 2" captionStartId="3.[151,250,1126,1150]" captionTargetBox="[161,1426,181,1101]" captionTargetId="figure-24@3.[161,1426,181,1101]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum and the base of elytra of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498067" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="2" pageNumber="86">2</figureCitation>
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||
).
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||
</paragraph>
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<paragraph id="8BBE950CFFB9F77CF298F871EF63D7C0" blockId="2.[151,1437,1625,2012]" pageId="2" pageNumber="86">
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Length (of the roughly anterior half of the individual):
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<quantity id="4CF938E9FFB9F77CF166F871EDECD7A4" box="[825,916,1949,1976]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="86" unit="mm" value="6.5">6.5 mm</quantity>
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. Width at humeri: 2.0 mm. Length of antenna ca.
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<quantity id="4CF938E9FFB9F77CF2E1F82DEF6FD7C0" box="[190,279,1985,2012]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="86" unit="mm" value="4.4">4.4 mm</quantity>
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.
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</paragraph>
|
||
<caption id="DF7EC584FFB8F77DF2C8FB8AEA2AD462" ID-DOI="http://doi.org/10.5281/zenodo.6498067" ID-Zenodo-Dep="6498067" box="[151,1106,1126,1150]" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="3" pageNumber="87" startId="3.[151,250,1126,1150]" targetBox="[161,1426,181,1101]" targetPageId="3">
|
||
<paragraph id="8BBE950CFFB8F77DF2C8FB8AEA2AD462" blockId="3.[151,1106,1126,1150]" box="[151,1106,1126,1150]" pageId="3" pageNumber="87">
|
||
<emphasis id="B975491EFFB8F77DF2C8FB8AEF75D462" bold="true" box="[151,269,1126,1150]" pageId="3" pageNumber="87">FIGURE 2</emphasis>
|
||
. Pronotum and the base of elytra of
|
||
<taxonomicName id="4C01EE8FFFB8F77DF021FB8AED8DD462" box="[638,1013,1126,1150]" class="Insecta" family="Lycidae" genus="Helcophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov." subGenus="Thibeteros">
|
||
<emphasis id="B975491EFFB8F77DF021FB8AED79D462" box="[638,769,1126,1150]" italics="true" pageId="3" pageNumber="87">Helcophorus</emphasis>
|
||
(
|
||
<emphasis id="B975491EFFB8F77DF14FFB8AED03D462" box="[784,891,1126,1150]" italics="true" pageId="3" pageNumber="87">Thibeteros</emphasis>
|
||
)
|
||
<emphasis id="B975491EFFB8F77DF1D5FB8AED8DD462" box="[906,1013,1126,1150]" italics="true" pageId="3" pageNumber="87">ekaterinae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A246F465FFB8F77DF65EFB8AEA2AD462" box="[1025,1106,1126,1150]" pageId="3" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBE950CFFB8F77DF298FB40EA93D4F6" blockId="3.[151,1436,1196,1547]" pageId="3" pageNumber="87">
|
||
<emphasis id="B975491EFFB8F77DF298FB40EF0AD4DA" bold="true" box="[199,370,1196,1222]" pageId="3" pageNumber="87">Syninclusions.</emphasis>
|
||
SIZK UA-28623: 2 Dolichopodaidae, 2
|
||
<taxonomicName id="4C01EE8FFFB8F77DF117FB40EDB0D4DA" box="[840,968,1196,1222]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
||
(1
|
||
<taxonomicName id="4C01EE8FFFB8F77DF1AEFB41EAB3D4DB" box="[1009,1227,1197,1223]" class="Insecta" family="Formicidae" genus="Yantaromyrmex" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B975491EFFB8F77DF1AEFB41EAD8D4DA" box="[1009,1184,1197,1222]" italics="true" pageId="3" pageNumber="87">Yantaromyrmex</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
), Aranei, cobweb; SIZK UA-28624: 2
|
||
<taxonomicName id="4C01EE8FFFB8F77DF327FB3CEF83D4F6" box="[376,507,1232,1258]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C01EE8FFFB8F77DF05AFB3CECD0D4F6" box="[517,680,1232,1258]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
|
||
, Cecydomyiidae, Collembola, Entomobryomorpha.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31BC687FFB8F77DF298FB18EAAAD52E" pageId="3" pageNumber="87" type="etymology">
|
||
<paragraph id="8BBE950CFFB8F77DF298FB18EAAAD52E" blockId="3.[151,1436,1196,1547]" pageId="3" pageNumber="87">
|
||
<emphasis id="B975491EFFB8F77DF298FB18EF36D512" bold="true" box="[199,334,1268,1294]" pageId="3" pageNumber="87">Etymology.</emphasis>
|
||
<taxonomicName id="4C01EE8FFFB8F77DF30BFB18EC88D512" box="[340,752,1268,1295]" class="Insecta" family="Lycidae" genus="Helcophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov." subGenus="Thibeteros">
|
||
<emphasis id="B975491EFFB8F77DF30BFB18EF9DD512" box="[340,485,1268,1294]" italics="true" pageId="3" pageNumber="87">Helcophorus</emphasis>
|
||
(
|
||
<emphasis id="B975491EFFB8F77DF3ABFB18EC13D512" box="[500,619,1268,1294]" italics="true" pageId="3" pageNumber="87">Thibeteros</emphasis>
|
||
)
|
||
<emphasis id="B975491EFFB8F77DF025FB18EC88D512" box="[634,752,1268,1294]" italics="true" pageId="3" pageNumber="87">ekaterinae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B975491EFFB8F77DF0A9FB18ED37D512" bold="true" box="[758,847,1268,1294]" pageId="3" pageNumber="87">
|
||
<taxonomicNameLabel id="A246F465FFB8F77DF0A9FB18ED37D512" box="[758,847,1268,1294]" pageId="3" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is named after our colleague Dr
|
||
<taxonomicName id="4C01EE8FFFB8F77DF6E7FB18EF75D52E" authority="Sidorchuk" authorityName="Sidorchuk" class="Insecta" family="Bethylidae" genus="Ekaterina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="genus">Ekaterina Sidorchuk (Moscow)</taxonomicName>
|
||
, a prominent paleoentomologist who lost her life in a diving accident in the
|
||
<collectingCountry id="F316D59CFFB8F77DF63CFAF4EAB7D52E" box="[1123,1231,1304,1330]" name="Maldives" pageId="3" pageNumber="87">Maldives</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31BC687FFB8F77DF298FAD0EAF7D616" pageId="3" pageNumber="87" type="diagnosis">
|
||
<paragraph id="8BBE950CFFB8F77DF298FAD0EAF7D616" blockId="3.[151,1436,1196,1547]" pageId="3" pageNumber="87">
|
||
<emphasis id="B975491EFFB8F77DF298FAD0EF3AD54A" bold="true" box="[199,322,1340,1366]" pageId="3" pageNumber="87">Diagnosis.</emphasis>
|
||
The new species may be distinguished from the extant members of the same subgenus,
|
||
<taxonomicName id="4C01EE8FFFB8F77DF71FFAD1EFD4D566" authority="Kazantsev, 2000" authorityName="Kazantsev" authorityYear="2000" class="Insecta" family="Lycidae" genus="Helcophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="species" species="tricolor" subGenus="Thibeteros">
|
||
<emphasis id="B975491EFFB8F77DF71FFAD1EB24D54A" box="[1344,1372,1341,1366]" italics="true" pageId="3" pageNumber="87">H.</emphasis>
|
||
(
|
||
<emphasis id="B975491EFFB8F77DF731FAD0EBEBD54A" box="[1390,1427,1340,1366]" italics="true" pageId="3" pageNumber="87">Th.</emphasis>
|
||
)
|
||
<emphasis id="B975491EFFB8F77DF2C8FA8CEE95D566" box="[151,237,1376,1402]" italics="true" pageId="3" pageNumber="87">tricolor</emphasis>
|
||
Kazantsev, 2000
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C01EE8FFFB8F77DF3BEFA8DEDEAD567" authority="Kazantsev, 2015" authorityName="Kazantsev" authorityYear="2015" box="[481,914,1376,1403]" class="Insecta" family="Lycidae" genus="Helcophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="species" species="sichuanensis" subGenus="Thibeteros">
|
||
<emphasis id="B975491EFFB8F77DF3BEFA8DEF85D566" box="[481,509,1377,1402]" italics="true" pageId="3" pageNumber="87">H.</emphasis>
|
||
(
|
||
<emphasis id="B975491EFFB8F77DF053FA8CEC49D566" box="[524,561,1376,1402]" italics="true" pageId="3" pageNumber="87">Th.</emphasis>
|
||
)
|
||
<emphasis id="B975491EFFB8F77DF01EFA8CECA8D566" box="[577,720,1376,1402]" italics="true" pageId="3" pageNumber="87">sichuanensis</emphasis>
|
||
<bibRefCitation id="EF90E8FDFFB8F77DF089FA8DEDEAD567" author="Kazantsev, S. V." box="[726,914,1376,1403]" pageId="3" pageNumber="87" pagination="383 - 389" refId="ref2761" refString="Kazantsev, S. V. (2015) New taxa of Helcophorus Fairmaire, 1891, with a key to species of the genus (Coleoptera: Lycidae). In: Hartmann, M. & Weipert, J. (Eds.), Biodiversitat und Naturausstattung im Himalaya. Band 5. Verein der Freunde & Forderer des Naturkundemuseums Erfurt e. V., Erfurt, pp. 383 - 389." type="book chapter" year="2015">Kazantsev, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, by the noticeably more transverse elytral cells between the primary costae and narrowly notched scutellum with broadly rounded distal lobes (
|
||
<figureCitation id="133A8989FFB8F77DF6E8FA68EA85D582" box="[1207,1277,1412,1438]" captionStart="FIGURE 1" captionStartId="2.[151,250,1555,1579]" captionTargetBox="[156,1432,728,1529]" captionTargetId="figure-137@2.[156,1432,728,1529]" captionTargetPageId="2" captionText="FIGURE 1. General view of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498052" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="3" pageNumber="87">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="133A8989FFB8F77DF755FA68EB62D582" box="[1290,1306,1412,1438]" captionStart="FIGURE 2" captionStartId="3.[151,250,1126,1150]" captionTargetBox="[161,1426,181,1101]" captionTargetId="figure-24@3.[161,1426,181,1101]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum and the base of elytra of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498067" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="3" pageNumber="87">2</figureCitation>
|
||
), compared to less transverse elytral cells and broadly notched scutellum with narrowed distal lobes in the extant species of the subgenus (
|
||
<figureCitation id="133A8989FFB8F77DF34FFA20EF2FD5FA" box="[272,343,1484,1510]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1792,1816]" captionTargetBox="[219,1367,540,1759]" captionTargetId="figure-179@4.[204,1384,525,1769]" captionTargetPageId="4" captionText="FIGURES 3–4. General view of extant Helcophorus: 3 – H. (s. str.) miniatus; 4 – H. (Thibeteros) tricolor (after Kazantsev, 2015)." figureDoi="http://doi.org/10.5281/zenodo.6498076" httpUri="https://zenodo.org/record/6498076/files/figure.png" pageId="3" pageNumber="87">Fig. 4</figureCitation>
|
||
). It differs from the only other fossil
|
||
<taxonomicName id="4C01EE8FFFB8F77DF15BFA20ED89D5FA" box="[772,1009,1484,1510]" pageId="3" pageNumber="87">
|
||
<emphasis id="B975491EFFB8F77DF15BFA20EDEDD5FA" box="[772,917,1484,1510]" italics="true" pageId="3" pageNumber="87">Helcophorus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C01EE8FFFB8F77DF1A1FA21EAADD5FA" authorityName="Winkler" authorityYear="1987" box="[1022,1237,1484,1511]" class="Insecta" family="Lycidae" genus="Hiekeolycus" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="87" phylum="Arthropoda" rank="species" species="berendti">
|
||
<emphasis id="B975491EFFB8F77DF1A1FA21EA61D5FA" box="[1022,1049,1485,1510]" italics="true" pageId="3" pageNumber="87">H.</emphasis>
|
||
(s. str.)
|
||
<emphasis id="B975491EFFB8F77DF626FA20EAADD5FA" box="[1145,1237,1484,1510]" italics="true" pageId="3" pageNumber="87">berendti</emphasis>
|
||
</taxonomicName>
|
||
, by the distinctly sparser elytral vestiture, relatively larger eyes, and more transverse elytral cells (
|
||
<figureCitation id="133A8989FFB8F77DF643FA1CEA1BD616" box="[1052,1123,1520,1546]" captionStart="FIGURE 1" captionStartId="2.[151,250,1555,1579]" captionTargetBox="[156,1432,728,1529]" captionTargetId="figure-137@2.[156,1432,728,1529]" captionTargetPageId="2" captionText="FIGURE 1. General view of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498052" httpUri="https://zenodo.org/record/6498052/files/figure.png" pageId="3" pageNumber="87">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="133A8989FFB8F77DF62EFA1CEAFAD616" box="[1137,1154,1520,1546]" captionStart="FIGURE 2" captionStartId="3.[151,250,1126,1150]" captionTargetBox="[161,1426,181,1101]" captionTargetId="figure-24@3.[161,1426,181,1101]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum and the base of elytra of Helcophorus (Thibeteros) ekaterinae, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6498067" httpUri="https://zenodo.org/record/6498067/files/figure.png" pageId="3" pageNumber="87">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |