740 lines
103 KiB
XML
740 lines
103 KiB
XML
<document id="956F06C3F7C1C05E8AD54A28CBB37248" ID-DOI="10.5252/zoosystema2020v42a6" ID-GBIF-Dataset="69e88778-6ab9-44ad-bce0-bb84eb35391b" ID-ISSN="1638-9387" ID-Zenodo-Dep="3695831" ID-ZooBank="urn:lsid:zoobank.org:pub:C87A10FB-E817-4293-96FD-00C2EF82D371" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1583259932333" checkinUser="felipe" docAuthor="Theil, Emma Palacios & Felder, Darryl L." docDate="2020" docId="DE24878AF651D62F30D7C77B58DCFA71" docLanguage="en" docName="zoosystema42.6.85-103.pdf.imf" docOrigin="Zoosystema 42 (6)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Glassella Campos & Wicksten 1997" docType="treatment" docVersion="6" lastPageNumber="96" masterDocId="221DFFF2F65AD6223355C30D5C69FFBE" masterDocTitle="Phylogeny of the genus Pinnixa White, 1846 (Crustacea, Brachyura, Pinnotheridae) and allies inferred from mitochondrial and nuclear molecular markers, with generic reassignment of twenty-one species" masterLastPageNumber="103" masterPageNumber="85" pageNumber="94" updateTime="1698724363688" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9263432C56310D1FB2FBAB37B9B376A9">Phylogeny of the genus Pinnixa White, 1846 (Crustacea, Brachyura, Pinnotheridae) and allies inferred from mitochondrial and nuclear molecular markers, with generic reassignment of twenty-one species</mods:title>
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<mods:namePart id="C0BBB2E1827F2CBFBEEDEAF41438EF1D">Theil, Emma Palacios</mods:namePart>
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<mods:namePart id="0BEE5F041B6ADF2943603E41ECCD1F53">Felder, Darryl L.</mods:namePart>
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<mods:title id="50F31F76762AE1F186A2BBB72F7DE9BD">Zoosystema</mods:title>
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<mods:date id="0EA82E8F6B5E764382B63213CB650AC6">2020</mods:date>
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<treatment id="DE24878AF651D62F30D7C77B58DCFA71" ID-DOI="http://doi.org/10.5281/zenodo.3703640" ID-GBIF-Taxon="162382197" ID-Zenodo-Dep="3703640" LSID="urn:lsid:plazi:treatment:DE24878AF651D62F30D7C77B58DCFA71" httpUri="http://treatment.plazi.org/id/DE24878AF651D62F30D7C77B58DCFA71" lastPageId="13" lastPageNumber="96" pageId="11" pageNumber="94">
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<subSubSection id="1E976517F651D62930D7C77B5930FB2E" box="[898,1369,1142,1168]" pageId="11" pageNumber="94" type="nomenclature">
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<paragraph id="5632369CF651D62930D7C77B5930FB2E" blockId="11.[898,1369,1142,1168]" box="[898,1369,1142,1168]" pageId="11" pageNumber="94">
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<heading id="0D7A81F0F651D62930D7C77B5930FB2E" box="[898,1369,1142,1168]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="94" reason="2">
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Genus
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<taxonomicName id="918D4D1FF651D629309AC77B5930FB2E" ID-CoL="7PGXH" authority="Campos & Wicksten, 1997" authorityName="Campos & Wicksten" authorityYear="1997" box="[975,1369,1142,1168]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF651D629309AC77B5841FB2E" box="[975,1064,1142,1168]" italics="true" pageId="11" pageNumber="94">Glassella</emphasis>
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<bibRefCitation id="321C4B6DF651D629377AC77B5930FB2E" author="Campos & Wicksten" box="[1071,1369,1142,1168]" firstAuthor="Campos" pageId="11" pageNumber="94" pagination="69 - 73" refId="ref17299" refString="CAMPOS E. & WICKSTEN M. K. 1997. - A new genus for the Central American crab Pinnixa costaricana Wicksten, 1982 (Crustacea: Brachyura: Pinnotheridae). Proceedings of the Biological Society of Washington 110 (1): 69 - 73." type="journal article" year="1997">Campos & Wicksten, 1997</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="1E976517F651D6293078C7A358D4FB78" box="[813,1213,1198,1222]" pageId="11" pageNumber="94" type="reference_group">
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<paragraph id="5632369CF651D6293078C7A358D4FB78" blockId="11.[813,1213,1198,1222]" box="[813,1213,1198,1222]" pageId="11" pageNumber="94">
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<treatmentCitationGroup id="769D11B2F651D6293078C7A358D4FB78" box="[813,1213,1198,1222]" pageId="11" pageNumber="94">
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<taxonomicName id="918D4D1FF651D6293078C7A358DEFB78" ID-CoL="7PGXH" authority="Campos & Wicksten, 1997: 69" authorityName="Campos & Wicksten" authorityPageNumber="69" authorityYear="1997" box="[813,1207,1198,1222]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF651D6293078C7A35F17FB78" box="[813,894,1198,1222]" italics="true" pageId="11" pageNumber="94">Glassella</emphasis>
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<treatmentCitation id="D72C108DF651D62930D1C7A358DEFB78" author="CAMPOS E. & WICKSTEN M. K." box="[900,1207,1198,1222]" page="69" pageId="11" pageNumber="94" year="1997">
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<bibRefCitation id="321C4B6DF651D62930D1C7A358DEFB78" author="Campos & Wicksten" box="[900,1207,1198,1222]" firstAuthor="Campos" pageId="11" pageNumber="94" pagination="69 - 73" refId="ref17299" refString="CAMPOS E. & WICKSTEN M. K. 1997. - A new genus for the Central American crab Pinnixa costaricana Wicksten, 1982 (Crustacea: Brachyura: Pinnotheridae). Proceedings of the Biological Society of Washington 110 (1): 69 - 73." type="journal article" year="1997">Campos & Wicksten, 1997: 69</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="1E976517F651D6293078C7D3594AF926" pageId="11" pageNumber="94" type="materials_examined">
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<paragraph id="5632369CF651D6293078C7D35FF2FA95" blockId="11.[813,1455,1246,1323]" pageId="11" pageNumber="94">
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<collectionCode id="309CAE59F651D6293078C7D35FC5FB4A" box="[813,940,1246,1270]" pageId="11" pageNumber="94">
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<typeStatus id="8936883EF651D6293078C7D35F34FB4D" box="[813,861,1246,1270]" pageId="11" pageNumber="94">TYPE</typeStatus>
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SPECIES
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</collectionCode>
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. —
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<taxonomicName id="918D4D1FF651D629308CC7D35925FB49" authority="(Wicksten, 1982)" baseAuthorityName="Wicksten" baseAuthorityYear="1982" box="[985,1356,1246,1271]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="94" phylum="Arthropoda" rank="species" species="costaricana">
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<emphasis id="64F9EA8EF651D629308CC7D358F1FB48" box="[985,1176,1246,1270]" italics="true" pageId="11" pageNumber="94">Glassella costaricana</emphasis>
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(Wicksten, 1982)
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</taxonomicName>
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[
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<taxonomicName id="918D4D1FF651D6293609C7D259CEFB48" authorityName="White" authorityYear="1846" box="[1372,1447,1247,1270]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="94" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF651D6293609C7D259CEFB48" box="[1372,1447,1247,1270]" italics="true" pageId="11" pageNumber="94">Pinnixa</emphasis>
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</taxonomicName>
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] assigned by monotypy when genus was erected (
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<bibRefCitation id="321C4B6DF651D6293654C7F45FE4FA95" author="Campos & Wicksten" firstAuthor="Campos" pageId="11" pageNumber="94" pagination="69 - 73" refId="ref17299" refString="CAMPOS E. & WICKSTEN M. K. 1997. - A new genus for the Central American crab Pinnixa costaricana Wicksten, 1982 (Crustacea: Brachyura: Pinnotheridae). Proceedings of the Biological Society of Washington 110 (1): 69 - 73." type="journal article" year="1997">Campos & Wicksten 1997</bibRefCitation>
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).
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</paragraph>
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<paragraph id="5632369CF651D6293078C64F594AF926" blockId="11.[813,1456,1346,1688]" pageId="11" pageNumber="94">
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<collectionCode id="309CAE59F651D6293078C64F582BFAE6" box="[813,1090,1346,1370]" pageId="11" pageNumber="94">ORIGINAL DESCRIPTION BY</collectionCode>
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<bibRefCitation id="321C4B6DF651D629371EC64F59ECFAE4" author="CAMPOS & WICKSTEN" box="[1099,1413,1346,1370]" firstAuthor="CAMPOS" pageId="11" pageNumber="94" pagination="69 - 73" refId="ref17299" refString="CAMPOS E. & WICKSTEN M. K. 1997. - A new genus for the Central American crab Pinnixa costaricana Wicksten, 1982 (Crustacea: Brachyura: Pinnotheridae). Proceedings of the Biological Society of Washington 110 (1): 69 - 73." type="journal article" year="1997">CAMPOS & WICKSTEN (1997)</bibRefCitation>
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. — “Carapace suboblong, dorsal surface pockmarked, wider than long, integument firm, regions not defined; cardiac ridge lacking; front truncated, with shallow median sulcus. MXP3 [= third maxilliped] with ischium-merus pyriform, fused, separated by faint line and distal margin truncated; palp as long as ischium-merus, 3-segmented, dactylus small, digitiform, inserted sub-distally on inner face of conical propodus; carpus stout, longer than combined length of propodus and dactylus; exopod with median lobe on outer margin, flagellum 2-segmented. WLl-4 [= walking leg] pockmarked, relative length 3> 2> 1> 4, WL3 considerably the longest. Abdomen of female with 6 somites and telson free, widest at third somite; tapering from fourth somite to triangular telson. Male unknown.”
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</paragraph>
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</subSubSection>
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<subSubSection id="1E976517F651D62E3078C5BD5C8FFE4D" lastPageId="12" lastPageNumber="95" pageId="11" pageNumber="94" type="diagnosis">
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<paragraph id="5632369CF651D62E3078C5BD5D94FE14" blockId="11.[813,1456,1712,2027]" lastBlockId="12.[132,775,217,426]" lastPageId="12" lastPageNumber="95" pageId="11" pageNumber="94">
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DIAGNOSIS. — (Modified from
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<bibRefCitation id="321C4B6DF651D6293730C5BD5907F976" author="Campos & Wicksten" box="[1125,1390,1712,1736]" firstAuthor="Campos" pageId="11" pageNumber="94" pagination="69 - 73" refId="ref17299" refString="CAMPOS E. & WICKSTEN M. K. 1997. - A new genus for the Central American crab Pinnixa costaricana Wicksten, 1982 (Crustacea: Brachyura: Pinnotheridae). Proceedings of the Biological Society of Washington 110 (1): 69 - 73." type="journal article" year="1997">Campos & Wicksten 1997</bibRefCitation>
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). Carapace transversely oblong, wider than long, dorsal surface smooth, punctate, integument firm, regions poorly defined, sometimes with blunt ridge across posterior portion of carapace, ridge not extending entirely across carapace. Third maxilliped with ischiomerus pyriform or subtrapezoidal, fused, sometimes separated by faint line; palp as long as or longer than ischiomerus, three-segmented; dactylus sometimes (
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<taxonomicName id="918D4D1FF651D62930F6C464580BF83F" baseAuthorityName="Wicksten" baseAuthorityYear="1982" box="[931,1122,1897,1921]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="94" phylum="Arthropoda" rank="species" species="costaricana">
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<emphasis id="64F9EA8EF651D62930F6C464580BF83F" box="[931,1122,1897,1921]" italics="true" pageId="11" pageNumber="94">Glassella costaricana</emphasis>
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</taxonomicName>
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) very small, inserting sub-distally on inner face of conical propodus, typically large, nearly as long as ischiomerus, inserting near base of propodus. Chelipeds small, subcylindrical to weakly compressed, setose; palm typically with one or more longitudinal ridges or lines of tubercles or setae on outer surface; fingers slender, dactylus superior margin typically with row of long setae. Walking pereopod articles heavy, stout, often marginally tuberculate or dentate, relative lengths P4> P3 ≥ P2> P5. Male pleon terminally broad, lobate to weakly polygonal; third pleonal somite typically bearing gonopodal plate or sheath extending between or against gonopods (
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<figureCitation id="CEB62A19F656D62E315CC2505E29FECB" box="[521,576,349,373]" captionStart="FIG" captionStartId="10.[133,144,1805,1823]" captionTargetBox="[300,1285,234,1752]" captionTargetId="figure@10.[296,1293,216,1758]" captionTargetPageId="10" captionText="FIG. 2. — Morphological characters of the type species of Pinnixa White,1846 s.s., P.cylindrica (Say,1818), along with those for five molecularly segregated genera formerly treated in Pinnixa s.l.: A-D, Pinnixa cylindrica: A, male dorsal view;B, male cheliped;C, third maxilliped (adapted from Rathbun 1918:160 fig.99a);D, male pleon; E-G: Glassella costaricana (Wicksten, 1982): E, female holotype dorsal view; F, female cheliped; G, third maxilliped (adapted from Campos & Wicksten 1997: fig. 1, fig. 2c, a, with permission from Allen Press); H, I, Glassella faxoni (Rathbun,1918) n. comb.: H, third maxilliped; I, male pleon (adapted from Rathbun 1918:133 fig. 77b, a); J-M: Rathbunixa sayana (Stimpson, 1960) n. comb.: J, male dorsal view;K, male cheliped; L, third maxilliped; M, male pleon (L, M adapted from Rathbun 1918:158 fig. 98a, b); N-Q: Sayixa monodactyla (Say, 1818) n. comb., male (ULLZ 8713, Fort Pierce, FL, USA); N, dorsal view; O, cheliped; P, third maxilliped; Q, pleon; R, T, U, Scleroplax granulata Rathbun, 1893; R, female carapace and pereopods 2-5; T, third maxilliped; U, male pleon (R, T adapted from Campos 2006:fig. 1a-c, with permission from Magnolia Press; U, adapted from Rathbun 1918:171 fig. 109a); S, Scleroplax littoralis (Holmes, 1894) n. comb., female and male chelipeds (adapted from Rathbun 1918:146 fig. 89a, b); V-Y, Tubicolixa chaetopterana (Stimpson, 1860) n. comb.: V, male dorsal view; W, female and male chelipeds; X, third maxilliped; Y, male pleon (X, Y, adapted from Rathbun 1918:152 fig. 94a, b)." figureDoi="http://doi.org/10.5281/zenodo.3695835" httpUri="https://zenodo.org/record/3695835/files/figure.png" pageId="12" pageNumber="95">Fig. 2</figureCitation>
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E-I). Male gonopods heavy, stout, terminally forming sharp angle, spinose tip, or distally to laterally directed corneous filament.
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</paragraph>
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<paragraph id="5632369CF656D62E33D1C2CC5C8FFE4D" blockId="12.[132,776,449,659]" pageId="12" pageNumber="95">
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INCLUDED SPECIES. —
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<taxonomicName id="918D4D1FF656D62E323DC2CC5ECFFE67" authority="(Glassell, 1935)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Glassell" baseAuthorityYear="1935" box="[360,678,449,473]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="abbotti" status="comb. nov.">
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<emphasis id="64F9EA8EF656D62E323DC2CC5E6DFE67" box="[360,516,449,473]" italics="true" pageId="12" pageNumber="95">Glassella abbotti</emphasis>
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(
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<bibRefCitation id="321C4B6DF656D62E3147C2CC5EC9FE67" author="GLASSELL S. A." box="[530,672,449,473]" pageId="12" pageNumber="95" pagination="13 - 14" refId="ref17605" refString="GLASSELL S. A. 1935 a. - Three new species of Pinnixa from the Gulf of California. Transactions of the San Diego Society of Natural History 8 (5): 13 - 14." type="journal article" year="1935">Glassell, 1935</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="7FCA57F5F656D62E31FBC2CF5F61FE67" box="[686,776,449,473]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
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[
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<taxonomicName id="918D4D1FF656D62E33D9C2D05CBEFE4A" authorityName="White" authorityYear="1846" box="[140,215,477,500]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
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<emphasis id="64F9EA8EF656D62E33D9C2D05CBEFE4A" box="[140,215,477,500]" italics="true" pageId="12" pageNumber="95">Pinnixa</emphasis>
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</taxonomicName>
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];
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</paragraph>
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</subSubSection>
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<subSubSection id="1E976517F656D62E33D1C2FB5EC8FDB0" box="[132,673,502,527]" pageId="12" pageNumber="95" type="description">
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<paragraph id="5632369CF656D62E33D1C2FB5EC8FDB0" blockId="12.[132,776,449,659]" box="[132,673,502,527]" pageId="12" pageNumber="95">
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<taxonomicName id="918D4D1FF656D62E33D1C2FB5DB5FDB1" authority="(Rathbun, 1922)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[132,476,502,527]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="arenicola" status="comb. nov.">
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<emphasis id="64F9EA8EF656D62E33D1C2FB5D46FDB0" box="[132,303,502,526]" italics="true" pageId="12" pageNumber="95">Glassella arenicola</emphasis>
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(Rathbun, 1922)
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</taxonomicName>
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<taxonomicNameLabel id="7FCA57F5F656D62E32B7C2FA5E53FDB0" box="[482,570,502,526]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
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[
|
||
<taxonomicName id="918D4D1FF656D62E3112C2FA5EFBFDB0" authorityName="White" authorityYear="1846" box="[583,658,503,526]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E3112C2FA5EFBFDB0" box="[583,658,503,526]" italics="true" pageId="12" pageNumber="95">Pinnixa</emphasis>
|
||
</taxonomicName>
|
||
];
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E976517F656D62E33D1C11C5F6EFD2C" pageId="12" pageNumber="95" type="reference_group">
|
||
<paragraph id="5632369CF656D62E33D1C11C5EC3FDFD" blockId="12.[132,776,449,659]" pageId="12" pageNumber="95">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C11C5DABFD97" ID-CoL="84GVC" authority="(Rathbun, 1918)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[132,450,529,553]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="faxoni" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C11C5D7CFD97" box="[132,277,529,553]" italics="true" pageId="12" pageNumber="95">Glassella faxoni</emphasis>
|
||
(
|
||
<bibRefCitation id="321C4B6DF656D62E3276C11C5DD2FD97" author="Rathbun" box="[291,443,529,553]" firstAuthor="Rathbun" pageId="12" pageNumber="95" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E329DC11C5E49FD97" box="[456,544,529,553]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
[
|
||
<taxonomicName id="918D4D1FF656D62E3178C11F5EA9FD97" ID-CoL="55aa1440-3f7e-4125-88b5-971723e568be" authorityName="McDermott" authorityYear="2014" box="[557,704,530,553]" class="Malacostraca" family="Pinnotheridae" genus="Laminapinnixa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E3178C11F5EA9FD97" box="[557,704,530,553]" italics="true" pageId="12" pageNumber="95">Laminapinnixa</emphasis>
|
||
</taxonomicName>
|
||
];
|
||
<taxonomicName id="918D4D1FF656D62E33D1C1265D8CFDFD" ID-CoL="84GVD" authority="(Rathtbun, 1918)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathtbun" baseAuthorityYear="1918" box="[132,485,555,579]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="floridana" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C1265D58FDFD" box="[132,305,555,579]" italics="true" pageId="12" pageNumber="95">Glassella floridana</emphasis>
|
||
(Rathtbun, 1918)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E32BEC1215E2AFDFD" box="[491,579,555,579]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
[
|
||
<taxonomicName id="918D4D1FF656D62E3105C1215EF2FDFD" ID-CoL="7PS58" authorityName="White" authorityYear="1846" box="[592,667,556,579]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E3105C1215EF2FDFD" box="[592,667,556,579]" italics="true" pageId="12" pageNumber="95">Pinnixa</emphasis>
|
||
</taxonomicName>
|
||
];
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62E33D1C14B5ECBFDC6" blockId="12.[132,776,449,659]" pageId="12" pageNumber="95">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C14B5E6AFDE0" ID-CoL="84GVH" authority="(McDermott, 2014)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[132,515,582,606]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="miamiensis" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C14B5D57FDE3" box="[132,318,582,606]" italics="true" pageId="12" pageNumber="95">Glassella miamiensis</emphasis>
|
||
(
|
||
<bibRefCitation id="321C4B6DF656D62E321CC14B5D95FDE0" author="McDermott" box="[329,508,582,606]" firstAuthor="McDermott" pageId="12" pageNumber="95" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott, 2014</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E315DC14B5E35FDE0" box="[520,604,582,606]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
[
|
||
<taxonomicName id="918D4D1FF656D62E313DC14B5E91FDE3" ID-CoL="55aa1440-3f7e-4125-88b5-971723e568be" authorityName="McDermott" authorityYear="2014" box="[616,760,582,605]" class="Malacostraca" family="Pinnotheridae" genus="Laminapinnixa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E313DC14B5E91FDE3" box="[616,760,582,605]" italics="true" pageId="12" pageNumber="95">Laminapinnixa</emphasis>
|
||
</taxonomicName>
|
||
];
|
||
<taxonomicName id="918D4D1FF656D62E33D1C16D5DB5FDC6" ID-CoL="84GVF" authority="(Wass, 1968)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Wass" baseAuthorityYear="1968" box="[132,476,608,632]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="leptosynaptae" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C16D5D3CFDC6" box="[132,341,608,632]" italics="true" pageId="12" pageNumber="95">Glassella leptosynaptae</emphasis>
|
||
(Wass, 1968)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E32B6C16C5E52FDC6" box="[483,571,608,632]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
[
|
||
<taxonomicName id="918D4D1FF656D62E311DC16C5EFAFDC6" ID-CoL="7PS58" authorityName="White" authorityYear="1846" box="[584,659,609,632]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E311DC16C5EFAFDC6" box="[584,659,609,632]" italics="true" pageId="12" pageNumber="95">Pinnixa</emphasis>
|
||
</taxonomicName>
|
||
];
|
||
</paragraph>
|
||
<caption id="02F26614F656D62E3078C162582CFCF3" ID-DOI="http://doi.org/10.5281/zenodo.3695839" ID-Zenodo-Dep="3695839" httpUri="https://zenodo.org/record/3695839/files/figure.png" pageId="12" pageNumber="95" startId="12.[813,824,623,641]" targetBox="[823,1438,225,564]" targetPageId="12">
|
||
<paragraph id="5632369CF656D62E3078C162582CFCF3" blockId="12.[813,1457,623,845]" pageId="12" pageNumber="95">
|
||
FIG. 4. — Reproduced thumbnail sketches of male gonopods and gonopodal plates on lost USNM specimens of
|
||
<taxonomicName id="918D4D1FF656D62E3700C18B5936FD26" ID-CoL="84GVC" authority="(Rathbun, 1918)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[1109,1375,646,664]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="faxoni" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E3700C18B58BCFD26" box="[1109,1237,646,664]" italics="true" pageId="12" pageNumber="95">Glassella faxoni</emphasis>
|
||
(
|
||
<bibRefCitation id="321C4B6DF656D62E37B5C18B5932FD26" author="Rathbun" box="[1248,1371,646,664]" firstAuthor="Rathbun" pageId="12" pageNumber="95" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3630C18B59D8FD26" box="[1381,1457,646,664]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="64F9EA8EF656D62E3064C1905F3DFD10" bold="true" box="[817,852,668,687]" pageId="12" pageNumber="95">
|
||
A
|
||
<emphasis id="64F9EA8EF656D62E306BC1915F2CFD10" bold="true" box="[830,837,668,686]" italics="true" pageId="12" pageNumber="95">-</emphasis>
|
||
C
|
||
</emphasis>
|
||
), by R. H. Gore, 1978-1979;
|
||
<taxonomicName id="918D4D1FF656D62E3719C19158F1FD10" ID-CoL="84GVC" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[1100,1176,668,686]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="faxoni" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E3719C19158F1FD10" box="[1100,1176,668,686]" italics="true" pageId="12" pageNumber="95">G. faxoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E37CBC1915882FD10" box="[1182,1259,668,686]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="64F9EA8EF656D62E37A0C190596AFD11" bold="true" box="[1269,1283,669,687]" pageId="12" pageNumber="95">D</emphasis>
|
||
), und
|
||
<taxonomicName id="918D4D1FF656D62E366FC191586BFD7B" ID-CoL="84GVK" authority="(Rathbun, 1922)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="vanderhorsti" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E366FC1915F1EFD7B" italics="true" pageId="12" pageNumber="95">Glassella vanderhorsti</emphasis>
|
||
(Rathbun, 1922)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E375DC1BE583EFD7B" box="[1032,1111,691,709]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
; (
|
||
<emphasis id="64F9EA8EF656D62E3736C1BE5806FD7B" bold="true" box="[1123,1135,691,709]" pageId="12" pageNumber="95">E</emphasis>
|
||
,
|
||
<emphasis id="64F9EA8EF656D62E372EC1BE58EFFD7B" bold="true" box="[1147,1158,691,709]" pageId="12" pageNumber="95">F</emphasis>
|
||
) by D. L. Felder, 1979
|
||
<emphasis id="64F9EA8EF656D62E3614C1BE5921FD7B" box="[1345,1352,691,709]" italics="true" pageId="12" pageNumber="95">-</emphasis>
|
||
1982.
|
||
<emphasis id="64F9EA8EF656D62E3628C1BE59E3FD7B" bold="true" box="[1405,1418,691,709]" pageId="12" pageNumber="95">A</emphasis>
|
||
, left gonopod, pleonal surface, paratype, USNM 23436;
|
||
<emphasis id="64F9EA8EF656D62E37B6C1C75899FD62" bold="true" box="[1251,1264,714,732]" pageId="12" pageNumber="95">B</emphasis>
|
||
, left gonopod, pleonal surface, holotype, USNM 7639;
|
||
<emphasis id="64F9EA8EF656D62E376AC1ED5824FD4C" bold="true" box="[1087,1101,736,754]" pageId="12" pageNumber="95">C</emphasis>
|
||
, gonopods and gonopodal plate, pleonal surface, holotype USNM 7639;
|
||
<emphasis id="64F9EA8EF656D62E3769C1FA5823FCB7" bold="true" box="[1084,1098,759,777]" pageId="12" pageNumber="95">D</emphasis>
|
||
, gonopods and gonopodal plate, pleonal surface,holotype,USNM 7639;
|
||
<emphasis id="64F9EA8EF656D62E3766C0035856FC9E" bold="true" box="[1075,1087,782,800]" pageId="12" pageNumber="95">E</emphasis>
|
||
, gonopodal plate, pleonal surface,topotypic material, USNM 56903;
|
||
<emphasis id="64F9EA8EF656D62E30ACC028586DFC89" bold="true" box="[1017,1028,805,823]" pageId="12" pageNumber="95">F</emphasis>
|
||
, gonopods and gonopodal plate, pleonal surface, topotypic material, USNM 56903.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5632369CF656D62E33D1C1765F6EFD2C" blockId="12.[132,776,449,659]" box="[132,775,635,659]" pageId="12" pageNumber="95">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C1765D93FD2D" ID-CoL="84GVK" authority="(Rathbun, 1922)" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[132,506,635,659]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="vanderhorsti" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C1765D26FD2D" box="[132,335,635,659]" italics="true" pageId="12" pageNumber="95">Glassella vanderhorsti</emphasis>
|
||
(Rathbun, 1922)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3155C1765E31FD2D" box="[512,600,635,659]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
[
|
||
<taxonomicName id="918D4D1FF656D62E3130C1765E91FD2C" ID-CoL="55aa1440-3f7e-4125-88b5-971723e568be" authorityName="McDermott" authorityYear="2014" box="[613,760,635,658]" class="Malacostraca" family="Pinnotheridae" genus="Laminapinnixa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E3130C1765E91FD2C" box="[613,760,635,658]" italics="true" pageId="12" pageNumber="95">Laminapinnixa</emphasis>
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E976517F656D62E33D1C1A75D18FC92" pageId="12" pageNumber="95" type="materials_examined">
|
||
<paragraph id="5632369CF656D62E33D1C1A75DE8FD49" blockId="12.[132,775,682,1315]" pageId="12" pageNumber="95">
|
||
MATERIAL EXAMINED. — In addition to the material included in the phylogenetic analyses (
|
||
<tableCitation id="1B0F0327F656D62E32C1C1C85D89FD63" box="[404,480,709,733]" captionStart="TABLE" captionStartId="5.[132,143,221,239]" captionText="TABLE 1. — Pinnotherid specimens used in phylogenetic analyses of the genus Pinnixa White, 1846 s.l. and allies. Names shown are in accord with Pinnixa s.l. prior to revisions in present paper. See “Material and Methods” for museum abbreviations." httpUri="http://table.plazi.org/id/02F26614F65FD62733D1C3D05F1CFEB8" pageId="12" pageNumber="95" tableUuid="02F26614F65FD62733D1C3D05F1CFEB8">Table 1</tableCitation>
|
||
) the following samples were available for examination:
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62E33D1C1F75D18FC92" blockId="12.[132,775,682,1315]" pageId="12" pageNumber="95">
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C1F75D07FC92" ID-GBIF-Occurrence="2573304528" collectionCode="ULLZ" country="Mexico" location="Mexico" pageId="12" pageNumber="95" specimenCode="ULLZ 5619, ULLZ 7392" specimenCount="1" stateProvince="Mexico">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C1F75D74FCAC" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Glassell" baseAuthorityYear="1935" box="[132,285,762,786]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="abbotti" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C1F75D74FCAC" box="[132,285,762,786]" italics="true" pageId="12" pageNumber="95">Glassella abbotti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3277C1F75D11FCAC" box="[290,376,762,786]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
—
|
||
<specimenCode id="062B9EE7F656D62E32CEC1F75E7BFCAC" box="[411,530,762,786]" pageId="12" pageNumber="95">ULLZ 5619</specimenCode>
|
||
(26),
|
||
<specimenCode id="062B9EE7F656D62E3119C1F75EAAFCAC" box="[588,707,762,786]" pageId="12" pageNumber="95">ULLZ 7392</specimenCode>
|
||
(Bahía de los Ángeles,
|
||
<collectingRegion id="9449F87EF656D62E324FC0195D0EFC92" box="[282,359,788,812]" country="Mexico" name="Mexico" pageId="12" pageNumber="95">Mexico</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E976517F656D62E33D1C0225D9CFA9D" pageId="12" pageNumber="95" type="description">
|
||
<paragraph id="5632369CF656D62E33D1C0225E75FCC2" blockId="12.[132,775,682,1315]" pageId="12" pageNumber="95">
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C0225D15FCDF" ID-GBIF-Occurrence="2573304530" collectionCode="ULLZ" country="Curaçao" location="Curacao" pageId="12" pageNumber="95" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C0225D44FCF9" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[132,301,815,839]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="arenicola" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C0225D44FCF9" box="[132,301,815,839]" italics="true" pageId="12" pageNumber="95">Glassella arenicola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3267C0225DE1FCF9" box="[306,392,815,839]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
— NCBN-ZMA De242240 (
|
||
<typeStatus id="8936883EF656D62E31FFC0225F6AFCF9" box="[682,771,815,839]" pageId="12" pageNumber="95" type="holotype">holotype</typeStatus>
|
||
, Spanish Harbor,
|
||
<collectingCountry id="2E9A760CF656D62E3275C0445D1BFCDF" box="[288,370,841,865]" name="Curaçao" pageId="12" pageNumber="95" value="Curaçao">Curaçao</collectingCountry>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="E6E53CC1F656D62E32D5C0475E2BFCDF" ID-GBIF-Occurrence="2573304561" box="[384,578,841,865]" collectionCode="ULLZ" country="Aruba" location="Aruba" pageId="12" pageNumber="95" specimenCode="ULLZ 6070" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="062B9EE7F656D62E32D5C0475D9BFCDF" box="[384,498,841,865]" pageId="12" pageNumber="95">ULLZ 6070</specimenCode>
|
||
(
|
||
<collectingCountry id="2E9A760CF656D62E32A8C0445E55FCDF" box="[509,572,841,865]" name="Aruba" pageId="12" pageNumber="95" value="Aruba">Aruba</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E6E53CC1F656D62E311FC0475CA9FCC5" ID-GBIF-Occurrence="2573304532" collectionCode="ULLZ" country="Puerto Rico" location="Puerto Rico" pageId="12" pageNumber="95" specimenCode="ULLZ 8989" specimenCount="1">
|
||
<specimenCode id="062B9EE7F656D62E311FC0475ED5FCDF" box="[586,700,841,865]" pageId="12" pageNumber="95">ULLZ 8989</specimenCode>
|
||
(
|
||
<collectingCountry id="2E9A760CF656D62E3192C0445CDCFCC5" name="Puerto Rico" pageId="12" pageNumber="95" value="Puerto Rico">Puerto Rico</collectingCountry>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="E6E53CC1F656D62E3393C0695E70FCC2" ID-GBIF-Occurrence="2573304533" box="[198,537,867,892]" collectionCode="FL, USA" country="Puerto Rico" pageId="12" pageNumber="95" specimenCode="ULLZ 9248" specimenCount="1">
|
||
<specimenCode id="062B9EE7F656D62E3393C0695D56FCC2" box="[198,319,868,892]" pageId="12" pageNumber="95">ULLZ 9248</specimenCode>
|
||
(Ft. Pierce,
|
||
<collectionCode id="309CAE59F656D62E32E3C0695DBDFCC5" box="[438,468,868,891]" pageId="12" pageNumber="95">FL</collectionCode>
|
||
,
|
||
<collectionCode id="309CAE59F656D62E328AC0695E7BFCC2" box="[479,530,868,892]" pageId="12" pageNumber="95">USA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62E33D1C0735D53FC5B" blockId="12.[132,775,682,1315]" pageId="12" pageNumber="95">
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C0735EA8FC28" ID-GBIF-Occurrence="2573304538" box="[132,705,894,918]" collectionCode="ULLZ" country="Iceland" location="Panama" pageId="12" pageNumber="95" specimenCount="1" stateProvince="Panama">
|
||
<emphasis id="64F9EA8EF656D62E33D1C0735D01FC28" box="[132,360,894,918]" italics="true" pageId="12" pageNumber="95">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C0735D2FFC28" baseAuthorityName="Wicksten" baseAuthorityYear="1982" box="[132,326,894,918]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="costaricana">Glassella costaricana</taxonomicName>
|
||
. —
|
||
</emphasis>
|
||
UF 18960 (
|
||
<collectingCountry id="2E9A760CF656D62E32B7C0735E6DFC28" box="[482,516,894,918]" name="Iceland" pageId="12" pageNumber="95" value="Iceland">Isla</collectingCountry>
|
||
Culebra,
|
||
<collectingRegion id="9449F87EF656D62E3130C0735EDFFC28" box="[613,694,894,918]" country="Panama" name="Panama" pageId="12" pageNumber="95">Panama</collectingRegion>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C0955F6EFC0E" ID-GBIF-Occurrence="2573304551" box="[132,775,920,945]" collectionCode="ULLZ" country="Panama" location="Campeche" pageId="12" pageNumber="95" specimenCode="ULLZ 14837" specimenCount="1" stateProvince="Campeche">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C0955D71FC0E" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[132,280,920,944]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="faxoni" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C0955D71FC0E" box="[132,280,920,944]" italics="true" pageId="12" pageNumber="95">Glassella faxoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3275C0945D13FC0E" box="[288,378,920,944]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
—
|
||
<specimenCode id="062B9EE7F656D62E32F7C0945E40FC0F" box="[418,553,921,945]" pageId="12" pageNumber="95">ULLZ 14837</specimenCode>
|
||
(
|
||
<collectingRegion id="9449F87EF656D62E3162C0955ECAFC0E" box="[567,675,920,944]" country="Mexico" name="Campeche" pageId="12" pageNumber="95">Campeche</collectingRegion>
|
||
,
|
||
<collectingRegion id="9449F87EF656D62E31FAC0955E95FC0E" box="[687,764,920,944]" country="Mexico" name="Mexico" pageId="12" pageNumber="95">Mexico</collectingRegion>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C0BE5E7FFC75" ID-GBIF-Occurrence="2573304546" box="[132,534,947,971]" collectionCode="ULLZ" country="Iceland" location="Margarita" pageId="12" pageNumber="95" specimenCode="ULLZ 14030" specimenCount="1" stateProvince="Campeche">
|
||
<specimenCode id="062B9EE7F656D62E33D1C0BE5D61FC75" box="[132,264,947,971]" pageId="12" pageNumber="95">ULLZ 14030</specimenCode>
|
||
(
|
||
<collectingCountry id="2E9A760CF656D62E3246C0BE5D5CFC75" box="[275,309,947,971]" name="Iceland" pageId="12" pageNumber="95" value="Iceland">Isla</collectingCountry>
|
||
<location id="53526047F656D62E326EC0BE5DF7FC75" LSID="urn:lsid:plazi:treatment:DE24878AF651D62F30D7C77B58DCFA71:53526047F656D62E326EC0BE5DF7FC75" box="[315,414,947,971]" country="Iceland" name="Margarita" pageId="12" pageNumber="95" stateProvince="Campeche">Margarita</location>
|
||
,
|
||
<collectingCountry id="2E9A760CF656D62E32F3C0BE5E65FC75" box="[422,524,947,971]" name="Venezuela" pageId="12" pageNumber="95" value="Venezuela">Venezuela</collectingCountry>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="E6E53CC1F656D62E314EC0BE5D5CFC5B" ID-GBIF-Occurrence="2573304576" collectionCode="ULLZ" country="Venezuela" location="Venezuela" pageId="12" pageNumber="95" specimenCode="ULLZ 14098" specimenCount="1">
|
||
<specimenCode id="062B9EE7F656D62E314EC0BE5EF7FC75" box="[539,670,947,971]" pageId="12" pageNumber="95">ULLZ 14098</specimenCode>
|
||
(2) (Punta Elvira,
|
||
<collectingCountry id="2E9A760CF656D62E339CC0C05D46FC5B" box="[201,303,973,997]" name="Venezuela" pageId="12" pageNumber="95" value="Venezuela">Venezuela</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62E33D1C0E55D91FBD4" blockId="12.[132,775,682,1315]" pageId="12" pageNumber="95">
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C0E55D44FBA4" ID-GBIF-Occurrence="2573304552" collectionCode="FL, USA" country="United States of America" pageId="12" pageNumber="95" specimenCode="ULLZ 5649, ULLZ 17733" specimenCount="1">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C0E55D5AFBBE" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathtbun" baseAuthorityYear="1918" box="[132,307,1000,1024]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="floridana" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C0E55D5AFBBE" box="[132,307,1000,1024]" italics="true" pageId="12" pageNumber="95">Glassella floridana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E326EC0E55DFCFBBE" box="[315,405,1000,1024]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
—
|
||
<specimenCode id="062B9EE7F656D62E32EBC0E55E53FBBE" box="[446,570,1000,1024]" pageId="12" pageNumber="95">ULLZ 5649</specimenCode>
|
||
,
|
||
<specimenCode id="062B9EE7F656D62E3113C0E55EA4FBBE" box="[582,717,1000,1024]" pageId="12" pageNumber="95">ULLZ 17733</specimenCode>
|
||
(Fort Pierce,
|
||
<collectionCode id="309CAE59F656D62E339EC70E5C80FBA4" box="[203,233,1027,1050]" pageId="12" pageNumber="95">FL</collectionCode>
|
||
,
|
||
<collectionCode id="309CAE59F656D62E33A6C70F5D4FFBA4" box="[243,294,1026,1050]" pageId="12" pageNumber="95">USA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="E6E53CC1F656D62E3262C70E5ED7FBA4" ID-GBIF-Occurrence="2573304527" box="[311,702,1026,1051]" collectionCode="FL, USA" country="United States of America" pageId="12" pageNumber="95" specimenCode="ULLZ 13888" specimenCount="1">
|
||
<specimenCode id="062B9EE7F656D62E3262C70E5DD2FBA4" box="[311,443,1026,1050]" pageId="12" pageNumber="95">ULLZ 13888</specimenCode>
|
||
(Content Keys,
|
||
<collectionCode id="309CAE59F656D62E3109C70E5E13FBA4" box="[604,634,1027,1050]" pageId="12" pageNumber="95">FL</collectionCode>
|
||
,
|
||
<collectionCode id="309CAE59F656D62E31D1C70F5EDEFBA4" box="[644,695,1026,1050]" pageId="12" pageNumber="95">USA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="E6E53CC1F656D62E319DC70E5DBFFB8B" ID-GBIF-Occurrence="2573304575" collectionCode="FL, USA" country="United States of America" pageId="12" pageNumber="95" specimenCode="ULLZ 14903" specimenCount="1">
|
||
<specimenCode id="062B9EE7F656D62E319DC70E5CA9FB8B" pageId="12" pageNumber="95">ULLZ 14903</specimenCode>
|
||
(2) (Alligator Point,
|
||
<collectionCode id="309CAE59F656D62E322EC7105DF1FB8A" box="[379,408,1053,1076]" pageId="12" pageNumber="95">FL</collectionCode>
|
||
,
|
||
<collectionCode id="309CAE59F656D62E32CAC7105DA6FB8B" box="[415,463,1053,1077]" pageId="12" pageNumber="95">USA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="E6E53CC1F656D62E328BC7105ED5FBF1" ID-GBIF-Occurrence="2573304547" collectionCode="FL, USA" country="United States of America" pageId="12" pageNumber="95" specimenCode="ULLZ 13096, ULLZ 14038, ULLZ 14181, ULLZ 15010" specimenCount="1">
|
||
<specimenCode id="062B9EE7F656D62E328BC7105E35FB8B" box="[478,604,1053,1077]" pageId="12" pageNumber="95">ULLZ 13096</specimenCode>
|
||
,
|
||
<specimenCode id="062B9EE7F656D62E3131C7105E88FB8B" box="[612,737,1053,1077]" pageId="12" pageNumber="95">ULLZ 14038</specimenCode>
|
||
(4),
|
||
<specimenCode id="062B9EE7F656D62E33D1C7355D67FBF1" box="[132,270,1079,1103]" pageId="12" pageNumber="95">ULLZ 14181</specimenCode>
|
||
,
|
||
<specimenCode id="062B9EE7F656D62E324FC7355DC8FBF1" box="[282,417,1079,1103]" pageId="12" pageNumber="95">ULLZ 15010</specimenCode>
|
||
(St. Joseph’s Bay,
|
||
<collectionCode id="309CAE59F656D62E3102C7355E1FFBF1" box="[599,630,1080,1103]" pageId="12" pageNumber="95">FL</collectionCode>
|
||
,
|
||
<collectionCode id="309CAE59F656D62E31D4C73A5EDDFBF1" box="[641,692,1079,1103]" pageId="12" pageNumber="95">USA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="E6E53CC1F656D62E3192C7355D9CFBD4" ID-GBIF-Occurrence="2573304566" collectionCode="ULLZ" country="United States of America" location="Mexico" pageId="12" pageNumber="95" specimenCode="ULLZ 17469" specimenCount="1" stateProvince="Mexico">
|
||
<specimenCode id="062B9EE7F656D62E3192C7355CAAFBD4" pageId="12" pageNumber="95">ULLZ 17469</specimenCode>
|
||
(northeastern Gulf of
|
||
<collectingRegion id="9449F87EF656D62E32F4C75F5D87FBD4" box="[417,494,1106,1130]" country="Mexico" name="Mexico" pageId="12" pageNumber="95">Mexico</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62E33D1C7615CABFB50" blockId="12.[132,775,682,1315]" pageId="12" pageNumber="95">
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C7615EBBFB07" ID-GBIF-Occurrence="2573304581" collectionCode="FL, USA" country="United States of America" pageId="12" pageNumber="95" specimenCode="ULLZ 5724, ULLZ 7398, ULLZ 11709, ULLZ 13338, ULLZ 14003, ULLZ 14011, ULLZ 14138" specimenCount="1" stateProvince="Florida">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C7615D5FFB3A" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[132,310,1132,1156]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="miamiensis" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C7615D5FFB3A" box="[132,310,1132,1156]" italics="true" pageId="12" pageNumber="95">Glassella miamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E326FC7605DE2FB3A" box="[314,395,1132,1156]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
—
|
||
<specimenCode id="062B9EE7F656D62E32FEC7605E74FB3A" box="[427,541,1132,1156]" pageId="12" pageNumber="95">ULLZ 5724</specimenCode>
|
||
, MNHN-IU-2017-9364 = former
|
||
<specimenCode id="062B9EE7F656D62E33B3C78A5D0BFB21" box="[230,354,1159,1183]" pageId="12" pageNumber="95">ULLZ 7398</specimenCode>
|
||
(2),
|
||
<specimenCode id="062B9EE7F656D62E32C2C78A5E4BFB21" box="[407,546,1159,1183]" pageId="12" pageNumber="95">ULLZ 11709</specimenCode>
|
||
,
|
||
<specimenCode id="062B9EE7F656D62E317AC78A5ED3FB21" box="[559,698,1159,1183]" pageId="12" pageNumber="95">ULLZ 13338</specimenCode>
|
||
,
|
||
<specimenCode id="062B9EE7F656D62E3192C78A5CAFFB07" pageId="12" pageNumber="95">ULLZ 14003</specimenCode>
|
||
,
|
||
<specimenCode id="062B9EE7F656D62E3385C7AF5D31FB07" box="[208,344,1185,1209]" pageId="12" pageNumber="95">ULLZ 14011</specimenCode>
|
||
,
|
||
<specimenCode id="062B9EE7F656D62E3237C7AF5D8FFB07" box="[354,486,1185,1209]" pageId="12" pageNumber="95">ULLZ 14138</specimenCode>
|
||
(Fort Pierce,
|
||
<collectionCode id="309CAE59F656D62E3139C7AF5EE3FB07" box="[620,650,1186,1209]" pageId="12" pageNumber="95">FL</collectionCode>
|
||
,
|
||
<collectionCode id="309CAE59F656D62E31C1C7AC5EAEFB07" box="[660,711,1185,1209]" pageId="12" pageNumber="95">USA</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C7B65CD7FB50" ID-GBIF-Occurrence="2573304571" collectionCode="FL, USA" country="United States of America" location="Florida" pageId="12" pageNumber="95" specimenCode="ULLZ 14834" specimenCount="1" stateProvince="Florida">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C7B65D3CFB6D" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Wass" baseAuthorityYear="1968" box="[132,341,1211,1235]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="leptosynaptae" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C7B65D3CFB6D" box="[132,341,1211,1235]" italics="true" pageId="12" pageNumber="95">Glassella leptosynaptae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E320EC7B15DDAFB6D" box="[347,435,1211,1235]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
—
|
||
<specimenCode id="062B9EE7F656D62E328DC7B15E35FB6A" box="[472,604,1212,1236]" pageId="12" pageNumber="95">ULLZ 14834</specimenCode>
|
||
(
|
||
<collectingRegion id="9449F87EF656D62E313CC7B65ED9FB6D" box="[617,688,1211,1235]" country="United States of America" name="Florida" pageId="12" pageNumber="95">Florida</collectingRegion>
|
||
Bay,
|
||
<collectionCode id="309CAE59F656D62E31B0C7B15F6AFB6D" box="[741,771,1212,1235]" pageId="12" pageNumber="95">FL</collectionCode>
|
||
,
|
||
<collectionCode id="309CAE59F656D62E33D1C7DB5CDEFB50" box="[132,183,1238,1262]" pageId="12" pageNumber="95">USA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62E33D1C7FD5D9CFA9D" blockId="12.[132,775,682,1315]" pageId="12" pageNumber="95">
|
||
<materialsCitation id="E6E53CC1F656D62E33D1C7FD5D9BFA9D" ID-GBIF-Occurrence="2573304543" country="Curaçao" location="Curacao" pageId="12" pageNumber="95" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="918D4D1FF656D62E33D1C7FD5D25FAB6" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[132,332,1264,1288]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="vanderhorsti" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E33D1C7FD5D25FAB6" box="[132,332,1264,1288]" italics="true" pageId="12" pageNumber="95">Glassella vanderhorsti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3204C7FC5DCEFAB6" box="[337,423,1264,1288]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
— NCBN-ZMA.C RUS.D 242234 (
|
||
<typeStatus id="8936883EF656D62E33DEC6065C8CFA9D" box="[139,229,1291,1315]" pageId="12" pageNumber="95" type="holotype">holotype</typeStatus>
|
||
, Spanish Harbor,
|
||
<collectingCountry id="2E9A760CF656D62E32C1C6065D83FA9D" box="[404,490,1291,1315]" name="Curaçao" pageId="12" pageNumber="95" value="Curaçao">Curaçao</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E976517F656D62F33D1C65858DCFA71" lastPageId="13" lastPageNumber="96" pageId="12" pageNumber="95" type="discussion">
|
||
<paragraph id="5632369CF656D62E33D1C6585C81FAD3" blockId="12.[132,775,1365,2030]" box="[132,232,1365,1391]" pageId="12" pageNumber="95">REMARKS</paragraph>
|
||
<paragraph id="5632369CF656D62E33D1C6785E78F950" blockId="12.[132,775,1365,2030]" pageId="12" pageNumber="95">
|
||
The morphological similarities among species transferred to this genus have in most cases been noted previously (
|
||
<bibRefCitation id="321C4B6DF656D62E31FDC6985CD4FA71" author="Rathbun" firstAuthor="Rathbun" pageId="12" pageNumber="95" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">Rathbun 1918</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="321C4B6DF656D62E3398C6B85D6FFA71" author="Rathbun" box="[205,262,1461,1487]" firstAuthor="Rathbun" pageId="12" pageNumber="95" pagination="13 - 21" refId="ref18632" refString="RATHBUN M. J. 1924. - Brachyuran crabs collected at Curacao. Bijdragen Tot de Kennis Der Faun van Curacao. Resultaten Eener Reis van Dr. C. J. van Der Horst in 1920, 23, 13 - 21." type="journal article" year="1924">1924</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="321C4B6DF656D62E3243C6B85DDFFA6E" author="Glassell" box="[278,438,1461,1488]" firstAuthor="Glassell" pageId="12" pageNumber="95" pagination="13 - 14" refId="ref17605" refString="GLASSELL S. A. 1935 a. - Three new species of Pinnixa from the Gulf of California. Transactions of the San Diego Society of Natural History 8 (5): 13 - 14." type="journal article" year="1935" yearSuffix="a">Glassell 1935a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="321C4B6DF656D62E3293C6B85EE4FA71" author="McDermott" box="[454,653,1461,1487]" firstAuthor="McDermott" pageId="12" pageNumber="95" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott 2014</bibRefCitation>
|
||
), although their resemblance to
|
||
<taxonomicName id="918D4D1FF656D62E320AC6D85DD1FA51" authorityName="Campos & Wicksten" authorityYear="1997" box="[351,440,1493,1519]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E320AC6D85DD1FA51" box="[351,440,1493,1519]" italics="true" pageId="12" pageNumber="95">Glassella</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<bibRefCitation id="321C4B6DF656D62E328DC6D85F6EFA51" author="Campos & Wicksten" box="[472,775,1493,1519]" firstAuthor="Campos" pageId="12" pageNumber="95" pagination="69 - 73" refId="ref17299" refString="CAMPOS E. & WICKSTEN M. K. 1997. - A new genus for the Central American crab Pinnixa costaricana Wicksten, 1982 (Crustacea: Brachyura: Pinnotheridae). Proceedings of the Biological Society of Washington 110 (1): 69 - 73." type="journal article" year="1997">Campos & Wicksten (1997)</bibRefCitation>
|
||
has likely remained unnoticed because of the weight given to differences in the third maxilliped (
|
||
<figureCitation id="CEB62A19F656D62E314BC5195EF1F991" box="[542,664,1556,1583]" captionStart="FIG" captionStartId="10.[133,144,1805,1823]" captionTargetBox="[300,1285,234,1752]" captionTargetId="figure@10.[296,1293,216,1758]" captionTargetPageId="10" captionText="FIG. 2. — Morphological characters of the type species of Pinnixa White,1846 s.s., P.cylindrica (Say,1818), along with those for five molecularly segregated genera formerly treated in Pinnixa s.l.: A-D, Pinnixa cylindrica: A, male dorsal view;B, male cheliped;C, third maxilliped (adapted from Rathbun 1918:160 fig.99a);D, male pleon; E-G: Glassella costaricana (Wicksten, 1982): E, female holotype dorsal view; F, female cheliped; G, third maxilliped (adapted from Campos & Wicksten 1997: fig. 1, fig. 2c, a, with permission from Allen Press); H, I, Glassella faxoni (Rathbun,1918) n. comb.: H, third maxilliped; I, male pleon (adapted from Rathbun 1918:133 fig. 77b, a); J-M: Rathbunixa sayana (Stimpson, 1960) n. comb.: J, male dorsal view;K, male cheliped; L, third maxilliped; M, male pleon (L, M adapted from Rathbun 1918:158 fig. 98a, b); N-Q: Sayixa monodactyla (Say, 1818) n. comb., male (ULLZ 8713, Fort Pierce, FL, USA); N, dorsal view; O, cheliped; P, third maxilliped; Q, pleon; R, T, U, Scleroplax granulata Rathbun, 1893; R, female carapace and pereopods 2-5; T, third maxilliped; U, male pleon (R, T adapted from Campos 2006:fig. 1a-c, with permission from Magnolia Press; U, adapted from Rathbun 1918:171 fig. 109a); S, Scleroplax littoralis (Holmes, 1894) n. comb., female and male chelipeds (adapted from Rathbun 1918:146 fig. 89a, b); V-Y, Tubicolixa chaetopterana (Stimpson, 1860) n. comb.: V, male dorsal view; W, female and male chelipeds; X, third maxilliped; Y, male pleon (X, Y, adapted from Rathbun 1918:152 fig. 94a, b)." figureDoi="http://doi.org/10.5281/zenodo.3695835" httpUri="https://zenodo.org/record/3695835/files/figure.png" pageId="12" pageNumber="95">Fig. 2G, H</figureCitation>
|
||
), often an important character in pinnotherids. However, there are other cases among pinnotherid genera for which striking variation in the third maxilliped has been observed among closely related species. In
|
||
<taxonomicName id="918D4D1FF656D62E33AFC5995E2EF910" authority="Campos, 1990" authorityName="Campos" authorityYear="1990" box="[250,583,1684,1710]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E33AFC5995DC8F910" box="[250,417,1684,1710]" italics="true" pageId="12" pageNumber="95">Calyptraeotheres</emphasis>
|
||
Campos, 1990
|
||
</taxonomicName>
|
||
some species lack a third maxilliped dactylus, whereas in others it is present (Hernández-Ávila &
|
||
<bibRefCitation id="321C4B6DF656D62E323DC5D95E68F950" author="Campos" box="[360,513,1748,1774]" firstAuthor="Campos" pageId="12" pageNumber="95" pagination="33 - 41" refId="ref17258" refString="CAMPOS E. 2006. - Systematics of the genus Scleroplax Rathbun, 1893 (Crustacea: Brachyura: Pinnotheridae). Zootaxa 1344: 33 - 41. https: // doi. org / 10.11646 / zootaxa. 1344.1.3" type="journal article" year="2006">Campos 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62E33CEC5F95840FA11" blockId="12.[132,775,1365,2030]" lastBlockId="12.[813,1456,919,2030]" pageId="12" pageNumber="95">
|
||
The genus
|
||
<taxonomicName id="918D4D1FF656D62E324CC5F85DADF8B0" authorityName="McDermott" authorityYear="2014" box="[281,452,1781,1806]" class="Malacostraca" family="Pinnotheridae" genus="Laminapinnixa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF656D62E324CC5F85DADF8B0" box="[281,452,1781,1806]" italics="true" pageId="12" pageNumber="95">Laminapinnixa</emphasis>
|
||
</taxonomicName>
|
||
was erected by
|
||
<bibRefCitation id="321C4B6DF656D62E312BC5F95CA7F890" author="McDermott" firstAuthor="McDermott" pageId="12" pageNumber="95" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott (2014)</bibRefCitation>
|
||
to accomodate
|
||
<taxonomicName id="918D4D1FF656D62E322CC4195E60F893" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[377,521,1812,1837]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="miamiensis">
|
||
<emphasis id="64F9EA8EF656D62E322CC4195E60F893" box="[377,521,1812,1837]" italics="true" pageId="12" pageNumber="95">L. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
, a newly described species for populations that we and colleagues had long regarded on morphological evidence as closely related to
|
||
<taxonomicName id="918D4D1FF656D62E31FFC45E5CAEF833" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="faxoni" status="n. comb. and G.">
|
||
<emphasis id="64F9EA8EF656D62E31FFC45E5CAEF833" italics="true" pageId="12" pageNumber="95">Glassella faxoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3384C4795DE7F830" box="[209,398,1907,1934]" pageId="12" pageNumber="95" rank="genus">
|
||
n. comb. and
|
||
<taxonomicName id="918D4D1FF656D62E3227C4795DE7F830" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[370,398,1908,1934]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="vanderhorsti" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E3227C4795DE7F830" box="[370,398,1908,1934]" italics="true" pageId="12" pageNumber="95">G.</emphasis>
|
||
</taxonomicName>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="64F9EA8EF656D62E32C2C47E5E75F833" box="[407,540,1907,1933]" italics="true" pageId="12" pageNumber="95">vanderhorsti</emphasis>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3170C4795EE2F833" box="[549,651,1907,1934]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
The genus was diagnosed in part by the presence in males of a plate (therein termed “abdominal plate” = our gonopodal plate) derived from the pleon (posterior to the gonopods), which in the description and discussion by
|
||
<bibRefCitation id="321C4B6DF656D62E37E0C09A59E0FC0F" author="McDermott" box="[1205,1417,919,945]" firstAuthor="McDermott" pageId="12" pageNumber="95" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott (2014)</bibRefCitation>
|
||
are portrayed as a structure that must be “lifted” to expose the gonopods during copulation. However, this structure is in all unmanipulated examples of
|
||
<taxonomicName id="918D4D1FF656D62E37D5C0FA597CFBAE" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[1152,1301,1015,1041]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="miamiensis" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E37D5C0FA597CFBAE" box="[1152,1301,1015,1041]" italics="true" pageId="12" pageNumber="95">G. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3649C0FA5914FBAE" box="[1308,1405,1014,1041]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
that we have examined positioned to extend from its more posterior origin obliquely between the gonopods (derived from the first pleonal somite) and then anterior to them, where it usually broadens terminally and separates the gonopods from the sternum when the pleon is flexed (as shown clearly by
|
||
<bibRefCitation id="321C4B6DF656D62E3078C7BB5F91FB6E" author="McDermott" box="[813,1016,1206,1232]" firstAuthor="McDermott" pageId="12" pageNumber="95" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott, 2014</bibRefCitation>
|
||
: fig. 7B). This would appear to not necessarily separate them from exposure for copulation when the pleon is extended (or, as they appear when illustrated with the pleon removed,
|
||
<figureCitation id="CEB62A19F656D62E3754C6185839FA8E" box="[1025,1104,1301,1328]" captionStart="FIG" captionStartId="16.[132,143,1170,1188]" captionTargetBox="[147,1441,233,1097]" captionTargetId="figure@16.[132,1455,198,1108]" captionTargetPageId="16" captionText="FIG. 5. — Illustrations of selected type and topotypic materials for Glassella spp., by Smithsonian artists MEH, Charisse Baker, and Jack Schroeder, predating loss of subject specimens: A, G. faxoni (Rathbun, 1918) n. comb., habitus,male paratype, cw 10.1 mm, USNM lot 7639; B, G. faxoni n. comb., left chela external surface, male holotype, cw 11.0 mm, USNM lot 7639; C, G. miamiensis (McDermott, 2014) n. comb., habitus, male, cw 4.7 mm, HBOI uncatalogued specimen from Indian River,Florida;D, G. floridana (Rathtbun,1918) n.comb.,habitus,male holotype,cw 6.7 mm, USNM 6996;E, G.vanderhorsti (Rathbun,1922) n. comb., habitus, male holotype,cw 6.0 mm, Zoological Museum Amsterdam,now Netherlands Naturalis Biodiversity Center;F, G. vanderhorsti n. comb.,gonopodal plate pleonal surface, male holotype, cw 6.0 mm, Amsterdam Museum." figureDoi="http://doi.org/10.5281/zenodo.3695841" httpUri="https://zenodo.org/record/3695841/files/figure.png" pageId="12" pageNumber="95">Fig. 5F</figureCitation>
|
||
). The broadened terminus of the plate in at least one related species can also perhaps move from anterior or posterior of the gonopods, provided the gonopods are flexed laterally, though this seems unlikely in
|
||
<taxonomicName id="918D4D1FF656D62E3078C6985FA8FA11" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[813,961,1429,1455]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="miamiensis" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E3078C6985FA8FA11" box="[813,961,1429,1455]" italics="true" pageId="12" pageNumber="95">G. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E309DC69B5840FA11" box="[968,1065,1429,1456]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="5632369CF656D62F3016C6B85D9EFE8A" blockId="12.[813,1456,919,2030]" lastBlockId="13.[132,776,217,2030]" lastPageId="13" lastPageNumber="96" pageId="12" pageNumber="95">
|
||
In the course of describing and investigating potential relationships of
|
||
<taxonomicName id="918D4D1FF656D62E30E4C6D8582CFA50" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[945,1093,1493,1519]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="95" phylum="Arthropoda" rank="species" species="miamiensis" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF656D62E30E4C6D8582CFA50" box="[945,1093,1493,1519]" italics="true" pageId="12" pageNumber="95">G. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F656D62E3719C6D858C5FA51" box="[1100,1196,1493,1519]" pageId="12" pageNumber="95" rank="species">n. comb.</taxonomicNameLabel>
|
||
and seeking evidence of other species having this gonopodal plate,
|
||
<bibRefCitation id="321C4B6DF656D62E37B4C6F859D9F9B1" author="McDermott" box="[1249,1456,1525,1551]" firstAuthor="McDermott" pageId="12" pageNumber="95" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott (2014)</bibRefCitation>
|
||
noted the unfortunate loss of
|
||
<typeStatus id="8936883EF656D62E3734C51858F1F991" box="[1121,1176,1557,1583]" pageId="12" pageNumber="95">types</typeStatus>
|
||
for potentially related species to which he had wished to make comparisons. By way of further explanation, most of these were among 98 pinnotherid specimens permanently lost to science when destroyed by the U.S. Postal Service, owing to the mishandling of a loan return shipment by a borrower in 2006. While this loss has also limited our own comparative efforts, one of us (DLF) and his late colleague Robert H. Gore had independently examined
|
||
<typeStatus id="8936883EF656D62E30CEC4195FBBF890" box="[923,978,1812,1838]" pageId="12" pageNumber="95">types</typeStatus>
|
||
and other now lost materials decades ago, at the time making rough-sketches of selected structures, several of which are herewith directly reproduced given the void they fill (
|
||
<figureCitation id="CEB62A19F656D62E3003C47E5FFFF833" box="[854,918,1907,1933]" captionStart="FIG" captionStartId="12.[813,824,623,641]" captionTargetBox="[823,1438,225,564]" captionTargetId="figure@12.[810,1453,219,579]" captionTargetPageId="12" captionText="FIG. 4. — Reproduced thumbnail sketches of male gonopods and gonopodal plates on lost USNM specimens of Glassella faxoni (Rathbun, 1918) n. comb. (A-C), by R. H. Gore, 1978-1979; G. faxoni n. comb. (D), und Glassella vanderhorsti (Rathbun, 1922) n. comb.; (E, F) by D. L. Felder, 1979-1982. A, left gonopod, pleonal surface, paratype, USNM 23436; B, left gonopod, pleonal surface, holotype, USNM 7639; C, gonopods and gonopodal plate, pleonal surface, holotype USNM 7639; D, gonopods and gonopodal plate, pleonal surface,holotype,USNM 7639;E, gonopodal plate, pleonal surface,topotypic material, USNM 56903; F, gonopods and gonopodal plate, pleonal surface, topotypic material, USNM 56903." figureDoi="http://doi.org/10.5281/zenodo.3695839" httpUri="https://zenodo.org/record/3695839/files/figure.png" pageId="12" pageNumber="95">Fig. 4</figureCitation>
|
||
). In addition, high-quality, previously unpublished illustrations by several Smithsonian Institution illustrators, contracted by Robert Gore or the late Waldo Schmitt (the latter for a never-published manuscript by W. L. Schmitt and E. S. Davidson), are in some cases annotated so as to be clearly identifiable with the now-lost
|
||
<typeStatus id="8936883EF657D62F315BC3F75E2CFEAA" box="[526,581,250,276]" pageId="13" pageNumber="96">types</typeStatus>
|
||
or other materials on which they were based (
|
||
<figureCitation id="CEB62A19F657D62F32FCC2145D83FE8A" box="[425,490,281,308]" captionStart="FIG" captionStartId="16.[132,143,1170,1188]" captionTargetBox="[147,1441,233,1097]" captionTargetId="figure@16.[132,1455,198,1108]" captionTargetPageId="16" captionText="FIG. 5. — Illustrations of selected type and topotypic materials for Glassella spp., by Smithsonian artists MEH, Charisse Baker, and Jack Schroeder, predating loss of subject specimens: A, G. faxoni (Rathbun, 1918) n. comb., habitus,male paratype, cw 10.1 mm, USNM lot 7639; B, G. faxoni n. comb., left chela external surface, male holotype, cw 11.0 mm, USNM lot 7639; C, G. miamiensis (McDermott, 2014) n. comb., habitus, male, cw 4.7 mm, HBOI uncatalogued specimen from Indian River,Florida;D, G. floridana (Rathtbun,1918) n.comb.,habitus,male holotype,cw 6.7 mm, USNM 6996;E, G.vanderhorsti (Rathbun,1922) n. comb., habitus, male holotype,cw 6.0 mm, Zoological Museum Amsterdam,now Netherlands Naturalis Biodiversity Center;F, G. vanderhorsti n. comb.,gonopodal plate pleonal surface, male holotype, cw 6.0 mm, Amsterdam Museum." figureDoi="http://doi.org/10.5281/zenodo.3695841" httpUri="https://zenodo.org/record/3695841/files/figure.png" pageId="13" pageNumber="96">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5632369CF657D62F33CEC2345E28FAF1" blockId="13.[132,776,217,2030]" pageId="13" pageNumber="96">
|
||
From this evidence, it is clear that both
|
||
<taxonomicName id="918D4D1FF657D62F3115C2345EC8FEED" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[576,673,313,339]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="faxoni" status="n. comb. and g.">
|
||
<emphasis id="64F9EA8EF657D62F3115C2345EC8FEED" box="[576,673,313,339]" italics="true" pageId="13" pageNumber="96">G. faxoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F657D62F31F2C2345CBBFECD" pageId="13" pageNumber="96" rank="genus">
|
||
n. comb. and
|
||
<taxonomicName id="918D4D1FF657D62F33E3C2545CBBFECD" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[182,210,345,371]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="vanderhorsti">
|
||
<emphasis id="64F9EA8EF657D62F33E3C2545CBBFECD" box="[182,210,345,371]" italics="true" pageId="13" pageNumber="96">G.</emphasis>
|
||
</taxonomicName>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="64F9EA8EF657D62F338EC2545D37FECD" box="[219,350,345,371]" italics="true" pageId="13" pageNumber="96">vanderhorsti</emphasis>
|
||
have forms of the gonopodal plate in mature males that we regard to be homologs of that in
|
||
<taxonomicName id="918D4D1FF657D62F3197C2745CB2FE0C" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="miamiensis">
|
||
<emphasis id="64F9EA8EF657D62F3197C2745CB2FE0C" italics="true" pageId="13" pageNumber="96">G. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
, in addition to their sharing a number of other characters that group them with
|
||
<taxonomicName id="918D4D1FF657D62F32EFC2B45E38FE6C" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[442,593,441,467]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="miamiensis">
|
||
<emphasis id="64F9EA8EF657D62F32EFC2B45E38FE6C" box="[442,593,441,467]" italics="true" pageId="13" pageNumber="96">G. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
and its herewith assigned congeners. The illustrated male gonopodal plate for
|
||
<taxonomicName id="918D4D1FF657D62F33FFC2F55DE7FDAC" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[170,398,504,530]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="vanderhorsti">
|
||
<emphasis id="64F9EA8EF657D62F33FFC2F55DE7FDAC" box="[170,398,504,530]" italics="true" pageId="13" pageNumber="96">Glassella vanderhorsti</emphasis>
|
||
</taxonomicName>
|
||
, published by
|
||
<bibRefCitation id="321C4B6DF657D62F317AC2F45F6EFDAD" author="McDermott" box="[559,775,504,531]" firstAuthor="McDermott" pageId="13" pageNumber="96" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott (2014)</bibRefCitation>
|
||
but reproduced here from the original figures with credit and voucher indicated (
|
||
<figureCitation id="CEB62A19F657D62F32D8C1355D88FDED" box="[397,481,568,595]" captionStart="FIG" captionStartId="16.[132,143,1170,1188]" captionTargetBox="[147,1441,233,1097]" captionTargetId="figure@16.[132,1455,198,1108]" captionTargetPageId="16" captionText="FIG. 5. — Illustrations of selected type and topotypic materials for Glassella spp., by Smithsonian artists MEH, Charisse Baker, and Jack Schroeder, predating loss of subject specimens: A, G. faxoni (Rathbun, 1918) n. comb., habitus,male paratype, cw 10.1 mm, USNM lot 7639; B, G. faxoni n. comb., left chela external surface, male holotype, cw 11.0 mm, USNM lot 7639; C, G. miamiensis (McDermott, 2014) n. comb., habitus, male, cw 4.7 mm, HBOI uncatalogued specimen from Indian River,Florida;D, G. floridana (Rathtbun,1918) n.comb.,habitus,male holotype,cw 6.7 mm, USNM 6996;E, G.vanderhorsti (Rathbun,1922) n. comb., habitus, male holotype,cw 6.0 mm, Zoological Museum Amsterdam,now Netherlands Naturalis Biodiversity Center;F, G. vanderhorsti n. comb.,gonopodal plate pleonal surface, male holotype, cw 6.0 mm, Amsterdam Museum." figureDoi="http://doi.org/10.5281/zenodo.3695841" httpUri="https://zenodo.org/record/3695841/files/figure.png" pageId="13" pageNumber="96">Fig. 5F</figureCitation>
|
||
), was in fact based on the Zoological Museum Amsterdam (now Naturalis Biodiversity Center, Netherlands)
|
||
<typeStatus id="8936883EF657D62F32CBC1755D96FD2C" box="[414,511,632,658]" pageId="13" pageNumber="96" type="holotype">holotype</typeStatus>
|
||
male (NCBN-ZMA.C RUS.D 242234). In a very similar but smaller topotypic male specimen (USNM 56903), also collected by van der Horst but now among the lost materials, the gonopodal plate was found to be somewhat longer but terminally very similar to the
|
||
<typeStatus id="8936883EF657D62F3277C01A5DEAFC8F" box="[290,387,791,817]" pageId="13" pageNumber="96" type="holotype">holotype</typeStatus>
|
||
male and like that of
|
||
<taxonomicName id="918D4D1FF657D62F3129C0155CF2FCEF" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="miamiensis">
|
||
<emphasis id="64F9EA8EF657D62F3129C0155CF2FCEF" italics="true" pageId="13" pageNumber="96">G. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
in that its terminal reaches were positioned anterior to the gonopods (
|
||
<figureCitation id="CEB62A19F657D62F327DC05A5DF3FCCC" box="[296,410,855,882]" captionStart="FIG" captionStartId="12.[813,824,623,641]" captionTargetBox="[823,1438,225,564]" captionTargetId="figure@12.[810,1453,219,579]" captionTargetPageId="12" captionText="FIG. 4. — Reproduced thumbnail sketches of male gonopods and gonopodal plates on lost USNM specimens of Glassella faxoni (Rathbun, 1918) n. comb. (A-C), by R. H. Gore, 1978-1979; G. faxoni n. comb. (D), und Glassella vanderhorsti (Rathbun, 1922) n. comb.; (E, F) by D. L. Felder, 1979-1982. A, left gonopod, pleonal surface, paratype, USNM 23436; B, left gonopod, pleonal surface, holotype, USNM 7639; C, gonopods and gonopodal plate, pleonal surface, holotype USNM 7639; D, gonopods and gonopodal plate, pleonal surface,holotype,USNM 7639;E, gonopodal plate, pleonal surface,topotypic material, USNM 56903; F, gonopods and gonopodal plate, pleonal surface, topotypic material, USNM 56903." figureDoi="http://doi.org/10.5281/zenodo.3695839" httpUri="https://zenodo.org/record/3695839/files/figure.png" pageId="13" pageNumber="96">Fig. 4E, F</figureCitation>
|
||
). Only in
|
||
<taxonomicName id="918D4D1FF657D62F315EC05A5E19FCCF" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[523,624,855,881]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="faxoni" status="comb. nov.">
|
||
<emphasis id="64F9EA8EF657D62F315EC05A5E19FCCF" box="[523,624,855,881]" italics="true" pageId="13" pageNumber="96">G. faxoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F657D62F3123C0555EB0FCCF" box="[630,729,855,882]" pageId="13" pageNumber="96" rank="species">n. comb.</taxonomicNameLabel>
|
||
was the male gonopodal plate found to be small enough to be positioned largely between the gonopods or to freely move its arched terminal lobe from anterior to posterior of the gonopods (
|
||
<figureCitation id="CEB62A19F657D62F33A8C0DB5D1DFC4F" box="[253,372,982,1009]" captionStart="FIG" captionStartId="12.[813,824,623,641]" captionTargetBox="[823,1438,225,564]" captionTargetId="figure@12.[810,1453,219,579]" captionTargetPageId="12" captionText="FIG. 4. — Reproduced thumbnail sketches of male gonopods and gonopodal plates on lost USNM specimens of Glassella faxoni (Rathbun, 1918) n. comb. (A-C), by R. H. Gore, 1978-1979; G. faxoni n. comb. (D), und Glassella vanderhorsti (Rathbun, 1922) n. comb.; (E, F) by D. L. Felder, 1979-1982. A, left gonopod, pleonal surface, paratype, USNM 23436; B, left gonopod, pleonal surface, holotype, USNM 7639; C, gonopods and gonopodal plate, pleonal surface, holotype USNM 7639; D, gonopods and gonopodal plate, pleonal surface,holotype,USNM 7639;E, gonopodal plate, pleonal surface,topotypic material, USNM 56903; F, gonopods and gonopodal plate, pleonal surface, topotypic material, USNM 56903." figureDoi="http://doi.org/10.5281/zenodo.3695839" httpUri="https://zenodo.org/record/3695839/files/figure.png" pageId="13" pageNumber="96">Fig. 4C, D</figureCitation>
|
||
). As evident, the cataloged specimens upon which our figures, sketches, and notes are based were the same as examined by
|
||
<bibRefCitation id="321C4B6DF657D62F32CDC71B5E2BFB8E" author="Rathbun" box="[408,578,1046,1072]" firstAuthor="Rathbun" pageId="13" pageNumber="96" pagination="123 - 184" refId="ref18600" refString="RATHBUN M. J. 1918. - The grapsoid crabs of America. Bulletin of the United States National Museum 103: 123 - 184, plates 54 - 66." type="journal article" year="1918">Rathbun (1918</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="321C4B6DF657D62F3105C71B5EE0FB8E" author="Rathbun" box="[592,649,1046,1072]" firstAuthor="Rathbun" pageId="13" pageNumber="96" pagination="13 - 21" refId="ref18632" refString="RATHBUN M. J. 1924. - Brachyuran crabs collected at Curacao. Bijdragen Tot de Kennis Der Faun van Curacao. Resultaten Eener Reis van Dr. C. J. van Der Horst in 1920, 23, 13 - 21." type="journal article" year="1924">1924</bibRefCitation>
|
||
). This confirms personal communications of E. S. Davidson regarding gonopodal plates in these species, as were mentioned by
|
||
<bibRefCitation id="321C4B6DF657D62F33D1C77A5D37FB2F" author="McDermott" box="[132,350,1142,1169]" firstAuthor="McDermott" pageId="13" pageNumber="96" pagination="367 - 390" refId="ref17961" refString="MCDERMOTT J. J. 2014. - A new genus and species of pinnotherid crab (Decapoda: Brachyura: Pinnotheridae: Pinnothereliinae) with a unique male abdominal appendage, and its symbiotic relationship with a sipunculan worm (Sipuncula) from Miami, Florida, USA. Proceedings of the Biological Society of Washington 127 (2): 367 - 390. https: // doi. org / 10.2988 / 0006 - 324 X- 127.2.367" type="journal article" year="2014">McDermott (2014)</bibRefCitation>
|
||
, who found the lack of Rathbun mentioning these plates in descriptions of
|
||
<taxonomicName id="918D4D1FF657D62F3166C79B5EF0FB0E" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[563,665,1174,1200]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="faxoni" status="n. comb. and g.">
|
||
<emphasis id="64F9EA8EF657D62F3166C79B5EF0FB0E" box="[563,665,1174,1200]" italics="true" pageId="13" pageNumber="96">G. faxoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCA57F5F657D62F31F6C79B5CB8FB6E" pageId="13" pageNumber="96" rank="genus">
|
||
n. comb. and
|
||
<taxonomicName id="918D4D1FF657D62F33E3C7BB5CB8FB6E" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[182,209,1206,1232]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="vanderhorsti">
|
||
<emphasis id="64F9EA8EF657D62F33E3C7BB5CB8FB6E" box="[182,209,1206,1232]" italics="true" pageId="13" pageNumber="96">G.</emphasis>
|
||
</taxonomicName>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="64F9EA8EF657D62F338CC7BB5D32FB6E" box="[217,347,1206,1232]" italics="true" pageId="13" pageNumber="96">vanderhorsti</emphasis>
|
||
as reason to question their congeneric assignment with
|
||
<taxonomicName id="918D4D1FF657D62F326EC7DB5DB8FB51" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[315,465,1238,1264]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="miamiensis">
|
||
<emphasis id="64F9EA8EF657D62F326EC7DB5DB8FB51" box="[315,465,1238,1264]" italics="true" pageId="13" pageNumber="96">G. miamiensis</emphasis>
|
||
</taxonomicName>
|
||
. We suspect that Rathbun at the time attached little importance to gonopods as characters, especially among pinnotherids in the early years while she was working primarily with a hand-lens.
|
||
</paragraph>
|
||
<paragraph id="5632369CF657D62F33CEC6585821FE0C" blockId="13.[132,776,217,2030]" lastBlockId="13.[813,1456,217,1487]" pageId="13" pageNumber="96">
|
||
While the male gonopodal plate, or some ramification of it, may prove to unite most if not all species that we here assign to
|
||
<taxonomicName id="918D4D1FF657D62F33BBC6985D20FA11" authorityName="Campos & Wicksten" authorityYear="1997" box="[238,329,1429,1455]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F33BBC6985D20FA11" box="[238,329,1429,1455]" italics="true" pageId="13" pageNumber="96">Glassella</emphasis>
|
||
</taxonomicName>
|
||
, our molecular phylogeny includes species in which it is at very least not reported to date, or for which intact male specimens are lacking. This remains the case for the generic
|
||
<typeStatus id="8936883EF657D62F320DC6F85DEEF9B1" box="[344,391,1525,1551]" pageId="13" pageNumber="96">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="918D4D1FF657D62F32B6C6F85E11F9B0" baseAuthorityName="Wicksten" baseAuthorityYear="1982" box="[483,632,1525,1551]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="costaricana">
|
||
<emphasis id="64F9EA8EF657D62F32B6C6F85E11F9B0" box="[483,632,1525,1551]" italics="true" pageId="13" pageNumber="96">G. costaricana</emphasis>
|
||
</taxonomicName>
|
||
, known only from a female at the time of description. Our phylogenetic analyses included two additional females, but we also attempted inclusion of tissues from a very small mutilated male specimen (UF 18960) that appeared to be this species. Its identity as
|
||
<taxonomicName id="918D4D1FF657D62F3275C5995DDFF910" baseAuthorityName="Wicksten" baseAuthorityYear="1982" box="[288,438,1684,1710]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="costaricana">
|
||
<emphasis id="64F9EA8EF657D62F3275C5995DDFF910" box="[288,438,1684,1710]" italics="true" pageId="13" pageNumber="96">G. costaricana</emphasis>
|
||
</taxonomicName>
|
||
was confirmed by clear match of its 16S mitochondrial sequence to those of the females, but it was not included in the final phylogenetic analysis for lack of additional sequence data. Unfortunately, damage to its pleon and sternum obliterated evidence that might have made obvious the presence or absence of a gonopodal plate, leaving that question unresolved. We strongly suspect that the “enclosing sheath” of the gonopod in
|
||
<taxonomicName id="918D4D1FF657D62F3117C4795EA8F833" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[578,705,1907,1934]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="arenicola">
|
||
<emphasis id="64F9EA8EF657D62F3117C4795EA8F833" box="[578,705,1907,1934]" italics="true" pageId="13" pageNumber="96">G. arenicola</emphasis>
|
||
</taxonomicName>
|
||
, as reported but only partially illustrated by
|
||
<bibRefCitation id="321C4B6DF657D62F317DC49E5F68F810" author="Thoma" box="[552,769,1939,1966]" etAl="et al." firstAuthor="Thoma" pageId="13" pageNumber="96" pagination="72 - 80" refId="ref19269" refString="THOMA B. P., HEARD W. R. & FELDER D. L. 2009. - Redescription of Pinnixa arenicola Rathbun, 1922 (Decapoda: Brachyura: Pinnotheridae) with new observations on its range and host. Proceedings of the Biological Society of Washington 122 (1): 72 - 80. https: // doi. org / 10.2988 / 08 - 25.1" type="journal article" year="2009">
|
||
Thoma
|
||
<emphasis id="64F9EA8EF657D62F3129C4995EC6F813" box="[636,687,1939,1965]" italics="true" pageId="13" pageNumber="96">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
, could represent yet another variation in this structure, and this species clearly groups with those that have the more obviously developed gonopodal plate.
|
||
<taxonomicName id="918D4D1FF657D62F3782C3D459C6FF4D" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="McDermott" baseAuthorityYear="2014" box="[1239,1455,217,243]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="miamiensis">
|
||
<emphasis id="64F9EA8EF657D62F3782C3D459C6FF4D" box="[1239,1455,217,243]" italics="true" pageId="13" pageNumber="96">Glassella miamiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="918D4D1FF657D62F300AC3F45FACFEAD" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1918" box="[863,965,249,275]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="faxoni">
|
||
<emphasis id="64F9EA8EF657D62F300AC3F45FACFEAD" box="[863,965,249,275]" italics="true" pageId="13" pageNumber="96">G. faxoni</emphasis>
|
||
</taxonomicName>
|
||
share the plate and are closely related both morphologically and genetically (
|
||
<figureCitation id="CEB62A19F657D62F37C3C21458BCFE8D" box="[1174,1237,281,307]" captionStart="FIG" captionStartId="8.[133,144,1919,1937]" captionTargetBox="[223,1405,231,1874]" captionTargetId="graphics@8.[290,1319,237,1858]" captionTargetPageId="8" captionText="FIG. 1. — Phylogeny for species of superfamily Pinnotheroidea De Haan,1833,emphasis on genus Pinnixa White, 1846 s.l. inferred from Randomized Accelerated Maximum Likelihood (RAxML) analysis of a 1445 bp long fragment concatenated from the mitochondrial complex 16S/tRNA-Leu/ NADH1 (776 bp),the mitochondrial 12S rRNA gene (340 bp) and the nuclear gene for the histone 3 subunit (327 bp). Bootstrap support values are shown at the nodes when higher than 50%. Collection number follows the species name to identify samples.For samples in the subfamily Pinnixinae Števčić,2005,abbreviations indicating geographic origin are defined in “Materials and Methods”. Species name combinations as shown are prior to revisions in present paper.Abbreviations as in Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.3695833" httpUri="https://zenodo.org/record/3695833/files/figure.png" pageId="13" pageNumber="96">Fig. 1</figureCitation>
|
||
), but we cannot yet determine if
|
||
<taxonomicName id="918D4D1FF657D62F30E0C2345834FEED" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Wass" baseAuthorityYear="1968" box="[949,1117,313,339]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="leptosynaptae">
|
||
<emphasis id="64F9EA8EF657D62F30E0C2345834FEED" box="[949,1117,313,339]" italics="true" pageId="13" pageNumber="96">G. leptosynaptae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="918D4D1FF657D62F37C5C2345958FEED" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Rathbun" baseAuthorityYear="1922" box="[1168,1329,313,339]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="vanderhorsti">
|
||
<emphasis id="64F9EA8EF657D62F37C5C2345958FEED" box="[1168,1329,313,339]" italics="true" pageId="13" pageNumber="96">G. vanderhorsti</emphasis>
|
||
</taxonomicName>
|
||
(at least the latter of which also has the plate) are included in that same well-supported molecular genetic clade for present lack of sequence quality material.
|
||
</paragraph>
|
||
<paragraph id="5632369CF657D62F3016C2B55850FBEE" blockId="13.[813,1456,217,1487]" pageId="13" pageNumber="96">
|
||
Thorough study of the first pleonal somite in mature males for all other suspected members and close relatives of
|
||
<taxonomicName id="918D4D1FF657D62F362DC2D55F31FDAC" authorityName="Campos & Wicksten" authorityYear="1997" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F362DC2D55F31FDAC" italics="true" pageId="13" pageNumber="96">Glassella</emphasis>
|
||
</taxonomicName>
|
||
is required to determine if any homologous ramification of the gonopodal plate may have also in those been thus far overlooked. This is to be undertaken in the course of coming descriptions of the American species “
|
||
<taxonomicName id="918D4D1FF657D62F37F2C154589FFDCC" authorityName="White" authorityYear="1846" box="[1191,1270,601,626]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F37F2C154589FFDCC" box="[1191,1270,601,626]" italics="true" pageId="13" pageNumber="96">Pinnixa</emphasis>
|
||
</taxonomicName>
|
||
sp. (ULLZ 13337 and ULLZ 14141)” from Panama (
|
||
<figureCitation id="CEB62A19F657D62F37FFC175588EFD2C" box="[1194,1255,632,658]" captionStart="FIG" captionStartId="8.[133,144,1919,1937]" captionTargetBox="[223,1405,231,1874]" captionTargetId="graphics@8.[290,1319,237,1858]" captionTargetPageId="8" captionText="FIG. 1. — Phylogeny for species of superfamily Pinnotheroidea De Haan,1833,emphasis on genus Pinnixa White, 1846 s.l. inferred from Randomized Accelerated Maximum Likelihood (RAxML) analysis of a 1445 bp long fragment concatenated from the mitochondrial complex 16S/tRNA-Leu/ NADH1 (776 bp),the mitochondrial 12S rRNA gene (340 bp) and the nuclear gene for the histone 3 subunit (327 bp). Bootstrap support values are shown at the nodes when higher than 50%. Collection number follows the species name to identify samples.For samples in the subfamily Pinnixinae Števčić,2005,abbreviations indicating geographic origin are defined in “Materials and Methods”. Species name combinations as shown are prior to revisions in present paper.Abbreviations as in Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.3695833" httpUri="https://zenodo.org/record/3695833/files/figure.png" pageId="13" pageNumber="96">Fig. 1</figureCitation>
|
||
) and at least three additional new western Atlantic species that clearly represent
|
||
<taxonomicName id="918D4D1FF657D62F3078C1BA5FEFFD6F" authorityName="Campos & Wicksten" authorityYear="1997" box="[813,902,695,721]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F3078C1BA5FEFFD6F" box="[813,902,695,721]" italics="true" pageId="13" pageNumber="96">Glassella</emphasis>
|
||
</taxonomicName>
|
||
in morphology, but for which we at present lack sequence quality materials. Further studies must also include molecular and morphological examinations of potentially related species, especially the eastern Pacific American species
|
||
<taxonomicName id="918D4D1FF657D62F3078C03558F0FCEC" authority="Garth, 1957" authorityName="Garth" authorityYear="1957" box="[813,1177,823,850]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="bahamondei">
|
||
<emphasis id="64F9EA8EF657D62F3078C035586EFCEF" box="[813,1031,823,849]" italics="true" pageId="13" pageNumber="96">Pinnixa bahamondei</emphasis>
|
||
Garth, 1957
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="918D4D1FF657D62F37F0C03559C2FCEF" authority=", Garth, 1960" authorityName="Garth" authorityYear="1960" box="[1189,1451,823,850]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="darwini">
|
||
<emphasis id="64F9EA8EF657D62F37F0C035597BFCEF" box="[1189,1298,823,849]" italics="true" pageId="13" pageNumber="96">P. darwini</emphasis>
|
||
, Garth, 1960
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="918D4D1FF657D62F3078C05558D2FCCC" authority="Salgado-Barragan, 2015" authorityName="Salgado-Barragan" authorityYear="2015" box="[813,1211,855,882]" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="hendrickxi">
|
||
<emphasis id="64F9EA8EF657D62F3078C0555FD8FCCF" box="[813,945,855,881]" italics="true" pageId="13" pageNumber="96">P. hendrickxi</emphasis>
|
||
Salgado-Barragán, 2015
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="918D4D1FF657D62F3790C0555F0FFC2C" authority="Glassell, 1935" authorityName="Glassell" authorityYear="1935" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="pembertoni">
|
||
<emphasis id="64F9EA8EF657D62F3790C0555939FCCF" box="[1221,1360,855,881]" italics="true" pageId="13" pageNumber="96">P. pembertoni</emphasis>
|
||
<bibRefCitation id="321C4B6DF657D62F3603C05A5F0FFC2C" author="GLASSELL S. A." pageId="13" pageNumber="96" pagination="13 - 14" refId="ref17605" refString="GLASSELL S. A. 1935 a. - Three new species of Pinnixa from the Gulf of California. Transactions of the San Diego Society of Natural History 8 (5): 13 - 14." type="journal article" year="1935">Glassell, 1935</bibRefCitation>
|
||
</taxonomicName>
|
||
, and perhaps
|
||
<taxonomicName id="918D4D1FF657D62F30A4C0755F18FC0F" authority="(H. Milne Edwards & Lucas, 1844)" baseAuthorityName="H. Milne Edwards & Lucas" baseAuthorityYear="1844" class="Malacostraca" family="Pinnotheridae" genus="Pinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="transversalis">
|
||
<emphasis id="64F9EA8EF657D62F30A4C0755817FC2F" box="[1009,1150,887,913]" italics="true" pageId="13" pageNumber="96">P.transversalis</emphasis>
|
||
(H. Milne Edwards & Lucas, 1844)
|
||
</taxonomicName>
|
||
. From our preliminary morphological observations of materials used in the present molecular study, we can state that a clear ramification of the male gonopodal plate is present in the eastern Pacific species
|
||
<taxonomicName id="918D4D1FF657D62F37CBC0FA5962FBAE" authorityName="Theil & Felder" authorityYear="2020" baseAuthorityName="Glassell" baseAuthorityYear="1935" box="[1182,1291,1014,1041]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="abbotti">
|
||
<emphasis id="64F9EA8EF657D62F37CBC0FA5962FBAE" box="[1182,1291,1014,1041]" italics="true" pageId="13" pageNumber="96">G. abbotti</emphasis>
|
||
</taxonomicName>
|
||
, underpinning our inclusion of it in
|
||
<taxonomicName id="918D4D1FF657D62F375CC71B5808FB8E" authorityName="Campos & Wicksten" authorityYear="1997" box="[1033,1121,1046,1072]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F375CC71B5808FB8E" box="[1033,1121,1046,1072]" italics="true" pageId="13" pageNumber="96">Glassella</emphasis>
|
||
</taxonomicName>
|
||
on the basis of more than solely molecular phylogenetics.
|
||
</paragraph>
|
||
<paragraph id="5632369CF657D62F3016C75B58DCFA71" blockId="13.[813,1456,217,1487]" pageId="13" pageNumber="96">
|
||
Further studies are also required to more thoroughly compare the Indo-West Pacific genus
|
||
<taxonomicName id="918D4D1FF657D62F37D1C77B596BFB2E" authorityName="Manning & Morton" authorityYear="1987" box="[1156,1282,1142,1168]" class="Malacostraca" family="Pinnotheridae" genus="Indopinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F37D1C77B596BFB2E" box="[1156,1282,1142,1168]" italics="true" pageId="13" pageNumber="96">Indopinnixa</emphasis>
|
||
</taxonomicName>
|
||
to the American
|
||
<taxonomicName id="918D4D1FF657D62F3078C79B5FEFFB0E" authorityName="Campos & Wicksten" authorityYear="1997" box="[813,902,1174,1200]" class="Malacostraca" family="Pinnotheridae" genus="Glassella" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F3078C79B5FEFFB0E" box="[813,902,1174,1200]" italics="true" pageId="13" pageNumber="96">Glassella</emphasis>
|
||
</taxonomicName>
|
||
. We retain separation of these genera as sister clades, though only two of the seven species assigned to
|
||
<taxonomicName id="918D4D1FF657D62F367AC7BB59C6FB6E" authorityName="Manning & Morton" authorityYear="1987" box="[1327,1455,1206,1232]" class="Malacostraca" family="Pinnotheridae" genus="Indopinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64F9EA8EF657D62F367AC7BB59C6FB6E" box="[1327,1455,1206,1232]" italics="true" pageId="13" pageNumber="96">Indopinnixa</emphasis>
|
||
</taxonomicName>
|
||
could for the present be represented in the molecular genetic analysis. Furthermore, only one of these two species was represented by an intact male specimen, the other represented only by a donated tissues sample. The intact male of
|
||
<taxonomicName id="918D4D1FF657D62F3602C63B5F34FAD1" authorityName="Naruse and Maenoso" authorityYear="2012" class="Malacostraca" family="Pinnotheridae" genus="Indopinnixa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="96" phylum="Arthropoda" rank="species" species="kumejima">
|
||
<emphasis id="64F9EA8EF657D62F3602C63B5F34FAD1" italics="true" pageId="13" pageNumber="96">I. kumejima</emphasis>
|
||
</taxonomicName>
|
||
was stained and carefully examined, and no evidence of a gonopodal plate or ramification thereof could be found, suggesting this could be a character of use in separating at least some species of the two genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |