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<document id="0F70AA922F99723874C7F0ACD4FBCDDC" ID-DOI="10.11646/zootaxa.4809.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3933799" ID-ZooBank="263939A4-E63E-4EE9-9AB6-0549794F9F24" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594125540984" checkinUser="plazi" docAuthor="Grimaldi, David A. &amp; Jones, Lance E." docDate="2020" docId="03FAC732FFF9066F0DC711B23392D2EB" docLanguage="en" docName="zootaxa.4809.1.1.pdf" docOrigin="Zootaxa 4809 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drosophila malagasy Grimaldi &amp; Jones 2020, new species" docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="FFC3BF4AFFF606600D50106F3362D534" masterDocTitle="A Revision of the Drosophila spinipes Species Group (Diptera: Drosophilidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="16" updateTime="1698778559937" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="79AE7EDD64058AF963B3709BF48ECE5F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6A8C2A4381B9DB57E63C726FEDCAE186">
<mods:title id="A20D9C3A10D5E9CB19D5FB525D479D2A">A Revision of the Drosophila spinipes Species Group (Diptera: Drosophilidae)</mods:title>
</mods:titleInfo>
<mods:name id="E48B9EA91F9FB42CAEDC71F380C5AB47" type="personal">
<mods:role id="B2EE98B347BB2C53AD7D0C455F4623AD">
<mods:roleTerm id="9CF038BF91B20435A26EBF62E72D856D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C04D6895F6128598D9FE2F850E6A6C55">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="C8F9B699FEDCD3D5E5272A3FB807A4F6">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 St., New York, NY 1002 &amp; grimaldi @ amnh. org; https: // orcid. org / 0000 - 0002 - 2271 - 0172</mods:affiliation>
</mods:name>
<mods:name id="6FB6EF80F72CA031A1AA896A81984FF6" type="personal">
<mods:role id="05A4FC072A450BC9636CF981A2585BAE">
<mods:roleTerm id="14A03689EE622C16AA80024EB500B95D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D9254799B06698A06B2DD309CA1596C">Jones, Lance E.</mods:namePart>
<mods:affiliation id="C709B02015A2B9CE6671184DB4092882">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 St., New York, NY 1002 &amp; ljones 1 @ amnh. org https: // orcid. org / 0000 - 0001 - 6368 - 9720</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7FAF372752CD8990146AC2881E83FD78">text</mods:typeOfResource>
<mods:relatedItem id="B17C82BD056D04051438E0085FF4A65C" type="host">
<mods:titleInfo id="63BFA67CB87A487763FF339D59A30913">
<mods:title id="13581349BB4E0888E5C5FD6B68399A5C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D9B383CB9BA7FC0C58F02B85A6BB4424">
<mods:date id="B474485663BE919CD93D9D7D4991328D">2020</mods:date>
<mods:detail id="00D47040762E453BCDFB88069BDDE505" type="pubDate">
<mods:number id="C71A3F14AC8010C4118DBA2B68B44F53">2020-07-06</mods:number>
</mods:detail>
<mods:detail id="E3C81418509A7BE45E5481C680F2CC7A" type="volume">
<mods:number id="754E6B0BA46FEBE5BB9531209DD023D3">4809</mods:number>
</mods:detail>
<mods:detail id="3666E521EE96F58FB9FED4031765E6C8" type="issue">
<mods:number id="966D599A6DC49DCFCCDF7697C5B3FBFD">1</mods:number>
</mods:detail>
<mods:extent id="725AC5782843A808F22E7CCCEA0542EF" unit="page">
<mods:start id="4A20923747CE4B2828F5ED3B2DA1D939">1</mods:start>
<mods:end id="AFA8138C9590E8E3EAC43EBB9A20B110">28</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="75A64C7CB517ACC0FA3D2C06EEEDD6B8">journal article</mods:classification>
<mods:identifier id="D5FB046CF07B4AD55DD94D3DDEC6F8EA" type="DOI">10.11646/zootaxa.4809.1.1</mods:identifier>
<mods:identifier id="6069F4EAC30D70454B77BB238F348670" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="03C6614ED34EDA041E62CC522B50F986" type="Zenodo-Dep">3933799</mods:identifier>
<mods:identifier id="ECF3B97DB80A705B2EC910D57A5EB7AD" type="ZooBank">263939A4-E63E-4EE9-9AB6-0549794F9F24</mods:identifier>
</mods:mods>
<treatment id="03FAC732FFF9066F0DC711B23392D2EB" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB" httpUri="http://treatment.plazi.org/id/03FAC732FFF9066F0DC711B23392D2EB" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="C34925AFFFF9066F0DC711B23154D4C3" box="[151,566,477,504]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BEC7624FFF9066F0DC711B23154D4C3" blockId="15.[151,1020,477,575]" box="[151,566,477,504]" pageId="15" pageNumber="16">
<heading id="D0A4C148FFF9066F0DC711B23154D4C3" bold="true" box="[151,566,477,504]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B927AA36FFF9066F0DC711B23154D4C3" bold="true" box="[151,566,477,504]" pageId="15" pageNumber="16">
<taxonomicName id="4C530DA7FFF9066F0DC711B232FAD4CC" ID-CoL="9H9MF" authority="Grimaldi &amp; Jones, 2020" authorityName="Grimaldi &amp; Jones" authorityYear="2020" box="[151,408,477,504]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="malagasy" status="sp. nov.">
<emphasis id="B927AA36FFF9066F0DC711B232FAD4CC" bold="true" box="[151,408,477,504]" italics="true" pageId="15" pageNumber="16">Drosophila malagasy</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A214174DFFF9066F0CF711B23154D4C3" box="[423,566,477,503]" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34925AFFFF9066F0DC7126E309ED70B" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEC7624FFF9066F0DC7126E30DDD72F" blockId="15.[151,1020,477,575]" box="[151,959,512,539]" pageId="15" pageNumber="16">
<figureCitation id="13686AA1FFF9066F0DC7126E3398D72F" box="[151,250,513,539]" captionStart="FIGURE 2" captionStartId="4.[151,250,1901,1926]" captionTargetBox="[151,1435,489,1876]" captionTargetId="figure@4.[151,1435,489,1877]" captionTargetPageId="4" captionText="FIGURE 2. Distinct or apomorphic features of the Drosophila spinipes species group. A. Lateral view, sp. C (HS-13). B. Same as A, with detail of mouthparts. C. D. nigrospinipes n. sp. (HS-12), dorsolateral view of thorax, showing setal tubercles and spiracular channel. D. Wing, D. malagasy n. sp. (HS-21). Arrow indicates end of dense costal spinules. E-F. SEM of protarsal spines, mesal view of specimen HS-14 (sp. B). Note the shallow longitudinal grooves." figureDoi="http://doi.org/10.5281/zenodo.3933803" httpUri="https://zenodo.org/record/3933803/files/figure.png" pageId="15" pageNumber="16">Figs. 2D</figureCitation>
,
<figureCitation id="13686AA1FFF9066F0C56126E324BD72F" box="[262,297,513,539]" captionStart="FIGURE 3" captionStartId="8.[151,250,1817,1842]" captionTargetBox="[151,1435,181,1792]" captionTargetId="figure@8.[151,1435,181,1792]" captionTargetPageId="8" captionText="FIGURE 3. Lateral and dorsal habitus of several Drosophila spinipes-group species. A. D. freidbergi n. sp. (HS-06). B. D. malagasy n. sp. (HS-18). C. D. nigrospinipes n. sp. (HS-24). D. D. phalloserra n. sp. (HS-08). E. Dorsal thorax, sp. C (HS-13). F. D. spinipes Lamb (NHMUK 014335955), dorsum. G. D. spinipes Lamb (NHMUK 014335956), lateral." figureDoi="http://doi.org/10.5281/zenodo.3933805" httpUri="https://zenodo.org/record/3933805/files/figure.png" pageId="15" pageNumber="16">3B</figureCitation>
,
<figureCitation id="13686AA1FFF9066F0C64126E3220D72F" box="[308,322,513,539]" captionStart="FIGURE 4" captionStartId="10.[151,250,1185,1210]" captionTargetBox="[151,1435,181,1160]" captionTargetId="figure@10.[151,1435,181,1160]" captionTargetPageId="10" captionText="FIGURE 4. Photomicrographs, dorsolateral view, of the abdomen of several Drosophila spinipes-group species. A-C. D. malagasy n. sp. A.HS-16. B. HS-22. C. HS-18. D. D. nigrospinipes n. sp. (HS-24). E. D. phalloserra n. sp. (HS-07). F. D. suma Burla (HS-01). G. D. spinipes Lamb (NHMUK 014335955)." figureDoi="http://doi.org/10.5281/zenodo.3933807" httpUri="https://zenodo.org/record/3933807/files/figure.png" pageId="15" pageNumber="16">4</figureCitation>
A–C, 5B–C, 6D, 7C–D, 8B, 9D–E, 10G, 11D–E, 12B–D
</paragraph>
<paragraph id="8BEC7624FFF9066F0DC7124A309ED70B" blockId="15.[151,1020,477,575]" box="[151,1020,548,575]" pageId="15" pageNumber="16">
LSID
<uri id="FFC27A26FFF9066F0D8C124B309ED70B" box="[220,1020,548,575]" pageId="15" pageNumber="16">
urn:lsid:zoobank.org:act:
<uuid id="FFF54CF1FFF9066F0CA5124B309ED70B" box="[501,1020,548,575]" pageId="15" pageNumber="16">3627E181-1B0E-4B78-8380-03B503D35E7B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34925AFFFF9066F0DC712033750D7FB" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BEC7624FFF9066F0DC712033750D7FB" blockId="15.[151,1437,620,2015]" pageId="15" pageNumber="16">
<emphasis id="B927AA36FFF9066F0DC712033276D7B3" bold="true" box="[151,276,620,647]" pageId="15" pageNumber="16">Diagnosis:</emphasis>
Usually with just 3 black spines (vs. 4) on protarsal segment 1 (the small spine no. 7 missing); apex of abdomen light (tergites
<quantity id="4CABDBC1FFF9066F0CCE12FE329AD79F" box="[414,504,657,683]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="15" pageNumber="16" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">6 to 8 in</quantity>
male, 7 and
<quantity id="4CABDBC1FFF9066F0FD312FE31CFD79F" box="[643,685,657,683]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="15" pageNumber="16" unit="in" value="8.0">8 in</quantity>
females); hypandrium small, anterior portion distinctly constricted; aedeagus without coarse inner scales but with finely serrate apical-lateral margins.
</paragraph>
</subSubSection>
<subSubSection id="C34925AFFFF9066F0D9712B7323BD1D8" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEC7624FFF9066F0D9712B7323BD1D8" blockId="15.[151,1437,620,2015]" pageId="15" pageNumber="16">
<emphasis id="B927AA36FFF9066F0D9712B73239D7C7" bold="true" box="[199,347,728,755]" pageId="15" pageNumber="16">Description:</emphasis>
Coloration typical for group, tergite
<quantity id="4CABDBC1FFF9066F0FAC12B6304AD7C7" box="[764,808,729,755]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="15" pageNumber="16" unit="in" value="7.0">7 in</quantity>
female light, paraprocts slightly darker; males with tergite 6, epandrium and cerci light. Protarsus with standard array of fang-shaped spines alternating in size, but with protarsomere 1 with usually 3 black spines instead of the typical 4 (small spine no. 7 often missing); in
<specimenCount id="9D55BDADFFF9066F0856134E36FFD60F" box="[1286,1437,801,827]" pageId="15" pageNumber="16" type="generic">14 specimens</specimenCount>
four spines were found in four individuals (HS18, 22, 28, and 30), spines on protarsomere 1 crowded onto distal half of segment. Male genitalia: Surstylus approximately triangular in full lateral view, prensisetae not crowded (e.g., touching); ventral lobe of epandrium with 2 short, spine-like setae; subepandrial sclerite with expanded anterior portion, posterior arms an inverted V-shape. Hypandrium small, anterior portion significantly narrowed to ~ 0.5x the posterior width; aedeagus narrowed apically in dorsoventral view, with dense, fine spicules medially and 5–6 very fine serrations at apex; laterally with apex bulbous. Oviscapt: Pegs in dorsal triad and ventral pair approximately equal in size; apical notch shallow to deep, ventral series of 9–10 smaller pegs evenly spaced to slightly crowded.
<emphasis id="B927AA36FFF9066F0C56142E314FD16F" bold="true" box="[262,557,1088,1115]" pageId="15" pageNumber="16">Measurements (N = 4–6)</emphasis>
:
<emphasis id="B927AA36FFF9066F0F6C142E31E7D16F" bold="true" box="[572,645,1089,1115]" pageId="15" pageNumber="16">Head:</emphasis>
HD/HW 0.80 (0.79–0.82); FL/LFW 1.27 (1.17–1.46); VB2/VB1 1.67 (0.61–0.84); EW/ED 0.70 (0.67–0.75); OR2/OR1 0.87 (0.82–0.92); OR3/OR1 1.13 (1.05–1.23); OC/POC 1.45 (1.30–1.50); FW/FD 0.55 (0.50–0.62); ODB/ODA 0.33 (0.27–0.40);
<emphasis id="B927AA36FFF9066F0EC614E6309AD197" bold="true" box="[918,1016,1161,1187]" pageId="15" pageNumber="16">Thorax:</emphasis>
ThL
<quantity id="4CABDBC1FFF9066F096314E637F4D197" box="[1075,1174,1161,1187]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="15" pageNumber="16" unit="mm" value="0.92">0.92 mm</quantity>
(0.80–1.14); UPS/LHS 0.75 (0.72–0.82); LHS/ThL 0.20 (0.17–0.22); ADC/PDC 0.46 (0.36–0.53); AKE/PKE 0.45 (0.38–0.50); LF1/LF2 0.79 (0.76–0.83).
</paragraph>
</subSubSection>
<subSubSection id="C34925AFFFF9066F0D97149A3067D233" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BEC7624FFF9066F0D97149A3390D04F" blockId="15.[151,1437,620,2015]" pageId="15" pageNumber="16">
<materialsCitation id="3B3B7C79FFF9066F0D97149A3390D04F" collectionCode="CAS" collectorName="M. Irwin &amp; M. Harin'Hala" country="Madagascar" elevation="1130" latitude="-21.250834" location="Male" longLatPrecision="12" longitude="47.407166" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B927AA36FFF9066F0D97149A325FD03B" bold="true" box="[199,317,1269,1295]" pageId="15" pageNumber="16">
<typeStatus id="54E8C886FFF9066F0D97149A325AD03B" box="[199,312,1269,1295]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="8E8C20FFFFF9066F0C13149A321DD03B" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0C13149A321DD03B" box="[323,383,1269,1295]" country="Madagascar" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" name="Male" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">Male</location>
, HS-16 “
<collectingCountry id="F34436B4FFF9066F0CB3149A310DD03B" box="[483,623,1269,1295]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
, Province
<collectingRegion id="4997B8C6FFF9066F0FB2149A3016D03B" box="[738,884,1269,1295]" country="Madagascar" name="Fianarantsoa" pageId="15" pageNumber="16">Fianarantsoa</collectingRegion>
, Parc National Ranomafana, radio tower, at forest edge, elev.
<quantity id="4CABDBC1FFF9066F0C4815763210D007" box="[280,370,1305,1331]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="16" unit="m" value="1130.0">
<elevation id="007E9117FFF9066F0C4815763210D007" box="[280,370,1305,1331]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="16" unit="m" value="1130.0">1130 m</elevation>
</quantity>
., 20 June–6 July, 2003 /
<geoCoordinate id="EE6710E3FFF9066F0FDC15763079D007" box="[652,795,1304,1331]" degrees="21" direction="south" minutes="15.05" orientation="latitude" pageId="15" pageNumber="16" precision="9" value="-21.250834">21° 15.05’ S</geoCoordinate>
,
<geoCoordinate id="EE6710E3FFF9066F0E79157630D8D007" box="[809,954,1305,1331]" degrees="47" direction="east" minutes="24.43" orientation="longitude" pageId="15" pageNumber="16" precision="9" value="47.407166">47° 24.43’ E</geoCoordinate>
, coll.
<collectorName id="26A613F2FFF9066F09511576370AD007" box="[1025,1128,1305,1331]" pageId="15" pageNumber="16">M. Irwin</collectorName>
,
<collectorName id="26A613F2FFF9066F09251576367CD007" box="[1141,1310,1305,1331]" pageId="15" pageNumber="16">M. Harin’Hala</collectorName>
, California Academy of Sciences, Malaise [trap] mixed tropical, MA-02-09B-64.” Point mounted, genitalia dissected by DG; in
<collectionCode id="ED42EEE1FFF9066F0DE4150E338CD04F" box="[180,238,1377,1403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="16" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEC7624FFF9066F0D9715EB3067D233" blockId="15.[151,1437,620,2015]" pageId="15" pageNumber="16">
<emphasis id="B927AA36FFF9066F0D9715EB32FCD0AB" bold="true" box="[199,414,1412,1439]" pageId="15" pageNumber="16">Other Specimens:</emphasis>
<materialsCitation id="3B3B7C79FFF9066F0CF615EA3211D233" collectingDate="1991-04" collectingDateMax="1991-04-12" collectingDateMin="1991-04" collectionCode="DG, TAU" collectorName="M. Irwin &amp; M. Harin'Hala &amp; A. Freidberg" country="Madagascar" county="Province" elevation="1130" latitude="-21.250834" location="June" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" pageId="15" pageNumber="16" specimenCount="807" specimenCount-female="6" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="54E8C886FFF9066F0CF615EA3175D0AB" box="[422,535,1413,1439]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
: HS-17, male “
<collectingCountry id="F34436B4FFF9066F0F9315EA302DD0AB" box="[707,847,1413,1439]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
,
<collectingCounty id="628D0EA8FFF9066F0E0B15EA30DDD0AB" box="[859,959,1413,1439]" pageId="15" pageNumber="16">Province</collectingCounty>
<collectingRegion id="4997B8C6FFF9066F0E9715EA373BD0AB" box="[967,1113,1413,1439]" country="Madagascar" name="Fianarantsoa" pageId="15" pageNumber="16">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="6B88EC5EFFF9066F093515EA36FAD0AB" box="[1125,1432,1413,1439]" pageId="15" pageNumber="16">Parc National Ranomafana</collectingMunicipality>
, radio tower, at forest edge, elev.
<quantity id="4CABDBC1FFF9066F0F4D15C63118D0F7" box="[541,634,1449,1475]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="16" unit="m" value="1130.0">
<elevation id="007E9117FFF9066F0F4D15C63118D0F7" box="[541,634,1449,1475]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="16" unit="m" value="1130.0">1130 m</elevation>
</quantity>
., 18–29
<location id="8E8C20FFFFF9066F0F8F15C63077D0F7" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0F8F15C63077D0F7" box="[735,789,1449,1475]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="June" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">June</location>
, 2003 /
<geoCoordinate id="EE6710E3FFF9066F0E2A15C6376DD0F7" box="[890,1039,1448,1475]" degrees="21" direction="south" minutes="15.05" orientation="latitude" pageId="15" pageNumber="16" precision="9" value="-21.250834">21° 15.05’ S</geoCoordinate>
,
<geoCoordinate id="EE6710E3FFF9066F097015C637D4D0F7" box="[1056,1206,1449,1475]" degrees="47" direction="east" minutes="24.43" orientation="longitude" pageId="15" pageNumber="16" precision="9" value="47.407166">47° 24.43’ E</geoCoordinate>
, coll.
<collectorName id="26A613F2FFF9066F085315C6360FD0F7" box="[1283,1389,1449,1475]" pageId="15" pageNumber="16">M. Irwin</collectorName>
,
<collectorName id="26A613F2FFF9066F082C15C6327AD0D3" pageId="15" pageNumber="16">M. Harin’Hala</collectorName>
,
<location id="8E8C20FFFFF9066F0C7215A231EFD0D3" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0C7215A231EFD0D3" box="[290,653,1485,1511]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="California Academy of Sciences" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">California Academy of Sciences</location>
,
<location id="8E8C20FFFFF9066F0FC715A23192D0D3" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0FC715A23192D0D3" box="[663,752,1485,1511]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="Malaise" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">Malaise</location>
[trap] mixed tropical, MA-02-09B-64” genitalia dissected by DG. HS-18, female “
<collectingCountry id="F34436B4FFF9066F0CD6159E3170D33F" box="[390,530,1521,1547]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
,
<location id="8E8C20FFFFF9066F0F4F159E31E1D33F" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0F4F159E31E1D33F" box="[543,643,1521,1547]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="Province" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">Province</location>
<collectingRegion id="4997B8C6FFF9066F0FDB159E307FD33F" box="[651,797,1521,1547]" country="Madagascar" name="Fianarantsoa" pageId="15" pageNumber="16">Fianarantsoa</collectingRegion>
,
<location id="8E8C20FFFFF9066F0E7A159E373FD33F" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0E7A159E373FD33F" box="[810,1117,1521,1547]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="Parc National Ranomafana" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">Parc National Ranomafana</location>
, radio tower, at forest edge, elev.
<quantity id="4CABDBC1FFF9066F0D83167A324FD31B" box="[211,301,1557,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="16" unit="m" value="1130.0">
<elevation id="007E9117FFF9066F0D83167A324FD31B" box="[211,301,1557,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="16" unit="m" value="1130.0">1130 m</elevation>
</quantity>
., 20 June–6
<location id="8E8C20FFFFF9066F0CE9167A328BD31B" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0CE9167A328BD31B" box="[441,489,1557,1583]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="July" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">July</location>
, 2003 /
<geoCoordinate id="EE6710E3FFF9066F0F15167A31B1D31B" box="[581,723,1556,1583]" degrees="21" direction="south" minutes="15.05" orientation="latitude" pageId="15" pageNumber="16" precision="9" value="-21.250834">21° 15.05’ S</geoCoordinate>
,
<geoCoordinate id="EE6710E3FFF9066F0FB1167A3013D31B" box="[737,881,1557,1583]" degrees="47" direction="east" minutes="24.43" orientation="longitude" pageId="15" pageNumber="16" precision="9" value="47.407166">47° 24.43’ E</geoCoordinate>
, coll.
<collectorName id="26A613F2FFF9066F0EE7167A377CD31B" box="[951,1054,1557,1583]" pageId="15" pageNumber="16">M. Irwin</collectorName>
,
<collectorName id="26A613F2FFF9066F097B167A37B6D31B" box="[1067,1236,1557,1583]" pageId="15" pageNumber="16">M. Harin’Hala</collectorName>
,
<location id="8E8C20FFFFF9066F09B0167A3233D367" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F09B0167A3233D367" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="California Academy of Sciences" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">California Academy of Sciences</location>
,
<location id="8E8C20FFFFF9066F0C0B165632D6D367" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0C0B165632D6D367" box="[347,436,1593,1619]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="Malaise" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">Malaise</location>
[trap] mixed tropical forest, MA-02-09B-64.”
<specimenCount id="9D55BDADFFF9066F0EEE16563743D367" box="[958,1057,1593,1619]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
, HS-23 “
<collectingCountry id="F34436B4FFF9066F09D516563671D367" box="[1157,1299,1593,1619]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
:
<location id="8E8C20FFFFF9066F084C165636FAD367" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F084C165636FAD367" box="[1308,1432,1593,1619]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="Ambohitra" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">Ambohitra</location>
,
<location id="8E8C20FFFFF9066F0DC716323218D343" LSID="urn:lsid:plazi:treatment:03FAC732FFF9066F0DC711B23392D2EB:8E8C20FFFFF9066F0DC716323218D343" box="[151,378,1629,1655]" country="Madagascar" county="Province" latitude="-21.250834" longLatPrecision="12" longitude="47.407166" municipality="Parc National Ranomafana" name="Montagne d'Ambre" pageId="15" pageNumber="16" stateProvince="Fianarantsoa">Montagne d’Ambre</location>
,
<quantity id="4CABDBC1FFF9066F0CD716323283D343" box="[391,481,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="1000.0">
<elevation id="007E9117FFF9066F0CD716323283D343" box="[391,481,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFED50E4FFF9066F0CBF1632313DD343" box="[495,607,1629,1655]" pageId="15" pageNumber="16" value="1991-04-09">
<collectingDate id="EFA9A90CFFF9066F0CBF1632313DD343" box="[495,607,1629,1655]" pageId="15" pageNumber="16" value="1991-04-09">9.IV.1991</collectingDate>
</date>
,
<collectorName id="26A613F2FFF9066F0F3B1632319ED343" box="[619,764,1629,1655]" pageId="15" pageNumber="16">A. Freidberg</collectorName>
&amp; Fini Kaplan” not dissected. HS-26 (female), HS-20, 21 (males, both dissected by
<collectionCode id="ED42EEE1FFF9066F0CE916EE3285D3AF" box="[441,487,1665,1691]" pageId="15" pageNumber="16">DG</collectionCode>
): “
<collectingCountry id="F34436B4FFF9066F0F5316EE31EFD3AF" box="[515,653,1665,1691]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
N., Ambohitra
<specimenCount id="9D55BDADFFF9066F0E6C16EE30EED3AF" box="[828,908,1665,1691]" pageId="15" pageNumber="16" type="generic">
<quantity id="4CABDBC1FFF9066F0E6C16EE30EAD3AF" box="[828,904,1665,1691]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="m" value="800.0">
<elevation id="007E9117FFF9066F0E6C16EE30EAD3AF" box="[828,904,1665,1691]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="m" value="800.0">800 m</elevation>
</quantity>
.
</specimenCount>
, Joffreville,
<date id="FFED50E4FFF9066F094816EE37D1D3AF" box="[1048,1203,1665,1691]" pageId="15" pageNumber="16" value="1991-04-09" valueMax="1991-04-12" valueMin="1991-04-09">
<collectingDate id="EFA9A90CFFF9066F094816EE37D1D3AF" box="[1048,1203,1665,1691]" pageId="15" pageNumber="16" value="1991-04-09" valueMax="1991-04-12" valueMin="1991-04-09">9–12.IV.1991</collectingDate>
</date>
,
<collectorName id="26A613F2FFF9066F09ED16EE3629D3AF" box="[1213,1355,1665,1691]" pageId="15" pageNumber="16">A. Freidberg</collectorName>
&amp; Fini Kaplan.”
<specimenCount id="9D55BDADFFF9066F0C5416CB3210D38B" box="[260,370,1700,1727]" pageId="15" pageNumber="16" type="female">5 females</specimenCount>
(HS-19, 25, 28, 29, 30 [19, 28, 30 dissected by DG]),
<specimenCount id="9D55BDADFFF9066F0EBA16CA3755D38B" box="[1002,1079,1701,1727]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
(HS-22, dissected by
<collectionCode id="ED42EEE1FFF9066F086716CA3601D38B" box="[1335,1379,1701,1727]" pageId="15" pageNumber="16">DG</collectionCode>
), all: “
<collectingCountry id="F34436B4FFF9066F0DF116A63249D3D7" box="[161,299,1737,1763]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
N., Perinet (Andasibe), 16.17.
<date id="FFED50E4FFF9066F0FD416A63183D3D7" box="[644,737,1737,1763]" pageId="15" pageNumber="16" value="1991-04">
<collectingDate id="EFA9A90CFFF9066F0FD416A63183D3D7" box="[644,737,1737,1763]" pageId="15" pageNumber="16" value="1991-04">IV.1991</collectingDate>
</date>
,
<collectorName id="26A613F2FFF9066F0FBB16A63018D3D7" box="[747,890,1737,1763]" pageId="15" pageNumber="16">A. Freidberg</collectorName>
&amp; Fini Kaplan.” All specimens from Ambohitra and Perinet in
<collectionCode id="ED42EEE1FFF9066F0C6916823211D233" box="[313,371,1773,1799]" country="Israel" httpUri="http://grbio.org/cool/ze71-842p" name="Tel-Aviv University" pageId="15" pageNumber="16">TAU</collectionCode>
</materialsCitation>
, except HS-19 and -22 (in
<collectionCode id="ED42EEE1FFF9066F0FCF16823198D233" box="[671,762,1773,1799]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="15" pageNumber="16">AMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34925AFFFF9066F0D97177E372BD21F" box="[199,1097,1809,1835]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BEC7624FFF9066F0D97177E372BD21F" blockId="15.[151,1437,620,2015]" box="[199,1097,1809,1835]" pageId="15" pageNumber="16">
<emphasis id="B927AA36FFF9066F0D97177E3230D21F" bold="true" box="[199,338,1809,1835]" pageId="15" pageNumber="16">Etymology:</emphasis>
In direct reference to the country of origin, as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34925AFFFF9066F0D97175B3392D2EB" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BEC7624FFF9066F0D97175B3392D2EB" blockId="15.[151,1437,620,2015]" pageId="15" pageNumber="16">
<emphasis id="B927AA36FFF9066F0D97175B3230D27B" bold="true" box="[199,338,1844,1871]" pageId="15" pageNumber="16">Comments:</emphasis>
This may be the species to which
<bibRefCitation id="EFC20BD5FFF9066F0F89175A30ADD264" author="Lachaise, D. &amp; Harry, M. &amp; Solignac, M." box="[729,975,1845,1872]" pageId="15" pageNumber="16" pagination="467 - 478" refId="ref16288" refString="Lachaise, D., Harry, M. &amp; Solignac, M. (1996) Affinites biogeographiques des Drosophilidae de Madagascar et des iles de l'Ocean Indien. In: Lourenco, W. R. (Ed.), Biogeographie de Madagascar. Ostrom, Paris, pp. 467 - 478." type="book chapter" year="1996">
Lachaise
<emphasis id="B927AA36FFF9066F0E1417593014D27B" box="[836,886,1845,1871]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. (1996)
</bibRefCitation>
mentioned as “
<taxonomicName id="4C530DA7FFF9066F092F175A3663D27B" authorityName="Lamb" authorityYear="1914" box="[1151,1281,1845,1871]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="spinipes">
cf.
<emphasis id="B927AA36FFF9066F09F517593663D27B" box="[1189,1281,1846,1871]" italics="true" pageId="15" pageNumber="16">spinipes</emphasis>
</taxonomicName>
” from
<collectingCountry id="F34436B4FFF9066F0803175A33BDD247" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
and
<collectingCountry id="F34436B4FFF9066F0C46173632E5D247" box="[278,391,1881,1907]" name="Mauritius" pageId="15" pageNumber="16">Mauritius</collectingCountry>
. We have not seen specimens from
<collectingCountry id="F34436B4FFF9066F0E46173630E5D247" box="[790,903,1881,1907]" name="Mauritius" pageId="15" pageNumber="16">Mauritius</collectingCountry>
. The species as we define it here may represent two or more morphocryptic species, based on the variation in oviscapt (cf. figs. 7C, D), though with minor variation in hypandrium size and shape (cf. figs. 9D, E), and the number of fine serrations at the tip and lateral margins of the phallus.
</paragraph>
</subSubSection>
</treatment>
</document>
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