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<mods:title id="0400E94EB87DC72FA68A543765077987">Jassa (Crustacea: Amphipoda): a new morphological and molecular assessment of the genus</mods:title>
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<treatment id="03B487DAFFC8D966C9C81B86FBDBFF2C" ID-DOI="http://doi.org/10.5281/zenodo.4580587" ID-GBIF-Taxon="181871898" ID-Zenodo-Dep="4580587" LSID="urn:lsid:plazi:treatment:03B487DAFFC8D966C9C81B86FBDBFF2C" httpUri="http://treatment.plazi.org/id/03B487DAFFC8D966C9C81B86FBDBFF2C" lastPageId="126" lastPageNumber="126" pageId="125" pageNumber="126">
<subSubSection id="C3076547FFC8D965C9C81B86FE65FF2C" pageId="125" pageNumber="126" type="nomenclature">
<paragraph id="8BA236CCFFC8D965C9C81B86FE18FF48" blockId="125.[151,492,151,213]" box="[151,492,151,178]" pageId="125" pageNumber="126">
<heading id="D0EA81A0FFC8D965C9C81B86FE18FF48" bold="true" box="[151,492,151,178]" fontSize="11" level="2" pageId="125" pageNumber="126" reason="4">
<taxonomicName id="4C1D4D4FFFC8D965C9C81B86FE18FF48" ID-CoL="3QMZB" authority="Conlan, 1990" authorityName="Conlan" authorityYear="1990" box="[151,492,151,178]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965C9C81B86FE18FF48" bold="true" box="[151,492,151,178]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9C81B86FEB4FF4B" bold="true" box="[151,320,151,178]" italics="true" pageId="125" pageNumber="126">Jassacarltoni</emphasis>
<bibRefCitation id="EF8C4B3DFFC8D965C8181B86FE18FF48" author="Conlan, K. E." box="[327,492,151,177]" pageId="125" pageNumber="126" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan, 1990</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BA236CCFFC8D965C9C81BAAFE65FF2C" blockId="125.[151,492,151,213]" box="[151,401,187,213]" pageId="125" pageNumber="126">
(
<tableCitation id="C69F0377FFC8D965C9C11BAAFEF6FF2C" box="[158,258,187,213]" captionStart="TABLE 12" captionStartId="102.[151,239,332,356]" captionTargetPageId="102" captionText="TABLE 12. Key character states of locally distributed North Pacific species. Trans-hemispheric species (Table 10) may occur there as well. X indicates the presence of the first character state described while — indicates the alternative character state given in italics. S and L indicate length.? indicates that the state is unknown. Character states apply to both sexes and all ages unless indicated otherwise." httpUri="http://table.plazi.org/id/DF626644FFD3D97EC9C81A5DFD89FE29" pageId="125" pageNumber="126" tableUuid="DF626644FFD3D97EC9C81A5DFD89FE29">Table 12</tableCitation>
,
<figureCitation id="13262A49FFC8D965C8521BAAFE7EFF2C" box="[269,394,187,213]" captionStart-0="FIGURE 78" captionStart-1="FIGURE 79" captionStart-2="FIGURE 80" captionStart-3="FIGURE 81" captionStart-4="FIGURE 82" captionStartId-0="126.[151,250,1606,1630]" captionStartId-1="127.[151,250,983,1007]" captionStartId-2="127.[151,250,1932,1956]" captionStartId-3="128.[151,250,1864,1888]" captionStartId-4="129.[151,250,1868,1892]" captionTargetBox-0="[279,1306,238,1577]" captionTargetBox-1="[276,1309,185,956]" captionTargetBox-2="[284,1300,1145,1904]" captionTargetBox-3="[167,1416,222,1840]" captionTargetBox-4="[158,1425,187,1844]" captionTargetId-0="figure-55@126.[274,1314,237,1583]" captionTargetId-1="figure-18@127.[271,1315,181,959]" captionTargetId-2="figure-67@127.[279,1308,1139,1908]" captionTargetId-3="figure-22@128.[151,1436,181,1840]" captionTargetId-4="figure-18@129.[151,1436,181,1844]" captionTargetPageId-0="126" captionTargetPageId-1="127" captionTargetPageId-2="127" captionTargetPageId-3="128" captionTargetPageId-4="129" captionText-0="FIGURE 78. Jassa carltoni Conlan, 1990. Holotype, adult male, major form, 3.0 mm, Corona del Mar, California, J. L. Barnard, coll., 911 December 1962, station 46-K-1, wash of 1/6 liter of ascidians and soft polychaete tubes collected at low tide (NHMLA 62-132.1). Paratype, juvenile male, 1.7 mm, La Jolla, California, 1113 Nov. 1962, J. L. Barnard, coll., station 45-R- 1, no further collection data, 62-135.1 (NHMLA); subadult male, 4.0 mm, Cable Beach, Vancouver Island, British Columbia, 14 July 1972, D. Kittle, coll., station 342, 1973-128 (CMN); paratype, adult male, minor form, 2.8 mm, La Jolla, California, 1113 Nov. 1962, J. L. Barnard, coll., station 45-W-5, scraping of calcareous sponge under ledge in surge channel at low tide, 62-134.1 (NHMLA); paratype, adult male 2, major form, 2.9 mm, Hazard Canyon Reef, Montana de Oro State Park, St. Louis Obispo County, California, 19 Dec. 1961, J. L. Barnard, coll., station 42-T-5, wash of Egregia holdfasts collected at low tide, 61-100.2 (NHMLA); adult male 3, major form, 4.0 mm, Sarita Bay, Vancouver Island, British Columbia, 2 July 1976, E. L. Bousfield, coll., station B13, 1976-157, bedrock, muddy sand, eelgrass, algae and fine gravel subtidally, hand and dip net collection at low tide, ~14 °C, 26 psu (CMN). Lateral view: whole body and gnathopod 2 views; dorsal view: telson; other views medial. Setae omitted from the juvenile male and major male 2 second gnathopods. Scale 0.1 mm. Illustration after Conlan (1990)." captionText-1="FIGURE 79. Jassa carltoni Conlan, 1990. Variation in thumb length relative to body length in males in a single collection at Wizard Islet, Vancouver Island, British Columbia, July 1972, D. Kittle, coll., station 712, NMNS 73-128 (CMN). Linear regression assumptions failed for all plots." captionText-2="FIGURE 80. Jassa carltoni Conlan, 1990. Variation in gnathopod 2 propodus length relative to body length in males and females in a single collection at Wizard Islet, Vancouver Island, British Columbia, July 1972, D. Kittle, coll., station 712, NMNS 73-128 (CMN). Linear regression assumptions failed for all plots." captionText-3="FIGURE 81. Jassa carltoni Conlan, 1990.Allotype, adult female, 2.8 mm, 62-132.2 (NHMLA). Corona del Mar, California, J. L. Barnard, coll., 911 December 1962, station 46-K-1, wash of 1/6 liter of ascidians and soft polychaete tubes collected at low tide. Scale 0.1 mm. Illustration after Conlan (1990)." captionText-4="FIGURE 82. Jassa carltoni Conlan, 1990. Holotype, adult male, major form, 3.0 mm, 62-132.1 (NHMLA). Corona del Mar, California, J. L. Barnard, coll., 911 December 1962, station 46-K-1, wash of 1/6 liter of ascidians and soft polychaete tubes collected at low tide. Mouthparts. Frontal view: upper lip; lateral view: maxilla 1; other views medial. Scale 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4580822" figureDoi-1="http://doi.org/10.5281/zenodo.4580824" figureDoi-2="http://doi.org/10.5281/zenodo.4580826" figureDoi-3="http://doi.org/10.5281/zenodo.4580828" figureDoi-4="http://doi.org/10.5281/zenodo.4580832" httpUri-0="https://zenodo.org/record/4580822/files/figure.png" httpUri-1="https://zenodo.org/record/4580824/files/figure.png" httpUri-2="https://zenodo.org/record/4580826/files/figure.png" httpUri-3="https://zenodo.org/record/4580828/files/figure.png" httpUri-4="https://zenodo.org/record/4580832/files/figure.png" pageId="125" pageNumber="126">Figs 7882</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3076547FFC8D965C9C81A12FF1EFBCC" pageId="125" pageNumber="126" type="diagnosis">
<paragraph id="8BA236CCFFC8D965C9C81A12FEE6FEE4" blockId="125.[151,1437,259,2014]" box="[151,274,259,285]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9C81A12FEE6FEE4" bold="true" box="[151,274,259,285]" pageId="125" pageNumber="126">Diagnosis.</emphasis>
</paragraph>
<paragraph id="8BA236CCFFC8D965C9981A36FEBFFEB8" blockId="125.[151,1437,259,2014]" box="[199,331,295,321]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9981A36FEBFFEB8" bold="true" box="[199,331,295,321]" pageId="125" pageNumber="126">Both sexes:</emphasis>
</paragraph>
<paragraph id="8BA236CCFFC8D965C9981A5AFC42FE9F" blockId="125.[151,1437,259,2014]" box="[199,950,331,358]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9981A5AFE72FE9C" box="[199,390,331,357]" italics="true" pageId="125" pageNumber="126">Mandibular palp</emphasis>
: article 2, dorsal margin without a fringe of setae.
</paragraph>
<paragraph id="8BA236CCFFC8D965C9981A7EFC19FE70" blockId="125.[151,1437,259,2014]" box="[199,1005,367,394]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9981A7EFEC7FE70" box="[199,307,367,393]" italics="true" pageId="125" pageNumber="126">Maxilla 1</emphasis>
: without a seta or setal cluster at the base of the palp article 1.
</paragraph>
<paragraph id="8BA236CCFFC8D965C9981A82FE8BFE0F" blockId="125.[151,1437,259,2014]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9981A82FEAEFE54" box="[199,346,403,429]" italics="true" pageId="125" pageNumber="126">Gnathopod 1</emphasis>
: basis, anterolateral margin with a few short setae along its length; carpus with a (usually) single or a small cluster of long setae at the anterodistal junction of the propodus (setae 65-70% of anterior margin length andslightly medial).
</paragraph>
<paragraph id="8BA236CCFFC8D965C9981AEEFAAEFDC7" blockId="125.[151,1437,259,2014]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9981AEEFEADFDE0" box="[199,345,511,537]" italics="true" pageId="125" pageNumber="126">Gnathopod 2</emphasis>
: basis with a few moderately long setae on the anterodistal margin (most setal lengths &lt;40% of the basis width); carpus and propodus, setae on the anterior margin short and simple (setal length &lt;basis width).
</paragraph>
<paragraph id="8BA236CCFFC8D965C9981956FD09FD98" blockId="125.[151,1437,259,2014]" box="[199,765,583,610]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9981956FE9BFD9B" box="[199,367,583,610]" italics="true" pageId="125" pageNumber="126">Pereopods 57</emphasis>
: propodus not expanded anteriorly.
</paragraph>
<paragraph id="8BA236CCFFC8D965C998197AFB59FD7F" blockId="125.[151,1437,259,2014]" box="[199,1197,619,646]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C998197AFEC6FD7C" box="[199,306,619,645]" italics="true" pageId="125" pageNumber="126">Uropod 1</emphasis>
: ventral peduncular spinous process underlying about 1/3 of the longest ramus.
</paragraph>
<paragraph id="8BA236CCFFC8D965C998199EFB64FD53" blockId="125.[151,1437,259,2014]" box="[199,1168,655,682]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C998199EFEC6FD50" box="[199,306,655,681]" italics="true" pageId="125" pageNumber="126">Uropod 3</emphasis>
: inner ramus without spines mid-dorsally (with only the single apical spine).
</paragraph>
<paragraph id="8BA236CCFFC8D965C99819A2FB8CFD37" blockId="125.[151,1437,259,2014]" box="[199,1144,691,718]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C99819A2FEFBFD34" box="[199,271,691,717]" italics="true" pageId="125" pageNumber="126">Telson</emphasis>
: tip without apical setae (only the usual short setae at each dorsolateral cusp).
</paragraph>
<paragraph id="8BA236CCFFC8D965C99819C6FE72FD08" blockId="125.[151,1437,259,2014]" box="[199,390,727,753]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C99819C6FE72FD08" bold="true" box="[199,390,727,753]" pageId="125" pageNumber="126">Thumbed male:</emphasis>
</paragraph>
<paragraph id="8BA236CCFFC8D965C99819EAFC01FCEC" blockId="125.[151,1437,259,2014]" box="[199,1013,763,790]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C99819EAFECEFCEC" box="[199,314,763,789]" italics="true" pageId="125" pageNumber="126">Antenna 2</emphasis>
: without plumose setae on peduncular article 5 and flagellum.
</paragraph>
<paragraph id="8BA236CCFFC8D965C998180EFB67FC5C" blockId="125.[151,1437,259,2014]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C998180EFEA8FCC0" box="[199,348,799,825]" italics="true" pageId="125" pageNumber="126">Gnathopod 2</emphasis>
: propodus large and stout, palmar defining spines not produced on a ledge, spines lost in large thumbed males. Major form: thumb about 30% the length of the propodus and curved posterodistally, dactyl sinuous, expanded proximally. Minor form: thumb about 15% the length of the propodus, dactyl expanded more distally than in the major form, into the palmar incision between the thumb and the proximal tooth.
</paragraph>
<paragraph id="8BA236CCFFC8D965C99818BEFE9FFC30" blockId="125.[151,1437,259,2014]" box="[199,363,943,969]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C99818BEFE9FFC30" bold="true" box="[199,363,943,969]" pageId="125" pageNumber="126">Adult female:</emphasis>
</paragraph>
<paragraph id="8BA236CCFFC8D965C99818C2FC11FC14" blockId="125.[151,1437,259,2014]" box="[199,997,979,1006]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C99818C2FECCFC14" box="[199,312,979,1005]" italics="true" pageId="125" pageNumber="126">Antenna 2</emphasis>
: without plumose setae on peduncular article 5 and flagellum.
</paragraph>
<paragraph id="8BA236CCFFC8D965C99818E6FF1EFBCC" blockId="125.[151,1437,259,2014]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C99818E6FEAFFBE8" box="[199,347,1015,1041]" italics="true" pageId="125" pageNumber="126">Gnathopod 2</emphasis>
: propodus, palm concave, palmar defining angle acute, palmar defining spines relatively widely spaced.
</paragraph>
</subSubSection>
<subSubSection id="C3076547FFC8D966C9981F2EFBDBFF2C" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126" type="discussion">
<paragraph id="8BA236CCFFC8D965C9981F2EFDB2FA38" blockId="125.[151,1437,259,2014]" pageId="125" pageNumber="126">
<emphasis id="B969EADEFFC8D965C9981F2EFECFFBA0" bold="true" box="[199,315,1087,1113]" pageId="125" pageNumber="126">Remarks.</emphasis>
<bibRefCitation id="EF8C4B3DFFC8D965C81C1F2EFE12FBA0" author="Conlan, K. E." box="[323,486,1087,1113]" pageId="125" pageNumber="126" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan (1990)</bibRefCitation>
first recognized
<taxonomicName id="4C1D4D4FFFC8D965CBFB1F51FCB7FBA0" authorityName="Conlan" authorityYear="1990" box="[676,835,1087,1113]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965CBFB1F51FCB7FBA0" box="[676,835,1087,1113]" italics="true" pageId="125" pageNumber="126">Jassa carltoni</emphasis>
</taxonomicName>
as a new species in Californian collections described by
<bibRefCitation id="EF8C4B3DFFC8D965C9E51F72FE92FB87" author="Barnard, J. L." box="[186,358,1123,1150]" journalOrPublisher="Bulletin of the United States National Museum" pageId="125" pageNumber="126" pagination="1 - 230" part="258" refId="ref87998" refString="Barnard, J. L. (1969) Gammaridean amphipoda of the rocky intertidal of California: Monterey Bay to La Jolla. Bulletin of the United States National Museum, 258, 1 - 230. https: // doi. org / 10.5479 / si. 03629236.258.1" title="Gammaridean amphipoda of the rocky intertidal of California: Monterey Bay to La Jolla" type="journal article" year="1969">Barnard (1969)</bibRefCitation>
as “
<emphasis id="B969EADEFFC8D965C8C61F75FDCFFB84" box="[409,571,1123,1149]" italics="true" pageId="125" pageNumber="126">
<taxonomicName id="4C1D4D4FFFC8D965C8C61F75FDC6FB84" authorityName=": Krapp-Schickel" authorityYear="1974" box="[409,562,1123,1149]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="falcata">Jassa falcata</taxonomicName>
</emphasis>
. The largest specimens available were chosen as the type lot (
<figureCitation id="13262A49FFC8D965CDAB1F72FABDFB84" box="[1268,1353,1123,1149]" captionStart="FIGURE 78" captionStartId="126.[151,250,1606,1630]" captionTargetBox="[279,1306,238,1577]" captionTargetId="figure-55@126.[274,1314,237,1583]" captionTargetPageId="126" captionText="FIGURE 78. Jassa carltoni Conlan, 1990. Holotype, adult male, major form, 3.0 mm, Corona del Mar, California, J. L. Barnard, coll., 911 December 1962, station 46-K-1, wash of 1/6 liter of ascidians and soft polychaete tubes collected at low tide (NHMLA 62-132.1). Paratype, juvenile male, 1.7 mm, La Jolla, California, 1113 Nov. 1962, J. L. Barnard, coll., station 45-R- 1, no further collection data, 62-135.1 (NHMLA); subadult male, 4.0 mm, Cable Beach, Vancouver Island, British Columbia, 14 July 1972, D. Kittle, coll., station 342, 1973-128 (CMN); paratype, adult male, minor form, 2.8 mm, La Jolla, California, 1113 Nov. 1962, J. L. Barnard, coll., station 45-W-5, scraping of calcareous sponge under ledge in surge channel at low tide, 62-134.1 (NHMLA); paratype, adult male 2, major form, 2.9 mm, Hazard Canyon Reef, Montana de Oro State Park, St. Louis Obispo County, California, 19 Dec. 1961, J. L. Barnard, coll., station 42-T-5, wash of Egregia holdfasts collected at low tide, 61-100.2 (NHMLA); adult male 3, major form, 4.0 mm, Sarita Bay, Vancouver Island, British Columbia, 2 July 1976, E. L. Bousfield, coll., station B13, 1976-157, bedrock, muddy sand, eelgrass, algae and fine gravel subtidally, hand and dip net collection at low tide, ~14 °C, 26 psu (CMN). Lateral view: whole body and gnathopod 2 views; dorsal view: telson; other views medial. Setae omitted from the juvenile male and major male 2 second gnathopods. Scale 0.1 mm. Illustration after Conlan (1990)." figureDoi="http://doi.org/10.5281/zenodo.4580822" httpUri="https://zenodo.org/record/4580822/files/figure.png" pageId="125" pageNumber="126">Fig. 78</figureCitation>
). However, the
<typeStatus id="54A6886EFFC8D965C9A01F96FE90FB58" box="[255,356,1159,1185]" pageId="125" pageNumber="126" type="holotype">holotype</typeStatus>
, a major form male, appears to have a damaged thumb, as the tip is short and apically indented compared to the major form
<typeStatus id="54A6886EFFC8D965C88C1FBDFDC7FB3F" box="[467,563,1196,1222]" pageId="125" pageNumber="126" type="paratype">paratype</typeStatus>
(
<figureCitation id="13262A49FFC8D965CB1E1FBAFD61FB3C" box="[577,661,1195,1221]" captionStart="FIGURE 79" captionStartId="127.[151,250,983,1007]" captionTargetBox="[276,1309,185,956]" captionTargetId="figure-18@127.[271,1315,181,959]" captionTargetPageId="127" captionText="FIGURE 79. Jassa carltoni Conlan, 1990. Variation in thumb length relative to body length in males in a single collection at Wizard Islet, Vancouver Island, British Columbia, July 1972, D. Kittle, coll., station 712, NMNS 73-128 (CMN). Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580824" httpUri="https://zenodo.org/record/4580824/files/figure.png" pageId="125" pageNumber="126">Fig. 79</figureCitation>
). The minor form
<typeStatus id="54A6886EFFC8D965CA021FBDFC49FB3F" box="[861,957,1196,1222]" pageId="125" pageNumber="126" type="paratype">paratype</typeStatus>
(
<figureCitation id="13262A49FFC8D965CA931FBAFBE8FB3C" box="[972,1052,1195,1221]" captionStart="FIGURE 79" captionStartId="127.[151,250,983,1007]" captionTargetBox="[276,1309,185,956]" captionTargetId="figure-18@127.[271,1315,181,959]" captionTargetPageId="127" captionText="FIGURE 79. Jassa carltoni Conlan, 1990. Variation in thumb length relative to body length in males in a single collection at Wizard Islet, Vancouver Island, British Columbia, July 1972, D. Kittle, coll., station 712, NMNS 73-128 (CMN). Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580824" httpUri="https://zenodo.org/record/4580824/files/figure.png" pageId="125" pageNumber="126">Fig. 79</figureCitation>
) has a similarly shaped but shorter thumb with the dactyl expanded into the palmar incision. Additional specimens from
<collectingRegion id="49D9F82EFFC8D965CD071FDEFAEEFB10" box="[1112,1306,1231,1257]" country="Canada" name="British Columbia" pageId="125" pageNumber="126">British Columbia</collectingRegion>
are figured in order to show the morphology of an undamaged major form male as well as a subadult male that had an obvious thumb inside the cuticle. Plots of thumb and propodus length on gnathopod 2 for specimens from a single population show that the major form thumbed male has a substantially longer thumb and gnathopod 2 propodus length than the minor form thumbed male (
<figureCitation id="13262A49FFC8D965C8AC1E4EFDBEFA80" box="[499,586,1375,1401]" captionStart="FIGURE 79" captionStartId="127.[151,250,983,1007]" captionTargetBox="[276,1309,185,956]" captionTargetId="figure-18@127.[271,1315,181,959]" captionTargetPageId="127" captionText="FIGURE 79. Jassa carltoni Conlan, 1990. Variation in thumb length relative to body length in males in a single collection at Wizard Islet, Vancouver Island, British Columbia, July 1972, D. Kittle, coll., station 712, NMNS 73-128 (CMN). Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580824" httpUri="https://zenodo.org/record/4580824/files/figure.png" pageId="125" pageNumber="126">Figs 79</figureCitation>
,
<figureCitation id="13262A49FFC8D965CB0A1E4EFD81FA80" box="[597,629,1375,1401]" captionStart="FIGURE 80" captionStartId="127.[151,250,1932,1956]" captionTargetBox="[284,1300,1145,1904]" captionTargetId="figure-67@127.[279,1308,1139,1908]" captionTargetPageId="127" captionText="FIGURE 80. Jassa carltoni Conlan, 1990. Variation in gnathopod 2 propodus length relative to body length in males and females in a single collection at Wizard Islet, Vancouver Island, British Columbia, July 1972, D. Kittle, coll., station 712, NMNS 73-128 (CMN). Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580826" httpUri="https://zenodo.org/record/4580826/files/figure.png" pageId="125" pageNumber="126">80</figureCitation>
). Propodus length of adult females is also much less than for major form males but more similar to minor form and subadult males. However there were insufficient specimens of similar body length for statistical comparison.
</paragraph>
<paragraph id="8BA236CCFFC8D965C9981EDDFC3DF8FC" blockId="125.[151,1437,259,2014]" pageId="125" pageNumber="126">
<taxonomicName id="4C1D4D4FFFC8D965C9981EDDFE97FA1C" authorityName="Conlan" authorityYear="1990" box="[199,355,1483,1509]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965C9981EDDFE97FA1C" box="[199,355,1483,1509]" italics="true" pageId="125" pageNumber="126">Jassa carltoni</emphasis>
</taxonomicName>
most resembles
<taxonomicName id="4C1D4D4FFFC8D965CB431EDDFD65FA1C" authorityName="Conlan" authorityYear="1990" box="[540,657,1483,1509]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965CB431EDDFD65FA1C" box="[540,657,1483,1509]" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
, both of which occur on the Pacific coast of North America. Although both have a 4-articled flagellum on antenna 2, the flagellum is slenderer in
<taxonomicName id="4C1D4D4FFFC8D965CA9D1EE1FBA8F9F0" authorityName="Conlan" authorityYear="1990" box="[962,1116,1519,1545]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965CA9D1EE1FBA8F9F0" box="[962,1116,1519,1545]" italics="true" pageId="125" pageNumber="126">Jassa carltoni</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C1D4D4FFFC8D965CDED1EE1FAD0F9F0" authorityName="Conlan" authorityYear="1990" box="[1202,1316,1519,1545]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965CDED1EE1FAD0F9F0" box="[1202,1316,1519,1545]" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
and unlike
<taxonomicName id="4C1D4D4FFFC8D965C9C81D05FEFFF9D4" authorityName="Conlan" authorityYear="1990" box="[151,267,1555,1581]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965C9C81D05FEFFF9D4" box="[151,267,1555,1581]" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
, it does not become plumose in large thumbed males. The palm of gnathopod 1 is convex while in
<taxonomicName id="4C1D4D4FFFC8D965CC0E1D05FF3DF9A8" authorityName="Conlan" authorityYear="1990" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965CC0E1D05FF3DF9A8" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
it is straight or shallowly concave. Both males and females of
<taxonomicName id="4C1D4D4FFFC8D965CADD1D29FC07F9A8" authorityName="Conlan" authorityYear="1990" box="[898,1011,1591,1617]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965CADD1D29FC07F9A8" box="[898,1011,1591,1617]" italics="true" pageId="125" pageNumber="126">J. carltoni</emphasis>
</taxonomicName>
have a sparser setation on the anterior margin of the basis and a wider propodus on gnathopod 2 (maximal width ~65% of maximal length) than for
<taxonomicName id="4C1D4D4FFFC8D965CC0E1D4DFF3DF960" authorityName="Conlan" authorityYear="1990" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965CC0E1D4DFF3DF960" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
(maximal width ~50% of maximal length). The thumb shape also differs: tip rounded and curved posteriorly in major form males of
<taxonomicName id="4C1D4D4FFFC8D965C8DF1DB5FE05F944" authorityName="Conlan" authorityYear="1990" box="[384,497,1699,1725]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965C8DF1DB5FE05F944" box="[384,497,1699,1725]" italics="true" pageId="125" pageNumber="126">J. carltoni</emphasis>
</taxonomicName>
and tip acute and not curved posteriorly in major form males of
<taxonomicName id="4C1D4D4FFFC8D965CDE41DB5FADBF944" authorityName="Conlan" authorityYear="1990" box="[1211,1327,1699,1725]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965CDE41DB5FADBF944" box="[1211,1327,1699,1725]" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
. In minor form males of
<taxonomicName id="4C1D4D4FFFC8D965C8671DD9FE5CF918" authorityName="Conlan" authorityYear="1990" box="[312,424,1735,1761]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965C8671DD9FE5CF918" box="[312,424,1735,1761]" italics="true" pageId="125" pageNumber="126">J. carltoni</emphasis>
</taxonomicName>
, the thumb is shorter and more acute than in the major forms, and more closely resembles
<taxonomicName id="4C1D4D4FFFC8D965CCD61DD9FF05F8FC" authorityName="Conlan" authorityYear="1990" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965CCD61DD9FF05F8FC" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
. The shape differences in the gnathopod 1 palm still hold, though.
</paragraph>
<paragraph id="8BA236CCFFC8D966C9981C01FBDBFF2C" blockId="125.[151,1437,259,2014]" lastBlockId="126.[151,1436,151,213]" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126">
Many (but not all) of the locations for
<taxonomicName id="4C1D4D4FFFC8D965CB281C01FD18F8D0" authorityName="Conlan" authorityYear="1990" box="[631,748,1807,1833]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">
<emphasis id="B969EADEFFC8D965CB281C01FD18F8D0" box="[631,748,1807,1833]" italics="true" pageId="125" pageNumber="126">J. slatteryi</emphasis>
</taxonomicName>
noted by
<bibRefCitation id="EF8C4B3DFFC8D965CA031C1EFC09F8D0" author="Conlan, K. E." box="[860,1021,1807,1833]" pageId="125" pageNumber="126" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan (1990)</bibRefCitation>
for
<collectingRegion id="49D9F82EFFC8D965CD721C1EFB18F8D0" box="[1069,1260,1807,1833]" country="Canada" name="British Columbia" pageId="125" pageNumber="126">British Columbia</collectingRegion>
are actually for
<taxonomicName id="4C1D4D4FFFC8D965C9C81C25FEFEF8B4" authorityName="Conlan" authorityYear="1990" box="[151,266,1843,1869]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965C9C81C25FEFEF8B4" box="[151,266,1843,1869]" italics="true" pageId="125" pageNumber="126">J. carltoni</emphasis>
</taxonomicName>
, as this latter species was not recognized at that time to occur outside of
<collectingRegion id="49D9F82EFFC8D965CD1D1C22FB41F8B4" box="[1090,1205,1843,1869]" country="United States of America" name="California" pageId="125" pageNumber="126">California</collectingRegion>
. It was only through recent molecular analysis that
<taxonomicName id="4C1D4D4FFFC8D965C8B71C49FDADF888" authorityName="Conlan" authorityYear="1990" box="[488,601,1879,1905]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965C8B71C49FDADF888" box="[488,601,1879,1905]" italics="true" pageId="125" pageNumber="126">J. carltoni</emphasis>
</taxonomicName>
was discovered to occur also in
<collectingRegion id="49D9F82EFFC8D965CA9B1C46FB72F888" box="[964,1158,1879,1905]" country="Canada" name="British Columbia" pageId="125" pageNumber="126">British Columbia</collectingRegion>
. Subsequent re-examination of collections found
<taxonomicName id="4C1D4D4FFFC8D965C8F31C6DFDE9F86C" authorityName="Conlan" authorityYear="1990" box="[428,541,1915,1941]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965C8F31C6DFDE9F86C" box="[428,541,1915,1941]" italics="true" pageId="125" pageNumber="126">J. carltoni</emphasis>
</taxonomicName>
also in
<collectingRegion id="49D9F82EFFC8D965CB2B1C6AFD3EF86C" box="[628,714,1915,1941]" country="United States of America" name="Oregon" pageId="125" pageNumber="126">Oregon</collectingRegion>
. A “
<emphasis id="B969EADEFFC8D965CBA71C6DFC88F86C" box="[760,892,1915,1941]" italics="true" pageId="125" pageNumber="126">
<taxonomicName id="4C1D4D4FFFC8D965CBA71C6DFC85F86C" authorityName="Conlan" authorityYear="1990" box="[760,881,1915,1941]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="slatteryi">J. slatteryi</taxonomicName>
</emphasis>
noted by
<bibRefCitation id="EF8C4B3DFFC8D965CAB61C6AFB7DF86C" author="Conlan, K. E." box="[1001,1161,1915,1941]" pageId="125" pageNumber="126" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan (1990)</bibRefCitation>
from
<collectingRegion id="49D9F82EFFC8D965CD931C6AFAECF86C" box="[1228,1304,1915,1941]" country="United States of America" name="Alaska" pageId="125" pageNumber="126">Alaska</collectingRegion>
(University of
<collectingRegion id="49D9F82EFFC8D965C9EC1C8EFF0BF840" box="[179,255,1951,1977]" country="United States of America" name="Alaska" pageId="125" pageNumber="126">Alaska</collectingRegion>
loan, Latouche T1, M75,
<date id="FFA3100CFFC8D965CB451C8EFD57F843" box="[538,675,1951,1978]" pageId="125" pageNumber="126" value="1975-09-05">5 Sept. 1975</date>
) may also be
<taxonomicName id="4C1D4D4FFFC8D965CA661CB1FC5DF840" authorityName="Conlan" authorityYear="1990" box="[825,937,1951,1977]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFC8D965CA661CB1FC5DF840" box="[825,937,1951,1977]" italics="true" pageId="125" pageNumber="126">J. carltoni</emphasis>
</taxonomicName>
but this specimen was no longer available for study and so is omitted from the distribution map. Given the similarity in environmental conditions between southeastern
<collectingRegion id="49D9F82EFFCBD966C9E01B86FEFFFF48" box="[191,267,151,177]" country="United States of America" name="Alaska" pageId="126" pageNumber="127">Alaska</collectingRegion>
and
<collectingRegion id="49D9F82EFFCBD966C81F1B86FDF4FF48" box="[320,512,151,177]" country="Canada" name="British Columbia" pageId="126" pageNumber="127">British Columbia</collectingRegion>
, it is likely that
<taxonomicName id="4C1D4D4FFFCBD966CBF21B86FCE9FF48" authorityName="Conlan" authorityYear="1990" box="[685,797,151,177]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFCBD966CBF21B86FCE9FF48" box="[685,797,151,177]" italics="true" pageId="126" pageNumber="127">J. carltoni</emphasis>
</taxonomicName>
ranges northwards at least into southeastern
<collectingRegion id="49D9F82EFFCBD966CC5A1B86FAA1FF48" box="[1285,1365,151,177]" country="United States of America" name="Alaska" pageId="126" pageNumber="127">Alaska</collectingRegion>
.
<taxonomicName id="4C1D4D4FFFCBD966CC001B86FF1AFF2C" authorityName="Conlan" authorityYear="1990" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B969EADEFFCBD966CC001B86FF1AFF2C" italics="true" pageId="126" pageNumber="127">Jassa carltoni</emphasis>
</taxonomicName>
has not been found outside of the Pacific coast of North America, however.
</paragraph>
</subSubSection>
</treatment>
</document>