305 lines
54 KiB
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305 lines
54 KiB
XML
<document id="EDF03DA5F16F4E00A1318BEE08ABC503" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FFF4E30DFF78FE52FD3D85D8" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Harpes boudibensis Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="46" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="44" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:title id="734FAD80867B62FE1D2B34C3964F1E11">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<mods:namePart id="38E13D7B023519DFEC56B64F9CF85FD5">Johnson, Robert G.</mods:namePart>
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<mods:typeOfResource id="59D98EA60C1DFB7F0369FA3ECB606766">text</mods:typeOfResource>
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<mods:title id="CD16BD41A2317F7F1378DC0C9B2028DB">Zootaxa</mods:title>
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<mods:part id="EFD34462001C04134089CDD9544E5969">
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<mods:date id="7F2C2240B004111C6DD113C0FCE038F0">2024</mods:date>
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<mods:number id="883C5790BCE5D804BC3EA46C6EECCB1D">2024-05-15</mods:number>
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<mods:number id="90F7DFD4631C5DB5ECAAB79A547520ED">5450</mods:number>
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<mods:url id="AD24B67DB9AE97341A4B9AF8188BAC1E">http://dx.doi.org/10.11646/zootaxa.5450.1.1</mods:url>
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<mods:classification id="4CF76395597E76FCC631198CA55CEEB1">journal article</mods:classification>
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<mods:identifier id="1839771FD091FCBC4801208763124034" type="DOI">10.11646/zootaxa.5450.1.1</mods:identifier>
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<mods:identifier id="56AA27028E913F158C3A616327D44569" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="4E0C24CB8729747B6F555B05FC4902AC" type="Zenodo-Dep">11232500</mods:identifier>
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<mods:identifier id="33992C67C97BA367D15EA741F9063DE5" type="ZooBank">1B5D192F-1D5B-4460-9133-9AEAE9C920BF</mods:identifier>
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<treatment id="03FD8227FFF4E30DFF78FE52FD3D85D8" ID-DOI="http://doi.org/10.5281/zenodo.11232537" ID-Zenodo-Dep="11232537" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8" httpUri="http://treatment.plazi.org/id/03FD8227FFF4E30DFF78FE52FD3D85D8" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
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||
<subSubSection id="C34E60BAFFF4E30BFF78FE52FE658139" box="[151,464,1581,1608]" pageId="43" pageNumber="44" type="nomenclature">
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||
<paragraph id="8BEB3331FFF4E30BFF78FE52FE658139" blockId="43.[151,464,1581,1644]" box="[151,464,1581,1608]" pageId="43" pageNumber="44">
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||
<heading id="D0A3845DFFF4E30BFF78FE52FE658139" bold="true" box="[151,464,1581,1608]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="4">
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||
<emphasis id="B920EF23FFF4E30BFF78FE52FE658139" bold="true" box="[151,464,1581,1608]" pageId="43" pageNumber="44">
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<taxonomicName id="4C5448B2FFF4E30BFF78FE52FE338139" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,390,1581,1608]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="boudibensis" status="sp. nov.">
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<emphasis id="B920EF23FFF4E30BFF78FE52FE338139" bold="true" box="[151,390,1581,1608]" italics="true" pageId="43" pageNumber="44">Harpes boudibensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2135258FFF4E30BFE62FE52FE658139" box="[397,464,1582,1608]" pageId="43" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFF4E30BFF78FE2EFEAF811D" box="[151,282,1618,1644]" pageId="43" pageNumber="44" type="description">
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<paragraph id="8BEB3331FFF4E30BFF78FE2EFEAF811D" blockId="43.[151,464,1581,1644]" box="[151,282,1618,1644]" pageId="43" pageNumber="44">
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<figureCitation id="136F2FB4FFF4E30BFF78FE2EFEAF811D" box="[151,282,1618,1644]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="43" pageNumber="44">Plate 1A–Q</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFF4E30BFF78FEE6FC498034" pageId="43" pageNumber="44" type="diagnosis">
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<paragraph id="8BEB3331FFF4E30BFF78FEE6FC498034" blockId="43.[151,1437,1690,2041]" pageId="43" pageNumber="44">
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<emphasis id="B920EF23FFF4E30BFF78FEE6FEA781C5" bold="true" box="[151,274,1690,1716]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
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Narrow cephalon, width (tr.) around 1.2 times sagittal length and widest (tr.) just posterior to eyes. Glabella cone-shaped, anteriorly rounded. Extension of axial furrow across ala distinct. Alae inflated and alar furrows proximally deep, becoming very shallow distally. Alar depression pronounced. Prolongations around same length (exsag.) as cephalon (sag.). In lateral view external rim posteriorly curving upwards and internal rim wave-like in shape and terminated by short genal spine. Over 25 thoracic segments.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFF4E30BFF28FF32FBFC8019" box="[199,1097,1870,1897]" pageId="43" pageNumber="44" type="etymology">
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<paragraph id="8BEB3331FFF4E30BFF28FF32FBFC8019" blockId="43.[151,1437,1690,2041]" box="[199,1097,1870,1897]" pageId="43" pageNumber="44">
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<emphasis id="B920EF23FFF4E30BFF28FF32FEFB8019" bold="true" box="[199,334,1870,1896]" pageId="43" pageNumber="44">Etymology.</emphasis>
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Named after the village bou Dîb, which is close to the
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<typeStatus id="54EF8D93FFF4E30BFC57FF33FC5D8018" box="[952,1000,1871,1897]" pageId="43" pageNumber="44">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFFF4E30CFF28FF0EFDD08630" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="materials_examined">
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<paragraph id="8BEB3331FFF4E30CFF28FF0EFC9187A4" blockId="43.[151,1437,1690,2041]" lastBlockId="44.[151,1437,151,2013]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
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<emphasis id="B920EF23FFF4E30BFF28FF0EFE4180FD" bold="true" box="[199,500,1906,1932]" pageId="43" pageNumber="44">Material and occurrence.</emphasis>
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<materialsCitation id="3B3C396CFFF4E30BFE15FF0EFAAF80C1" collectionCode="NHMUK" collectorName="It & Section & El Otfal Formation & Eifelian & Jbel Issoumour" pageId="43" pageNumber="44" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8D93FFF4E30BFE15FF0EFDD280FD" box="[506,615,1906,1932]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED45ABF4FFF4E30BFD9FFF0FFD6F80FC" box="[624,730,1907,1933]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="43" pageNumber="44" type="Museum">NHMUK</collectionCode>
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<collectorName id="26A156E7FFF4E30BFD0FFF0FFD4480FC" box="[736,753,1907,1933]" pageId="43" pageNumber="44">It</collectorName>
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29227,
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<figureCitation id="136F2FB4FFF4E30BFCA4FF0EFC0380FD" box="[843,950,1906,1932]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="43" pageNumber="44">Pl. 1K–Q</figureCitation>
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, from horizon 1,
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<collectorName id="26A156E7FFF4E30BFB9AFF0EFB7C80FD" box="[1141,1225,1906,1932]" pageId="43" pageNumber="44">Section</collectorName>
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2 (
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<figureCitation id="136F2FB4FFF4E30BFB04FF0EFA9A80FD" box="[1259,1327,1906,1932]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[159,1431,194,1849]" captionTargetId="figure-15@13.[151,1435,181,1857]" captionTargetPageId="13" captionText="FIGURE 7. Section 2 Jbel Issoumour" figureDoi="http://doi.org/10.5281/zenodo.11232517" httpUri="https://zenodo.org/record/11232517/files/figure.png" pageId="43" pageNumber="44">Fig. 7</figureCitation>
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),
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<collectorName id="26A156E7FFF4E30BFAABFF0EFEBA80C1" pageId="43" pageNumber="44">El Otfal Formation</collectorName>
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,
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<collectorName id="26A156E7FFF4E30BFEF3FFEAFECD80C1" box="[284,376,1942,1968]" pageId="43" pageNumber="44">Eifelian</collectorName>
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,
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<collectorName id="26A156E7FFF4E30BFE6AFFEAFD8780C0" box="[389,562,1942,1969]" pageId="43" pageNumber="44">Jbel Issoumour</collectorName>
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(
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<figureCitation id="136F2FB4FFF4E30BFDABFFEAFD2B80C1" box="[580,670,1942,1968]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="43" pageNumber="44">Fig. 2B</figureCitation>
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, Map 7, site 4), enrolled articulated dorsal exoskeleton
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</materialsCitation>
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.
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<materialsCitation id="3B3C396CFFF4E30BFAC7FFEBFCDA80A5" collectionCode="NHMUK" pageId="43" pageNumber="44" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54EF8D93FFF4E30BFAC7FFEBFA2C80C0" box="[1320,1433,1943,1969]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED45ABF4FFF4E30BFF78FFC7FEB480A4" box="[151,257,1979,2005]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="43" pageNumber="44" type="Museum">NHMUK</collectionCode>
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It 29226,
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<figureCitation id="136F2FB4FFF4E30BFE99FFC6FE5680A5" box="[374,483,1978,2004]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="43" pageNumber="44">Pl. 1A–D</figureCitation>
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, articulated dorsal exoskeleton and
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</materialsCitation>
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<collectionCode id="ED45ABF4FFF4E30BFC98FFC7FC5480A4" box="[887,993,1979,2005]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="43" pageNumber="44" type="Museum">NHMUK</collectionCode>
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It 29228,
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<figureCitation id="136F2FB4FFF4E30BFBB9FFC6FB0280A5" box="[1110,1207,1978,2004]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="43" pageNumber="44">Pl. 1E–J</figureCitation>
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, dorsal exoskeleton, pygidium enrolled.
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<collectionCode id="ED45ABF4FFF4E30BFE9AFFA3FE6A8088" box="[373,479,2015,2041]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="43" pageNumber="44" type="Museum">NHMUK</collectionCode>
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It 29226 and
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<collectionCode id="ED45ABF4FFF4E30BFD92FFA3FD528088" box="[637,743,2015,2041]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="43" pageNumber="44" type="Museum">NHMUK</collectionCode>
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It 29228 from the type locality and horizon, site 1 and site 5, respectively.
|
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<taxonomicName id="4C5448B2FFF3E30CFEC2F8EBFDBA87C0" authorityName="Johnson" authorityYear="2024" box="[301,527,151,177]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="boudibensis">
|
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<emphasis id="B920EF23FFF3E30CFEC2F8EBFDBA87C0" box="[301,527,151,177]" italics="true" pageId="44" pageNumber="45">Harpes boudibensis</emphasis>
|
||
</taxonomicName>
|
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is almost certainly present at other Jbel Issoumour Eifelian localities, many of which have been extensively mined for harpetid trilobites.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF3E30CFF28F8A3FDD08630" blockId="44.[151,1437,151,2013]" pageId="44" pageNumber="45">
|
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Other material:
|
||
<taxonomicName id="4C5448B2FFF3E30CFE97F8A3FD048788" authority=", NHMUK It" authorityName="NHMUK It" box="[376,689,223,249]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
cf.
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<emphasis id="B920EF23FFF3E30CFE73F8A3FD9D8788" box="[412,552,223,249]" italics="true" pageId="44" pageNumber="45">boudibensis,</emphasis>
|
||
NHMUK It
|
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</taxonomicName>
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29225 from horizon 7, Section 1 (
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<figureCitation id="136F2FB4FFF3E30CFBDCF8A3FBC28788" box="[1075,1143,223,249]" captionStart="FIGURE 6" captionStartId="12.[151,250,1857,1881]" captionTargetBox="[154,1432,185,1829]" captionTargetId="figure-20@12.[151,1435,181,1833]" captionTargetPageId="12" captionText="FIGURE 6. Section 1 Jbel Ou-Driss" figureDoi="http://doi.org/10.5281/zenodo.11232515" httpUri="https://zenodo.org/record/11232515/files/figure.png" pageId="44" pageNumber="45">Fig. 6</figureCitation>
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), El Otfal Formation, Jbel Ou-Driss (
|
||
<figureCitation id="136F2FB4FFF3E30CFEE1F97FFED3866C" box="[270,358,259,285]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="44" pageNumber="45">Fig. 2B</figureCitation>
|
||
, Map 4, site 11),
|
||
<figureCitation id="136F2FB4FFF3E30CFDC4F97FFD2D866C" box="[555,664,259,285]" captionStart="PLATE 2" captionStartId="108.[151,234,1756,1780]" captionTargetBox="[202,1386,181,1732]" captionTargetId="figure-20@108.[202,1386,181,1732]" captionTargetPageId="108" captionText="PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232532" httpUri="https://zenodo.org/record/11232532/files/figure.png" pageId="44" pageNumber="45">Pl. 2A–D</figureCitation>
|
||
, an enrolled dorsal exoskeleton. Horizon 7 of a similar age to the
|
||
<taxonomicName id="4C5448B2FFF3E30CFA6EF97FFEA88630" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF3E30CFA6EF97FFEA88630" italics="true" pageId="44" pageNumber="45">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
horizons on Jebel Issoumour
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFF3E30CFF28F936FC668074" pageId="44" pageNumber="45" type="description">
|
||
<paragraph id="8BEB3331FFF3E30CFF28F936FAFF8568" blockId="44.[151,1437,151,2013]" pageId="44" pageNumber="45">
|
||
<emphasis id="B920EF23FFF3E30CFF28F936FEEC8615" bold="true" box="[199,345,330,356]" pageId="44" pageNumber="45">Description.</emphasis>
|
||
In dorsal view, cephalon widest just posterior to level of eyes, where length (sag.) ≈ 80% of width (tr.). Cephalon and glabella strongly vaulted. Glabella cone-shaped, rounded anteriorly and sometimes with sparse fine tubercles along posterior two thirds of top of glabella. L1 weakly inflated. S1, S2 and S3 poorly marked by change in tone of cuticle (
|
||
<figureCitation id="136F2FB4FFF3E30CFE51F9CBFDB086A0" box="[446,517,439,465]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pl. 1P</figureCitation>
|
||
). Shallow furrow which rises 2/3 of way up flank of glabella delineates posterior edge of S1 muscle attachment scar area (
|
||
<figureCitation id="136F2FB4FFF3E30CFDB6F9A7FD168684" box="[601,675,475,501]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pl. 1A</figureCitation>
|
||
). S1 tulip-shaped, S2 and S3 subcircular and about 1/4 size of S1. S0 oval and about two thirds of area of S1. S1 with subparallel intermittent dark vertical stripes and dark edge.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF3E30CFF28FA5FFBC584B8" blockId="44.[151,1437,151,2013]" pageId="44" pageNumber="45">
|
||
<materialsCitation id="3B3C396CFFF3E30CFF28FA5FFBDD84CD" collectionCode="NHMUK" county="Genal" location="It" municipality="Broad" pageId="44" pageNumber="45" specimenCount="2" typeStatus="holotype">
|
||
Preglabellar furrow very shallow and preglabellar genal area length (sag.) around 45% of width (tr.) of glabella at S1. Axial furrows deepening posteriorly and alar depression prominent. Alae large, reaching more than 70% of way across (tr.) genal area. Low rounded ridge crosses (exsag.) ala close to axial furrow, visible on
|
||
<typeStatus id="54EF8D93FFF3E30CFB18FA17FAEC85F4" box="[1271,1369,619,645]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
|
||
in
|
||
<figureCitation id="136F2FB4FFF3E30CFA91FA17FF0B85D8" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pl. 1M</figureCitation>
|
||
(lateral view) and in dorsal view on
|
||
<typeStatus id="54EF8D93FFF3E30CFD8EFAF3FD7485D8" box="[609,705,655,681]" pageId="44" pageNumber="45" type="paratype">paratype</typeStatus>
|
||
<collectionCode id="ED45ABF4FFF3E30CFD25FAF3FC8185D8" box="[714,820,655,681]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="44" pageNumber="45" type="Museum">NHMUK</collectionCode>
|
||
<location id="8E8B65EAFFF3E30CFCD4FAF3FCF985D8" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFCD4FAF3FCF985D8" box="[827,844,655,681]" county="Genal" municipality="Broad" name="It" pageId="44" pageNumber="45">It</location>
|
||
29226 (
|
||
<figureCitation id="136F2FB4FFF3E30CFC44FAF3FC4285D8" box="[939,1015,655,681]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pl. 1A</figureCitation>
|
||
).
|
||
<location id="8E8B65EAFFF3E30CFBE4FAF3FB8885D8" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFBE4FAF3FB8885D8" box="[1035,1085,655,681]" county="Genal" municipality="Broad" name="Alar" pageId="44" pageNumber="45">Alar</location>
|
||
furrows shallow, well-defined, becoming shallow distally.
|
||
<location id="8E8B65EAFFF3E30CFE20FACFFD8385BC" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFE20FACFFD8385BC" box="[463,566,691,717]" county="Genal" municipality="Broad" name="Occipital" pageId="44" pageNumber="45">Occipital</location>
|
||
furrow broad, occipital ring narrow (sag. and exsag.) and covered with fine granular tubercles.
|
||
<location id="8E8B65EAFFF3E30CFE83FAABFE0D8580" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFE83FAABFE0D8580" box="[364,440,727,753]" county="Genal" municipality="Broad" name="Larger" pageId="44" pageNumber="45">Larger</location>
|
||
, low axial tubercle located on anterior edge of ring.
|
||
<location id="8E8B65EAFFF3E30CFC1BFAABFBEC8580" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFC1BFAABFBEC8580" box="[1012,1113,727,753]" county="Genal" municipality="Broad" name="Posterior" pageId="44" pageNumber="45">Posterior</location>
|
||
border very short (tr.), ≈30% of width (tr.) of occipital ring.
|
||
<location id="8E8B65EAFFF3E30CFE09FA87FDEB8464" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFE09FA87FDEB8464" box="[486,606,763,789]" county="Genal" municipality="Broad" name="Genal area" pageId="44" pageNumber="45">Genal area</location>
|
||
narrow, width (tr.) around 50% of length (exsag.).
|
||
<location id="8E8B65EAFFF3E30CFB62FA87FB0C8464" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFB62FA87FB0C8464" box="[1165,1209,763,789]" county="Genal" municipality="Broad" name="Eye" pageId="44" pageNumber="45">Eye</location>
|
||
lobes nearly conical in shape, reaching across to near inner margin of fringe, with eye position ratio about 0.85.
|
||
<location id="8E8B65EAFFF3E30CFB61FB63FB0A8448" LSID="urn:lsid:plazi:treatment:03FD8227FFF4E30DFF78FE52FD3D85D8:8E8B65EAFFF3E30CFB61FB63FB0A8448" box="[1166,1215,799,825]" county="Genal" municipality="Broad" name="Two" pageId="44" pageNumber="45">Two</location>
|
||
horizontal oval eye lenses stadium-shape and well separated, face outwards anterolaterally and posterolaterally, respectively (
|
||
<figureCitation id="136F2FB4FFF3E30CFAA8FB3FFA22842C" box="[1351,1431,835,861]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pls 1Q</figureCitation>
|
||
,
|
||
<figureCitation id="136F2FB4FFF3E30CFF78FB1BFF7784F0" box="[151,194,871,897]" captionStart="PLATE 2" captionStartId="108.[151,234,1756,1780]" captionTargetBox="[202,1386,181,1732]" captionTargetId="figure-20@108.[202,1386,181,1732]" captionTargetPageId="108" captionText="PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232532" httpUri="https://zenodo.org/record/11232532/files/figure.png" pageId="44" pageNumber="45">2M</figureCitation>
|
||
).
|
||
<collectingMunicipality id="6B8FA94BFFF3E30CFF38FB1BFEAE84F0" box="[215,283,871,897]" pageId="44" pageNumber="45">Broad</collectingMunicipality>
|
||
low ridge reaching down from between the two lenses onto genal area, where it narrows and forms a very faint genal ridge, extending posterolaterally to cross inner margin of fringe level with ala.
|
||
<collectingCounty id="628A4BBDFFF3E30CFB24FBF7FABB84D4" box="[1227,1294,907,933]" pageId="44" pageNumber="45">Genal</collectingCounty>
|
||
roll convex, wide and sloping gently anteriorly at under
|
||
<superScript id="7C219E79FFF3E30CFD6FFBD3FD1184CD" attach="left" box="[640,676,942,969]" fontSize="6" pageId="44" pageNumber="45">400</superScript>
|
||
and laterally more steeply at around
|
||
<superScript id="7C219E79FFF3E30CFBABFBD2FBDD84CD" attach="none" box="[1092,1128,942,969]" fontSize="6" pageId="44" pageNumber="45">550</superScript>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF3E30CFF28FBAFFB498398" blockId="44.[151,1437,151,2013]" pageId="44" pageNumber="45">
|
||
Brim sloping moderately anteriorly; width (sag.) about 30% of cephalic length (sag.) and brim width ratio around 0.50. A row of large perforations lie either side of girder, away from which perforations quickly decrease in size to those on main part of genal roll and brim respectively. Average standardised diameter of perforations on central part of brim around 150 μm (
|
||
<figureCitation id="136F2FB4FFF3E30CFDB4FC43FD518328" box="[603,740,1087,1113]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pl. 1D, J, O</figureCitation>
|
||
). Perforations on main part of genal roll finer (
|
||
<figureCitation id="136F2FB4FFF3E30CFB16FC43FAF68328" box="[1273,1347,1087,1113]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pl. 1N</figureCitation>
|
||
) except where inner margin of fringe crosses in front of glabella, where there is a row of larger perforations. Around external margin of brim, usual row of coarser perforations. External rim raised and broad with dorsal rim suture central. Marginal band straight, vertical and without tubercles. Material described includes one poorly preserved hypostome (
|
||
<figureCitation id="136F2FB4FFF3E30CFF4FFCB3FEB48398" box="[160,257,1231,1257]" captionStart="PLATE 1" captionStartId="107.[151,234,1710,1734]" captionTargetBox="[226,1360,181,1685]" captionTargetId="figure-15@107.[225,1361,181,1687]" captionTargetPageId="107" captionText="PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232530" httpUri="https://zenodo.org/record/11232530/files/figure.png" pageId="44" pageNumber="45">Pl. 1H, I</figureCitation>
|
||
), which has an inverted cone shape with no discernible middle furrow and narrow borders.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF3E30CFF28FC8FFC148178" blockId="44.[151,1437,151,2013]" pageId="44" pageNumber="45">
|
||
Prolongations around 1.05 times sagittal cephalic length and internal rims set at about
|
||
<superScript id="7C219E79FFF3E30CFB62FC8FFB0A8271" attach="left" box="[1165,1215,1266,1293]" fontSize="6" pageId="44" pageNumber="45">1400</superScript>
|
||
to posterior border, curving inwards posteriorly so that distal ends point obliquely adaxially. In lateral view, external brim narrows and curves steadily upwards posteriorly to meet internal rim at end of prolongation. Internal rim rises gently from posterior border then curves down towards distal end, before curving up to meet external rim to form very short stout spine. Lateral profile of internal rim wave-like. In dorsal view, internal rim broad, rounded and covered by small tubercles. Extension of girder on prolongations rises to run just under internal rim before joining it at about midpoint. Perforations on extension of genal roll on prolongation increase in size towards internal rim. Below internal rim a row of large perforations becoming smaller posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF3E30CFF28FE6FFC668074" blockId="44.[151,1437,151,2013]" pageId="44" pageNumber="45">
|
||
Thorax has 26 to 28 thoracic segments. Axis strongly vaulted and with axial rings becoming narrower (tr.) posteriorly, last having width (tr.) of about 30% of first. Postannuli wider (sag. and exsag.) than preannuli and not narrowing medially. Pleurae widening (tr.) posteriorly until 7
|
||
<superScript id="7C219E79FFF3E30CFC99FE26FC368119" attach="left" box="[886,899,1626,1640]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
|
||
segment and then narrowing until last segment which is just over 20% of 7
|
||
<superScript id="7C219E79FFF3E30CFE23FE02FE6C81FD" attach="none" box="[460,473,1662,1676]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
|
||
. Pleurae narrow (exsag.) with anterior and posterior bands approximately same width (exsag.) and a relatively deep pleural furrow. Pleural furrows cross onto outer portion of pleurae, which are turned downwards, straight and terminate in rounded end. Pygidium very short (sag.) with two axial rings and pleural furrows extending to distal edge. Terminal piece short (sag.) and rounded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFFF3E30DFF28FF73FD3D85D8" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="discussion">
|
||
<paragraph id="8BEB3331FFF3E30DFF28FF73FD6587A4" blockId="44.[151,1437,151,2013]" lastBlockId="45.[151,1437,151,681]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
|
||
<emphasis id="B920EF23FFF3E30CFF28FF73FE8E8058" bold="true" box="[199,315,1807,1833]" pageId="44" pageNumber="45">Remarks.</emphasis>
|
||
<taxonomicName id="4C5448B2FFF3E30CFEABFF73FD918058" box="[324,548,1807,1833]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF3E30CFEABFF73FD918058" box="[324,548,1807,1833]" italics="true" pageId="44" pageNumber="45">Harpes boudibensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C5448B2FFF3E30CFD52FF73FC3F8058" authorityName="Goldfuss" authorityYear="1839" box="[701,906,1807,1833]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="macrocephalus">
|
||
<emphasis id="B920EF23FFF3E30CFD52FF73FC3F8058" box="[701,906,1807,1833]" italics="true" pageId="44" pageNumber="45">H. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
in having a cephalon which is widest (tr.) just posterior to the eye lobes rather than the anterior of the alae and is ovoid rather than U-shaped. Its glabella anterior to S0 lacks tubercles and S1 furrow is shallower, becoming effaced before reaching the top of muscle attachment area. Preglabellar furrow of
|
||
<taxonomicName id="4C5448B2FFF3E30CFE75FF07FDF680E4" box="[410,579,1915,1941]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF3E30CFE75FF07FDF680E4" box="[410,579,1915,1941]" italics="true" pageId="44" pageNumber="45">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
is very shallow as is the extension of the axial furrow across the ala. Genal area and genal roll not so strongly convex and anterior slope of genal roll is under
|
||
<superScript id="7C219E79FFF3E30CFBD1FFE3FBD780DD" attach="left" box="[1086,1122,1950,1977]" fontSize="6" pageId="44" pageNumber="45">400</superScript>
|
||
compared with over
|
||
<superScript id="7C219E79FFF3E30CFABCFFE2FAC280DD" attach="left" box="[1363,1399,1950,1977]" fontSize="6" pageId="44" pageNumber="45">500</superScript>
|
||
for that of
|
||
<emphasis id="B920EF23FFF3E30CFF09FFBFFE0F80AC" box="[230,442,1987,2013]" italics="true" pageId="44" pageNumber="45">
|
||
<taxonomicName id="4C5448B2FFF3E30CFF09FFBFFE0380AC" authorityName="Goldfuss" authorityYear="1839" box="[230,438,1987,2013]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="macrocephalus">H. macrocephalus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The prolongations are a third longer (exsag.) in relation to length (sag.) of cephalon than those of
|
||
<taxonomicName id="4C5448B2FFF2E30DFF18F8EBFE7687C0" authorityName="Goldfuss" authorityYear="1839" box="[247,451,151,177]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="macrocephalus">
|
||
<emphasis id="B920EF23FFF2E30DFF18F8EBFE7687C0" box="[247,451,151,177]" italics="true" pageId="45" pageNumber="46">H. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
. In dorsal view, internal rims curve adaxially rather than being subparallel and in lateral view, their profile is wave-like rather than straight.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF2E30DFF28F8A3FC258614" blockId="45.[151,1437,151,681]" pageId="45" pageNumber="46">
|
||
<taxonomicName id="4C5448B2FFF2E30DFF28F8A3FDF08788" authority=", NHMUK" authorityName="NHMUK" box="[199,581,223,249]" class="Trilobita" family="Harpetidae" genus="Harpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF2E30DFF28F8A3FEAC8789" box="[199,281,223,248]" italics="true" pageId="45" pageNumber="46">Harpes</emphasis>
|
||
cf.
|
||
<emphasis id="B920EF23FFF2E30DFEA9F8A3FE798788" box="[326,460,223,249]" italics="true" pageId="45" pageNumber="46">boudibensis</emphasis>
|
||
, NHMUK
|
||
</taxonomicName>
|
||
it 29225, (
|
||
<figureCitation id="136F2FB4FFF2E30DFD2EF8A3FC9B8788" box="[705,814,223,249]" captionStart="PLATE 2" captionStartId="108.[151,234,1756,1780]" captionTargetBox="[202,1386,181,1732]" captionTargetId="figure-20@108.[202,1386,181,1732]" captionTargetPageId="108" captionText="PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232532" httpUri="https://zenodo.org/record/11232532/files/figure.png" pageId="45" pageNumber="46">Pl. 2A–D</figureCitation>
|
||
) is very similar to
|
||
<taxonomicName id="4C5448B2FFF2E30DFBEAF8A3FB1A8788" box="[1029,1199,223,249]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF2E30DFBEAF8A3FB1A8788" box="[1029,1199,223,249]" italics="true" pageId="45" pageNumber="46">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
but differs in having fine tubercles scattered along top of the glabella, a wider occipital furrow which is medially bent forward, an occipital ring which is narrower, a slightly shorter (tr.) posterior border, more inflated alae and a steeper-sloping genal roll (sag.). Some of these features are shared with
|
||
<taxonomicName id="4C5448B2FFF2E30DFCE2F937FC3C8614" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[781,905,331,357]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="174" phylum="Arthropoda" rank="species" species="lahceni" status="sp. nov.">
|
||
<emphasis id="B920EF23FFF2E30DFCE2F937FC3C8614" box="[781,905,331,357]" italics="true" pageId="45" pageNumber="46">H. Lahceni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FFF2E30DFF28F913FD3D85D8" blockId="45.[151,1437,151,681]" pageId="45" pageNumber="46">
|
||
<taxonomicName id="4C5448B2FFF2E30DFF28F913FE1C86F8" box="[199,425,367,393]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF2E30DFF28F913FE1C86F8" box="[199,425,367,393]" italics="true" pageId="45" pageNumber="46">Harpes boudibensis</emphasis>
|
||
</taxonomicName>
|
||
has been widely mistaken for
|
||
<taxonomicName id="4C5448B2FFF2E30DFCE3F913FAE786F8" authority="(Richter & Richter, 1943)" baseAuthorityName="Richter & Richter" baseAuthorityYear="1943" box="[780,1362,367,393]" class="Trilobita" family="Harpetidae" genus="Helioharpes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="perradiatus">
|
||
<emphasis id="B920EF23FFF2E30DFCE3F913FB9786F8" box="[780,1058,367,393]" italics="true" pageId="45" pageNumber="46">Helioharpes perradiatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC54EC0FFF2E30DFBDCF913FAFF86F8" author="Richter, R. & Richter, E." box="[1075,1354,367,393]" pageId="45" pageNumber="46" refId="ref90337" refString="Richter, R. & Richter, E. (1943) Studien im Palaozoikum der Mittelmeer-Lander. 4 a. Trilobiten aus dem Devon von Marokko. Senckenbergiana, 26,116 - 199." type="book" year="1943">Richter & Richter, 1943</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in the Moroccan trilobite mining industry, and by
|
||
<bibRefCitation id="EFC54EC0FFF2E30DFD90F9EFFC0D86DC" author="Ebach, M. C. & McNamara, K. J." box="[639,952,403,429]" pageId="45" pageNumber="46" pagination="235 - 267" refId="ref86447" refString="Ebach, M. C. & McNamara, K. J. (2002) A systematic revision of the family Harpetida (Trilobita). Records of the Western Australian Museum, 21, 235 - 267. https: // doi. org / 10.18195 / issn. 0312 - 3162.21 (3). 2002.235 - 267" type="journal article" year="2002">Ebach & McNamara (2002)</bibRefCitation>
|
||
and most likely
|
||
<bibRefCitation id="EFC54EC0FFF2E30DFB9DF9EFFA2D86DC" author="Beech, J. D. & Lamsdell, J. C." box="[1138,1432,403,429]" pageId="45" pageNumber="46" pagination="1 - 21" refId="ref85520" refString="Beech, J. D. & Lamsdell, J. C. (2021) Phylogeny, disparity and mass extinctionresponse in the trilobite order Harpetida. The Palaeontological Associations. Papers in Palaeontology, 2021, 1 - 21. https: // doi. org / 10.1002 / spp 2.1399" type="journal article" year="2021">Beech & Lamsdell (2021)</bibRefCitation>
|
||
. The
|
||
<typeStatus id="54EF8D93FFF2E30DFF25F9CBFF4F86A0" box="[202,250,439,465]" pageId="45" pageNumber="46">type</typeStatus>
|
||
locality of
|
||
<emphasis id="B920EF23FFF2E30DFE93F9CBFD2D86A0" box="[380,664,439,465]" italics="true" pageId="45" pageNumber="46">
|
||
<taxonomicName id="4C5448B2FFF2E30DFE93F9CBFD2186A0" baseAuthorityName="Richter & Richter" baseAuthorityYear="1943" box="[380,660,439,465]" class="Trilobita" family="Harpetidae" genus="Helioharpes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="perradiatus">Helioharpes perradiatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
Dechra-ait-Abdallah, is some
|
||
<quantity id="4CAC9ED4FFF2E30DFC1EF9CAFBF386A0" box="[1009,1094,438,465]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="45" pageNumber="46" unit="km" value="250.0">250 km</quantity>
|
||
North-North-West of bou Dîb on the southern edge of the Moroccan Central Meseta.
|
||
<taxonomicName id="4C5448B2FFF2E30DFCE4F9A7FC5C8684" authorityName="Richter & Richter" authorityYear="1943" box="[779,1001,475,501]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="perradiatus">
|
||
<emphasis id="B920EF23FFF2E30DFCE4F9A7FC5C8684" box="[779,1001,475,501]" italics="true" pageId="45" pageNumber="46">Harpes perradiatus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C5448B2FFF2E30DFB6FF9A7FA9F8684" box="[1152,1322,475,501]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF2E30DFB6FF9A7FA9F8684" box="[1152,1322,475,501]" italics="true" pageId="45" pageNumber="46">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
in having a less vaulted cephalon and less steep genal roll; a less inflated glabella which is subcylindrical rather than oval or cone-shaped and is without tubercles. Eye lobe is small and is not cone-shaped and the brim does not narrow posterolaterally. The brim width of
|
||
<taxonomicName id="4C5448B2FFF2E30DFDC3FA3BFD638510" box="[556,726,583,609]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="boudibensis">
|
||
<emphasis id="B920EF23FFF2E30DFDC3FA3BFD638510" box="[556,726,583,609]" italics="true" pageId="45" pageNumber="46">H. boudibensis</emphasis>
|
||
</taxonomicName>
|
||
reduces (tr.) posterolaterally to become one half of its sagittal width at posterior border. Also, there is a strong web-like caeca on the brim, forming pseudo perforations and caeca on genal roll, not present on
|
||
<emphasis id="B920EF23FFF2E30DFE37FAF3FD3D85D8" box="[472,648,655,681]" italics="true" pageId="45" pageNumber="46">
|
||
<taxonomicName id="4C5448B2FFF2E30DFE37FAF3FD3185D8" box="[472,644,655,681]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="boudibensis">H. boudibensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |