252 lines
40 KiB
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252 lines
40 KiB
XML
<document id="5CFBF4663E1DF7B2CD785C30493858FF" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FF98E368FF78FC42FB8D8159" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Maghroharpes azmamarensis Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="73" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="72" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:titleInfo id="2DA0FE9C880DF53795476A408527277C">
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<mods:title id="5A034D67213237F0AE5D5117321C692A">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<mods:namePart id="D89F4A49D35361DE2ACD33A3FF946C49">Johnson, Robert G.</mods:namePart>
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<mods:typeOfResource id="5D4015400BED3E8B13618AA7A0E03EDD">text</mods:typeOfResource>
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<mods:title id="C6FFB7FDD1C6DC463D048467D3C8B1A0">Zootaxa</mods:title>
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<mods:part id="D5A4BB6357D2A1136314ABFC89FFCE3F">
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<mods:date id="9A95F0F9D9DE039AF8996E0A22322717">2024</mods:date>
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<mods:number id="B8815FB19BC8A8FBBDA1337B0AD384B0">2024-05-15</mods:number>
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<mods:number id="14E02136583FF5D72889049E7F1A807C">5450</mods:number>
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<mods:url id="6415DC7A01A04F7C0F1AE8E3E04B84B4">http://dx.doi.org/10.11646/zootaxa.5450.1.1</mods:url>
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<mods:classification id="B762CC8B699843983F50279F61A4E767">journal article</mods:classification>
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<mods:identifier id="E36D32143B1A11FBAA7A5BCB6F523F1C" type="DOI">10.11646/zootaxa.5450.1.1</mods:identifier>
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<mods:identifier id="3E1598A66F1B3D1A7A3975D2EA9CA7E7" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03FD8227FF98E368FF78FC42FB8D8159" ID-DOI="http://doi.org/10.5281/zenodo.11232588" ID-Zenodo-Dep="11232588" LSID="urn:lsid:plazi:treatment:03FD8227FF98E368FF78FC42FB8D8159" httpUri="http://treatment.plazi.org/id/03FD8227FF98E368FF78FC42FB8D8159" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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<subSubSection id="C34E60BAFF98E367FF78FC42FDFD8328" box="[151,584,1086,1113]" pageId="71" pageNumber="72" type="nomenclature">
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<paragraph id="8BEB3331FF98E367FF78FC42FDFD8328" blockId="71.[151,584,1086,1149]" box="[151,584,1086,1113]" pageId="71" pageNumber="72">
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<heading id="D0A3845DFF98E367FF78FC42FDFD8328" bold="true" box="[151,584,1086,1113]" fontSize="11" level="2" pageId="71" pageNumber="72" reason="4">
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||
<emphasis id="B920EF23FF98E367FF78FC42FDFD8328" bold="true" box="[151,584,1086,1113]" pageId="71" pageNumber="72">
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||
<taxonomicName id="4C5448B2FF98E367FF78FC42FE4A8328" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,511,1086,1113]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="azmamarensis" status="sp. nov.">
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<emphasis id="B920EF23FF98E367FF78FC42FE4A8328" bold="true" box="[151,511,1086,1113]" italics="true" pageId="71" pageNumber="72">Maghroharpes azmamarensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2135258FF98E367FDE9FC43FDFD8328" box="[518,584,1087,1113]" pageId="71" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E60BAFF98E367FF78FC1FFE12830C" box="[151,423,1123,1149]" pageId="71" pageNumber="72" type="description">
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||
<paragraph id="8BEB3331FF98E367FF78FC1FFE12830C" blockId="71.[151,584,1086,1149]" box="[151,423,1123,1149]" pageId="71" pageNumber="72">
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<figureCitation id="136F2FB4FF98E367FF78FC1FFEFE830C" box="[151,331,1123,1149]" captionStart="PLATE 29" captionStartId="135.[151,234,1652,1676]" captionTargetBox="[247,1340,181,1628]" captionTargetId="figure-15@135.[247,1340,181,1628]" captionTargetPageId="135" captionText="PLATE 29. A–D Maghroharpes forteyi n. sp., from top of Pragian, Ihandar Formation, El Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29288 (Fig. 2B, Map 8, site 16). A, dorsal B, anterior and C, lateral view. D, dorsal view of brim perforations. E–J,O Maghroharpes azmamarensis n. sp., from top of Pragian, Ihandar Formation, Oued el Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29281 (Fig. 2B, Map 8, site 18). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe, respectively. J, dorsal view of brim perforations. O, from type locality partial dorsal exoskeleton of lower Lamella, paratype NHMUK It 29283 (Fig. 2B, Map 8, site 19)). K–N Maghroharpes forteyi, from top of Pragian, Ihandar Formation, bou Lachrhal, dorsal exoskeleton of partial cephalon only, paratype, NHMUK It 29289 (Fig. 2B, Map 9, Site 7). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232628" httpUri="https://zenodo.org/record/11232628/files/figure.png" pageId="71" pageNumber="72">Plates 29E–J, O</figureCitation>
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,
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<figureCitation id="136F2FB4FF98E367FEB8FC1FFE12830C" box="[343,423,1123,1149]" captionStart="PLATE 30" captionStartId="136.[151,234,1698,1722]" captionTargetBox="[236,1351,181,1672]" captionTargetId="figure-20@136.[236,1351,181,1672]" captionTargetPageId="136" captionText="PLATE 30. A–E Maghroharpes azmamarensis n. sp., from type locality and horizon, Ihandar Formation, Pragian, dorsal exoskeleton of cephalon, paratype, NHMUK It 29282 (Fig., 2B, Map 8, site 19). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. 1 azmamarensis, from top of Pragian, Ihandar Formation, ElAtchane Jbel Oufatene, dorsal exoskeleton of partial cephalon, NHMUK It 29285 (Fig. 2B, Map 8, site 17). F, dorsal, G, lateral and H, anterior view. I, lateral and J, dorsal view of eye lobe, respectively. K, dorsal view of brim perforations. L–O Maghroharpes cf. 2 azmamarensis, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib, dorsal exoskeleton of partial cephalon, NHMUK It 29286 (Fig. 2B, Map 9, site 14). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232632" httpUri="https://zenodo.org/record/11232632/files/figure.png" pageId="71" pageNumber="72">30A–E</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34E60BAFF98E367FF78FCD6FA398240" pageId="71" pageNumber="72" type="diagnosis">
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<paragraph id="8BEB3331FF98E367FF78FCD6FA398240" blockId="71.[151,1437,1194,2011]" pageId="71" pageNumber="72">
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<emphasis id="B920EF23FF98E367FF78FCD6FEA783B5" bold="true" box="[151,274,1194,1220]" pageId="71" pageNumber="72">Diagnosis.</emphasis>
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Cephalon nearly ovoid; glabella inset, broad, strongly vaulted and with convex axial furrows, converging anteriorly. Genal roll gently sloping (sag. and exsag.) and without caeca. Axial and alar furrows deep; preaxial furrows forming circular depressions, crossing inner margin of fringe. Eye lobes inflated and with three eye lenses each. Brim width ratio 0.80 to 0.85, and diameter of standardised brim perforations moderate to large (>145 μm).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFF98E367FF28FD47FD428224" box="[199,759,1339,1365]" pageId="71" pageNumber="72" type="etymology">
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<paragraph id="8BEB3331FF98E367FF28FD47FD428224" blockId="71.[151,1437,1194,2011]" box="[199,759,1339,1365]" pageId="71" pageNumber="72">
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<emphasis id="B920EF23FF98E367FF28FD47FEFB8224" bold="true" box="[199,334,1339,1365]" pageId="71" pageNumber="72">Etymology.</emphasis>
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Named from
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<typeStatus id="54EF8D93FF98E367FE05FD47FDAF8224" box="[490,538,1339,1365]" pageId="71" pageNumber="72">type</typeStatus>
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locality: Azmamar.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFF98E367FF28FD22FEB181CD" pageId="71" pageNumber="72" type="materials_examined">
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<paragraph id="8BEB3331FF98E367FF28FD22FE9A8178" blockId="71.[151,1437,1194,2011]" pageId="71" pageNumber="72">
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<emphasis id="B920EF23FF98E367FF28FD22FE408208" bold="true" box="[199,501,1374,1401]" pageId="71" pageNumber="72">Material and occurrence.</emphasis>
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<materialsCitation id="3B3C396CFF98E367FE13FD23FD5D82B0" collectionCode="NHMUK" collectorName="It & Pragian & Ihandar Formation & Azmamar & Oued el Atchane & Jbel Oufatene" pageId="71" pageNumber="72" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8D93FF98E367FE13FD23FDDC8208" box="[508,617,1375,1401]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED45ABF4FF98E367FD9CFD23FD688208" box="[627,733,1375,1401]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="71" pageNumber="72" type="Museum">NHMUK</collectionCode>
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||
<collectorName id="26A156E7FF98E367FD0CFD23FD418208" box="[739,756,1375,1401]" pageId="71" pageNumber="72">It</collectorName>
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29281,
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<figureCitation id="136F2FB4FF98E367FCA0FD23FC088208" box="[847,957,1375,1401]" captionStart="PLATE 29" captionStartId="135.[151,234,1652,1676]" captionTargetBox="[247,1340,181,1628]" captionTargetId="figure-15@135.[247,1340,181,1628]" captionTargetPageId="135" captionText="PLATE 29. A–D Maghroharpes forteyi n. sp., from top of Pragian, Ihandar Formation, El Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29288 (Fig. 2B, Map 8, site 16). A, dorsal B, anterior and C, lateral view. D, dorsal view of brim perforations. E–J,O Maghroharpes azmamarensis n. sp., from top of Pragian, Ihandar Formation, Oued el Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29281 (Fig. 2B, Map 8, site 18). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe, respectively. J, dorsal view of brim perforations. O, from type locality partial dorsal exoskeleton of lower Lamella, paratype NHMUK It 29283 (Fig. 2B, Map 8, site 19)). K–N Maghroharpes forteyi, from top of Pragian, Ihandar Formation, bou Lachrhal, dorsal exoskeleton of partial cephalon only, paratype, NHMUK It 29289 (Fig. 2B, Map 9, Site 7). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232628" httpUri="https://zenodo.org/record/11232628/files/figure.png" pageId="71" pageNumber="72">Pl. 29E–J</figureCitation>
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, from coral horizon at or near the “Couche Rouge” horizon at the top of the
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<collectorName id="26A156E7FF98E367FDE4FDFFFDD682EC" box="[523,611,1411,1437]" pageId="71" pageNumber="72">Pragian</collectorName>
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,
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<collectorName id="26A156E7FF98E367FD80FDFFFCF182EC" box="[623,836,1411,1437]" pageId="71" pageNumber="72">Ihandar Formation</collectorName>
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,
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<collectorName id="26A156E7FF98E367FCA1FDFFFC0B82EC" box="[846,958,1411,1437]" pageId="71" pageNumber="72">Azmamar</collectorName>
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,
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<collectorName id="26A156E7FF98E367FC25FDFFFB3D82EC" box="[970,1160,1411,1437]" pageId="71" pageNumber="72">Oued el Atchane</collectorName>
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,
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<collectorName id="26A156E7FF98E367FB7BFDFFFA8582EC" box="[1172,1328,1411,1437]" pageId="71" pageNumber="72">Jbel Oufatene</collectorName>
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(
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<figureCitation id="136F2FB4FF98E367FAAFFDFFFA2D82EC" box="[1344,1432,1411,1437]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="71" pageNumber="72">Fig. 2B</figureCitation>
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, Map 8, site 18), dorsal exoskeleton of cephalon only
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</materialsCitation>
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.
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<materialsCitation id="3B3C396CFF98E367FD1DFDDBFE998178" collectionCode="NHMUK" pageId="71" pageNumber="72" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54EF8D93FF98E367FD1DFDDBFCD682B0" box="[754,867,1447,1473]" pageId="71" pageNumber="72" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED45ABF4FF98E367FC81FDDBFC6D82B0" box="[878,984,1447,1473]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="71" pageNumber="72" type="Museum">NHMUK</collectionCode>
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It 29282 (1 and 2),
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<figureCitation id="136F2FB4FF98E367FB56FDDBFA9B82B0" box="[1209,1326,1447,1473]" captionStart="PLATE 30" captionStartId="136.[151,234,1698,1722]" captionTargetBox="[236,1351,181,1672]" captionTargetId="figure-20@136.[236,1351,181,1672]" captionTargetPageId="136" captionText="PLATE 30. A–E Maghroharpes azmamarensis n. sp., from type locality and horizon, Ihandar Formation, Pragian, dorsal exoskeleton of cephalon, paratype, NHMUK It 29282 (Fig., 2B, Map 8, site 19). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. 1 azmamarensis, from top of Pragian, Ihandar Formation, ElAtchane Jbel Oufatene, dorsal exoskeleton of partial cephalon, NHMUK It 29285 (Fig. 2B, Map 8, site 17). F, dorsal, G, lateral and H, anterior view. I, lateral and J, dorsal view of eye lobe, respectively. K, dorsal view of brim perforations. L–O Maghroharpes cf. 2 azmamarensis, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib, dorsal exoskeleton of partial cephalon, NHMUK It 29286 (Fig. 2B, Map 9, site 14). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232632" httpUri="https://zenodo.org/record/11232632/files/figure.png" pageId="71" pageNumber="72">Pl. 30A–E</figureCitation>
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and
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<figureCitation id="136F2FB4FF98E367FA8AFDDBFA2282B0" box="[1381,1431,1447,1473]" captionStart="PLATE 39" captionStartId="145.[151,234,1737,1761]" captionTargetBox="[213,1375,181,1713]" captionTargetId="figure-15@145.[213,1375,181,1713]" captionTargetPageId="145" captionText="PLATE 39. A–M Pinnuloharpes apteros n. sp., El Otfal Formation, upper Eifelian. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29356 from Ksar Hannabou (Fig. 2B, Map 6, site 8). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–K dorsal exoskeleton with thorax and pygidium disarticulated, paratype, NHMUK It 29359, from type locality and horizon (Fig. 2B, Map 6, site 19). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, lateral view of eye lobe, J, anterior lateral view of rim and marginal band, and K, dorsal view of pleurae close to pygidium. L–M dorsal exoskeleton of a partial cephalon, paratype, NHMUK It 29357, from Ksar Hannabou (Fig. 2B, Map 6, site 15). L, dorsal view and M, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232664" httpUri="https://zenodo.org/record/11232664/files/figure.png" pageId="71" pageNumber="72">39O</figureCitation>
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, from type locality and horizon (
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||
<figureCitation id="136F2FB4FF98E367FE16FDB7FDE48294" box="[505,593,1483,1509]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="71" pageNumber="72">Fig. 2B</figureCitation>
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, Map 8, site 19), A–E dorsal exoskeleton of cephalon and, O, lower lamella on same slab
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BEB3331FF98E367FF28FE6EFEB181CD" blockId="71.[151,1437,1194,2011]" pageId="71" pageNumber="72">
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Other material:
|
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<taxonomicName id="4C5448B2FF98E367FE6EFE6EFD4C815D" authorityName="Johnson" authorityYear="2024" box="[385,761,1554,1580]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="azmamarensis">
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<emphasis id="B920EF23FF98E367FE6EFE6EFD93815D" box="[385,550,1554,1580]" italics="true" pageId="71" pageNumber="72">Maghroharpes</emphasis>
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cf.
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<emphasis id="B920EF23FF98E367FDB5FE6FFD4C815D" box="[602,761,1555,1580]" italics="true" pageId="71" pageNumber="72">azmamarensis</emphasis>
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||
</taxonomicName>
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||
:
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<collectionCode id="ED45ABF4FF98E367FCE3FE6FFCC3815C" box="[780,886,1555,1581]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="71" pageNumber="72" type="Museum">NHMUK</collectionCode>
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It 29285,
|
||
<figureCitation id="136F2FB4FF98E367FC19FE6EFBC5815C" box="[1014,1136,1554,1581]" captionStart="PLATE 30" captionStartId="136.[151,234,1698,1722]" captionTargetBox="[236,1351,181,1672]" captionTargetId="figure-20@136.[236,1351,181,1672]" captionTargetPageId="136" captionText="PLATE 30. A–E Maghroharpes azmamarensis n. sp., from type locality and horizon, Ihandar Formation, Pragian, dorsal exoskeleton of cephalon, paratype, NHMUK It 29282 (Fig., 2B, Map 8, site 19). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. 1 azmamarensis, from top of Pragian, Ihandar Formation, ElAtchane Jbel Oufatene, dorsal exoskeleton of partial cephalon, NHMUK It 29285 (Fig. 2B, Map 8, site 17). F, dorsal, G, lateral and H, anterior view. I, lateral and J, dorsal view of eye lobe, respectively. K, dorsal view of brim perforations. L–O Maghroharpes cf. 2 azmamarensis, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib, dorsal exoskeleton of partial cephalon, NHMUK It 29286 (Fig. 2B, Map 9, site 14). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232632" httpUri="https://zenodo.org/record/11232632/files/figure.png" pageId="71" pageNumber="72">Pl. 30F–K</figureCitation>
|
||
, from coral horizon at or near the “Couche Rouge” horizon at the top of the Pragian, Ihandar Formation, Azmamar, Oued el Atchane, Jbel Oufatene (
|
||
<figureCitation id="136F2FB4FF98E367FEE1FE26FED38105" box="[270,358,1626,1652]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="71" pageNumber="72">Fig. 2B</figureCitation>
|
||
, Map 8, site 17), dorsal exoskeleton of cephalon.
|
||
<collectionCode id="ED45ABF4FF98E367FC76FE27FBB68104" box="[921,1027,1627,1653]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="71" pageNumber="72" type="Museum">NHMUK</collectionCode>
|
||
It 29286,
|
||
<figureCitation id="136F2FB4FF98E367FB97FE26FB458105" box="[1144,1264,1626,1652]" captionStart="PLATE 30" captionStartId="136.[151,234,1698,1722]" captionTargetBox="[236,1351,181,1672]" captionTargetId="figure-20@136.[236,1351,181,1672]" captionTargetPageId="136" captionText="PLATE 30. A–E Maghroharpes azmamarensis n. sp., from type locality and horizon, Ihandar Formation, Pragian, dorsal exoskeleton of cephalon, paratype, NHMUK It 29282 (Fig., 2B, Map 8, site 19). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. 1 azmamarensis, from top of Pragian, Ihandar Formation, ElAtchane Jbel Oufatene, dorsal exoskeleton of partial cephalon, NHMUK It 29285 (Fig. 2B, Map 8, site 17). F, dorsal, G, lateral and H, anterior view. I, lateral and J, dorsal view of eye lobe, respectively. K, dorsal view of brim perforations. L–O Maghroharpes cf. 2 azmamarensis, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib, dorsal exoskeleton of partial cephalon, NHMUK It 29286 (Fig. 2B, Map 9, site 14). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232632" httpUri="https://zenodo.org/record/11232632/files/figure.png" pageId="71" pageNumber="72">Pl. 30L–O</figureCitation>
|
||
, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib (
|
||
<figureCitation id="136F2FB4FF98E367FCAFFE02FC2D81E9" box="[832,920,1662,1688]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="71" pageNumber="72">Fig. 2B</figureCitation>
|
||
, Map 9, site 14), dorsal exoskeleton of partial cephalon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFF98E368FF28FEBAFCF38447" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="description">
|
||
<paragraph id="8BEB3331FF98E367FF28FEBAFAD580C6" blockId="71.[151,1437,1194,2011]" pageId="71" pageNumber="72">
|
||
<emphasis id="B920EF23FF98E367FF28FEBAFEEC8191" bold="true" box="[199,345,1734,1760]" pageId="71" pageNumber="72">Description.</emphasis>
|
||
Cephalon ovoid, widest at midpoint between eyes and alae. Genicranium broad, widest at posterior border. Glabella inset, strongly vaulted, very broad with convex flanks, rounded crest and tubercles. Axial furrows convex and converging strongly anteriorly. L1 inflated, S1 sloping posteriorly and effaced before curving anteriorly near crest. No furrows at S2–S4. Preglabellar furrow straight in anterior view, and preglabellar field narrow and well defined. Occipital furrow narrow and medially straight. In lateral view, occipital ring sloping posteriorly and same height as glabella anterior to S0; not medially broadened and has sculptural tubercles. Occipital node large and well inflated. Alae small (tr.) and laterally directed. Alar depression present and posterior border moderately short.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF98E368FF28FFBDFD9C86BE" blockId="71.[151,1437,1194,2011]" lastBlockId="72.[151,1437,151,1576]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
|
||
Genal area moderately broad, with no caeca or tubercles. Large eyes (exsag.), set close to inner margin of fringe. Eye location ratio around 0.90, and tubercles and pitting on eye lobes. Each eye lobe with 3 eye lenses, anterior and posterior largest and small central lens located high up between other two lenses. Eye ridge not well defined and internal margin of fringe convex across anterior of genal area. Preaxial furrows broad, not converging and forming depressions where they cross at inner margin of fringe. Glabella, preglabellar field, genal area and dorsal surface of eye lobes have fine, dense pitting. Anterior boss barely inflated, reaching down to weak girder kink. Genal roll sloping very gently anteriorly (≈
|
||
<superScript id="7C219E79FF97E368FD34F936FD4A8629" attach="none" box="[731,767,330,357]" fontSize="6" pageId="72" pageNumber="73">350</superScript>
|
||
) and moderately steeply laterally (≈
|
||
<superScript id="7C219E79FF97E368FB5FF936FB618629" attach="none" box="[1200,1236,330,357]" fontSize="6" pageId="72" pageNumber="73">550</superScript>
|
||
). In lateral view, fringe narrows strongly posteriorly. Perforations on genal roll smaller than on brim, with row of larger perforations immediately above girder and across anterior boss at inner margin of fringe. No caeca. Girder moderately wide (≈ 2.6% of width (tr.) of genicranium).
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF97E368FF28F9A5FB608463" blockId="72.[151,1437,151,1576]" pageId="72" pageNumber="73">Brim concave, not steeply sloping (≈15%) and with width (sag.) 30% of cephalic length (sag.). Brim width ratio 0.80. Standardised brim perforations 147µm, with row of larger perforations at proximal and distal edges of brim. Density of perforations on brim around 15 per square mm on standardised brim. Light caeca across brim. External rim stout, with rim suture running around centre of dorsal surface. Marginal band near vertical, concave, with tubercles and ridge around top and bottom. Length (exsag.) of prolongations around 75% of cephalic length (sag.). Internal and external rims curve adaxially and internal rims are concave in lateral view. Posterior border and internal rims thicker than exterior rim in dorsal view. Dorsal surface of internal rim sparsely covered by small tubercles. Girder extends onto prolongation, to meet internal rim halfway down prolongation. Proximal edge of brim steepening on prolongations, and prolongations terminate in spine carried on lower lamella.</paragraph>
|
||
<paragraph id="8BEB3331FF97E368FF28FB61FCF38447" blockId="72.[151,1437,151,1576]" box="[199,838,796,823]" pageId="72" pageNumber="73">No hypostomes, thoracic or pygidial sclerites are known.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFF97E368FF28FB3CFB8D8159" pageId="72" pageNumber="73" type="discussion">
|
||
<paragraph id="8BEB3331FF97E368FF28FB3CFBF1840E" blockId="72.[151,1437,151,1576]" pageId="72" pageNumber="73">
|
||
<emphasis id="B920EF23FF97E368FF28FB3CFE8E842B" bold="true" box="[199,315,832,858]" pageId="72" pageNumber="73">Remarks.</emphasis>
|
||
<taxonomicName id="4C5448B2FF97E368FEACFB3CFD3A842B" authorityName="Johnson" authorityYear="2024" box="[323,655,832,858]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="azmamarensis">
|
||
<emphasis id="B920EF23FF97E368FEACFB3CFD3A842B" box="[323,655,832,858]" italics="true" pageId="72" pageNumber="73">Maghroharpes azmamarensis</emphasis>
|
||
</taxonomicName>
|
||
lacks one of the diagnostic characters of its genus as shown in
|
||
<tableCitation id="C6D6068AFF97E368FAB1FB3CFF00840E" captionStart="TABLE 5" captionStartId="182.[151,239,503,529]" captionText="TABLE 5b. Proposed diagnostic characters for Devonian harpetid genera (continued)" pageId="72" pageNumber="73">Table 5b</tableCitation>
|
||
, having an alar depression, which is lacking in most other members of the genus.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF97E368FF28FBF4FCE584B7" blockId="72.[151,1437,151,1576]" pageId="72" pageNumber="73">
|
||
<materialsCitation id="3B3C396CFF97E368FF28FBF4FCF984B7" collectionCode="NHMUK" pageId="72" pageNumber="73" specimenCount="1" typeStatus="holotype">
|
||
<collectionCode id="ED45ABF4FF97E368FF28FBF4FE8484D3" box="[199,305,904,930]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="72" pageNumber="73" type="Museum">NHMUK</collectionCode>
|
||
It 29282 is very similar to the
|
||
<typeStatus id="54EF8D93FF97E368FD78FBF4FD4C84D3" box="[663,761,904,930]" pageId="72" pageNumber="73" type="holotype">holotype</typeStatus>
|
||
but lacks bilateral symmetry, with one prolongation longer than the other, which may be the result of a teratological fault
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF97E368FF28FBB3FDA18220" blockId="72.[151,1437,151,1576]" pageId="72" pageNumber="73">
|
||
<taxonomicName id="4C5448B2FF97E368FF28FBB3FDA38498" authorityName="Johnson" authorityYear="2024" box="[199,534,975,1001]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="azmamarensis">
|
||
<emphasis id="B920EF23FF97E368FF28FBB3FDA38498" box="[199,534,975,1001]" italics="true" pageId="72" pageNumber="73">Maghroharpes azmamarensis</emphasis>
|
||
</taxonomicName>
|
||
closely resembles
|
||
<taxonomicName id="4C5448B2FF97E368FD1BFBACFC1C849B" authorityName="Johnson" authorityYear="2024" box="[756,937,976,1002]" class="Trilobita" family="Harpetidae" genus="Maghrohapes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="oufatenensis">
|
||
<emphasis id="B920EF23FF97E368FD1BFBACFC1C849B" box="[756,937,976,1002]" italics="true" pageId="72" pageNumber="73">M. oufatenensis</emphasis>
|
||
</taxonomicName>
|
||
but differs in having: an ovoid rather than oval cephalon, which is widest at midpoint between the eye lobes and the alae, not at the alae as in the case of
|
||
<emphasis id="B920EF23FF97E368FA91FB88FE988340" italics="true" pageId="72" pageNumber="73">
|
||
<taxonomicName id="4C5448B2FF97E368FA91FB88FE908340" authorityName="Johnson" authorityYear="2024" class="Trilobita" family="Harpetidae" genus="Maghrohapes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="oufatenensis">M. oufatenensis</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
an inset, carinate glabella with tubercles on its dorsal surface; eye lobes with pitting and tubercles on their dorsal surfaces and 3 lenses each the anterior and posterior lenses of equal size and the third lens between the other two, much smaller, while
|
||
<taxonomicName id="4C5448B2FF97E368FE16FC1CFD1F8308" authorityName="Johnson" authorityYear="2024" box="[505,682,1119,1145]" class="Trilobita" family="Harpetidae" genus="Maghrohapes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="oufatenensis">
|
||
<emphasis id="B920EF23FF97E368FE16FC1CFD1F8308" box="[505,682,1119,1145]" italics="true" pageId="72" pageNumber="73">M. oufatenensis</emphasis>
|
||
</taxonomicName>
|
||
’s eye lobes lack tubercles and pitting and have only two lenses, the anterior lens being the largest; a low poorly defined eye ridge, which is absent on
|
||
<emphasis id="B920EF23FF97E368FBC3FCF8FB5383EC" box="[1068,1254,1155,1181]" italics="true" pageId="72" pageNumber="73">
|
||
<taxonomicName id="4C5448B2FF97E368FBC3FCF8FB6A83EC" authorityName="Johnson" authorityYear="2024" box="[1068,1247,1155,1181]" class="Trilobita" family="Harpetidae" genus="Maghrohapes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="oufatenensis">M. oufatenensis</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
a genal roll that slopes gently anteriorly at around
|
||
<superScript id="7C219E79FF97E368FDF8FCDBFD8E83C4" attach="left" box="[535,571,1191,1217]" fontSize="6" pageId="72" pageNumber="73">360</superScript>
|
||
and laterally at around
|
||
<superScript id="7C219E79FF97E368FCA9FCDBFCDF83C4" attach="none" box="[838,874,1191,1218]" fontSize="6" pageId="72" pageNumber="73">540</superScript>
|
||
, compared to
|
||
<superScript id="7C219E79FF97E368FBE4FCDBFB9A83C4" attach="left" box="[1035,1071,1191,1218]" fontSize="6" pageId="72" pageNumber="73">550</superScript>
|
||
and
|
||
<superScript id="7C219E79FF97E368FB89FCDBFB3F83C4" attach="left" box="[1126,1162,1191,1217]" fontSize="6" pageId="72" pageNumber="73">720</superScript>
|
||
respectively, in the case of that of
|
||
<taxonomicName id="4C5448B2FF97E368FEE7FCB7FE0F8394" authorityName="Johnson" authorityYear="2024" box="[264,442,1227,1253]" class="Trilobita" family="Harpetidae" genus="Maghrohapes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="oufatenensis">
|
||
<emphasis id="B920EF23FF97E368FEE7FCB7FE0F8394" box="[264,442,1227,1253]" italics="true" pageId="72" pageNumber="73">M. oufatenensis</emphasis>
|
||
</taxonomicName>
|
||
; the brim sloping at around
|
||
<superScript id="7C219E79FF97E368FD12FCB6FC9483A9" attach="left" box="[765,801,1226,1253]" fontSize="6" pageId="72" pageNumber="73">150</superScript>
|
||
with caeca that stretch right across it, while the brim of
|
||
<taxonomicName id="4C5448B2FF97E368FF78FC93FEFD8278" authorityName="Johnson" authorityYear="2024" box="[151,328,1263,1289]" class="Trilobita" family="Harpetidae" genus="Maghrohapes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="oufatenensis">
|
||
<emphasis id="B920EF23FF97E368FF78FC93FEFD8278" box="[151,328,1263,1289]" italics="true" pageId="72" pageNumber="73">M. oufatenensis</emphasis>
|
||
</taxonomicName>
|
||
slopes at
|
||
<superScript id="7C219E79FF97E368FE59FC93FE79838D" attach="left" box="[438,460,1262,1289]" fontSize="6" pageId="72" pageNumber="73">70</superScript>
|
||
and has caeca does not reach far across it; internal rims that curve adaxially and are not subparallel; and genal spines that follow the curve of external rim rather than being set at an oblique angle, as they are in the case of
|
||
<taxonomicName id="4C5448B2FF97E368FEB3FD4BFDB88221" authorityName="Johnson" authorityYear="2024" box="[348,525,1334,1360]" class="Trilobita" family="Harpetidae" genus="Maghrohapes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="oufatenensis">
|
||
<emphasis id="B920EF23FF97E368FEB3FD4BFDB88221" box="[348,525,1334,1360]" italics="true" pageId="72" pageNumber="73">M. oufatenensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF97E368FF28FD26FD4F82CD" blockId="72.[151,1437,151,1576]" pageId="72" pageNumber="73">
|
||
<taxonomicName id="4C5448B2FF97E368FF28FD26FED98205" authorityName="Johnson" authorityYear="2024" box="[199,364,1370,1396]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EF23FF97E368FF28FD26FED98205" box="[199,364,1370,1396]" italics="true" pageId="72" pageNumber="73">Maghroharpes</emphasis>
|
||
</taxonomicName>
|
||
cf.1
|
||
<taxonomicName id="4C5448B2FF97E368FE44FD27FDFF8205" authorityName="Johnson" authorityYear="2024" box="[427,586,1371,1396]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="azmamarensis">
|
||
<emphasis id="B920EF23FF97E368FE44FD27FDFF8205" box="[427,586,1371,1396]" italics="true" pageId="72" pageNumber="73">azmamarensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136F2FB4FF97E368FDB2FD26FD6B8205" box="[605,734,1370,1396]" captionStart="PLATE 40" captionStartId="146.[151,234,1704,1728]" captionTargetBox="[225,1363,181,1681]" captionTargetId="figure-20@146.[225,1363,181,1681]" captionTargetPageId="146" captionText="PLATE 40.A–N Pinnuloharpes hannabouensis n. sp., from an horizon 5 m down slope of “Radiaspis Couche”, upper Eifelian/ Givetian boundary, Ksar Hannabou. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29334 (Fig. 2B, Map 6, site 12). A, dorsal and B, lateral view. C, lateral view of eye lobe (uncoated) and D, dorsal view of brim perforations. E–I dorsal exoskeleton cephalon and thorax only, holotype, NHMUK It 29333 (Fig. 2B, Map 6, site 11). E, dorsal, F, anterior and G, lateral view. H, lateral view of eye lobes, I, dorsal view of brim perforations. J–N dorsal exoskeleton pygidium incomplete, paratype, NHMUK It 29335 (Fig. 2B, Map 6, site 13). J, dorsal, K, anterior and L, lateral view. M, lateral view of eye lobe and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232666" httpUri="https://zenodo.org/record/11232666/files/figure.png" pageId="72" pageNumber="73">Pl. 40 A–E</figureCitation>
|
||
) specimen NHMUK It 29285 differs from
|
||
<taxonomicName id="4C5448B2FF97E368FB3AFD27FA298205" authorityName="Johnson" authorityYear="2024" box="[1237,1436,1371,1396]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="azmamarensis">
|
||
<emphasis id="B920EF23FF97E368FB3AFD27FA298205" box="[1237,1436,1371,1396]" italics="true" pageId="72" pageNumber="73">M. azmamarensis</emphasis>
|
||
</taxonomicName>
|
||
in having a wider (tr.) cephalon and genicranium, and shorter (exsag.) prolongations. The exoskeleton is not well preserved but otherwise seems similar to the
|
||
<typeStatus id="54EF8D93FF97E368FD7FFDDEFD4382CD" box="[656,758,1442,1468]" pageId="72" pageNumber="73" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF97E368FF28FDBAFB8D8159" blockId="72.[151,1437,151,1576]" pageId="72" pageNumber="73">
|
||
<taxonomicName id="4C5448B2FF97E368FF28FDBAFED98291" authorityName="Johnson" authorityYear="2024" box="[199,364,1478,1504]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EF23FF97E368FF28FDBAFED98291" box="[199,364,1478,1504]" italics="true" pageId="72" pageNumber="73">Maghroharpes</emphasis>
|
||
</taxonomicName>
|
||
cf.2 azmamarensis (
|
||
<figureCitation id="136F2FB4FF97E368FDBDFDBAFD7C8291" box="[594,713,1478,1504]" captionStart="PLATE 30" captionStartId="136.[151,234,1698,1722]" captionTargetBox="[236,1351,181,1672]" captionTargetId="figure-20@136.[236,1351,181,1672]" captionTargetPageId="136" captionText="PLATE 30. A–E Maghroharpes azmamarensis n. sp., from type locality and horizon, Ihandar Formation, Pragian, dorsal exoskeleton of cephalon, paratype, NHMUK It 29282 (Fig., 2B, Map 8, site 19). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. 1 azmamarensis, from top of Pragian, Ihandar Formation, ElAtchane Jbel Oufatene, dorsal exoskeleton of partial cephalon, NHMUK It 29285 (Fig. 2B, Map 8, site 17). F, dorsal, G, lateral and H, anterior view. I, lateral and J, dorsal view of eye lobe, respectively. K, dorsal view of brim perforations. L–O Maghroharpes cf. 2 azmamarensis, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib, dorsal exoskeleton of partial cephalon, NHMUK It 29286 (Fig. 2B, Map 9, site 14). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232632" httpUri="https://zenodo.org/record/11232632/files/figure.png" pageId="72" pageNumber="73">Pl. 30L–O</figureCitation>
|
||
) specimen NHMUK It 29286 is similar to
|
||
<taxonomicName id="4C5448B2FF97E368FB48FDBAFADE82AE" authorityName="Johnson" authorityYear="2024" box="[1191,1387,1478,1503]" class="Trilobita" family="Harpetidae" genus="Maghroharpes" kingdom="Animalia" order="Harpetida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="azmamarensis">
|
||
<emphasis id="B920EF23FF97E368FB48FDBAFADE82AE" box="[1191,1387,1478,1503]" italics="true" pageId="72" pageNumber="73">M. azmamarensis</emphasis>
|
||
</taxonomicName>
|
||
, but differs in having a broader and less vaulted glabella that is not inset and has more tectiform flanks. Also, it lacks alar depressions and has a brim that is wider (sag.) and has slightly bigger perforations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |