156 lines
19 KiB
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156 lines
19 KiB
XML
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<mods:title id="AB261BD547FEEB21F8D1CE841824B0B0">The upper dentition and relationships of the enigmatic Australian Cretaceous mammal Kollikodon ritchiei</mods:title>
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<mods:namePart id="5813FEB09DE7F18909152FD7B224F5EF">Pian, Rebecca</mods:namePart>
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<mods:namePart id="AC51BABD4D3D452707995DDA1A455E7B">Archer, Michael</mods:namePart>
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<mods:namePart id="3BAE8CE32AC6F2A5299CDB799088E5F8">Hand, Suzanne J.</mods:namePart>
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<mods:namePart id="E4B91978DB3B6D25F11A7347A635E406">Beck, Robin M. D.</mods:namePart>
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<mods:namePart id="F1F87A9064CC4E9DEF1EA0632579383F">Cody, Andrew</mods:namePart>
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<mods:title id="BC193B46A99E4D321710B46A720CFB2F">Memoirs of Museum Victoria</mods:title>
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<treatment id="03E1AE36FF88FFE4FC869379126B2655" ID-DOI="http://doi.org/10.5281/zenodo.12214089" ID-Zenodo-Dep="12214089" LSID="urn:lsid:plazi:treatment:03E1AE36FF88FFE4FC869379126B2655" httpUri="http://treatment.plazi.org/id/03E1AE36FF88FFE4FC869379126B2655" lastPageNumber="98" pageId="1" pageNumber="98">
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<subSubSection id="C3524CABFF88FFE4FC86937912EA231F" box="[817,1420,525,548]" pageId="1" pageNumber="98" type="nomenclature">
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<paragraph id="8BF71F20FF88FFE4FC86937912EA231F" blockId="1.[817,1420,525,548]" box="[817,1420,525,548]" pageId="1" pageNumber="98">
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<heading id="D0BFA84CFF88FFE4FC86937912EA231F" box="[817,1420,525,548]" fontSize="9" level="2" pageId="1" pageNumber="98" reason="5">
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<taxonomicName id="4C4864A3FF88FFE4FC86937912EA231F" authority="Flannery, Archer, Rich and Jones, 1995" authorityName="Flannery, Archer, Rich and Jones" authorityYear="1995" box="[817,1420,525,548]" class="Mammalia" family="Kollikodontidae" genus="Kollikodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Monotremata" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="ritchiei">
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<emphasis id="B93CC332FF88FFE4FC869379149E2318" bold="true" box="[817,1016,525,547]" italics="true" pageId="1" pageNumber="98">Kollikodon ritchiei</emphasis>
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<bibRefCitation id="EFD962D1FF88FFE4FC49937A12EA231F" author="Flannery, T. F. & Archer, M. & Rich, T. H. & Jones, R." box="[1022,1420,526,548]" pageId="1" pageNumber="98" pagination="418 - 420" refId="ref5668" refString="Flannery, T. F., Archer, M., Rich, T. H. and Jones, R. 1995. A new family of monotremes from the Cretaceous of Australia. Nature 377 6548: 418 - 420. doi: 10.1038 / 377418 a 0." type="journal article" year="1995">Flannery, Archer, Rich and Jones, 1995</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3524CABFF88FFE4FC86934E126B2655" pageId="1" pageNumber="98" type="materials_examined">
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<paragraph id="8BF71F20FF88FFE4FC86934E123A23B2" blockId="1.[817,1447,570,650]" pageId="1" pageNumber="98">
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<materialsCitation id="3B20157DFF88FFE4FC86934E123A23B2" collectionCode="AM" country="Australia" location="Australian Museum Palaeontological Collection" municipality="Sydney" pageId="1" pageNumber="98" specimenCode="F96602" specimenCount="1" typeStatus="holotype">
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<emphasis id="B93CC332FF88FFE4FC86934E14E82374" box="[817,910,570,591]" italics="true" pageId="1" pageNumber="98">
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<typeStatus id="54F3A182FF88FFE4FC86934E14E82374" box="[817,910,570,591]" pageId="1" pageNumber="98" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<collectionCode id="ED5987E5FF88FFE4FC2A934E14A02374" box="[925,966,570,591]" country="Australia" name="Australian Museum" pageId="1" pageNumber="98" type="Museum">AM</collectionCode>
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<specimenCode id="DBEEB75BFF88FFE4FC78934E137B236B" box="[975,1053,570,592]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="1" pageNumber="98" type="Museum">F96602</specimenCode>
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(
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<location id="8E9749FBFF88FFE4FB9B934E14FA2356" LSID="urn:lsid:plazi:treatment:03E1AE36FF88FFE4FC869379126B2655:8E9749FBFF88FFE4FB9B934E14FA2356" country="Australia" municipality="Sydney" name="Australian Museum Palaeontological Collection" pageId="1" pageNumber="98">Australian Museum Palaeontological Collection</location>
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,
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<collectingMunicipality id="6B93855AFF88FFE4FC00932313642356" box="[951,1026,599,621]" pageId="1" pageNumber="98">Sydney</collectingMunicipality>
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,
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<collectingCountry id="F35F5FB0FF88FFE4FBAB932313182356" box="[1052,1150,599,621]" name="Australia" pageId="1" pageNumber="98">Australia</collectingCountry>
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,), right dentary fragment preserving m1-3 and alveoli for two premolars and m4.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BF71F20FF88FFE4FC8693D413A72550" blockId="1.[815,1450,672,1131]" pageId="1" pageNumber="98">
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<emphasis id="B93CC332FF88FFE4FC8693D41489238D" box="[817,1007,672,694]" italics="true" pageId="1" pageNumber="98">Referred specimen</emphasis>
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. Opalised right skull fragment preserving part of the maxilla, which retains the posterior premolar (possibly P4) and M1-4, and possibly also part of the palatine (
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<figureCitation id="137303A5FF88FFE4FA9A93AF12E223CB" box="[1325,1412,731,753]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="2.[140,200,1246,1265]" captionStartId-1="3.[147,207,1857,1876]" captionTargetBox-0="[140,1440,227,1229]" captionTargetBox-1="[229,1367,227,1837]" captionTargetId-0="figure-358@2.[137,1440,224,1232]" captionTargetId-1="figure-14@3.[229,1367,224,1837]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figure 1. Kollikodon ritchiei right maxillary fragment and molar row preserving Px, M1, M2, M3 and RM4. Stereopair occlusal view. ant, anterior; bucc, buccal." captionText-1="Figure 2. Kollikodon ritchiei right maxillary fragment and molar row preserving Px, M1, M2, M3 and M4. (A) Stereopair oblique-occlusal view. (B) Stereopair lingual view. (C) Stereopair buccal view. ant, anterior." figureDoi-0="http://doi.org/10.5281/zenodo.12213819" figureDoi-1="http://doi.org/10.5281/zenodo.12213821" httpUri-0="https://zenodo.org/record/12213819/files/figure.png" httpUri-1="https://zenodo.org/record/12213821/files/figure.png" pageId="1" pageNumber="98">Figs 1–2</figureCitation>
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). A 35μm voxel Xradia microCT data file of the specimen has been lodged with the Museum of
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<collectingRegion id="498CD1C2FF88FFE4FBF0926113FE2210" box="[1095,1176,789,811]" country="Australia" name="Victoria" pageId="1" pageNumber="98">Victoria</collectingRegion>
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in Melbourne. Detailed 3D prints of this specimen can be made from the scan data. Solid casts taken from a mould of the complete upper dentition are also available; one (
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<collectionCode id="ED5987E5FF88FFE4FC709218149622BA" box="[967,1008,876,897]" country="Australia" name="Australian Museum" pageId="1" pageNumber="98" type="Museum">AM</collectionCode>
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F140201) is registered in the collections of the Australian Museum. Although the original specimen, which is a natural glass cast without internal structure of any kind, is less informative than the microCT scan data (given that it reveals structures in undercut areas not visible via conventional microscopy) and no more informative than 3D prints and hard casts, this specimen is available for further examination as part of the National Opal Collection, on application to its Director, Andrew Cody (andrew@codyopal.com).
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</paragraph>
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<paragraph id="8BF71F20FF88FFE4FC8695F513B02413" blockId="1.[817,1448,1153,1320]" pageId="1" pageNumber="98">
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<emphasis id="B93CC332FF88FFE4FC8695F514B825AD" box="[817,990,1153,1174]" italics="true" pageId="1" pageNumber="98">Locality and age</emphasis>
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. Griman Creek Formation; Early Cretaceous (Middle Albian) (
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<bibRefCitation id="EFD962D1FF88FFE4FC5E95EA13AE258F" author="Flannery, T. F. & Archer, M. & Rich, T. H. & Jones, R." box="[1001,1224,1182,1204]" pageId="1" pageNumber="98" pagination="418 - 420" refId="ref5668" refString="Flannery, T. F., Archer, M., Rich, T. H. and Jones, R. 1995. A new family of monotremes from the Cretaceous of Australia. Nature 377 6548: 418 - 420. doi: 10.1038 / 377418 a 0." type="journal article" year="1995">Flannery et al., 1995</bibRefCitation>
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). The
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<typeStatus id="54F3A182FF88FFE4FABB95EA125E2588" box="[1292,1336,1182,1203]" pageId="1" pageNumber="98">type</typeStatus>
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locality is claim 30226, Moonshine area of the Cocoran opal field, Lightning Ridge,
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<collectingRegion id="498CD1C2FF88FFE4FC5495AD13FF25D5" box="[995,1177,1240,1262]" country="Australia" name="New South Wales" pageId="1" pageNumber="98">New South Wales</collectingRegion>
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,
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<collectingCountry id="F35F5FB0FF88FFE4FB1495AC126625D5" box="[1187,1280,1240,1262]" name="Australia" pageId="1" pageNumber="98">Australia</collectingCountry>
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(
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<bibRefCitation id="EFD962D1FF88FFE4FABA95AC14002430" author="Flannery, T. F. & Archer, M. & Rich, T. H. & Jones, R." pageId="1" pageNumber="98" pagination="418 - 420" refId="ref5668" refString="Flannery, T. F., Archer, M., Rich, T. H. and Jones, R. 1995. A new family of monotremes from the Cretaceous of Australia. Nature 377 6548: 418 - 420. doi: 10.1038 / 377418 a 0." type="journal article" year="1995">Flannery et al., 1995</bibRefCitation>
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). The new skull fragment described here comes from an unnamed mine on the Cocoran opal field.
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</paragraph>
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<paragraph id="8BF71F20FF88FFE4FC86944B12EC2706" blockId="1.[813,1448,1342,1597]" pageId="1" pageNumber="98">
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<emphasis id="B93CC332FF88FFE4FC86944B13E5246F" box="[817,1155,1342,1364]" italics="true" pageId="1" pageNumber="98">New diagnosis of clade containing</emphasis>
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<taxonomicName id="4C4864A3FF88FFE4FB32944A1391246F" authorityName="Flannery, Archer, Rich and Jones" authorityYear="1995" box="[1157,1271,1342,1364]" class="Mammalia" family="Kollikodontidae" genus="Kollikodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Monotremata" pageId="1" pageNumber="103" phylum="Chordata" rank="genus">Kollikodon</taxonomicName>
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<emphasis id="B93CC332FF88FFE4FB49944A12C4246F" box="[1278,1442,1342,1364]" italics="true" pageId="1" pageNumber="98">and monotremes</emphasis>
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.
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<taxonomicName id="4C4864A3FF88FFE4FC86942F137E244B" authorityName="Flannery, Archer, Rich and Jones" authorityYear="1995" box="[817,1048,1371,1392]" class="Mammalia" family="Kollikodontidae" genus="Kollikodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Monotremata" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="ritchiei">
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<emphasis id="B93CC332FF88FFE4FC86942F137E244B" box="[817,1048,1371,1392]" italics="true" pageId="1" pageNumber="98">Kollikodon ritchiei</emphasis>
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</taxonomicName>
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and definitive monotremes (ornithorhynchids, tachyglossids,
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<taxonomicName id="4C4864A3FF88FFE4FB34940D139024B5" box="[1155,1270,1401,1422]" class="Mammalia" family="Ornithorhynchidae" genus="Steropodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Monotremata" pageId="1" pageNumber="98" phylum="Chordata" rank="genus">
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<emphasis id="B93CC332FF88FFE4FB34940D139024B5" box="[1155,1270,1401,1422]" italics="true" pageId="1" pageNumber="98">Steropodon</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4864A3FF88FFE4FA91940D12C624B5" authorityName="Rich, Vickers-Rich, Constantine, Flannery, Kool & van Klaveren" authorityYear="1999" box="[1318,1440,1401,1422]" class="Mammalia" family="Ornithorhynchidae" genus="Teinolophos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Monotremata" pageId="1" pageNumber="98" phylum="Chordata" rank="genus">
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<emphasis id="B93CC332FF88FFE4FA91940D12C624B5" box="[1318,1440,1401,1422]" italics="true" pageId="1" pageNumber="98">Teinolophos</emphasis>
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</taxonomicName>
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) differ from other groups variously regarded as australosphenidans (ausktribosphenids,
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<taxonomicName id="4C4864A3FF88FFE4FC4D94C7130524F3" box="[1018,1123,1459,1480]" class="Mammalia" family="Ausktribosphenidae" genus="Ambondro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ausktribosphenida" pageId="1" pageNumber="98" phylum="Chordata" rank="genus">
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<emphasis id="B93CC332FF88FFE4FC4D94C7130524F3" box="[1018,1123,1459,1480]" italics="true" pageId="1" pageNumber="98">Ambondro</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C4864A3FF88FFE4FBD994C7139624F3" box="[1134,1264,1459,1480]" class="Mammalia" family="Ausktribosphenidae" genus="Asfaltomylos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ausktribosphenida" pageId="1" pageNumber="98" phylum="Chordata" rank="genus">
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<emphasis id="B93CC332FF88FFE4FBD994C7139624F3" box="[1134,1264,1459,1480]" italics="true" pageId="1" pageNumber="98">Asfaltomylos</emphasis>
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</taxonomicName>
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and shuotheriids), in so far as they are known, in having no paraconid on the first lower molar, an extremely abrupt discontinuity in size between the ultimate premolar and the first molar in the upper and lower dentitions, and in the presence of an enlarged dentary canal.
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</paragraph>
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<paragraph id="8BF71F20FF88FFE4FC869727126B2655" blockId="1.[816,1449,1619,1903]" pageId="1" pageNumber="98">
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<emphasis id="B93CC332FF88FFE4FC86972713262753" box="[817,1088,1619,1640]" italics="true" pageId="1" pageNumber="98">Revised specific diagnosis</emphasis>
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.
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<taxonomicName id="4C4864A3FF88FFE4FBF8972712742753" authorityName="Flannery, Archer, Rich and Jones" authorityYear="1995" box="[1103,1298,1619,1640]" class="Mammalia" family="Kollikodontidae" genus="Kollikodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Monotremata" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="ritchiei">
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<emphasis id="B93CC332FF88FFE4FBF8972712742753" box="[1103,1298,1619,1640]" italics="true" pageId="1" pageNumber="98">Kollikodon ritchiei</emphasis>
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</taxonomicName>
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is a large (by Mesozoic standards) mammal (estimated body mass approximately
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<quantity id="4CB0B2C5FF88FFE4FC7F97F9136A2798" box="[968,1036,1677,1699]" metricMagnitude="0" metricUnit="kg" metricValue="1.935" pageId="1" pageNumber="98" unit="g" value="1935.0">1935 g</quantity>
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based on m1 area (
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<bibRefCitation id="EFD962D1FF88FFE4FB7997F9120A2798" author="Legendre, S." box="[1230,1388,1677,1699]" pageId="1" pageNumber="98" pagination="191 - 212" refId="ref5941" refString="Legendre, S. 1986. Analysis of mammalian communities from the Late Eocene and Oligocene of southern France. Palaeovertebrata 16: 191 - 212." type="journal article" year="1986">Legendre, 1986</bibRefCitation>
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)) that differs from definitive monotremes that retain a functional dentition in exhibiting the following combination of features: bunodont molars with no vertical blades (lophs or crests) of any kind; broadly crescentic upper molars with unique cusp arrangement; reduced or absent posterior cingula/cingulids on all molars; markedly convex curve of the buccal edge of the upper and (to a lesser extent) lower molar rows.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |