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<document id="AC2A4A2026F88F969FA40E27CCA272EA" ID-CLB-Dataset="199797" ID-DOI="10.11646/zootaxa.5214.3.5" ID-GBIF-Dataset="9eef089e-6399-479b-9200-b2e091ccbb12" ID-ISSN="1175-5326" ID-Zenodo-Dep="7389401" ID-ZooBank="B9AD99D4-7061-4280-BFA2-F5339A9B9A55" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669970497842" checkinUser="plazi" docAuthor="Reynoso-Velasco, Daniel &amp; Sites, Robert W." docDate="2022" docId="03D6B32E8200D4243ECAF9A574FAFEA2" docLanguage="en" docName="zootaxa.5214.3.5.pdf" docOrigin="Zootaxa 5214 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ambrysus morelosensis Reynoso-Velasco &amp; Sites 2022, NEW SPECIES" docType="treatment" docVersion="2" lastPageNumber="433" masterDocId="FFEFCB568208D4283E5DFFB77524FFE3" masterDocTitle="Revision of the Ambrysus signoreti Stål species complex (Heteroptera: Naucoridae Ambrysinae). Section 1." masterLastPageNumber="439" masterPageNumber="421" pageNumber="429" updateTime="1699413971213" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4B2081167B7A40505ADA65762C963DE3">Revision of the Ambrysus signoreti Stål species complex (Heteroptera: Naucoridae Ambrysinae). Section 1.</mods:title>
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<mods:namePart id="38841FE4F1E571C2CC6B08B7BB3BCE78">Reynoso-Velasco, Daniel</mods:namePart>
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<mods:namePart id="2CB6B992358F3E35B7922C5A9ADAD51D">Sites, Robert W.</mods:namePart>
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<mods:affiliation id="FC45F30054B868A9F74029A4320A4A73">Enns Entomology Museum, Division of Plant Sciences, University of Missouri, Columbia, Missouri 65211, U. S. A. sitesr @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<treatment id="03D6B32E8200D4243ECAF9A574FAFEA2" ID-DOI="http://doi.org/10.5281/zenodo.7391986" ID-GBIF-Taxon="217325069" ID-Zenodo-Dep="7391986" LSID="urn:lsid:plazi:treatment:03D6B32E8200D4243ECAF9A574FAFEA2" httpUri="http://treatment.plazi.org/id/03D6B32E8200D4243ECAF9A574FAFEA2" lastPageId="12" lastPageNumber="433" pageId="8" pageNumber="429">
<subSubSection id="C36551B38200D4203ECAF9A57671F9CF" box="[151,853,1554,1581]" pageId="8" pageNumber="429" type="nomenclature">
<paragraph id="8BC002388200D4203ECAF9A57671F9CF" blockId="8.[151,853,1554,1617]" box="[151,853,1554,1581]" pageId="8" pageNumber="429">
<heading id="D088B5548200D4203ECAF9A57671F9CF" bold="true" box="[151,853,1554,1581]" fontSize="11" level="1" pageId="8" pageNumber="429" reason="1">
<emphasis id="B90BDE2A8200D4203ECAF9A57671F9CF" bold="true" box="[151,853,1554,1581]" pageId="8" pageNumber="429">
<taxonomicName id="4C7F79BB8200D4203ECAF9A577A3F9CF" authority="Reynoso &amp; Sites" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[151,647,1554,1581]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="429" phylum="Arthropoda" rank="species" species="morelosensis" status="sp. nov.">
<emphasis id="B90BDE2A8200D4203ECAF9A57497F9CE" bold="true" box="[151,435,1554,1581]" italics="true" pageId="8" pageNumber="429">Ambrysus morelosensis</emphasis>
Reynoso &amp; Sites
</taxonomicName>
<taxonomicNameLabel id="A23863518200D4203CD3F9A47671F9CF" box="[654,853,1554,1581]" pageId="8" pageNumber="429" rank="species">NEW SPECIES</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BC002388200D4203ECAF9807436F9B2" blockId="8.[151,853,1554,1617]" box="[151,274,1590,1617]" pageId="8" pageNumber="429">
(
<figureCitation id="13441EBD8200D4203EC2F98075C8F9B2" box="[159,236,1590,1617]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="8" pageNumber="429">Figs. 5</figureCitation>
7)
</paragraph>
<paragraph id="8BC002388200D4203ECAF9C9744BF8CA" blockId="8.[151,1437,1662,2013]" pageId="8" pageNumber="429">
<emphasis id="B90BDE2A8200D4203ECAF9C9771CF97B" bold="true" box="[151,568,1662,1689]" pageId="8" pageNumber="429">Description. Macropterous female.</emphasis>
<typeStatus id="54C4BC9A8200D4203C1DF9C877F3F97A" box="[576,727,1663,1689]" pageId="8" pageNumber="429" type="holotype">HOLOTYPE</typeStatus>
, length 12.87; maximum width 8.12.
<typeStatus id="54C4BC9A8200D4203ADEF9C871D4F97A" box="[1155,1264,1663,1689]" pageId="8" pageNumber="429" type="paratype">Paratypes</typeStatus>
(n = 1), length 11.75; maximum width 7.63. General shape elliptical; widest across embolia. Overall dorsal coloration of hemelytra dark-brown with light-brown marks; head light-brown; pronotum brown with dark-brown marks; legs brown (
<figureCitation id="13441EBD8200D4203B32F9707599F8E6" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="8" pageNumber="429">Fig. 5A</figureCitation>
). Dorsal surface coarsely punctate. Ventral coloration of head and thorax light-brown, abdomen with yellowishbrown pubescence.
</paragraph>
<paragraph id="8BC002388200D4213E9AF88475EFFF1A" blockId="8.[151,1437,1662,2013]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="430" pageId="8" pageNumber="429">
<emphasis id="B90BDE2A8200D4203E9AF8847420F8AE" box="[199,260,1843,1869]" italics="true" pageId="8" pageNumber="429">Head</emphasis>
. Head length 1.73; maximum width 3.41. Mostly light-brown with dark-brown marks along midline and posteriorly; coarsely punctate. Eyes convergent anteriorly, synthlipsis 1.73; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes convex, extending anteriorly in front of eyes 10.71% of head length; posterior margin between eyes strongly convex, extending posteriorly 28.57% of head length. Labrum width 2.00× length, evenly rounded. Labium with three visible light-brown segments, darkening distally, extending 0.76 beyond labrum not including extruded stylets. Antennal proportions 5:14:19:9, length 0.94, not reaching lateral margin of eye, elongate hairs on segment 4 and distal half of 3.
</paragraph>
<paragraph id="8BC002388201D4213E9AFEB471E7FCDA" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="430">
<emphasis id="B90BDE2A8201D4213E9AFEB4743CFEFE" box="[199,280,259,285]" italics="true" pageId="9" pageNumber="430">Thorax</emphasis>
. Pronotum coarsely punctate, ground color brown, dark-brown marking medially, dark-brown line behind eye; transverse sulcus marking anterior border of transverse band in posterior 1/3; transverse band brown; lateral margins light-brown, convergent, evenly convex, explanate; posterior margin almost straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 2.86× length; length at midline 2.35; maximum width at posterolateral corners 6.75. Prothorax ventrally pruinose throughout, except laterally; apices of propleura meeting at midline, not appressed to prosternellum; propleuron light-brown, medial 2/3 of posterior margin with golden setae. Probasisternum with sharp median carina and row of setae lateral to carina. Prosternellum not covered by apices of propleura. Scutellum coarsely punctate, triangular, brown, two reniform to ovate dark-brown marks medially, width 2.02× length, width 4.52, length 2.23. Hemelytra densely punctate, mostly dark-brown, length 9.20 (chord measurement); corium dark-brown, lightbrown irregularly shaped and checkmark-shaped markings at lateral and posterior margins, respectively. Clavus dark-brown, light-brown at mid-length and apically, length 3.80× width, length 4.71, width 1.24; claval commissure black, length 1.24. Embolium length 3.53, greatest width 1.30; lateral margin convex, yellowish-brown in anterior 2/3, dark-brown posteriorly. Membrane dark-brown (
<figureCitation id="13441EBD8201D4213CB6FD607662FD12" box="[747,838,726,753]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="9" pageNumber="430">Fig. 5A</figureCitation>
). Hindwings extending to posterior margin of tergum VI. Mesobasisternum midventral tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina.
</paragraph>
<paragraph id="8BC002388201D4213E9AFCF47722FAEE" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="430">
<emphasis id="B90BDE2A8201D4213E9AFCF475D8FCBF" box="[199,252,835,860]" italics="true" pageId="9" pageNumber="430">Legs</emphasis>
. All legs segments brown. Profemur posterior margin with row of tightly arranged setae in basal 2/3, row of short brown spines along middle third; anterior margin with dense pad of setae without associated spines. Protibia and tarsus with occlusal inner surface flattened; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibiae with ventrolateral and dorsolateral rows of stout reddishbrown spines; mesolateral rows intermixed with combs of spines. Metatibia with ventrolateral and dorsolateral rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apex of dorsal and mesal margins; two comb rows of stout spines near apex of ventral margin. Meso- and metatibiae and metatarsus with long, yellow to brown swimming hairs; hairs profuse on metatibia and -tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 3.10, tibia 2.41, tarsus 0.55; middle leg, femur 2.97, tibia 2.79, tarsomeres 13, 0.18, 0.54, 0.66; hind leg, femur 3.90, tibia 4.65, tarsomeres 13, 0.31, 1.00, 0.90.
</paragraph>
<paragraph id="8BC002388201D4213E9AFAA071C4F9B2" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="430">
<emphasis id="B90BDE2A8201D4213E9AFAA07415FAD2" box="[199,305,1303,1329]" italics="true" pageId="9" pageNumber="430">Abdomen</emphasis>
. Dorsally with lateral margins of IIIVIII exposed, IVVI each dark-brown in anterior 1/3 and yellowish-brown posteriorly; lateral margins of II to basal half of III smooth, distal half of IIIVI finely serrate, marginal row of profuse and short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corner of II (visible ventrally) right angled, IIIVI acute. Ventrally yellowish-brown to brown, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches clearly visible around spiracles on laterosternites IIVI. Mediosternite VII (subgenital plate) asymmetrical, width 1.36× length; length at midline 1.26; maximum width 1.72; left margin with broadly rounded lobe occupying its entire length; right margin with small apical lobe; posterolateral corners short, pointed, subtriangular, extending posteriorly further than the central lobe; central lobe poorly developed, inconspicuous, shallowly concave medially (
<figureCitation id="13441EBD8201D4213A26F98071F7F9B2" box="[1147,1235,1590,1617]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="9" pageNumber="430">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BC002388201D4213E9AF9EC76E0F83E" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="430">
<emphasis id="B90BDE2A8201D4213E9AF9EC749EF996" bold="true" box="[199,442,1627,1653]" pageId="9" pageNumber="430">Macropterous male.</emphasis>
<typeStatus id="54C4BC9A8201D4213F9EF9EC7714F996" box="[451,560,1627,1653]" pageId="9" pageNumber="430" type="paratype">Paratypes</typeStatus>
(n = 1), length 11.88; maximum width 7.75. Coloration, setation, pronotum proportions, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for
<typeStatus id="54C4BC9A8201D4213ECAF91475D9F95E" box="[151,253,1699,1725]" pageId="9" pageNumber="430" type="holotype">holotype</typeStatus>
. Otherwise hindwings extending to posterior margin of tergum V. Accessory genitalic process of tergum VI gently curved to right close to base, distal 1/3 angled to right to continue perpendicular to body axis (
<figureCitation id="13441EBD8201D4213B6BF97070B4F902" box="[1334,1424,1734,1761]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="9" pageNumber="430">Fig. 5C</figureCitation>
). Medial lobes of tergum VIII (pseudoparameres) symmetrical, posterolateral corners narrowly rounded, posteromesal corners broadly rounded, posterior margin concave (
<figureCitation id="13441EBD8201D4213C81F8B87611F8CA" box="[732,821,1806,1833]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="9" pageNumber="430">Fig. 5D</figureCitation>
). Phallosoma elongate, linear, slender, broadly rounded dorsoapically; left endosomal sclerite reduced, with few small and inconspicuous denticles; right endosomal sclerite vestigial, rounded, without denticles (
<figureCitation id="13441EBD8201D4213C66F8E077B5F892" box="[571,657,1878,1905]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="9" pageNumber="430">Fig. 5E</figureCitation>
). Parameres symmetrical, slightly longer than maximum width, mesal margin rounded, setae emanating from distal half of dorsal surface. Proctiger almost as long as width at base. Pygophore with anterior margin obtuse to subobtuse medially, elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (
<figureCitation id="13441EBD8201D4213D3FF8747693F83E" box="[866,951,1986,2013]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="9" pageNumber="430">Fig. 5F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0052B08202D4223EC5F9D077C5F908" ID-DOI="http://doi.org/10.5281/zenodo.7389421" ID-Zenodo-Dep="7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="10" pageNumber="431" startId="10.[152,255,1639,1663]" targetBox="[151,1435,181,1615]" targetPageId="10">
<paragraph id="8BC002388202D4223EC5F9D077C5F908" blockId="10.[151,1437,1639,1771]" pageId="10" pageNumber="431">
<emphasis id="B90BDE2A8202D4223EC5F9D0743DF99C" bold="true" box="[152,281,1639,1663]" pageId="10" pageNumber="431">FIGURE 5.</emphasis>
Structures of
<taxonomicName id="4C7F79BB8202D4223FF7F9D0766FF99C" authority="Reynoso &amp; Sites" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[426,843,1639,1664]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="431" phylum="Arthropoda" rank="species" species="morelosensis" status="sp. nov.">
<emphasis id="B90BDE2A8202D4223FF7F9D077BCF99C" box="[426,664,1639,1663]" italics="true" pageId="10" pageNumber="431">Ambrysus morelosensis</emphasis>
Reynoso &amp; Sites
</taxonomicName>
<emphasis id="B90BDE2A8202D4223D0EF9D076A9F99C" bold="true" box="[851,909,1639,1663]" pageId="10" pageNumber="431">
<taxonomicNameLabel id="A23863518202D4223D0EF9D076A9F99C" box="[851,909,1639,1663]" pageId="10" pageNumber="431" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite.
</paragraph>
</caption>
<subSubSection id="C36551B38202D4223E9AF8AE77D3F843" pageId="10" pageNumber="431" type="diagnosis">
<paragraph id="8BC002388202D4223E9AF8AE77D3F843" blockId="10.[151,1436,1817,2024]" pageId="10" pageNumber="431">
<emphasis id="B90BDE2A8202D4223E9AF8AE7466F8D0" bold="true" box="[199,322,1817,1843]" pageId="10" pageNumber="431">Diagnosis.</emphasis>
This species can be distinguished from the other three in this section by the female SGP with a large, plate-like, rounded lobe on the left margin and symmetrical subtriangular posterolateral corners (
<figureCitation id="13441EBD8202D4223B68F88A70ABF8BB" box="[1333,1423,1853,1880]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="10" pageNumber="431">Fig. 5B</figureCitation>
). Additionally, the male AGP tapers distally to a pointed and laterally directed apex (
<figureCitation id="13441EBD8202D4223A76F8D671A4F89F" box="[1067,1152,1889,1916]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="10" pageNumber="431">Fig. 5C</figureCitation>
) and the pseudoparamere posterolateral corners are narrowly rounded (
<figureCitation id="13441EBD8202D4223CCDF83277CEF843" box="[656,746,1925,1952]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="10" pageNumber="431">Fig. 5D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36551B38202D4233E9AF81E76E4FE86" lastPageId="11" lastPageNumber="432" pageId="10" pageNumber="431" type="discussion">
<paragraph id="8BC002388202D4233E9AF81E76E4FE86" blockId="10.[151,1436,1817,2024]" lastBlockId="11.[151,1437,151,897]" lastPageId="11" lastPageNumber="432" pageId="10" pageNumber="431">
<emphasis id="B90BDE2A8202D4223E9AF81E7497F827" bold="true" box="[199,435,1961,1988]" pageId="10" pageNumber="431">Comparative notes.</emphasis>
<taxonomicName id="4C7F79BB8202D4223FE1F81E77E1F820" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[444,709,1961,1987]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="431" phylum="Arthropoda" rank="species" species="morelosensis">
<emphasis id="B90BDE2A8202D4223FE1F81E77E1F820" box="[444,709,1961,1987]" italics="true" pageId="10" pageNumber="431">Ambrysus morelosensis</emphasis>
</taxonomicName>
presents a unique female SGP; none of the other three species in this group exhibits a similar large lobe on the left margin. The only other two species in the complex with a &quot;comparable&quot; feature are
<taxonomicName id="4C7F79BB8203D4233FEDFF207784FF52" authority="La Rivers" authorityName="La Rivers" authorityYear="1953" box="[432,672,151,177]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="scalenus">
<emphasis id="B90BDE2A8203D4233FEDFF20770AFF52" box="[432,558,151,177]" italics="true" pageId="11" pageNumber="432">A. scalenus</emphasis>
La Rivers
</taxonomicName>
and
<taxonomicName id="4C7F79BB8203D4233C8EFF2076B2FF52" authority="Reynoso" authorityName="Reynoso" authorityYear="2019" box="[723,918,151,177]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="sitesi">
<emphasis id="B90BDE2A8203D4233C8EFF20760FFF53" box="[723,811,151,177]" italics="true" pageId="11" pageNumber="432">A. sitesi</emphasis>
Reynoso
</taxonomicName>
; nonetheless, the lobe is truncate in
<taxonomicName id="4C7F79BB8203D4233B43FF2070B8FF52" authorityName="La Rivers" authorityYear="1953" box="[1310,1436,151,177]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="scalenus">
<emphasis id="B90BDE2A8203D4233B43FF2070B8FF52" box="[1310,1436,151,177]" italics="true" pageId="11" pageNumber="432">A. scalenus</emphasis>
</taxonomicName>
and in both species it is located at mid-length, not occupying the entire margin as in the new species. Additionally, these two species present the central lobe almost at the same level as the posterolateral corners, whereas in
<taxonomicName id="4C7F79BB8203D4233BD9FF68740CFEFE" authority="Reynoso-Velasco &amp; Sites, 2022" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="morelosensis" status="sp. nov.">
<emphasis id="B90BDE2A8203D4233BD9FF68740CFEFE" italics="true" pageId="11" pageNumber="432">A. morelosensis</emphasis>
</taxonomicName>
<emphasis id="B90BDE2A8203D4233F72FEB47449FEFE" bold="true" box="[303,365,259,285]" pageId="11" pageNumber="432">
<taxonomicNameLabel id="A23863518203D4233F72FEB47449FEFE" box="[303,365,259,285]" pageId="11" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the central lobe is poorly expressed, almost flat, and clearly below the level of the posterolateral corners. This new species is superficially similar to
<taxonomicName id="4C7F79BB8203D4233C82FE90761CFEA3" authorityName="Reynoso" authorityYear="2019" box="[735,824,295,321]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="sitesi">
<emphasis id="B90BDE2A8203D4233C82FE90761CFEA3" box="[735,824,295,321]" italics="true" pageId="11" pageNumber="432">A. sitesi</emphasis>
</taxonomicName>
; this similarity together with the &quot;comparable&quot; female SGP and the similar male pseudoparameres suggest a close relationship.
</paragraph>
</subSubSection>
<subSubSection id="C36551B38203D4233E9AFED97111FE4E" pageId="11" pageNumber="432" type="description">
<paragraph id="8BC002388203D4233E9AFED97111FE4E" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="432">
<emphasis id="B90BDE2A8203D4233E9AFED9741BFE6B" bold="true" box="[199,319,366,392]" pageId="11" pageNumber="432">Variation.</emphasis>
The single female
<typeStatus id="54C4BC9A8203D4233C41FED87758FE6A" box="[540,636,367,393]" pageId="11" pageNumber="432" type="paratype">paratype</typeStatus>
exhibits the posterior margin of the SGP evenly concave between the blunt posterolateral corners and the small apical lobe on the right margin is absent.
</paragraph>
</subSubSection>
<subSubSection id="C36551B38203D4233E9AFE017622FD12" pageId="11" pageNumber="432" type="distribution">
<paragraph id="8BC002388203D4233E9AFE01778EFD82" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="432">
<emphasis id="B90BDE2A8203D4233E9AFE017492FE33" bold="true" box="[199,438,438,464]" pageId="11" pageNumber="432">Habitat description.</emphasis>
The
<typeStatus id="54C4BC9A8203D4233FB0FE007739FE32" box="[493,541,439,465]" pageId="11" pageNumber="432">type</typeStatus>
locality, Río Amacuzac at Huajintlán, is in the southwestern area of the Mexican state of
<collectingRegion id="49BBCCDA8203D4233EB2FE6C7468FE16" box="[239,332,475,501]" country="Mexico" name="Morelos" pageId="11" pageNumber="432">Morelos</collectingRegion>
at an elevation of 931 meters. The first author visited the
<typeStatus id="54C4BC9A8203D4233D92FE6C76DBFE16" box="[975,1023,475,501]" pageId="11" pageNumber="432">type</typeStatus>
locality (L-1989) three decades after the
<typeStatus id="54C4BC9A8203D4233EE2FE4875CBFDFA" box="[191,239,511,537]" pageId="11" pageNumber="432">type</typeStatus>
specimens were collected; unfortunately, no more specimens of
<taxonomicName id="4C7F79BB8203D4233DE5FE487142FDFA" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[952,1126,511,537]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="morelosensis">
<emphasis id="B90BDE2A8203D4233DE5FE487142FDFA" box="[952,1126,511,537]" italics="true" pageId="11" pageNumber="432">A. morelosensis</emphasis>
</taxonomicName>
were found, only numerous specimens of
<taxonomicName id="4C7F79BB8203D4233F6FFD947716FDDE" authority="La Rivers. At" authorityName="La Rivers. At" authorityYear="1953" box="[306,562,547,573]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="hydor">
<emphasis id="B90BDE2A8203D4233F6FFD9474B7FDDE" box="[306,403,547,573]" italics="true" pageId="11" pageNumber="432">A. hydor</emphasis>
La Rivers. At
</taxonomicName>
the time, the river at the
<typeStatus id="54C4BC9A8203D4233D0DFD9476A4FDDE" box="[848,896,547,573]" pageId="11" pageNumber="432">type</typeStatus>
locality was approximately 40 meters wide and shallow, exposing boulders and cobble (
<figureCitation id="13441EBD8203D4233C0AFDF077B8FD82" box="[599,668,583,609]" captionStart="FIGURE 6" captionStartId="11.[152,255,1889,1913]" captionTargetBox="[164,1423,921,1866]" captionTargetId="figure-343@11.[164,1423,921,1866]" captionTargetPageId="11" captionText="FIGURE 6. Type locality of Ambrysus morelosensis Reynoso &amp; Sites n. sp. at Río Amacuzac, in the Mexican central state of Morelos" figureDoi="http://doi.org/10.5281/zenodo.7389423" httpUri="https://zenodo.org/record/7389423/files/figure.png" pageId="11" pageNumber="432">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BC002388203D4233E9AFDDD7622FD12" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="432">
<emphasis id="B90BDE2A8203D4233E9AFDDD7445FD67" bold="true" box="[199,353,618,644]" pageId="11" pageNumber="432">Distribution.</emphasis>
<taxonomicName id="4C7F79BB8203D4233F3AFDDC7749FD66" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[359,621,619,645]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="morelosensis">
<emphasis id="B90BDE2A8203D4233F3AFDDC7749FD66" box="[359,621,619,645]" italics="true" pageId="11" pageNumber="432">Ambrysus morelosensis</emphasis>
</taxonomicName>
is known from only two localities, one in the state of
<collectingRegion id="49BBCCDA8203D4233A95FDDC7060FD66" box="[1224,1348,619,645]" country="Mexico" name="Michoacan de Ocampo" pageId="11" pageNumber="432">Michoacán</collectingRegion>
and the other in
<collectingRegion id="49BBCCDA8203D4233EAFFD387476FD4A" box="[242,338,655,681]" country="Mexico" name="Morelos" pageId="11" pageNumber="432">Morelos</collectingRegion>
, in central-western and central
<collectingCountry id="F36842A88203D4233CF4FD387625FD4A" box="[681,769,655,681]" name="Mexico" pageId="11" pageNumber="432">Mexico</collectingCountry>
, respectively (Fig. 7). Both localities are at elevations below 1,000 meters and within the limits of the Balsas Basin biogeographical province; more specifically, in the middle (
<collectingRegion id="49BBCCDA8203D4233EC2FD607424FD12" box="[159,256,727,753]" country="Mexico" name="Morelos" pageId="11" pageNumber="432">Morelos</collectingRegion>
) and lower (
<collectingRegion id="49BBCCDA8203D4233FD1FD607728FD12" box="[396,524,727,753]" country="Mexico" name="Michoacan de Ocampo" pageId="11" pageNumber="432">Michoacán</collectingRegion>
) sections of the basin.
</paragraph>
</subSubSection>
<subSubSection id="C36551B38203D4233E9AFD4C710FFCDA" pageId="11" pageNumber="432" type="etymology">
<paragraph id="8BC002388203D4233E9AFD4C710FFCDA" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="432">
<emphasis id="B90BDE2A8203D4233E9AFD4C746AFCF6" bold="true" box="[199,334,763,789]" pageId="11" pageNumber="432">Etymology.</emphasis>
The specific epithet &quot;
<taxonomicName id="4C7F79BB8203D4233C18FD4C77F2FCF6" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[581,726,763,789]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="morelosensis">
<emphasis id="B90BDE2A8203D4233C18FD4C77F2FCF6" box="[581,726,763,789]" italics="true" pageId="11" pageNumber="432">morelosensis</emphasis>
</taxonomicName>
&quot; is in reference to the central Mexican state of
<collectingRegion id="49BBCCDA8203D4233AB1FD4C7068FCF6" box="[1260,1356,763,789]" country="Mexico" name="Morelos" pageId="11" pageNumber="432">Morelos</collectingRegion>
, where the
<typeStatus id="54C4BC9A8203D4233E9DFCA875D4FCDA" box="[192,240,799,825]" pageId="11" pageNumber="432">type</typeStatus>
locality is located. The species name means
<emphasis id="B90BDE2A8203D4233CBAFCA87100FCDA" box="[743,1060,799,825]" italics="true" pageId="11" pageNumber="432">
The
<taxonomicName id="4C7F79BB8203D4233D45FCA876A2FCDA" authorityName="Stal" authorityYear="1861" box="[792,902,799,825]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="genus">Ambrysus</taxonomicName>
from
<collectingRegion id="49BBCCDA8203D4233D95FCA87100FCDA" box="[968,1060,799,825]" country="Mexico" name="Morelos" pageId="11" pageNumber="432">Morelos</collectingRegion>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C36551B38203D4233E9AFCF574BDFC62" pageId="11" pageNumber="432" type="multiple">
<paragraph id="8BC002388203D4233E9AFCF574BDFC62" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="432">
<emphasis id="B90BDE2A8203D4233E9AFCF57475FCBE" bold="true" box="[199,337,834,861]" pageId="11" pageNumber="432">Repository.</emphasis>
The
<typeStatus id="54C4BC9A8203D4233FCFFCF474D0FCBE" box="[402,500,835,861]" pageId="11" pageNumber="432" type="holotype">holotype</typeStatus>
and
<typeStatus id="54C4BC9A8203D4233C6FFCF477B9FCBE" box="[562,669,835,861]" pageId="11" pageNumber="432" type="paratype">paratypes</typeStatus>
will be deposited in the
<location id="8EA054E38203D4233D95FCF475DFFC62" LSID="urn:lsid:plazi:treatment:03D6B32E8200D4243ECAF9A574FAFEA2:8EA054E38203D4233D95FCF475DFFC62" name="Coleccion Entomologica del Instituto de Ecologia" pageId="11" pageNumber="432">
Colección Entomológica del
<collectorName id="268A67EE8203D4233B47FCF475DFFC62" pageId="11" pageNumber="432">Instituto de Ecología</collectorName>
</location>
A.C. (IEXA).
</paragraph>
</subSubSection>
<caption id="DF0052B08203D4233EC5F8D675CFF87E" ID-DOI="http://doi.org/10.5281/zenodo.7389423" ID-Zenodo-Dep="7389423" httpUri="https://zenodo.org/record/7389423/files/figure.png" pageId="11" pageNumber="432" startId="11.[152,255,1889,1913]" targetBox="[164,1423,921,1866]" targetPageId="11">
<paragraph id="8BC002388203D4233EC5F8D675CFF87E" blockId="11.[151,1436,1889,1949]" pageId="11" pageNumber="432">
<emphasis id="B90BDE2A8203D4233EC5F8D6743CF89A" bold="true" box="[152,280,1889,1913]" pageId="11" pageNumber="432">FIGURE 6.</emphasis>
Type locality of
<taxonomicName id="4C7F79BB8203D4233F9AF8D67642F89A" authority="Reynoso &amp; Sites" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[455,870,1889,1914]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="432" phylum="Arthropoda" rank="species" species="morelosensis" status="sp. nov.">
<emphasis id="B90BDE2A8203D4233F9AF8D67790F89A" box="[455,692,1889,1913]" italics="true" pageId="11" pageNumber="432">Ambrysus morelosensis</emphasis>
Reynoso &amp; Sites
</taxonomicName>
<emphasis id="B90BDE2A8203D4233D30F8D67682F89A" bold="true" box="[877,934,1889,1913]" pageId="11" pageNumber="432">
<taxonomicNameLabel id="A23863518203D4233D30F8D67682F89A" box="[877,934,1889,1913]" pageId="11" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
at Río Amacuzac, in the Mexican central state of Morelos
</paragraph>
</caption>
<subSubSection id="C36551B38204D4243E9AFF2074FAFEA2" pageId="12" pageNumber="433" type="type_taxon">
<paragraph id="8BC002388204D4243E9AFF2074FAFEA2" blockId="12.[151,1437,150,321]" pageId="12" pageNumber="433">
<emphasis id="B90BDE2A8204D4243E9AFF2074D3FF53" bold="true" box="[199,503,150,177]" pageId="12" pageNumber="433">Type material examined.</emphasis>
<typeStatus id="54C4BC9A8204D4243C5FFF2077B2FF52" box="[514,662,151,177]" pageId="12" pageNumber="433" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D79C9B18204D4243CFCFF207793FF52" box="[673,695,151,177]" pageId="12" pageNumber="433" type="female"></specimenCount>
.
<materialsCitation id="3B1708658204D4243C95FF21740FFF1A" ID-GBIF-Occurrence="3986688356" collectingDate="1985-03-09" collectorName="M. Albores" country="Mexico" elevation="931" latitude="18.609879" location="Cuernavaca-Taxco" longLatPrecision="1" longitude="-99.42788" pageId="12" pageNumber="433" specimenCount="1" stateProvince="Morelos" typeStatus="holotype">
<emphasis id="B90BDE2A8204D4243C95FF2176CFFF53" bold="true" box="[712,1003,150,176]" pageId="12" pageNumber="433">
<collectingCountry id="F36842A88204D4243C95FF217667FF53" box="[712,835,150,176]" name="Mexico" pageId="12" pageNumber="433">MEXICO</collectingCountry>
:
<collectingRegion id="49BBCCDA8204D4243D0EFF2176C3FF53" box="[851,999,150,176]" country="Mexico" name="Morelos" pageId="12" pageNumber="433">MORELOS</collectingRegion>
:
</emphasis>
[Mpio. Amacuzac, Río Amacuzac at Huajintlán], km 50 [on carr. 95]
<location id="8EA054E38204D4243C7CFF0C77D1FF36" LSID="urn:lsid:plazi:treatment:03D6B32E8200D4243ECAF9A574FAFEA2:8EA054E38204D4243C7CFF0C77D1FF36" box="[545,757,187,213]" country="Mexico" latitude="18.609879" longLatPrecision="1" longitude="-99.42788" name="Cuernavaca-Taxco" pageId="12" pageNumber="433" stateProvince="Morelos">Cuernavaca-Taxco</location>
, [
<geoCoordinate id="EE4B64FF8204D4243D51FF0D7690FF36" box="[780,948,186,213]" degrees="18" direction="north" minutes="36" orientation="latitude" pageId="12" pageNumber="433" precision="1" seconds="35.56" value="18.609879">18º36'35.56''N</geoCoordinate>
,
<geoCoordinate id="EE4B64FF8204D4243D98FF0D714AFF36" box="[965,1134,186,213]" degrees="99" direction="west" minutes="25" orientation="longitude" pageId="12" pageNumber="433" precision="1" seconds="40.38" value="-99.42788">99º25'40.38''W</geoCoordinate>
,
<quantity id="4C87AFDD8204D4243A22FF0C71EBFF36" box="[1151,1231,187,213]" metricMagnitude="2" metricUnit="m" metricValue="9.309999999999999" pageId="12" pageNumber="433" unit="m" value="931.0">
<elevation id="0052E50B8204D4243A22FF0C71EBFF36" box="[1151,1231,187,213]" metricMagnitude="2" metricUnit="m" metricValue="9.309999999999999" pageId="12" pageNumber="433" unit="m" value="931.0">931 m</elevation>
</quantity>
],
<date id="FFC124F88204D4243ABAFF0D7048FF36" box="[1255,1388,186,213]" pageId="12" pageNumber="433" value="1985-03-09">
<collectingDate id="EF85DD108204D4243ABAFF0D7048FF36" box="[1255,1388,186,213]" pageId="12" pageNumber="433" value="1985-03-09">09-III-1985</collectingDate>
</date>
,
<collectorName id="268A67EE8204D4243B21FF0C75D4FF1A" pageId="12" pageNumber="433">M. Albores</collectorName>
coll.
</materialsCitation>
<materialsCitation id="3B1708658204D4243F69FF68764BFF1A" ID-GBIF-Occurrence="3986688337" box="[308,879,223,249]" collectingDate="1985-03-09" collectionCode="IEXA" collectorName="M. Albores" country="Mexico" elevation="931" latitude="18.609879" location="Cuernavaca-Taxco" longLatPrecision="1" longitude="-99.42788" pageId="12" pageNumber="433" specimenCount="1" specimenCount-male="1" stateProvince="Morelos" typeStatus="paratype">
<typeStatus id="54C4BC9A8204D4243F69FF6874F7FF1A" box="[308,467,223,249]" pageId="12" pageNumber="433" type="paratype">PARATYPES</typeStatus>
: same data as holotype (
<specimenCount id="9D79C9B18204D4243CAEFF687632FF1A" box="[755,790,223,249]" pageId="12" pageNumber="433" type="male">1♂</specimenCount>
<collectionCode id="ED6E9AFD8204D4243D42FF687643FF1A" box="[799,871,223,249]" pageId="12" pageNumber="433">IEXA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1708658204D4243D21FF5774FEFEA2" ID-GBIF-Occurrence="3986688352" collectingDate="2012-12-24" collectionCode="IEXA" collectorName="ReynosoVelasco, D." country="Mexico" county="Mpio. Buenavista" elevation="440" latitude="19.216057" location="Buenavista Tomatlan" longLatPrecision="1" longitude="-102.57506" municipality="Buenavista River" pageId="12" pageNumber="433" specimenCount="1" specimenCount-female="1" stateProvince="Michoacan de Ocampo" typeStatus="paratype">
<emphasis id="B90BDE2A8204D4243D21FF57711FFF1F" bold="true" box="[892,1083,224,252]" pageId="12" pageNumber="433">
<collectingRegion id="49BBCCDA8204D4243D21FF577112FF1F" box="[892,1078,224,252]" country="Mexico" name="Michoacan de Ocampo" pageId="12" pageNumber="433">MICHOACÁN</collectingRegion>
:
</emphasis>
<collectingCounty id="62A17AB48204D4243A18FF687036FF1A" box="[1093,1298,223,249]" pageId="12" pageNumber="433">Mpio. Buenavista</collectingCounty>
,
<collectingMunicipality id="6BA498428204D4243B7DFF6875F1FEFE" pageId="12" pageNumber="433">Buenavista River</collectingMunicipality>
at
<location id="8EA054E38204D4243EA7FEB474CEFEFE" LSID="urn:lsid:plazi:treatment:03D6B32E8200D4243ECAF9A574FAFEA2:8EA054E38204D4243EA7FEB474CEFEFE" box="[250,490,259,285]" country="Mexico" county="Mpio. Buenavista" latitude="19.216057" longLatPrecision="1" longitude="-102.57506" municipality="Buenavista River" name="Buenavista Tomatlan" pageId="12" pageNumber="433" stateProvince="Michoacan de Ocampo">Buenavista Tomatlán</location>
, below dam,
<geoCoordinate id="EE4B64FF8204D4243CDFFEB5763DFEFE" box="[642,793,258,285]" degrees="19" direction="north" minutes="12" orientation="latitude" pageId="12" pageNumber="433" precision="1" seconds="57.8" value="19.216057">19º12'57.8''N</geoCoordinate>
,
<geoCoordinate id="EE4B64FF8204D4243D7BFEB476EBFEFE" box="[806,975,259,285]" degrees="102" direction="west" minutes="34" orientation="longitude" pageId="12" pageNumber="433" precision="1" seconds="30.2" value="-102.57506">102º34'30.2''W</geoCoordinate>
,
<quantity id="4C87AFDD8204D4243D81FEB47102FEFE" box="[988,1062,259,285]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="12" pageNumber="433" unit="m" value="440.0">
<elevation id="0052E50B8204D4243D81FEB47102FEFE" box="[988,1062,259,285]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="12" pageNumber="433" unit="m" value="440.0">440 m</elevation>
</quantity>
,
<date id="FFC124F88204D4243A6EFEB471E7FEFE" box="[1075,1219,259,285]" pageId="12" pageNumber="433" value="2012-12-24">
<collectingDate id="EF85DD108204D4243A6EFEB471E7FEFE" box="[1075,1219,259,285]" pageId="12" pageNumber="433" value="2012-12-24">24-XII-2012</collectingDate>
</date>
, L-1481,
<collectorName id="268A67EE8204D4243B6CFEB4743CFEA2" pageId="12" pageNumber="433">ReynosoVelasco, D.</collectorName>
coll. (
<specimenCount id="9D79C9B18204D4243F3FFE9074A7FEA2" box="[354,387,295,321]" pageId="12" pageNumber="433" type="female">1♀</specimenCount>
<collectionCode id="ED6E9AFD8204D4243FD7FE9074F6FEA2" box="[394,466,295,321]" pageId="12" pageNumber="433">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>