494 lines
69 KiB
XML
494 lines
69 KiB
XML
<document id="3460B70B10DB9C4C0121D71F5F8CCC65" ID-DOI="10.11646/zootaxa.4750.1.6" ID-GBIF-Dataset="7c160556-e0e1-4fd6-85e1-1f6fa0ab87d7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3702903" ID-ZooBank="575B1970-C7B5-4688-9B41-99F046AFD3F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583849335187" checkinUser="plazi" docAuthor="Lee, Damin & Kim, Won" docDate="2020" docId="03D687D47C0A4132FF51F450FAF4D6CE" docLanguage="en" docName="zootaxa.4750.1.6.pdf" docOrigin="Zootaxa 4750 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pycnogonum (Nulloviger) granulatum Lee & Kim 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="128" masterDocId="FFEFFFAC7C0B4134FFC6F619FFD7D619" masterDocTitle="New species of Pycnogonum (Pycnogonida: Pycnogonidae) from Green Island Taiwan, with an additional note on the holotype of P. spatium" masterLastPageNumber="130" masterPageNumber="122" pageNumber="123" updateTime="1698726705764" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B7AE90EE65B022938E34171918B2A2BB">
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<mods:title id="45CC195218025ADC426E82A16866ECD1">New species of Pycnogonum (Pycnogonida: Pycnogonidae) from Green Island Taiwan, with an additional note on the holotype of P. spatium</mods:title>
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<mods:name id="4F5C2B47627D14B58EAF4B3448577E86" type="personal">
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<mods:namePart id="FF7C5F8F302D8496A35D8F1B22E1B7D5">Lee, Damin</mods:namePart>
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<mods:namePart id="B9BC8CD447043251A1A18A4A266A9119">Kim, Won</mods:namePart>
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<mods:typeOfResource id="6C059B596FAD46E0A862970B25297B13">text</mods:typeOfResource>
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<mods:title id="22548DFAF6E7D99ADFDC655702BC1633">Zootaxa</mods:title>
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<mods:part id="128EDBB9CF6D90C1FA9333A307643DDC">
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<mods:date id="D57E455E0C798AF1BFE417EA9632A162">2020</mods:date>
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<mods:number id="5619A53F84AC83EE24896329E0435685">2020-03-10</mods:number>
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<mods:number id="71F259395F67DA6DF7B196C63D8FAC97">4750</mods:number>
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<mods:detail id="75981978A80F1870C797092C98B96874" type="issue">
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<mods:number id="46DC0AEED09C0522F482E12846B8BECD">1</mods:number>
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<treatment id="03D687D47C0A4132FF51F450FAF4D6CE" ID-DOI="http://doi.org/10.5281/zenodo.3706313" ID-GBIF-Taxon="162481367" ID-Zenodo-Dep="3706313" LSID="urn:lsid:plazi:treatment:03D687D47C0A4132FF51F450FAF4D6CE" httpUri="http://treatment.plazi.org/id/03D687D47C0A4132FF51F450FAF4D6CE" lastPageId="6" lastPageNumber="128" pageId="1" pageNumber="123">
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<subSubSection id="C36565497C0A4135FF51F450FD19D47A" box="[151,718,585,612]" pageId="1" pageNumber="123" type="nomenclature">
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<paragraph id="8BC036C27C0A4135FF51F450FD19D47A" blockId="1.[151,718,585,647]" box="[151,718,585,612]" pageId="1" pageNumber="123">
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<heading id="D08881AE7C0A4135FF51F450FD19D47A" bold="true" box="[151,718,585,612]" fontSize="11" level="1" pageId="1" pageNumber="123" reason="1">
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<emphasis id="B90BEAD07C0A4135FF51F450FD19D47A" bold="true" box="[151,718,585,612]" pageId="1" pageNumber="123">
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<taxonomicName id="4C7F4D417C0A4135FF51F450FDBED47D" ID-CoL="8YKBK" authority="Lee & Kim, 2020" authorityName="Lee & Kim" authorityYear="2020" box="[151,617,585,612]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="1" pageNumber="123" phylum="Arthropoda" rank="species" species="granulatum" status="sp. nov." subGenus="Nulloviger">
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<emphasis id="B90BEAD07C0A4135FF51F450FEE0D47A" bold="true" box="[151,311,585,611]" italics="true" pageId="1" pageNumber="123">Pycnogonum</emphasis>
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(
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<emphasis id="B90BEAD07C0A4135FE8EF450FE1ED47D" bold="true" box="[328,457,585,612]" italics="true" pageId="1" pageNumber="123">Nulloviger</emphasis>
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)
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<emphasis id="B90BEAD07C0A4135FE1CF450FDBED47D" bold="true" box="[474,617,585,612]" italics="true" pageId="1" pageNumber="123">granulatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23857AB7C0A4135FDB7F450FD19D47A" box="[625,718,585,611]" pageId="1" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36565497C0A4135FF51F474FDF2D49E" box="[151,549,621,647]" pageId="1" pageNumber="123" type="description">
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<paragraph id="8BC036C27C0A4135FF51F474FDF2D49E" blockId="1.[151,718,585,647]" box="[151,549,621,647]" pageId="1" pageNumber="123">
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<figureCitation id="13442A477C0A4135FF51F474FF2CD49E" box="[151,251,621,647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1981,2006]" captionTargetBox="[165,1423,342,1952]" captionTargetId="figure@2.[164,1423,342,1953]" captionTargetPageId="2" captionText="FIGURE 1. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male. A, trunk, dorsal view; B, trunk, lateral view; C, trunk, anterodorsal view; D, third leg. Scale bars: 1 mm (A–C); 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.3702905" httpUri="https://zenodo.org/record/3702905/files/figure.png" pageId="1" pageNumber="123">Figures 1</figureCitation>
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A–D, 2A–F, 3A–B, 4A–D
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</paragraph>
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</subSubSection>
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<subSubSection id="C36565497C0A4135FF51F4ADFC02D4EA" pageId="1" pageNumber="123" type="materials_examined">
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<paragraph id="8BC036C27C0A4135FF51F4ADFC02D4EA" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">
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<emphasis id="B90BEAD07C0A4135FF51F4ADFE5FD4D6" bold="true" box="[151,392,692,719]" pageId="1" pageNumber="123">Material examined.</emphasis>
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<materialsCitation id="3B173C9F7C0A4135FE51F4ACFC02D4EA" ID-GBIF-Occurrence="2575116366" collectingDate="2016-05-24" collectionCode="SCUBA" country="Taiwan" elevation="18" latitude="22.681944" location="Gong Guan Bi" longLatPrecision="1" longitude="121.488304" municipality="Green Island" pageId="1" pageNumber="123" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54C488607C0A4135FE51F4ACFDD5D4D6" box="[407,514,693,719]" pageId="1" pageNumber="123" type="holotype">Holotype</typeStatus>
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, male, ASIZCH000073,
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<location id="8EA060197C0A4135FCE9F4ACFC36D4D6" LSID="urn:lsid:plazi:treatment:03D687D47C0A4132FF51F450FAF4D6CE:8EA060197C0A4135FCE9F4ACFC36D4D6" box="[815,993,693,719]" country="Taiwan" latitude="22.681944" longLatPrecision="1" longitude="121.488304" municipality="Green Island" name="Gong Guan Bi" pageId="1" pageNumber="123">Gong Guan Bi</location>
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diving point,
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<collectingMunicipality id="6BA4ACB87C0A4135FB50F4ACFAE7D4D6" box="[1174,1328,693,719]" pageId="1" pageNumber="123">Green Island</collectingMunicipality>
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,
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<collectingCountry id="F36876527C0A4135FA85F4ACFA4FD4D6" box="[1347,1432,693,719]" name="Taiwan" pageId="1" pageNumber="123">Taiwan</collectingCountry>
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,
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<geoCoordinate id="EE4B50057C0A4135FF51F4C1FEEFD4EA" box="[151,312,728,755]" degrees="22" direction="north" minutes="40" orientation="latitude" pageId="1" pageNumber="123" precision="1" seconds="55.0" value="22.681944">22°40’55.0”N</geoCoordinate>
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,
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<geoCoordinate id="EE4B50057C0A4135FE85F4C0FE39D4EA" box="[323,494,729,755]" degrees="121" direction="east" minutes="29" orientation="longitude" pageId="1" pageNumber="123" precision="1" seconds="17.9" value="121.488304">121°29’17.9”E</geoCoordinate>
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,
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<elevation id="0052D1F17C0A4135FE3CF4C0FDE2D4EA" box="[506,565,729,755]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="123" unit="m" value="18.0">18 m</elevation>
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</quantity>
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,
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<collectionCode id="ED6EAE077C0A4135FD87F4C0FD76D4EA" box="[577,673,729,755]" pageId="1" pageNumber="123">SCUBA</collectionCode>
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, Damin Lee,
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<date id="FFC110027C0A4135FCFEF4C0FC06D4EA" box="[824,977,729,755]" pageId="1" pageNumber="123" value="2016-05-24">
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<collectingDate id="EF85E9EA7C0A4135FCFEF4C0FC06D4EA" box="[824,977,729,755]" pageId="1" pageNumber="123" value="2016-05-24">24 May 2016</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36565497C0A4135FF01F4E5FEC6D156" pageId="1" pageNumber="123" type="description">
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<paragraph id="8BC036C27C0A4135FF01F4E5FCA2D546" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">
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<emphasis id="B90BEAD07C0A4135FF01F4E5FE8ED50E" bold="true" box="[199,345,764,791]" pageId="1" pageNumber="123">Description.</emphasis>
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Male
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<typeStatus id="54C488607C0A4135FE66F4E4FDD1D50E" box="[416,518,765,791]" pageId="1" pageNumber="123" type="holotype">holotype</typeStatus>
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. Trunk granulated (
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<figureCitation id="13442A477C0A4135FD26F4E4FCEED50E" box="[736,825,765,791]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="1" pageNumber="123">Fig. 2A</figureCitation>
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,
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<figureCitation id="13442A477C0A4135FC83F4E4FCBDD50E" box="[837,874,765,791]" captionStart="FIGURE 3" captionStartId="4.[151,250,1297,1322]" captionTargetBox="[177,1411,345,1272]" captionTargetId="figure@4.[177,1411,345,1272]" captionTargetPageId="4" captionText="FIGURE 3. Integument. Pycnogonum (Nulloviger) granulatum sp. nov.; A–B. A, granulated integument of abdomen, dorsal view; B, granulated integument of femur, distal surface. P. spatium; C–D. C, pitted integument of abdomen, dorsal view; D, granulated and fitted integument of femur, distal surface. All images are observed under the light microscope (Olympus BX51, Japan). Scale bars: 0.1 mm (A, C)." figureDoi="http://doi.org/10.5281/zenodo.3702911" httpUri="https://zenodo.org/record/3702911/files/figure.png" pageId="1" pageNumber="123">3A</figureCitation>
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); segment 1, 2, and 3 articulated; segment 3 and 4 fused. Anterior three segments with transverse ridge, each 1/4 times as high as trunk height (
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<figureCitation id="13442A477C0A4135FB65F538FAFFD522" box="[1187,1320,801,827]" captionStart="FIGURE 1" captionStartId="2.[151,250,1981,2006]" captionTargetBox="[165,1423,342,1952]" captionTargetId="figure@2.[164,1423,342,1953]" captionTargetPageId="2" captionText="FIGURE 1. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male. A, trunk, dorsal view; B, trunk, lateral view; C, trunk, anterodorsal view; D, third leg. Scale bars: 1 mm (A–C); 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.3702905" httpUri="https://zenodo.org/record/3702905/files/figure.png" pageId="1" pageNumber="123">Fig. 1B, 1C</figureCitation>
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,
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<figureCitation id="13442A477C0A4135FAF3F538FA8FD522" box="[1333,1368,801,827]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="1" pageNumber="123">2B</figureCitation>
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); segment
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<quantity id="4C879B277C0A4135FF10F55CFED6D546" box="[214,257,837,863]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="1" pageNumber="123" unit="in" value="4.0">4 in</quantity>
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trapezoidal shape, expanding posteriorly (
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<figureCitation id="13442A477C0A4135FD1BF55CFCE1D546" box="[733,822,837,863]" captionStart="FIGURE 1" captionStartId="2.[151,250,1981,2006]" captionTargetBox="[165,1423,342,1952]" captionTargetId="figure@2.[164,1423,342,1953]" captionTargetPageId="2" captionText="FIGURE 1. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male. A, trunk, dorsal view; B, trunk, lateral view; C, trunk, anterodorsal view; D, third leg. Scale bars: 1 mm (A–C); 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.3702905" httpUri="https://zenodo.org/record/3702905/files/figure.png" pageId="1" pageNumber="123">Fig. 1A</figureCitation>
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,
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<figureCitation id="13442A477C0A4135FC84F55CFCBFD546" box="[834,872,837,863]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="1" pageNumber="123">2A</figureCitation>
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).
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</paragraph>
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<paragraph id="8BC036C27C0A4135FF01F570FDADD5BE" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">Lateral processes half as long as broad, separated by 1/4 to 1/3 of base width; dorsodistal tubercle about 1/4 times as high as transverse ridges on trunk.</paragraph>
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<paragraph id="8BC036C27C0A4135FF01F5A8FC5AD5F6" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">Ocular tubercle on anterior margin of cephalon, truncated cone shape with round tip, height half of base width, slightly taller than transverse ridges on trunk. Four eyes pigmented.</paragraph>
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<paragraph id="8BC036C27C0A4135FF01F5E0FCBBD242" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">Proboscis cylindrical, 1.7 times as long as base width, with slight constriction at half length; ventral surface slightly longer than dorsal surface; dorsal tubercle present at 2/3 length from base, rising steeply on anterior surface, sloping down gently on posterior surface. Oral glands not found.</paragraph>
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<paragraph id="8BC036C27C0A4135FF01F27CFDB3D2BA" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">
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Abdomen (
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<figureCitation id="13442A477C0A4135FE83F27CFE49D266" box="[325,414,1125,1151]" captionStart="FIGURE 1" captionStartId="2.[151,250,1981,2006]" captionTargetBox="[165,1423,342,1952]" captionTargetId="figure@2.[164,1423,342,1953]" captionTargetPageId="2" captionText="FIGURE 1. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male. A, trunk, dorsal view; B, trunk, lateral view; C, trunk, anterodorsal view; D, third leg. Scale bars: 1 mm (A–C); 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.3702905" httpUri="https://zenodo.org/record/3702905/files/figure.png" pageId="1" pageNumber="123">Fig. 1A</figureCitation>
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,
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<figureCitation id="13442A477C0A4135FE6FF27CFE19D266" box="[425,462,1125,1151]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="1" pageNumber="123">2A</figureCitation>
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,
|
||
<figureCitation id="13442A477C0A4135FE1FF27CFE2AD266" box="[473,509,1125,1151]" captionStart="FIGURE 3" captionStartId="4.[151,250,1297,1322]" captionTargetBox="[177,1411,345,1272]" captionTargetId="figure@4.[177,1411,345,1272]" captionTargetPageId="4" captionText="FIGURE 3. Integument. Pycnogonum (Nulloviger) granulatum sp. nov.; A–B. A, granulated integument of abdomen, dorsal view; B, granulated integument of femur, distal surface. P. spatium; C–D. C, pitted integument of abdomen, dorsal view; D, granulated and fitted integument of femur, distal surface. All images are observed under the light microscope (Olympus BX51, Japan). Scale bars: 0.1 mm (A, C)." figureDoi="http://doi.org/10.5281/zenodo.3702911" httpUri="https://zenodo.org/record/3702911/files/figure.png" pageId="1" pageNumber="123">3A</figureCitation>
|
||
) articulated at base, spindle shaped directing horizontally, reaching to distal end of coxa 2 of fourth legs, slightly bifid at tip.
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C0A4135FF01F2B4FD88D2DE" blockId="1.[151,1437,692,2015]" box="[199,607,1197,1223]" pageId="1" pageNumber="123">Palps, chelifores and ovigers absent.</paragraph>
|
||
<paragraph id="8BC036C27C0A4135FF01F2C8FA55D082" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">
|
||
Third leg (
|
||
<figureCitation id="13442A477C0A4135FE84F2C8FE4BD2F2" box="[322,412,1233,1259]" captionStart="FIGURE 1" captionStartId="2.[151,250,1981,2006]" captionTargetBox="[165,1423,342,1952]" captionTargetId="figure@2.[164,1423,342,1953]" captionTargetPageId="2" captionText="FIGURE 1. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male. A, trunk, dorsal view; B, trunk, lateral view; C, trunk, anterodorsal view; D, third leg. Scale bars: 1 mm (A–C); 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.3702905" httpUri="https://zenodo.org/record/3702905/files/figure.png" pageId="1" pageNumber="123">Fig. 1D</figureCitation>
|
||
,
|
||
<figureCitation id="13442A477C0A4135FE6DF2C8FE19D2F2" box="[427,462,1233,1259]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="1" pageNumber="123">2C</figureCitation>
|
||
,
|
||
<figureCitation id="13442A477C0A4135FE1AF2C8FE2AD2F2" box="[476,509,1233,1259]" captionStart="FIGURE 4" captionStartId="5.[151,250,1135,1160]" captionTargetBox="[177,1411,181,1110]" captionTargetId="figure@5.[177,1411,181,1110]" captionTargetPageId="5" captionText="FIGURE 4. Legs of Pycnogonum (Nulloviger) granulatum sp. nov. A, first leg; B, second leg; C, third leg; D, fourth leg. Scale bars: 0.5 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.3702913" httpUri="https://zenodo.org/record/3702913/files/figure.png" pageId="1" pageNumber="123">4C</figureCitation>
|
||
) stout, granulated, having intermittent small spines. Coxa 1 with two dorsodistal tubercles, without ventral spines. Coxa 2 slightly longer than coxa 1 and coxa 3, with two minute dorsodistal tubercles, several spines on ventral surface. Coxa 3 slightly longer than coxa 1, with two minute dorsodistal tubercles, several spines ventrally. Femur about 3 times as long as base width, bearing two heavy dorsodistal lobes, two dorsolateral digitiform processes, and ventroproximal swelling; mid-dorsal projection large, with two tubercles of which anterior tubercle larger, nodulous, leaning slightly forward with short spine and posterior tubercle smaller, bearing spine. Tibia 1 3.6 times as long as base width, with pair of dorsodistal tubercles, several irregular dorsal tubercles, bearing ventrodistal swelling with two small spines on ventrodistal surface. Tibia 2 1.7 times as long as base width, half as long as tibia 1, with pair of dorsodistal tubercles, having nodulous tubercles on dorsal surface, bearing several spines on ventrodistal surface. Each femur and tibia with long, dorsodistal spine. Tarsus 0.25 times as long as propodus; dorsal surface concave; ventral surface convex, with several spines. Propodus moderately curved, with eight spines dorsally and eight sole spines ventrally, heel spines absent. Spines on ventral surface of tibia, tarsus, and propodus with cleft tips (
|
||
<figureCitation id="13442A477C0A4135FE19F098FDE0D082" box="[479,567,1665,1691]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="1" pageNumber="123">Fig. 2E</figureCitation>
|
||
). Main claw curved, 0.6 times as long as propodus. Auxiliary claws absent.
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C0A4135FF01F0BCFD46D0FA" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">
|
||
Male gonopores present on ventral surface of coxa 2 of fourth legs (
|
||
<figureCitation id="13442A477C0A4135FC77F0BCFBD2D0A6" box="[945,1029,1701,1727]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="1" pageNumber="123">Fig. 2F</figureCitation>
|
||
). Coxal pellicula (
|
||
<emphasis id="B90BEAD07C0A4135FB16F0BCFB28D0A6" box="[1232,1279,1701,1727]" italics="true" pageId="1" pageNumber="123">vide</emphasis>
|
||
<bibRefCitation id="EFEE4B337C0A4135FAC2F0BCFA42D0A6" author="Staples, D. A." box="[1284,1429,1701,1727]" pageId="1" pageNumber="123" pagination="79 - 94" refId="ref5442" refString="Staples, D. A. (2007) A new species of Colossendeis (Pycnogonida: Colossendeidae) together with records from Australian and New Zealand waters. Memoirs of Museum Victoria, 64, 79 - 94. https: // doi. org / 10.24199 / j. mmv. 2007.64.8" type="journal article" year="2007">Staples 2007</bibRefCitation>
|
||
) absent on dorsal surface of coxa 2 of all legs.
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C0A4135FF01F0F4FEC6D156" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">
|
||
<emphasis id="B90BEAD07C0A4135FF01F0F4FE10D11E" bold="true" box="[199,455,1772,1799]" pageId="1" pageNumber="123">Measurements (mm).</emphasis>
|
||
<typeStatus id="54C488607C0A4135FE16F0F4FDEBD11E" box="[464,572,1773,1799]" pageId="1" pageNumber="123" type="holotype">Holotype</typeStatus>
|
||
. Trunk length, 2.03; width, 0.94; proboscis, 0.66; abdomen, 0.48. Third leg; coxa 1, 0.20; coxa 2, 0.26; coxa 3, 0.22; femur, 0.77; tibia 1, 0.45; tibia 2, 0.24; tarsus, 0.10; propodus, 0.40; main claw, 0.21.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565497C0A4135FF01F141FC60D16A" box="[199,951,1880,1907]" pageId="1" pageNumber="123" type="distribution">
|
||
<paragraph id="8BC036C27C0A4135FF01F141FC60D16A" blockId="1.[151,1437,692,2015]" box="[199,951,1880,1907]" pageId="1" pageNumber="123">
|
||
<emphasis id="B90BEAD07C0A4135FF01F141FEB7D16A" bold="true" box="[199,352,1880,1907]" pageId="1" pageNumber="123">Distribution.</emphasis>
|
||
At present only recorded from Green Island,
|
||
<collectingCountry id="F36876527C0A4135FC9BF140FC64D16A" box="[861,947,1881,1907]" name="Taiwan" pageId="1" pageNumber="123">Taiwan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565497C0A4135FF01F164FE13D1A2" pageId="1" pageNumber="123" type="etymology">
|
||
<paragraph id="8BC036C27C0A4135FF01F164FE13D1A2" blockId="1.[151,1437,692,2015]" pageId="1" pageNumber="123">
|
||
<emphasis id="B90BEAD07C0A4135FF01F164FE99D18E" bold="true" box="[199,334,1917,1943]" pageId="1" pageNumber="123">Etymology.</emphasis>
|
||
The specific name is from the Latin
|
||
<taxonomicName id="4C7F4D417C0A4135FD31F164FCADD18E" authorityName="Lee & Kim" authorityYear="2020" box="[759,890,1917,1943]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="1" pageNumber="123" phylum="Arthropoda" rank="species" species="granulatum">
|
||
<emphasis id="B90BEAD07C0A4135FD31F164FCADD18E" box="[759,890,1917,1943]" italics="true" pageId="1" pageNumber="123">granulatum</emphasis>
|
||
</taxonomicName>
|
||
(granulated), referring to the granulated integument of the trunk and legs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36565497C0A4132FF01F1DCFAF4D6CE" lastPageId="6" lastPageNumber="128" pageId="1" pageNumber="123" type="discussion">
|
||
<paragraph id="8BC036C27C0A4136FF01F1DCFD02D75A" blockId="1.[151,1437,692,2015]" lastBlockId="2.[151,1437,153,323]" lastPageId="2" lastPageNumber="124" pageId="1" pageNumber="123">
|
||
<emphasis id="B90BEAD07C0A4135FF01F1DCFEECD1C6" bold="true" box="[199,315,1989,2015]" pageId="1" pageNumber="123">Remarks.</emphasis>
|
||
The legs of the present species were examined in detail. The femur and tibia of the first leg were longer than those of the other legs, but the ratio did not differ significantly (
|
||
<tableCitation id="C6FD03797C094136FC2CF680FBE9D6AA" box="[1002,1086,153,179]" captionStart="TABLE 1" captionStartId="5.[151,239,1244,1268]" captionTargetPageId="5" captionText="TABLE 1. Measurements of length and ratio of femur and tibiae and number of sole spines on propodi of Pycnogonum (Nulloviger) granulatum sp. nov." httpUri="http://table.plazi.org/id/DF00664A7C0E4131FF51F2C5FE2FD30E" pageId="2" pageNumber="124" tableUuid="DF00664A7C0E4131FF51F2C5FE2FD30E">Table 1</tableCitation>
|
||
). Comparing only the absolute length seems insufficient, taking the ratio of length of leg segments seems more appropriate. Since the number of sole spines in the propodus vary depending on the order of the legs, species diagnosis based on the exact number seems inappropriate (
|
||
<tableCitation id="C6FD03797C094136FE41F71CFE0CD706" box="[391,475,261,287]" captionStart="TABLE 1" captionStartId="5.[151,239,1244,1268]" captionTargetPageId="5" captionText="TABLE 1. Measurements of length and ratio of femur and tibiae and number of sole spines on propodi of Pycnogonum (Nulloviger) granulatum sp. nov." httpUri="http://table.plazi.org/id/DF00664A7C0E4131FF51F2C5FE2FD30E" pageId="2" pageNumber="124" tableUuid="DF00664A7C0E4131FF51F2C5FE2FD30E">Table 1</tableCitation>
|
||
). The mid-dorsal projection of all legs has two tubercles, the posterior tubercle of the first leg being the largest and most distinct (
|
||
<figureCitation id="13442A477C094136FD45F730FD1FD75A" box="[643,712,297,323]" captionStart="FIGURE 4" captionStartId="5.[151,250,1135,1160]" captionTargetBox="[177,1411,181,1110]" captionTargetId="figure@5.[177,1411,181,1110]" captionTargetPageId="5" captionText="FIGURE 4. Legs of Pycnogonum (Nulloviger) granulatum sp. nov. A, first leg; B, second leg; C, third leg; D, fourth leg. Scale bars: 0.5 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.3702913" httpUri="https://zenodo.org/record/3702913/files/figure.png" pageId="2" pageNumber="124">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF00664A7C094136FF51F1A4FC49D1E3" ID-DOI="http://doi.org/10.5281/zenodo.3702905" ID-Zenodo-Dep="3702905" httpUri="https://zenodo.org/record/3702905/files/figure.png" pageId="2" pageNumber="124" startId="2.[151,250,1981,2006]" targetBox="[165,1423,342,1952]" targetPageId="2">
|
||
<paragraph id="8BC036C27C094136FF51F1A4FC49D1E3" blockId="2.[151,1436,1981,2042]" pageId="2" pageNumber="124">
|
||
<emphasis id="B90BEAD07C094136FF51F1A4FEC2D1CF" bold="true" box="[151,277,1981,2006]" pageId="2" pageNumber="124">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C7F4D417C094136FEDBF1A6FD73D1CF" authority="Lee & Kim, 2020" authorityName="Lee & Kim" authorityYear="2020" box="[285,676,1982,2006]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="2" pageNumber="124" phylum="Arthropoda" rank="species" species="granulatum" status="sp. nov." subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C094136FEDBF1A6FE76D1CF" box="[285,417,1983,2006]" italics="true" pageId="2" pageNumber="124">Pycnogonum</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C094136FE77F1A7FDCAD1CF" box="[433,541,1982,2006]" italics="true" pageId="2" pageNumber="124">Nulloviger</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C094136FDE8F1A7FD73D1CF" box="[558,676,1982,2006]" italics="true" pageId="2" pageNumber="124">granulatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857AB7C094136FD6BF1A7FD28D1CC" box="[685,767,1982,2005]" pageId="2" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, male. A, trunk, dorsal view; B, trunk, lateral view; C, trunk, anterodorsal view; D, third leg. Scale bars: 1 mm (A–C); 0.5 mm (D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00664A7C084137FF51F061FBBDD0C0" ID-DOI="http://doi.org/10.5281/zenodo.3702907" ID-Zenodo-Dep="3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="3" pageNumber="125" startId="3.[151,250,1656,1681]" targetBox="[151,1436,181,1631]" targetPageId="3">
|
||
<paragraph id="8BC036C27C084137FF51F061FBBDD0C0" blockId="3.[151,1436,1656,1753]" pageId="3" pageNumber="125">
|
||
<emphasis id="B90BEAD07C084137FF51F061FEC4D088" bold="true" box="[151,275,1656,1681]" pageId="3" pageNumber="125">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7F4D417C084137FEDFF063FD4AD088" authority="Lee & Kim, 2020" authorityName="Lee & Kim" authorityYear="2020" box="[281,669,1657,1681]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="granulatum" status="sp. nov." subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C084137FEDFF063FE4AD088" box="[281,413,1658,1681]" italics="true" pageId="3" pageNumber="125">Pycnogonum</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C084137FE6AF060FDCFD088" box="[428,536,1657,1681]" italics="true" pageId="3" pageNumber="125">Nulloviger</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C084137FDE1F060FD4AD088" box="[551,669,1657,1681]" italics="true" pageId="3" pageNumber="125">granulatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857AB7C084137FD65F060FD23D089" box="[675,756,1657,1680]" pageId="3" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, male. A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036C27C084137FF01F111FC44D15F" blockId="3.[151,1437,1799,2042]" pageId="3" pageNumber="125">
|
||
In the absence of ovigers and the presence of gonopores on the ventral surface of coxa 2 of the fourth pair of legs, this species belongs to the subgenus
|
||
<taxonomicName id="4C7F4D417C084137FDABF132FC59D15F" authority="(Stock 1968 b)" baseAuthorityName="Stock" baseAuthorityYear="1968" box="[621,910,1835,1862]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="subGenus" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C084137FDABF132FD32D15C" box="[621,741,1835,1861]" italics="true" pageId="3" pageNumber="125">Nulloviger</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEE4B337C084137FD35F132FC51D15F" author="Stock, J. H." box="[755,902,1835,1862]" pageId="3" pageNumber="125" pagination="7 - 65" refId="ref5672" refString="Stock, J. H. (1968 b) Pycnogonida collected by the Galathea and Anton Bruun in the Indian and Pacific Oceans. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjφbenhavn, 131, 7 - 65." type="journal article" year="1968">Stock 1968b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C084137FF01F156FCCFD1A8" blockId="3.[151,1437,1799,2042]" pageId="3" pageNumber="125">
|
||
The present species is morphologically similar to
|
||
<taxonomicName id="4C7F4D417C084137FD35F149FB88D170" authority="Stock, 1992" authorityName="Stock" authorityYear="1992" box="[755,1119,1871,1898]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="moolenbeeki" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C084137FD35F149FCDED170" box="[755,777,1872,1897]" italics="true" pageId="3" pageNumber="125">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C084137FCDEF149FCE5D170" box="[792,818,1872,1897]" italics="true" pageId="3" pageNumber="125">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C084137FC84F156FC07D170" box="[834,976,1871,1897]" italics="true" pageId="3" pageNumber="125">moolenbeeki</emphasis>
|
||
<bibRefCitation id="EFEE4B337C084137FC10F156FB88D170" author="Stock, J. H." box="[982,1119,1871,1897]" pageId="3" pageNumber="125" pagination="81 - 98" refId="ref5742" refString="Stock, J. H. (1992) Littoral Pycnogonida from Oman. Bijdragen tot de Dierkunde, 62, 81 - 98. https: // doi. org / 10.1163 / 26660644 - 06202003" type="journal article" year="1992">Stock, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D417C084137FBACF149FA40D170" authority="Stock, 1991" authorityName="Stock" authorityYear="1991" box="[1130,1431,1871,1898]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="lobipes" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C084137FBACF149FBA8D170" box="[1130,1151,1872,1897]" italics="true" pageId="3" pageNumber="125">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C084137FB48F149FB7FD170" box="[1166,1192,1872,1897]" italics="true" pageId="3" pageNumber="125">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C084137FB7EF156FADED170" box="[1208,1289,1871,1897]" italics="true" pageId="3" pageNumber="125">lobipes</emphasis>
|
||
<bibRefCitation id="EFEE4B337C084137FAD6F156FA40D170" author="Stock, J. H." box="[1296,1431,1871,1897]" pageId="3" pageNumber="125" pagination="125 - 212" refId="ref5712" refString="Stock, J. H. (1991) Deep-water Pycnogonida from the surroundings of New Caledonia. Resultats des Campagnes MUSOR- STOM, 8, 125 - 212." type="journal article" year="1991">Stock, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D417C084137FF51F16DFDD0D194" authority="Clark, 1963" authorityName="Clark" authorityYear="1963" box="[151,519,1907,1934]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="tuberculatum" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C084137FF51F16DFF7AD194" box="[151,173,1908,1933]" italics="true" pageId="3" pageNumber="125">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C084137FF7AF16DFF01D194" box="[188,214,1908,1933]" italics="true" pageId="3" pageNumber="125">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C084137FF23F16AFEAED194" box="[229,377,1907,1933]" italics="true" pageId="3" pageNumber="125">tuberculatum</emphasis>
|
||
<bibRefCitation id="EFEE4B337C084137FE46F16AFDD0D194" author="Clark, W. C." box="[384,519,1907,1933]" pageId="3" pageNumber="125" pagination="1 - 82" refId="ref5068" refString="Clark, W. C. (1963) Australian Pycnogonida. Records of the Australian Museum, 26, 1 - 82. https: // doi. org / 10.3853 / j. 0067 - 1975.26.1963.669" type="journal article" year="1963">Clark, 1963</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D417C084137FDD4F16DFCE9D194" authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" box="[530,830,1907,1933]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="nodulosum">
|
||
<emphasis id="B90BEAD07C084137FDD4F16DFD7FD194" box="[530,680,1907,1933]" italics="true" pageId="3" pageNumber="125">P. nodulosum</emphasis>
|
||
<bibRefCitation id="EFEE4B337C084137FD69F16AFCE9D194" author="Dohrn, A." box="[687,830,1907,1933]" pageId="3" pageNumber="125" pagination="1 - 252" refId="ref5143" refString="Dohrn, A. (1881) Die Pantopoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Monographie der Fauna und Flora des Golfes von Neapel, 3, 1 - 252. https: // doi. org / 10.5962 / bhl. title. 16340" type="journal article" year="1881">Döhrn, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7F4D417C084137FCBFF16DFB3FD194" authority="Takahashi et al., 2007" authorityName="Takahashi" authorityYear="2007" box="[889,1256,1907,1934]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="spatium">
|
||
<emphasis id="B90BEAD07C084137FCBFF16DFC3AD194" box="[889,1005,1908,1933]" italics="true" pageId="3" pageNumber="125">P. spatium</emphasis>
|
||
<bibRefCitation id="EFEE4B337C084137FC35F16AFB3FD194" author="Takahashi, Y. & Dick, M. H. & Mawatari, S. F." box="[1011,1256,1907,1934]" pageId="3" pageNumber="125" pagination="61 - 79" refId="ref5809" refString="Takahashi, Y., Dick, M. H. & Mawatari, S. F. (2007) Sea spiders (Arthropoda: Pycnogonida) from waters adjacent to the Nansei Islands of Japan. Journal of Natural History, 41, 61 - 79. https: // doi. org / 10.1080 / 00222930601121783" type="journal article" year="2007">
|
||
Takahashi
|
||
<emphasis id="B90BEAD07C084137FBADF16DFB4BD194" box="[1131,1180,1907,1933]" italics="true" pageId="3" pageNumber="125">et al</emphasis>
|
||
., 2007
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
by the presence of a cylindrical proboscis and distinct features of femora.
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C084130FF01F1A2FE28D75D" blockId="3.[151,1437,1799,2042]" lastBlockId="4.[151,1437,152,324]" lastPageId="4" lastPageNumber="126" pageId="3" pageNumber="125">
|
||
Compared with the descriptions of
|
||
<taxonomicName id="4C7F4D417C084137FDA7F1A5FC93D1CC" authorityName="Stock" authorityYear="1992" box="[609,836,1979,2006]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="moolenbeeki" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C084137FDA7F1A5FDA0D1CC" box="[609,631,1980,2005]" italics="true" pageId="3" pageNumber="125">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C084137FD4FF1A5FD73D1CC" box="[649,676,1980,2005]" italics="true" pageId="3" pageNumber="125">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C084137FD70F1A2FC93D1CC" box="[694,836,1979,2005]" italics="true" pageId="3" pageNumber="125">moolenbeeki</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFEE4B337C084137FC90F1A2FC0BD1CC" author="Stock, J. H." box="[854,988,1979,2005]" pageId="3" pageNumber="125" pagination="81 - 98" refId="ref5742" refString="Stock, J. H. (1992) Littoral Pycnogonida from Oman. Bijdragen tot de Dierkunde, 62, 81 - 98. https: // doi. org / 10.1163 / 26660644 - 06202003" type="journal article" year="1992">Stock 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B337C084137FC2DF1A2FB55D1CC" author="Staples, D. A." box="[1003,1154,1979,2005]" pageId="3" pageNumber="125" pagination="541 - 553" refId="ref5391" refString="Staples, D. A. (2002) Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria, 59, 541 - 553. https: // doi. org / 10.24199 / j. mmv. 2002.59.12" type="journal article" year="2002">Staples 2002</bibRefCitation>
|
||
), the following different characteristics are found in the new species. (1) Distinct transverse ridges on the dorsal surface of trunk (incon- spicuous in
|
||
<taxonomicName id="4C7F4D417C0F4130FEDBF683FE2FD6AA" authorityName="Stock" authorityYear="1992" box="[285,504,153,179]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="moolenbeeki" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0F4130FEDBF683FEE5D6AA" box="[285,306,154,179]" italics="true" pageId="4" pageNumber="126">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0F4130FE87F683FE8BD6AA" box="[321,348,154,179]" italics="true" pageId="4" pageNumber="126">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0F4130FEADF680FE2FD6AA" box="[363,504,153,179]" italics="true" pageId="4" pageNumber="126">moolenbeeki</emphasis>
|
||
</taxonomicName>
|
||
). (2) 33% in length ratio of the proboscis to the trunk (compared to 42 to 45%). (3) Two tubercles on mid-dorsal projection of femur arranged in proximodistal axis (
|
||
<figureCitation id="13442A477C0F4130FBFFF6A4FB45D6CE" box="[1081,1170,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1656,1681]" captionTargetBox="[151,1436,181,1631]" captionTargetId="figure@3.[151,1436,181,1631]" captionTargetPageId="3" captionText="FIGURE 2. Pycnogonum (Nulloviger) granulatum sp. nov., holotype, male.A, trunk, dorsal view; B, trunk, anterodorsal view; C, third leg, long spine on femur and tibia 1 broken, lateral view; D, femur of fourth leg, dorsal view; E, spine on tarsus; F, fourth leg, arrow indicating gonopore. Scale bars: 1 mm (A); 0.5 mm (B–C); 0.01 mm (E); 0.1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.3702907" httpUri="https://zenodo.org/record/3702907/files/figure.png" pageId="4" pageNumber="126">Fig. 2D</figureCitation>
|
||
) (anteroposterior axis). (4) Long spine on dorsal surface of femur without dorsal tubercle (spine on dorsal tubercle). (5) Moderately curved propodus (compared to less curved and swollen). (6) Cleft tips of the spines on the ventral surface of tibia, tarsus, and propodus (molariform tips).
|
||
</paragraph>
|
||
<caption id="DF00664A7C0F4130FF51F308FE21D38F" ID-DOI="http://doi.org/10.5281/zenodo.3702911" ID-Zenodo-Dep="3702911" httpUri="https://zenodo.org/record/3702911/files/figure.png" pageId="4" pageNumber="126" startId="4.[151,250,1297,1322]" targetBox="[177,1411,345,1272]" targetPageId="4">
|
||
<paragraph id="8BC036C27C0F4130FF51F308FE21D38F" blockId="4.[151,1436,1297,1430]" pageId="4" pageNumber="126">
|
||
<emphasis id="B90BEAD07C0F4130FF51F308FEC2D333" bold="true" box="[151,277,1297,1322]" pageId="4" pageNumber="126">FIGURE 3.</emphasis>
|
||
Integument.
|
||
<emphasis id="B90BEAD07C0F4130FE58F30BFCEFD330" box="[414,824,1298,1322]" italics="true" pageId="4" pageNumber="126">
|
||
<taxonomicName id="4C7F4D417C0F4130FE58F30BFCF1D333" authorityName="Lee & Kim" authorityYear="2020" box="[414,806,1298,1322]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="granulatum" subGenus="Nulloviger">Pycnogonum (Nulloviger) granulatum</taxonomicName>
|
||
s
|
||
</emphasis>
|
||
p. nov.; A–B. A, granulated integument of abdomen, dorsal view; B, granulated integument of femur, distal surface.
|
||
<emphasis id="B90BEAD07C0F4130FD27F32FFC83D354" box="[737,852,1334,1357]" italics="true" pageId="4" pageNumber="126">
|
||
<taxonomicName id="4C7F4D417C0F4130FD27F32FFC9AD354" box="[737,845,1334,1357]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="spatium">P. spatium</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
C–D. C, pitted integument of abdomen, dorsal view; D, granulated and fitted integument of femur, distal surface. All images are observed under the light microscope (Olympus BX51, Japan). Scale bars: 0.1 mm (A, C).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036C27C0F4130FF01F3DDFDBDD076" blockId="4.[151,1437,1476,2043]" pageId="4" pageNumber="126">
|
||
Compared with the description figures (
|
||
<bibRefCitation id="EFEE4B337C0F4130FD55F3DDFCCED3C7" author="Stock, J. H." box="[659,793,1476,1502]" pageId="4" pageNumber="126" pagination="125 - 212" refId="ref5712" refString="Stock, J. H. (1991) Deep-water Pycnogonida from the surroundings of New Caledonia. Resultats des Campagnes MUSOR- STOM, 8, 125 - 212." type="journal article" year="1991">Stock 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B337C0F4130FCEEF3DDFC69D3C7" author="Staples, D. A." box="[808,958,1476,1502]" pageId="4" pageNumber="126" pagination="541 - 553" refId="ref5391" refString="Staples, D. A. (2002) Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria, 59, 541 - 553. https: // doi. org / 10.24199 / j. mmv. 2002.59.12" type="journal article" year="2002">Staples 2002</bibRefCitation>
|
||
), the following features are the main differences between the present species and
|
||
<taxonomicName id="4C7F4D417C0F4130FDB3F3F0FCCBD01B" authorityName="Stock" authorityYear="1991" box="[629,796,1512,1539]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="lobipes" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0F4130FDB3F3F0FD5DD01B" box="[629,650,1513,1538]" italics="true" pageId="4" pageNumber="126">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0F4130FD5AF3F0FD61D01B" box="[668,694,1513,1538]" italics="true" pageId="4" pageNumber="126">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0F4130FD0EF3F1FCCBD01B" box="[712,796,1512,1538]" italics="true" pageId="4" pageNumber="126">lobipes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90BEAD07C0F4130FCDAF3F1FCF7D01B" box="[796,800,1512,1538]" italics="true" pageId="4" pageNumber="126">.</emphasis>
|
||
(1) Trunk segment 3 and 4 fused (articulated in
|
||
<taxonomicName id="4C7F4D417C0F4130FA94F3F0FF3FD03F" authorityName="Stock" authorityYear="1991" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="lobipes" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0F4130FA94F3F0FABFD01B" box="[1362,1384,1513,1538]" italics="true" pageId="4" pageNumber="126">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0F4130FABFF3F0FA43D01B" box="[1401,1428,1513,1538]" italics="true" pageId="4" pageNumber="126">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0F4130FF51F015FF3FD03F" box="[151,232,1548,1574]" italics="true" pageId="4" pageNumber="126">lobipes</emphasis>
|
||
</taxonomicName>
|
||
). (2) Tip of ocular tubercle round (acuminate). (3) Dorsal tubercle present on the proboscis (absent). (4) Abdomen distally round in dorsal view (truncated). (5) Abdomen articulated at base (non articulated). (6) 8 to 16 sole spines on the propodus (four spines).
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C0F4130FF01F060FCEBD15E" blockId="4.[151,1437,1476,2043]" pageId="4" pageNumber="126">
|
||
<taxonomicName id="4C7F4D417C0F4130FF01F060FD5ED08B" authorityName="Clark" authorityYear="1963" box="[199,649,1656,1683]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="tuberculatum" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0F4130FF01F060FE8ED08B" box="[199,345,1657,1682]" italics="true" pageId="4" pageNumber="126">Pycnogonum</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0F4130FEADF061FE34D08B" box="[363,483,1656,1682]" italics="true" pageId="4" pageNumber="126">Nulloviger</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0F4130FE33F061FD5ED08B" box="[501,649,1656,1682]" italics="true" pageId="4" pageNumber="126">tuberculatum</emphasis>
|
||
</taxonomicName>
|
||
was well described by
|
||
<bibRefCitation id="EFEE4B337C0F4130FC5EF061FBFDD08A" author="Clark, W. C." box="[920,1066,1656,1683]" pageId="4" pageNumber="126" pagination="1 - 82" refId="ref5068" refString="Clark, W. C. (1963) Australian Pycnogonida. Records of the Australian Museum, 26, 1 - 82. https: // doi. org / 10.3853 / j. 0067 - 1975.26.1963.669" type="journal article" year="1963">Clark (1963)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEE4B337C0F4130FBA2F061FADCD08B" author="Staples, D. A." box="[1124,1291,1656,1682]" pageId="4" pageNumber="126" pagination="541 - 553" refId="ref5391" refString="Staples, D. A. (2002) Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria, 59, 541 - 553. https: // doi. org / 10.24199 / j. mmv. 2002.59.12" type="journal article" year="2002">Staples (2002)</bibRefCitation>
|
||
. The present species is distinguished from the
|
||
<taxonomicName id="4C7F4D417C0F4130FDCDF084FD39D0AF" authorityName="Clark" authorityYear="1963" box="[523,750,1692,1719]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="tuberculatum" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0F4130FDCDF084FDF7D0AF" box="[523,544,1693,1718]" italics="true" pageId="4" pageNumber="126">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0F4130FDF6F084FD9DD0AF" box="[560,586,1693,1718]" italics="true" pageId="4" pageNumber="126">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0F4130FD9CF085FD39D0AF" box="[602,750,1692,1718]" italics="true" pageId="4" pageNumber="126">tuberculatum</emphasis>
|
||
</taxonomicName>
|
||
by following characteristics. (1) Trunk length
|
||
<quantity id="4C879B277C0F4130FB3EF085FA8CD0AE" box="[1272,1371,1692,1719]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="4" pageNumber="126" unit="mm" value="2.03">2.03 mm</quantity>
|
||
(
|
||
<quantity id="4C879B277C0F4130FAACF082FEC0D0C2" metricMagnitude="-3" metricUnit="m" metricValue="5.115" metricValueMax="5.65" metricValueMin="4.58" pageId="4" pageNumber="126" unit="mm" value="5.115" valueMax="5.65" valueMin="4.58">4.58 to 5.65 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C7F4D417C0F4130FEFFF0D8FDCDD0C3" authorityName="Clark" authorityYear="1963" box="[313,538,1728,1755]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="tuberculatum" subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0F4130FEFFF0D8FE99D0C3" box="[313,334,1729,1754]" italics="true" pageId="4" pageNumber="126">P.</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0F4130FE9BF0D8FEA0D0C3" box="[349,375,1729,1754]" italics="true" pageId="4" pageNumber="126">N.</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0F4130FE40F0D9FDCDD0C3" box="[390,538,1728,1754]" italics="true" pageId="4" pageNumber="126">tuberculatum</emphasis>
|
||
</taxonomicName>
|
||
). (2) Lateral processes separated by 1/4 to 1/3 of base width (slightly separated). (3) 33% in length ratio of the proboscis to the trunk (compared to 25 to 27%). (4) Dorsal tubercle on the proboscis prominent (small). (5) Abdomen reaching to the distal end of coxa 2 (half of coxa 3). (6) Cleft tips of the spines on the ventral surface of tibia, tarsus, and propodus (blunt tips).
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C0F4130FF01F149FB76D1E3" blockId="4.[151,1437,1476,2043]" pageId="4" pageNumber="126">
|
||
Through the descriptions and figures (
|
||
<bibRefCitation id="EFEE4B337C0F4130FD47F149FCD8D173" author="Dohrn, A." box="[641,783,1872,1898]" pageId="4" pageNumber="126" pagination="1 - 252" refId="ref5143" refString="Dohrn, A. (1881) Die Pantopoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Monographie der Fauna und Flora des Golfes von Neapel, 3, 1 - 252. https: // doi. org / 10.5962 / bhl. title. 16340" type="journal article" year="1881">Döhrn 1881</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B337C0F4130FCD8F149FBE3D173" author="Kocak, C. & Katagan, T." box="[798,1076,1872,1899]" pageId="4" pageNumber="126" pagination="110 - 112" refId="ref5238" refString="Kocak, C. & Katagan, T. (2007) A pycnogonid species new for the Turkish fauna: Pycnogonum nodulosum Dohrn, 1881 (Arthropoda: Pycnogonidae). Zoology in the Middle East, 42, 110 - 112. https: // doi. org / 10.1080 / 09397140.2007.10638258" type="journal article" year="2007">Kocak & Katagan 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B337C0F4130FB84F149FAE1D173" author="Lehmann, T. & Hess, M. & Melzer, R." box="[1090,1334,1872,1899]" pageId="4" pageNumber="126" pagination="163 - 224" refId="ref5344" refString="Lehmann, T., Hess, M. & Melzer, R. (2014) Common littoral pycnogonids of the Mediterranean Sea. Zoosystematics and Evolution, 90, 163 - 224. https: // doi. org / 10.3897 / zse. 90.7520" type="journal article" year="2014">Lehmann et al. 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B337C0F4130FA82F149FE69D197" author="Munilla, T. & Soler-Membrives, A." pageId="4" pageNumber="126" pagination="1 - 293" refId="ref5292" refString="Munilla, T. & Soler-Membrives, A. (2014) Pycnogonida. In: Sanchez, M. A. R. (Ed.), Fauna Iberica. Vol. 39. Museo Nacional de Ciencias Naturales. CSIC, Madrid, pp. 1 - 293." type="book chapter" year="2014">Munilla & Soler-Membrives 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B337C0F4130FE0CF16DFD8DD197" author="Stock, J. H." box="[458,602,1908,1934]" pageId="4" pageNumber="126" pagination="1 - 38" refId="ref5636" refString="Stock, J. H. (1968 a) Faune marine des Pyrenees-Orientales. Fascicule 6: Pycnogonides. Vie et Milieu, Supplement 19 (1 - A), 1 - 38." type="journal article" year="1968">Stock 1968a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B337C0F4130FDAEF16DFD3ED197" author="Stock, J. H." box="[616,745,1908,1934]" pageId="4" pageNumber="126" pagination="17 - 77" refId="ref5779" refString="Stock, J. H. (1994) Indo-west pacific Pycnogonida collected by some major oceanographic expeditions. Beaufortia, 44 (3), 17 - 77." type="journal article" year="1994">Stock 1994</bibRefCitation>
|
||
), the present species is different from
|
||
<taxonomicName id="4C7F4D417C0F4130FB64F16CFAEDD197" authorityName="Dohrn" authorityYear="1881" box="[1186,1338,1908,1934]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="nodulosum">
|
||
<emphasis id="B90BEAD07C0F4130FB64F16CFAEDD197" box="[1186,1338,1908,1934]" italics="true" pageId="4" pageNumber="126">P. nodulosum</emphasis>
|
||
</taxonomicName>
|
||
in some characters. (1) Trunk segment 3 and 4 fused (articulated in
|
||
<taxonomicName id="4C7F4D417C0F4130FCEBF180FC14D1AB" authorityName="Dohrn" authorityYear="1881" box="[813,963,1944,1970]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="nodulosum">
|
||
<emphasis id="B90BEAD07C0F4130FCEBF180FC14D1AB" box="[813,963,1944,1970]" italics="true" pageId="4" pageNumber="126">P. nodulosum</emphasis>
|
||
</taxonomicName>
|
||
). (2) Posterior surface of lateral processes smooth instead of enlarged or swollen. (3) Two tubercles on the dorsal surface of femur instead of three. (4) Cleft tips of the spines on the ventral surface of tibia, tarsus, and propodus instead of club shaped.
|
||
</paragraph>
|
||
<caption id="DF00664A7C0E4131FF51F276FEBBD2B5" ID-DOI="http://doi.org/10.5281/zenodo.3702913" ID-Zenodo-Dep="3702913" httpUri="https://zenodo.org/record/3702913/files/figure.png" pageId="5" pageNumber="127" startId="5.[151,250,1135,1160]" targetBox="[177,1411,181,1110]" targetPageId="5">
|
||
<paragraph id="8BC036C27C0E4131FF51F276FEBBD2B5" blockId="5.[151,1436,1135,1196]" pageId="5" pageNumber="127">
|
||
<emphasis id="B90BEAD07C0E4131FF51F276FEC4D291" bold="true" box="[151,275,1135,1160]" pageId="5" pageNumber="127">FIGURE 4.</emphasis>
|
||
Legs of
|
||
<taxonomicName id="4C7F4D417C0E4131FEAAF269FD38D291" authority="Lee & Kim, 2020" authorityName="Lee & Kim" authorityYear="2020" box="[364,751,1136,1160]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="granulatum" status="sp. nov." subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0E4131FEAAF269FE27D29E" box="[364,496,1136,1159]" italics="true" pageId="5" pageNumber="127">Pycnogonum</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0E4131FE38F269FDBDD291" box="[510,618,1136,1160]" italics="true" pageId="5" pageNumber="127">Nulloviger</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0E4131FDBFF269FD38D291" box="[633,751,1136,1160]" italics="true" pageId="5" pageNumber="127">granulatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857AB7C0E4131FD30F269FC93D29E" box="[758,836,1136,1159]" pageId="5" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A, first leg; B, second leg; C, third leg; D, fourth leg. Scale bars: 0.5 mm (A–D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00664A7C0E4131FF51F2C5FE2FD30E" ID-Table-UUID="DF00664A7C0E4131FF51F2C5FE2FD30E" httpUri="http://table.plazi.org/id/DF00664A7C0E4131FF51F2C5FE2FD30E" pageId="5" pageNumber="127" targetBox="[213,1329,1319,1768]" targetIsTable="true" targetPageId="5">
|
||
<paragraph id="8BC036C27C0E4131FF51F2C5FE2FD30E" blockId="5.[151,1436,1244,1304]" pageId="5" pageNumber="127">
|
||
<emphasis id="B90BEAD07C0E4131FF51F2C5FEDCD2ED" bold="true" box="[151,267,1244,1268]" pageId="5" pageNumber="127">TABLE 1.</emphasis>
|
||
Measurements of length and ratio of femur and tibiae and number of sole spines on propodi of
|
||
<taxonomicName id="4C7F4D417C0E4131FAD7F2C4FE49D301" authority="Lee & Kim, 2020" authorityName="Lee & Kim" authorityYear="2020" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="granulatum" status="sp. nov." subGenus="Nulloviger">
|
||
<emphasis id="B90BEAD07C0E4131FAD7F2C4FA4BD2ED" box="[1297,1436,1245,1268]" italics="true" pageId="5" pageNumber="127">Pycnogonum</emphasis>
|
||
(
|
||
<emphasis id="B90BEAD07C0E4131FF66F319FEC5D301" box="[160,274,1280,1304]" italics="true" pageId="5" pageNumber="127">Nulloviger</emphasis>
|
||
)
|
||
<emphasis id="B90BEAD07C0E4131FEE4F319FE49D301" box="[290,414,1280,1304]" italics="true" pageId="5" pageNumber="127">granulatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857AB7C0E4131FE63F319FE2FD30E" box="[421,504,1280,1303]" pageId="5" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036C27C0E4131FF20F343FB11D0DA" pageId="5" pageNumber="127">
|
||
<table id="F97FC4627C0EBECBFF13F33EFAE6D0F1" box="[213,1329,1319,1768]" gridcols="7" gridrows="5" pageId="5" pageNumber="127">
|
||
<tr id="354F34807C0EBECBFF13F33EFAE6D391" box="[213,1329,1319,1416]" gridrow="0" pageId="5" pageNumber="127">
|
||
<th id="769E5DFC7C0EBECBFE4DF33EFE1AD391" box="[395,461,1319,1416]" gridcol="1" gridrow="0" pageId="5" pageNumber="127">Femur (mm)</th>
|
||
<th id="769E5DFC7C0EBECBFE38F33EFDEDD391" box="[510,570,1319,1416]" gridcol="2" gridrow="0" pageId="5" pageNumber="127">Ratio</th>
|
||
<th id="769E5DFC7C0EBECBFDB2F33EFD6BD391" box="[628,700,1319,1416]" gridcol="3" gridrow="0" pageId="5" pageNumber="127">Tibia 1 (mm)</th>
|
||
<th id="769E5DFC7C0EBECBFCC4F33EFCEED391" box="[770,825,1319,1416]" gridcol="4" gridrow="0" pageId="5" pageNumber="127">Ratio</th>
|
||
<th id="769E5DFC7C0EBECBFCA6F33EFC7FD391" box="[864,936,1319,1416]" gridcol="5" gridrow="0" pageId="5" pageNumber="127">Tibia 2 (mm)</th>
|
||
<th id="769E5DFC7C0EBECBFB88F33EFAE6D391" box="[1102,1329,1319,1416]" gridcol="6" gridrow="0" pageId="5" pageNumber="127">Number of sole spines on propodus</th>
|
||
</tr>
|
||
<tr id="354F34807C0EBECBFF13F382FAE6D3F9" box="[213,1329,1435,1504]" gridrow="1" pageId="5" pageNumber="127">
|
||
<th id="769E5DFC7C0EBECBFF13F382FEC7D3F9" box="[213,272,1435,1504]" gridcol="0" gridrow="1" pageId="5" pageNumber="127">Leg 1</th>
|
||
<td id="769E5DFC7C0EBECBFE4DF382FE1AD3F9" box="[395,461,1435,1504]" gridcol="1" gridrow="1" pageId="5" pageNumber="127">0.88</td>
|
||
<td id="769E5DFC7C0EBECBFE38F382FDEDD3F9" box="[510,570,1435,1504]" gridcol="2" gridrow="1" pageId="5" pageNumber="127">57%</td>
|
||
<td id="769E5DFC7C0EBECBFDB2F382FD6BD3F9" box="[628,700,1435,1504]" gridcol="3" gridrow="1" pageId="5" pageNumber="127">0.5</td>
|
||
<td id="769E5DFC7C0EBECBFCC4F382FCEED3F9" box="[770,825,1435,1504]" gridcol="4" gridrow="1" pageId="5" pageNumber="127">53%</td>
|
||
<td id="769E5DFC7C0EBECBFCA6F382FC7FD3F9" box="[864,936,1435,1504]" gridcol="5" gridrow="1" pageId="5" pageNumber="127">0.26</td>
|
||
<td id="769E5DFC7C0EBECBFB88F382FAE6D3F9" box="[1102,1329,1435,1504]" gridcol="6" gridrow="1" pageId="5" pageNumber="127">12</td>
|
||
</tr>
|
||
<tr id="354F34807C0EBECBFF13F3EAFAE6D021" box="[213,1329,1523,1592]" gridrow="2" pageId="5" pageNumber="127">
|
||
<th id="769E5DFC7C0EBECBFF13F3EAFEC7D021" box="[213,272,1523,1592]" gridcol="0" gridrow="2" pageId="5" pageNumber="127">Leg 2</th>
|
||
<td id="769E5DFC7C0EBECBFE4DF3EAFE1AD021" box="[395,461,1523,1592]" gridcol="1" gridrow="2" pageId="5" pageNumber="127">0.81</td>
|
||
<td id="769E5DFC7C0EBECBFE38F3EAFDEDD021" box="[510,570,1523,1592]" gridcol="2" gridrow="2" pageId="5" pageNumber="127">55%</td>
|
||
<td id="769E5DFC7C0EBECBFDB2F3EAFD6BD021" box="[628,700,1523,1592]" gridcol="3" gridrow="2" pageId="5" pageNumber="127">0.45</td>
|
||
<td id="769E5DFC7C0EBECBFCC4F3EAFCEED021" box="[770,825,1523,1592]" gridcol="4" gridrow="2" pageId="5" pageNumber="127">54%</td>
|
||
<td id="769E5DFC7C0EBECBFCA6F3EAFC7FD021" box="[864,936,1523,1592]" gridcol="5" gridrow="2" pageId="5" pageNumber="127">0.24</td>
|
||
<td id="769E5DFC7C0EBECBFB88F3EAFAE6D021" box="[1102,1329,1523,1592]" gridcol="6" gridrow="2" pageId="5" pageNumber="127">16</td>
|
||
</tr>
|
||
<tr id="354F34807C0EBECBFF13F052FAE6D089" box="[213,1329,1611,1680]" gridrow="3" pageId="5" pageNumber="127">
|
||
<th id="769E5DFC7C0EBECBFF13F052FEC7D089" box="[213,272,1611,1680]" gridcol="0" gridrow="3" pageId="5" pageNumber="127">Leg 3</th>
|
||
<td id="769E5DFC7C0EBECBFE4DF052FE1AD089" box="[395,461,1611,1680]" gridcol="1" gridrow="3" pageId="5" pageNumber="127">0.77</td>
|
||
<td id="769E5DFC7C0EBECBFE38F052FDEDD089" box="[510,570,1611,1680]" gridcol="2" gridrow="3" pageId="5" pageNumber="127">59%</td>
|
||
<td id="769E5DFC7C0EBECBFDB2F052FD6BD089" box="[628,700,1611,1680]" gridcol="3" gridrow="3" pageId="5" pageNumber="127">0.45</td>
|
||
<td id="769E5DFC7C0EBECBFCC4F052FCEED089" box="[770,825,1611,1680]" gridcol="4" gridrow="3" pageId="5" pageNumber="127">53%</td>
|
||
<td id="769E5DFC7C0EBECBFCA6F052FC7FD089" box="[864,936,1611,1680]" gridcol="5" gridrow="3" pageId="5" pageNumber="127">0.24</td>
|
||
<td id="769E5DFC7C0EBECBFB88F052FAE6D089" box="[1102,1329,1611,1680]" gridcol="6" gridrow="3" pageId="5" pageNumber="127">8</td>
|
||
</tr>
|
||
<tr id="354F34807C0EBECBFF13F0BAFAE6D0F1" box="[213,1329,1699,1768]" gridrow="4" pageId="5" pageNumber="127">
|
||
<th id="769E5DFC7C0EBECBFF13F0BAFEC7D0F1" box="[213,272,1699,1768]" gridcol="0" gridrow="4" pageId="5" pageNumber="127">Leg 4</th>
|
||
<td id="769E5DFC7C0EBECBFE4DF0BAFE1AD0F1" box="[395,461,1699,1768]" gridcol="1" gridrow="4" pageId="5" pageNumber="127">0.79</td>
|
||
<td id="769E5DFC7C0EBECBFE38F0BAFDEDD0F1" box="[510,570,1699,1768]" gridcol="2" gridrow="4" pageId="5" pageNumber="127">58%</td>
|
||
<td id="769E5DFC7C0EBECBFDB2F0BAFD6BD0F1" box="[628,700,1699,1768]" gridcol="3" gridrow="4" pageId="5" pageNumber="127">0.46</td>
|
||
<td id="769E5DFC7C0EBECBFCC4F0BAFCEED0F1" box="[770,825,1699,1768]" gridcol="4" gridrow="4" pageId="5" pageNumber="127">50%</td>
|
||
<td id="769E5DFC7C0EBECBFCA6F0BAFC7FD0F1" box="[864,936,1699,1768]" gridcol="5" gridrow="4" pageId="5" pageNumber="127">0.23</td>
|
||
<td id="769E5DFC7C0EBECBFB88F0BAFAE6D0F1" box="[1102,1329,1699,1768]" gridcol="6" gridrow="4" pageId="5" pageNumber="127">8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C0E4131FF01F100FA44D1FE" blockId="5.[151,1436,1817,2023]" pageId="5" pageNumber="127">
|
||
Among the congeners,
|
||
<taxonomicName id="4C7F4D417C0E4131FE0BF103FD94D12A" box="[461,579,1818,1843]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="spatium">
|
||
<emphasis id="B90BEAD07C0E4131FE0BF103FD94D12A" box="[461,579,1818,1843]" italics="true" pageId="5" pageNumber="127">P. spatium</emphasis>
|
||
</taxonomicName>
|
||
is geographically closest to the present species, but can be distinguished by the following: (1) Trunk integument granulated (pitted in
|
||
<taxonomicName id="4C7F4D417C0E4131FCE1F127FC4AD14E" box="[807,925,1854,1879]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="spatium">
|
||
<emphasis id="B90BEAD07C0E4131FCE1F127FC4AD14E" box="[807,925,1854,1879]" italics="true" pageId="5" pageNumber="127">P. spatium</emphasis>
|
||
</taxonomicName>
|
||
). (2) Trunk segment 4 expanding posteriorly without parallel part (expanding posteriorly after the parallel part in
|
||
<taxonomicName id="4C7F4D417C0E4131FC52F17BFBDFD162" box="[916,1032,1890,1915]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="spatium">
|
||
<emphasis id="B90BEAD07C0E4131FC52F17BFBDFD162" box="[916,1032,1890,1915]" italics="true" pageId="5" pageNumber="127">P. spatium</emphasis>
|
||
</taxonomicName>
|
||
). (3) Lateral processes separated by 1/4 to 1/3 of base width, wider in
|
||
<taxonomicName id="4C7F4D417C0E4131FDD1F19FFD5AD186" box="[535,653,1926,1951]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="spatium">
|
||
<emphasis id="B90BEAD07C0E4131FDD1F19FFD5AD186" box="[535,653,1926,1951]" italics="true" pageId="5" pageNumber="127">P. spatium</emphasis>
|
||
</taxonomicName>
|
||
. (4) Femur with larger ventroproximal swelling. (5) Scarce spines on the ventral surface of the femur compared to several spines in
|
||
<taxonomicName id="4C7F4D417C0E4131FC83F1B3FC6ED1DA" box="[837,953,1962,1987]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="spatium">
|
||
<emphasis id="B90BEAD07C0E4131FC83F1B3FC6ED1DA" box="[837,953,1962,1987]" italics="true" pageId="5" pageNumber="127">P. spatium</emphasis>
|
||
</taxonomicName>
|
||
. (6) Tibia 2 1.7 times as long as base width, not 2.4 times. (7) Cleft tips of the spines on the ventral surface of tibia, tarsus, and propodus instead of round tips.
|
||
</paragraph>
|
||
<paragraph id="8BC036C27C0D4132FF01F680FAF4D6CE" blockId="6.[151,1436,153,215]" pageId="6" pageNumber="128">Experiments were carried out to attempt to add DNA data to the species determination but non-degraded DNA could not be obtained. With only one specimen collected it was difficult to conduct further experiments.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |