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<mods:title id="36604DB25E46443B6A1D808FBABF709D">On the taxonomic status of the water mite Hydryphantes hellichi Thon, 1899 (Acari, Hydrachnidia, Hydryphantidae)</mods:title>
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<mods:namePart id="AF74A58B89B4202486DFC3452522D71B">Tuzovskij, Petr V.</mods:namePart>
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<mods:title id="44ADD89E5A8320B0C8AC6C5B1DF8F922">Ecologica Montenegrina</mods:title>
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<mods:date id="B60930B0A2D353DE2D1CACC1AE282683">2014</mods:date>
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<treatment id="03D687C0FFAEFFB7FF1AFC84FD7CF81E" ID-DOI="http://doi.org/10.5281/zenodo.8031923" ID-Zenodo-Dep="8031923" LSID="urn:lsid:plazi:treatment:03D687C0FFAEFFB7FF1AFC84FD7CF81E" httpUri="http://treatment.plazi.org/id/03D687C0FFAEFFB7FF1AFC84FD7CF81E" lastPageId="7" lastPageNumber="241" pageId="1" pageNumber="235">
<subSubSection id="C365655DFFAEFFB1FF1AFC84FDF0FC95" box="[151,562,770,796]" pageId="1" pageNumber="235" type="nomenclature">
<paragraph id="8BC036D6FFAEFFB1FF1AFC84FDF0FC95" blockId="1.[151,562,770,830]" box="[151,562,770,796]" pageId="1" pageNumber="235">
<heading id="D08881BAFFAEFFB1FF1AFC84FDF0FC95" box="[151,562,770,796]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="235" reason="2">
<taxonomicName id="4C7F4D55FFAEFFB1FF1AFC84FDF0FC95" authority="Thon, 1899" authorityName="Thon" authorityYear="1899" box="[151,562,770,796]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="235" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFAEFFB1FF1AFC84FE61FC95" bold="true" box="[151,419,770,796]" italics="true" pageId="1" pageNumber="235">Hydryphantes hellichi</emphasis>
<bibRefCitation id="EFEE4B27FFAEFFB1FE26FC84FDF0FC95" author="Thon, K." box="[427,562,770,796]" pageId="1" pageNumber="235" pagination="55 - 67" refId="ref4413" refString="Thon, K. (1899) Monographie der bohmischen Hydryphantes-Arten. Bulletin International Academie des Sciences, Prague, 8, 55 - 67." type="journal article" year="1899">Thon, 1899</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365655DFFAEFFB1FF1AFCA2FEDDFCB7" box="[151,287,804,830]" pageId="1" pageNumber="235" type="description">
<paragraph id="8BC036D6FFAEFFB1FF1AFCA2FEDDFCB7" blockId="1.[151,562,770,830]" box="[151,287,804,830]" pageId="1" pageNumber="235">
(
<figureCitation id="13442A53FFAEFFB1FF12FCA2FED4FCB7" box="[159,278,804,830]" captionStart-0="Figures 12" captionStart-1="Figures 39" captionStart-2="Figures 1014" captionStart-3="Figures 1516" captionStart-4="Figures 1724" captionStart-5="Figures 2529" captionStartId-0="1.[151,237,1862,1886]" captionStartId-1="2.[151,237,1561,1585]" captionStartId-2="3.[151,237,1637,1661]" captionStartId-3="4.[151,237,1640,1664]" captionStartId-4="5.[151,237,1960,1984]" captionStartId-5="6.[151,237,1768,1792]" captionTargetBox-0="[167,1428,1038,1854]" captionTargetBox-1="[171,1391,360,1528]" captionTargetBox-2="[176,1423,208,1599]" captionTargetBox-3="[158,1423,808,1624]" captionTargetBox-4="[196,1410,200,1946]" captionTargetBox-5="[154,1421,210,1746]" captionTargetId-0="figure-239@1.[151,1434,999,1856]" captionTargetId-1="figure-70@2.[151,1436,570,1554]" captionTargetId-2="figure-13@3.[151,1436,590,1631]" captionTargetId-3="figure-310@4.[151,1436,808,1634]" captionTargetId-4="figure-13@5.[168,1418,591,1954]" captionTargetId-5="figure-13@6.[151,1436,600,1749]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionText-0="Figures 12. Hydryphantes hellichi Thon, 1899, larva: 1­ dorsal view, 2­ ventral view. Scale bar: 50µm." captionText-1="Figures 39.Hydryphantes hellichi Thon, 1899, larva: 36 excretory pore plate, 7 capitulum, 8 chelicerae, dorsal view, 9 pedipalp. Scale bars: 36, 9 = 50µm, 78 = 100µm." captionText-2="Figures 1014. Hydryphantes hellichi Thon, 1899, larva: 10 ­ ILeg­46; 11 ­ IILeg­46; 12 ­ IIILeg­46; 13 ­ claws of leg I; 14 ­ claws of leg III. Simple setae on IIIILeg­46 are not shown." captionText-3="Figures 1516. Hydryphantes hellichi Thon, 1899, deutonymph: 16 dorsal view, 17 ventral view. Scale bar: 50µm." captionText-4="Figures 1724. Hydryphantes hellichi Thon, 1899, deutonymph: 17 frontal plate; 18 genital field; 19 fragment of integument; 20 capitulum; 21 chelicera; 22 pedipalp; 23 ­ IILeg­45; 24 claw. Scale bars: 17, 23 = 200µm, 18, 20­21 = 100µm, 19, 22, 24 = 50µm." captionText-5="Figures 2529. Hydryphantes hellichi Thon, 1899, adults: 25 ­ frontal plate; 26 coxal plates; 27 genital field; 28 capitulum; 29 pedipalp. Scale bars: 2526, 28 = 200µm, 27, 29 = 100µm." figureDoi-0="http://doi.org/10.5281/zenodo.8031925" figureDoi-1="http://doi.org/10.5281/zenodo.8031927" figureDoi-2="http://doi.org/10.5281/zenodo.8031929" figureDoi-3="http://doi.org/10.5281/zenodo.8031931" figureDoi-4="http://doi.org/10.5281/zenodo.8031933" figureDoi-5="http://doi.org/10.5281/zenodo.8031935" httpUri-0="https://zenodo.org/record/8031925/files/figure.png" httpUri-1="https://zenodo.org/record/8031927/files/figure.png" httpUri-2="https://zenodo.org/record/8031929/files/figure.png" httpUri-3="https://zenodo.org/record/8031931/files/figure.png" httpUri-4="https://zenodo.org/record/8031933/files/figure.png" httpUri-5="https://zenodo.org/record/8031935/files/figure.png" pageId="1" pageNumber="235">Figs 129</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C365655DFFAEFFB1FF1AFCE1FBE1FC6F" pageId="1" pageNumber="235" type="materials_examined">
<paragraph id="8BC036D6FFAEFFB1FF1AFCE1FBE1FC6F" blockId="1.[151,1444,871,998]" pageId="1" pageNumber="235">
<emphasis id="B90BEAC4FFAEFFB1FF1AFCE1FE44FC08" bold="true" box="[151,390,871,897]" pageId="1" pageNumber="235">Material examined</emphasis>
.
<specimenCount id="9D79FD5FFFAEFFB1FE15FCE1FE3AFC08" box="[408,504,871,897]" pageId="1" pageNumber="235" type="male">8 males</specimenCount>
,
<specimenCount id="9D79FD5FFFAEFFB1FD8AFCE1FDB9FC08" box="[519,635,871,897]" pageId="1" pageNumber="235" type="female">7 females</specimenCount>
<materialsCitation id="3B173C8BFFAEFFB1FD08FCE1FEE3FC4C" collectionCode="V" collectorName="Postyltsevo &amp; Tuzovskij. Larvae" country="Russia" location="April-July" municipality="Nekouz District" pageId="1" pageNumber="235" specimenCount="1" stateProvince="Yaroslavl'">
and 4 deutonymphs,
<collectingCountry id="F3687646FFAEFFB1FC0FFCE1FC11FC08" box="[898,979,871,897]" name="Russia" pageId="1" pageNumber="235">Russia</collectingCountry>
,
<collectingRegion id="49BBF834FFAEFFB1FC6CFCE1FB0BFC08" box="[993,1225,871,897]" country="Russia" name="Yaroslavl'" pageId="1" pageNumber="235">Yaroslavl Province</collectingRegion>
,
<collectingMunicipality id="6BA4ACACFFAEFFB1FB55FCE1FA5DFC08" box="[1240,1439,871,897]" pageId="1" pageNumber="235">Nekouz District</collectingMunicipality>
,
<location id="8EA0600DFFAEFFB1FF1AFC0FFED2FC2A" LSID="urn:lsid:plazi:treatment:03D687C0FFAEFFB7FF1AFC84FD7CF81E:8EA0600DFFAEFFB1FF1AFC0FFED2FC2A" box="[151,272,905,931]" country="Russia" municipality="Nekouz District" name="April-July" pageId="1" pageNumber="235" stateProvince="Yaroslavl'">April­July</location>
2000­2002, 2004, sedge bogs near village
<collectorName id="268A5300FFAEFFB1FC9CFC0FFC59FC2A" box="[785,923,905,931]" pageId="1" pageNumber="235">Postyltsevo</collectorName>
, leg. P.
<collectionCode id="ED6EAE13FFAEFFB1FC79FC0FFBCFFC2A" box="[1012,1037,905,931]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="235" type="Museum">V</collectionCode>
.
<collectorName id="268A5300FFAEFFB1FB94FC0FFB32FC2A" box="[1049,1264,905,931]" pageId="1" pageNumber="235">Tuzovskij. Larvae</collectorName>
(n = 60) were reared from
</materialsCitation>
<specimenCount id="9D79FD5FFFAEFFB1FEA5FC2DFE79FC4C" box="[296,443,939,965]" pageId="1" pageNumber="235" type="female">four females</specimenCount>
in laboratory conditions,
<specimenCount id="9D79FD5FFFAEFFB1FD67FC2DFCAEFC4C" box="[746,876,939,965]" pageId="1" pageNumber="235" type="female">one female</specimenCount>
<date id="FFC11016FFAEFFB1FCFEFC2DFBD1FC4C" box="[883,1043,939,965]" pageId="1" pageNumber="235" value="2000-05-28">28 May 2000</date>
,
<specimenCount id="9D79FD5FFFAEFFB1FB93FC2DFB62FC4C" box="[1054,1184,939,965]" pageId="1" pageNumber="235" type="female">one female</specimenCount>
<date id="FFC11016FFAEFFB1FB2AFC2DFAF0FC4C" box="[1191,1330,939,965]" pageId="1" pageNumber="235" value="2001-07-04">4 July 2001</date>
, and
<specimenCount id="9D79FD5FFFAEFFB1FAFDFC2DFF30FC6F" pageId="1" pageNumber="235" type="female">two females</specimenCount>
<date id="FFC11016FFAEFFB1FF77FC4AFE58FC6F" box="[250,410,972,998]" pageId="1" pageNumber="235" value="2002-05-27">27 May 2002</date>
.
<materialsCitation id="3B173C8BFFAEFFB1FE28FC4AFBDDFC6F" box="[421,1055,972,998]" country="Russia" location="The" pageId="1" pageNumber="235" specimenCount="1" stateProvince="Yaroslavl'">
<location id="8EA0600DFFAEFFB1FE28FC4AFE11FC6F" LSID="urn:lsid:plazi:treatment:03D687C0FFAEFFB7FF1AFC84FD7CF81E:8EA0600DFFAEFFB1FE28FC4AFE11FC6F" box="[421,467,972,998]" country="Russia" name="The" pageId="1" pageNumber="235" stateProvince="Yaroslavl'">The</location>
duration of the embryonic period was 12­15 days
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF00665EFFAEFFB1FF1AF8C0FB3AF8D4" ID-DOI="http://doi.org/10.5281/zenodo.8031925" ID-Zenodo-Dep="8031925" box="[151,1272,1862,1886]" httpUri="https://zenodo.org/record/8031925/files/figure.png" pageId="1" pageNumber="235" startId="1.[151,237,1862,1886]" targetBox="[167,1428,1038,1854]" targetPageId="1" targetType="figure">
<paragraph id="8BC036D6FFAEFFB1FF1AF8C0FB3AF8D4" blockId="1.[151,1272,1862,1886]" box="[151,1272,1862,1886]" pageId="1" pageNumber="235">
<emphasis id="B90BEAC4FFAEFFB1FF1AF8C0FEE0F8D7" bold="true" box="[151,290,1862,1886]" pageId="1" pageNumber="235">Figures 12.</emphasis>
<taxonomicName id="4C7F4D55FFAEFFB1FEA4F8C1FD58F8D4" authority="Thon, 1899" authorityName="Thon" authorityYear="1899" box="[297,666,1862,1886]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="235" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFAEFFB1FEA4F8C1FDD7F8D7" box="[297,533,1863,1886]" italics="true" pageId="1" pageNumber="235">Hydryphantes hellichi</emphasis>
<bibRefCitation id="EFEE4B27FFAEFFB1FD91F8C0FD58F8D4" author="Thon, K." box="[540,666,1862,1885]" pageId="1" pageNumber="235" pagination="55 - 67" refId="ref4413" refString="Thon, K. (1899) Monographie der bohmischen Hydryphantes-Arten. Bulletin International Academie des Sciences, Prague, 8, 55 - 67." type="journal article" year="1899">Thon, 1899</bibRefCitation>
</taxonomicName>
, larva: 1­ dorsal view, 2­ ventral view. Scale bar: 50µm.
</paragraph>
</caption>
<subSubSection id="C365655DFFAEFFB2FF1AF801FE73FECD" lastPageId="2" lastPageNumber="236" pageId="1" pageNumber="235" type="diagnosis">
<paragraph id="8BC036D6FFAEFFB2FF1AF801FE73FECD" blockId="1.[151,1444,1927,2055]" lastBlockId="2.[151,1444,197,324]" lastPageId="2" lastPageNumber="236" pageId="1" pageNumber="235">
<emphasis id="B90BEAC4FFAEFFB1FF1AF801FE4DF828" bold="true" box="[151,399,1927,1953]" pageId="1" pageNumber="235">Diagnosis. Larva: d</emphasis>
orsal plate wider than long, distance between bases of trichobothria
<emphasis id="B90BEAC4FFAEFFB1FB34F80EFB14F828" box="[1209,1238,1928,1953]" italics="true" pageId="1" pageNumber="235">Oi</emphasis>
larger than their length; all dorsal hysterosomal setae subequal; excretory pore plate wider than long; basal segment of chelicera with wide strips; I Leg­4 solenidion and eupathidium equal in length, ILeg­6
<emphasis id="B90BEAC4FFAEFFB1FB4EF84DFB24F86D" box="[1219,1254,1995,2020]" italics="true" pageId="1" pageNumber="235">de</emphasis>
=
<emphasis id="B90BEAC4FFAEFFB1FA8DF84DFAD8F86D" box="[1280,1306,1995,2020]" italics="true" pageId="1" pageNumber="235">ds</emphasis>
; II Leg­4 solenidion longer than eupathidium;
<emphasis id="B90BEAC4FFAEFFB1FDD8F86BFD71F78E" bold="true" box="[597,691,2029,2055]" pageId="1" pageNumber="235">adults:</emphasis>
frontal plate elongate (L/W ratio 1.17­1.35), with rather long posterior projections, median eye situated at level of anterior setae of plate; capitulum long (base of capitulum/rostrum L ratio 2.0­2.7); P­3 with 710 setae;
<emphasis id="B90BEAC4FFADFFB2FCB9FF60FC10FE89" bold="true" box="[820,978,230,256]" pageId="2" pageNumber="236">deutonymph</emphasis>
: frontal plate elongate, capitulum with long rostrum; P­3 with four to six long, thin setae, genital field with two pairs subequal acetabula and six to eight pairs of thin setae.
</paragraph>
</subSubSection>
<caption id="DF00665EFFADFFB2FF1AF99FFCFBF9C6" ID-DOI="http://doi.org/10.5281/zenodo.8031927" ID-Zenodo-Dep="8031927" httpUri="https://zenodo.org/record/8031927/files/figure.png" pageId="2" pageNumber="236" startId="2.[151,237,1561,1585]" targetBox="[171,1391,360,1528]" targetPageId="2" targetType="figure">
<paragraph id="8BC036D6FFADFFB2FF1AF99FFCFBF9C6" blockId="2.[151,1443,1561,1616]" pageId="2" pageNumber="236">
<emphasis id="B90BEAC4FFADFFB2FF1AF99FFEDFF9B8" bold="true" box="[151,285,1561,1585]" pageId="2" pageNumber="236">Figures 39</emphasis>
.
<taxonomicName id="4C7F4D55FFADFFB2FEA6F99CFD5DF9B8" authority="Thon, 1899" authorityName="Thon" authorityYear="1899" box="[299,671,1562,1585]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="236" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFADFFB2FEA6F99CFDDBF9B8" box="[299,537,1562,1585]" italics="true" pageId="2" pageNumber="236">Hydryphantes hellichi</emphasis>
<bibRefCitation id="EFEE4B27FFADFFB2FDACF99CFD5DF9B8" author="Thon, K." box="[545,671,1562,1585]" pageId="2" pageNumber="236" pagination="55 - 67" refId="ref4413" refString="Thon, K. (1899) Monographie der bohmischen Hydryphantes-Arten. Bulletin International Academie des Sciences, Prague, 8, 55 - 67." type="journal article" year="1899">Thon, 1899</bibRefCitation>
</taxonomicName>
, larva: 36 excretory pore plate, 7 capitulum, 8 chelicerae, dorsal view, 9 pedipalp. Scale bars: 36, 9 = 50µm, 78 = 100µm.
</paragraph>
</caption>
<subSubSection id="C365655DFFADFFB7FF1AF9FFFE18FA68" lastPageId="7" lastPageNumber="241" pageId="2" pageNumber="236" type="description">
<paragraph id="8BC036D6FFADFFB2FF1AF9FFFE87F91A" blockId="2.[151,1444,1657,2021]" box="[151,325,1657,1683]" pageId="2" pageNumber="236">
<emphasis id="B90BEAC4FFADFFB2FF1AF9FFFE87F91A" bold="true" box="[151,325,1657,1683]" pageId="2" pageNumber="236">Redescription</emphasis>
</paragraph>
<paragraph id="8BC036D6FFADFFB2FF1AF91DFDCDF86C" blockId="2.[151,1444,1657,2021]" pageId="2" pageNumber="236">
<emphasis id="B90BEAC4FFADFFB2FF1AF91DFF26F93C" bold="true" box="[151,228,1691,1717]" pageId="2" pageNumber="236">Larva</emphasis>
. Color red. Idiosoma oval, all setae not associated with glandularia (figures 12). Proterosoma with two pairs of trichobothria (
<emphasis id="B90BEAC4FFADFFB2FE6AF93BFD80F95F" box="[487,578,1725,1750]" italics="true" pageId="2" pageNumber="236">Fp, Oi)</emphasis>
and three pairs of simple setae:
<emphasis id="B90BEAC4FFADFFB2FC6CF93BFBB0F95F" box="[993,1138,1725,1750]" italics="true" pageId="2" pageNumber="236">Fch, Vi, Oe</emphasis>
. Anterior pair of dorsal platelets triangular or oval, trichobothria
<emphasis id="B90BEAC4FFADFFB2FD02F959FD72F971" box="[655,688,1759,1784]" italics="true" pageId="2" pageNumber="236">Fp</emphasis>
long and well extending to posterior margin of dorsal plate. Posterior plate wider than long (L/W ratio 0.750.90), pentagonal, narrows anteriorly and widens posteriorly; median eye slightly developed (rarely absent) and situated slightly posterior to setae
<emphasis id="B90BEAC4FFADFFB2FB18F8A4FB6DF8B2" box="[1173,1199,1826,1851]" italics="true" pageId="2" pageNumber="236">Vi</emphasis>
;
<emphasis id="B90BEAC4FFADFFB2FB4CF8A4FB2DF8B2" box="[1217,1263,1826,1851]" italics="true" pageId="2" pageNumber="236">Fch</emphasis>
shorter
<emphasis id="B90BEAC4FFADFFB2FADEF8A4FAAFF8B2" box="[1363,1389,1826,1851]" italics="true" pageId="2" pageNumber="236">Vi</emphasis>
;
<emphasis id="B90BEAC4FFADFFB2FAF2F8A4FA61F8B2" box="[1407,1443,1826,1851]" italics="true" pageId="2" pageNumber="236">Oi</emphasis>
short, distance between
<emphasis id="B90BEAC4FFADFFB2FE36F8C2FDC9F8D4" box="[443,523,1860,1885]" italics="true" pageId="2" pageNumber="236">OiOi</emphasis>
larger than their length. Hysterosomal setae
<emphasis id="B90BEAC4FFADFFB2FBA7F8C2FA8CF8D4" box="[1066,1358,1860,1885]" italics="true" pageId="2" pageNumber="236">Hi, He, Sci, Sce, Li, Le</emphasis>
and
<emphasis id="B90BEAC4FFADFFB2FA08F8C2FA61F8D4" box="[1413,1443,1860,1885]" italics="true" pageId="2" pageNumber="236">Si</emphasis>
subequal, their bases situated on very small sclerites. All coxal plates separated on each side. Coxal plates II triangular, coxal plates I and III more or less trapezoidal and broadly rounded medially, all coxal setae short and subequal. Urstigma oval and wider than long. Setae
<emphasis id="B90BEAC4FFADFFB2FCB6F82FFC77F84B" box="[827,949,1961,1986]" italics="true" pageId="2" pageNumber="236">Se, Ci, Pi</emphasis>
and
<emphasis id="B90BEAC4FFADFFB2FC64F82FFBCAF84B" box="[1001,1032,1961,1986]" italics="true" pageId="2" pageNumber="236">Pe</emphasis>
subequal and slightly longer than both pairs of anal setae (
<emphasis id="B90BEAC4FFADFFB2FE3AF84DFE3CF86D" box="[439,510,1995,2020]" italics="true" pageId="2" pageNumber="236">Ai, Ae</emphasis>
).
</paragraph>
<caption id="DF00665EFFACFFB3FF1AF9E3FC7CF912" ID-DOI="http://doi.org/10.5281/zenodo.8031929" ID-Zenodo-Dep="8031929" httpUri="https://zenodo.org/record/8031929/files/figure.png" pageId="3" pageNumber="237" startId="3.[151,237,1637,1661]" targetBox="[176,1423,208,1599]" targetPageId="3">
<paragraph id="8BC036D6FFACFFB3FF1AF9E3FC7CF912" blockId="3.[151,1439,1637,1691]" pageId="3" pageNumber="237">
<emphasis id="B90BEAC4FFACFFB3FF1AF9E3FEFFF9F4" bold="true" box="[151,317,1637,1661]" pageId="3" pageNumber="237">Figures 1014.</emphasis>
<taxonomicName id="4C7F4D55FFACFFB3FEC9F9E0FD76F9F4" authority="Thon, 1899" authorityName="Thon" authorityYear="1899" box="[324,692,1638,1661]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="237" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFACFFB3FEC9F9E0FDF2F9F4" box="[324,560,1638,1661]" italics="true" pageId="3" pageNumber="237">Hydryphantes hellichi</emphasis>
<bibRefCitation id="EFEE4B27FFACFFB3FDBAF9E0FD76F9F4" author="Thon, K." box="[567,692,1638,1661]" pageId="3" pageNumber="237" pagination="55 - 67" refId="ref4413" refString="Thon, K. (1899) Monographie der bohmischen Hydryphantes-Arten. Bulletin International Academie des Sciences, Prague, 8, 55 - 67." type="journal article" year="1899">Thon, 1899</bibRefCitation>
</taxonomicName>
, larva: 10 ­ ILeg­46; 11 ­ IILeg­46; 12 ­ IIILeg­46; 13 ­ claws of leg I; 14 ­ claws of leg III. Simple setae on IIIILeg­46 are not shown.
</paragraph>
</caption>
<paragraph id="8BC036D6FFACFFB3FF1AF961FAC2F8EF" blockId="3.[151,1444,1767,2029]" pageId="3" pageNumber="237">Excretory pore plate small, wider than long (L/W ratio 0.570.64), its shape variable (figures 3­6). Capitulum (figure 7) with short, wide base, ventral setae slightly shorter than dorsal ones. The mouth opening surrounded by numerous papillae. Basal part of capitulum without distinct reticulations. Basal segment of chelicera with wide strips, cheliceral stylet heavy and without apical teeth (figure 8).</paragraph>
<paragraph id="8BC036D6FFACFFB3FF70F8E8FB1EF864" blockId="3.[151,1444,1767,2029]" pageId="3" pageNumber="237">Pedipalps moderately long (figure 9): P1 short, without setae; P2 large with convex dorsal margin and single dorsal seta proximally to middle of segment; P3 with two unequal setae (long, thick proximal and relatively short, thin distal); P4 with three unequal setae and large dorsodistal bifurcate claw; P5 small with moderately long solenidion, five long, thick and two relatively short, thin unequal setae.</paragraph>
<paragraph id="8BC036D6FFABFFB4FF1AFF43FC9DFE42" blockId="4.[151,1444,197,796]" pageId="4" pageNumber="238">Leg 6­segmented. Leg II distinctly shorter than anterior and posterior ones. Number of leg setae (specialized setae indicated in parenthesis): I­Leg­1­6 1, 2, 5, 6(s, e), 13 (2s, e), 22 (s, e, ac); II­Leg­1­6 1, 2, 5, 6 (s, e), 12 (2s), 22 (s, e, ac); III­Leg­1­6 1, 2, 5, 5 (s, e), 11 (s), 20. Shape and arrangement of specialized setae on terminal legs segments as shown in figures 1012. All simple setae heavy and usually with long serrations. ILeg­4 solenidion and eupathidium equal in length, ILeg­5 with subequal long proximal solenidia and short distal eupathidium, ILeg­6 de = ds; IILeg­4 solenidion longer than eupathidiumm, II Leg­5 with unequal proximal solenidia, IILeg­6 solenidion is proximal and eupathidium is submedial; III Leg­4 proximal solenidion longer than IIILeg­5 solenidion.</paragraph>
<paragraph id="8BC036D6FFABFFB4FF7BFE55FEDFFD87" blockId="4.[151,1444,197,796]" pageId="4" pageNumber="238">Empodium large and crescent on all tarsi, ambulacra I (figure 13) shorter than ambulacra II and III (figure 14).</paragraph>
<paragraph id="8BC036D6FFABFFB4FF7BFD90FAE4FC95" blockId="4.[151,1444,197,796]" pageId="4" pageNumber="238">
Measurements, n=10. Dorsal plate L 4148, W 5158; setae
<emphasis id="B90BEAC4FFABFFB4FC5FFD90FBC5FDA6" box="[978,1031,534,559]" italics="true" pageId="4" pageNumber="238">Fch</emphasis>
L 2532, setae
<emphasis id="B90BEAC4FFABFFB4FB4FFD90FB28FDA6" box="[1218,1258,534,559]" italics="true" pageId="4" pageNumber="238">Fp</emphasis>
L 4452, setae
<emphasis id="B90BEAC4FFABFFB4FF1AFDBEFF7AFDD8" box="[151,184,568,593]" italics="true" pageId="4" pageNumber="238">Vi</emphasis>
L 3238, setae
<emphasis id="B90BEAC4FFABFFB4FEFDFDBEFE57FDD8" box="[368,405,568,593]" italics="true" pageId="4" pageNumber="238">Oi</emphasis>
L 1013, setae
<emphasis id="B90BEAC4FFABFFB4FDC0FDBEFC52FDD8" box="[589,912,568,593]" italics="true" pageId="4" pageNumber="238">Oe, Hi, He, Sci, Sce, Li, Le</emphasis>
and
<emphasis id="B90BEAC4FFABFFB4FC41FDBEFC28FDD8" box="[972,1002,568,593]" italics="true" pageId="4" pageNumber="238">Si</emphasis>
L 2832; setae
<emphasis id="B90BEAC4FFABFFB4FB2FFDBEFB7CFDD8" box="[1186,1214,568,593]" italics="true" pageId="4" pageNumber="238">Ci</emphasis>
,
<emphasis id="B90BEAC4FFABFFB4FB43FDBEFA8BFDD8" box="[1230,1353,568,593]" italics="true" pageId="4" pageNumber="238">Se, Pi, Pe</emphasis>
1619; setae
<emphasis id="B90BEAC4FFABFFB4FF57FDDCFF39FDFA" box="[218,251,602,627]" italics="true" pageId="4" pageNumber="238">Ai</emphasis>
and
<emphasis id="B90BEAC4FFABFFB4FEBDFDDCFE8DFDFA" box="[304,335,602,627]" italics="true" pageId="4" pageNumber="238">Ae</emphasis>
1315; distance between setae
<emphasis id="B90BEAC4FFABFFB4FD4AFDDCFCD3FDFA" box="[711,785,602,627]" italics="true" pageId="4" pageNumber="238">ViVi</emphasis>
4045, distance between setae
<emphasis id="B90BEAC4FFABFFB4FB0FFDDCFB09FDFA" box="[1154,1227,602,627]" italics="true" pageId="4" pageNumber="238">OiOi</emphasis>
1523; excretory pore plate L 911, W 1216; urstigma L 57, W 812; basal segments of chelicerae L 8599, cheliceral stylet L 2126; strips on basal segment of chelicera
<geoCoordinate id="EE4B5011FFABFFB4FC8BFD1BFC8CFD3E" box="[774,846,669,695]" degrees="2.5" direction="west" orientation="longitude" pageId="4" pageNumber="238" precision="5555" value="-2.5">W 2.5</geoCoordinate>
4.0, distance between strips on basal segment of chelicera 2.55.0; pedipalpal segments (P15) L: 68, 2632, 1726, 1319, 913; legs segments L: ILeg­ 16: 2126, 16­19, 1316, 2229, 2735, 5364; IILeg­16: 2225, 1219, 1014, 1618, 2632, 4148; IIILeg­16: 2430, 1216, 914, 1723, 2733, 4148; de1= 6­8, ds1­6­8; de2­ 20­24, ds2= 6­10.
</paragraph>
<caption id="DF00665EFFABFFB4FF1AF9EEFA58F909" ID-DOI="http://doi.org/10.5281/zenodo.8031931" ID-Zenodo-Dep="8031931" box="[151,1434,1640,1665]" httpUri="https://zenodo.org/record/8031931/files/figure.png" pageId="4" pageNumber="238" startId="4.[151,237,1640,1664]" targetBox="[158,1423,808,1624]" targetPageId="4" targetType="figure">
<paragraph id="8BC036D6FFABFFB4FF1AF9EEFA58F909" blockId="4.[151,1434,1640,1665]" box="[151,1434,1640,1665]" pageId="4" pageNumber="238">
<emphasis id="B90BEAC4FFABFFB4FF1AF9EEFEFFF909" bold="true" box="[151,317,1640,1664]" pageId="4" pageNumber="238">Figures 1516.</emphasis>
<taxonomicName id="4C7F4D55FFABFFB4FEC9F9EFFD76F909" authority="Thon, 1899" authorityName="Thon" authorityYear="1899" box="[324,692,1641,1664]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="238" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFABFFB4FEC9F9EFFDF2F909" box="[324,560,1641,1664]" italics="true" pageId="4" pageNumber="238">Hydryphantes hellichi</emphasis>
<bibRefCitation id="EFEE4B27FFABFFB4FDBAF9EFFD76F909" author="Thon, K." box="[567,692,1641,1664]" pageId="4" pageNumber="238" pagination="55 - 67" refId="ref4413" refString="Thon, K. (1899) Monographie der bohmischen Hydryphantes-Arten. Bulletin International Academie des Sciences, Prague, 8, 55 - 67." type="journal article" year="1899">Thon, 1899</bibRefCitation>
</taxonomicName>
, deutonymph: 16 dorsal view, 17 ventral view. Scale bar: 50µm.
</paragraph>
</caption>
<paragraph id="8BC036D6FFABFFB4FF1AF94DFE20F87D" blockId="4.[151,1444,1739,2036]" pageId="4" pageNumber="238">
<emphasis id="B90BEAC4FFABFFB4FF1AF94DFE80F96C" bold="true" box="[151,322,1739,1765]" pageId="4" pageNumber="238">Deutonymph.</emphasis>
Color red. Idiosoma oval and some what flattened dorsoventrally. Trichobothria
<emphasis id="B90BEAC4FFABFFB4FA95F94AFAA5F96C" box="[1304,1383,1740,1765]" italics="true" pageId="4" pageNumber="238">Fp, Oi</emphasis>
and setae
<emphasis id="B90BEAC4FFABFFB4FF51F968FF3FF88E" box="[220,253,1774,1799]" italics="true" pageId="4" pageNumber="238">Pi</emphasis>
not associated with glandularia, other idiosomal setae associated with glandularia (figures 15­16). Frontal plate (figure 17) elongate (L/W ratio 1.181.30), anterior margin obtuse­angled or slightly convex, posterior margin concave, posterior projections rather long, their length equal to 1/3­1/4 length of basal portion of plate; frontal eye situated posterior trichobothria
<emphasis id="B90BEAC4FFABFFB4FCECF8D5FC40F8E5" box="[865,898,1875,1900]" italics="true" pageId="4" pageNumber="238">Fp</emphasis>
. Coxal plates in four groups, all coxal plates with a few fine setae each. Coxal plates I+II with small subcutaneous posteromedial extension on each side. Genital field (figure 18) with two pairs of acetabula and six to eight pairs of thin setae; anterior and posterior pairs of acetabula subequal. Excretory pore surrounded by sclerotized ring. Integument with short papillae distally rounded (figure 19).
</paragraph>
<caption id="DF00665EFFAAFFB5FF1AF82EFDE1F874" ID-DOI="http://doi.org/10.5281/zenodo.8031933" ID-Zenodo-Dep="8031933" httpUri="https://zenodo.org/record/8031933/files/figure.png" pageId="5" pageNumber="239" startId="5.[151,237,1960,1984]" targetBox="[196,1410,200,1946]" targetPageId="5" targetType="figure">
<paragraph id="8BC036D6FFAAFFB5FF1AF82EFDE1F874" blockId="5.[151,1437,1960,2046]" pageId="5" pageNumber="239">
<emphasis id="B90BEAC4FFAAFFB5FF1AF82EFEFFF849" bold="true" box="[151,317,1960,1984]" pageId="5" pageNumber="239">Figures 1724.</emphasis>
<taxonomicName id="4C7F4D55FFAAFFB5FEC9F82FFD76F836" authority="Thon, 1899" authorityName="Thon" authorityYear="1899" box="[324,692,1960,1984]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="239" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFAAFFB5FEC9F82FFDF2F849" box="[324,560,1961,1984]" italics="true" pageId="5" pageNumber="239">Hydryphantes hellichi</emphasis>
<bibRefCitation id="EFEE4B27FFAAFFB5FDBAF82EFD76F836" author="Thon, K." box="[567,692,1960,1983]" pageId="5" pageNumber="239" pagination="55 - 67" refId="ref4413" refString="Thon, K. (1899) Monographie der bohmischen Hydryphantes-Arten. Bulletin International Academie des Sciences, Prague, 8, 55 - 67." type="journal article" year="1899">Thon, 1899</bibRefCitation>
</taxonomicName>
, deutonymph: 17 frontal plate; 18 genital field; 19 fragment of integument; 20 capitulum; 21 chelicera; 22 pedipalp; 23 ­ IILeg­45; 24 claw. Scale bars: 17, 23 = 200µm, 18, 20­21 = 100µm, 19, 22, 24 = 50µm.
</paragraph>
</caption>
<caption id="DF00665EFFA9FFB6FF1AF96EFC05F897" ID-DOI="http://doi.org/10.5281/zenodo.8031935" ID-Zenodo-Dep="8031935" httpUri="https://zenodo.org/record/8031935/files/figure.png" pageId="6" pageNumber="240" startId="6.[151,237,1768,1792]" targetBox="[154,1421,210,1746]" targetPageId="6" targetType="figure">
<paragraph id="8BC036D6FFA9FFB6FF1AF96EFC05F897" blockId="6.[151,1436,1768,1823]" pageId="6" pageNumber="240">
<emphasis id="B90BEAC4FFA9FFB6FF1AF96EFEFDF889" bold="true" box="[151,319,1768,1792]" pageId="6" pageNumber="240">Figures 2529.</emphasis>
<taxonomicName id="4C7F4D55FFA9FFB6FECAF96FFD79F976" authority="Thon, 1899" authorityName="Thon" authorityYear="1899" box="[327,699,1768,1792]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="240" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFA9FFB6FECAF96FFDF6F889" box="[327,564,1769,1792]" italics="true" pageId="6" pageNumber="240">Hydryphantes hellichi</emphasis>
<bibRefCitation id="EFEE4B27FFA9FFB6FDB1F96EFD79F976" author="Thon, K." box="[572,699,1768,1791]" pageId="6" pageNumber="240" pagination="55 - 67" refId="ref4413" refString="Thon, K. (1899) Monographie der bohmischen Hydryphantes-Arten. Bulletin International Academie des Sciences, Prague, 8, 55 - 67." type="journal article" year="1899">Thon, 1899</bibRefCitation>
</taxonomicName>
, adults: 25 ­ frontal plate; 26 coxal plates; 27 genital field; 28 capitulum; 29 pedipalp. Scale bars: 2526, 28 = 200µm, 27, 29 = 100µm.
</paragraph>
</caption>
<paragraph id="8BC036D6FFA9FFB6FF1AF8CEFEA7F860" blockId="6.[151,1444,1864,2025]" pageId="6" pageNumber="240">Capitulum (figure 20) with long rostrum (base of capitulum/rostrum L ratio 2.2­3.0) and convex basal part. Chelicera (figures 21) rather slender, basal segment with large dorsal hump near middle, cheliceral stylet moderately long. Pedipalp compact (figures 22): P­1 with one to two setae, P­2 with 10­13 short, thick setae, P­3 with four to six long, thin setae; P­4 slightly tapering distally, with three distal setae and short, thick dorsodistal spine.</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFF43FE58FEAB" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">II­Leg­4­5 (figures 23) and III/IV­Leg­3­5 with long swimming setae. Number of swimming setae: II­Leg­4­5, 0­2, 9­14; III­Leg­3­5, 2­3, 6­9, 9­14; IV­Leg­3­5, 3­4, 9­13, 10­14. All legs with simple hooklike claws (figure 24).</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFEACFE6EFD87" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">Measurements (n=4). Idiosoma L 1000­1250; coxal plates I+II L 175­190, W 200­210; Coxal plates III+IV L 250­285, W 210­240; genital plate L 78­100, W 42­66; anterior genital acetabula D 30­33, posterior genital acetabula D 35­38; capitulum L 215­240; chelicera L 210­240, cheliceral stylet L 95­115; pedipalpal segments (P15) L: 50­57, 100­135, 55­72, 130­150, 24­30; leg segments L: ILeg16: 75­90, 85­115, 85­ 115, 135­165, 160­200, 185­215; IILeg16: 75­90, 85­115, 115­140, 185­225, 215­275, 250­290; III­Leg­ 1­6: 75­90, 85­115, 100­145, 185­250, 225­290, 250­295; IVLeg16: 150­165, 125­165, 160­215, 210­ 340, 260­340, 250­300.</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFD90FD2CFD1C" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">
<emphasis id="B90BEAC4FFA8FFB7FF7BFD90FE9AFDB9" bold="true" box="[246,344,534,560]" pageId="7" pageNumber="241">Adults.</emphasis>
Males and females are similar to deutonymph, but differ in structure of external genital organs, large sizes, number of idiosomal glandularia (setae
<emphasis id="B90BEAC4FFA8FFB7FCF4FDBEFC58FDD8" box="[889,922,568,593]" italics="true" pageId="7" pageNumber="241">Pi</emphasis>
associated with glandularia) and by more numerous setae on all segments of appendages. Males and females do not exhibit external sexual dimorphism, but mature females larger than males.
</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF70FD1BFD22FD73" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">Frontal shield (figure 25) elongate (L/W ratio 1.17­1.35), anterior margin convex, lateral margins concave, posterior projections rather long. Median eye small and situated almost at level of anterior setae. All coxal plates with numerous setae (figures 26).</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFC84FBC9FCB7" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">Genital field (figure 27) with three pairs of acetabula and of thin setae; anterior pair of acetabula large than second pair but slightly smaller than posterior pairs of acetabula.</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF1AFCC0FBD9FCE9" blockId="7.[151,1444,197,1505]" box="[151,1051,838,864]" pageId="7" pageNumber="241">Acetabular plate elongate (L/W ratio 2.0­2.5), with 20­30 pairs medial setae.</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFCE1FC2AFC4C" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">
Capitulum (figures 28) with long rostrum (base of capitulum/rostrum L ratio 2.0­2.7) and convex basal part. Pedipalp compact (
<figureCitation id="13442A53FFA8FFB7FE77FC0FFD91FC2A" box="[506,595,905,931]" captionStart="Figures 2529" captionStartId="6.[151,237,1768,1792]" captionTargetBox="[154,1421,210,1746]" captionTargetId="figure-13@6.[151,1436,600,1749]" captionTargetPageId="6" captionText="Figures 2529. Hydryphantes hellichi Thon, 1899, adults: 25 ­ frontal plate; 26 coxal plates; 27 genital field; 28 capitulum; 29 pedipalp. Scale bars: 2526, 28 = 200µm, 27, 29 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031935" httpUri="https://zenodo.org/record/8031935/files/figure.png" pageId="7" pageNumber="241">Fig. 29</figureCitation>
): P1 with 3­4 dorsodistal setae; P2 with 1318 setae; P3 with 710 setae; P4 slender with four thin setae and short, thick dorsodistal spine.
</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFC4AFEF4FB81" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">Number of swimming setae: II­Leg­4­5, 3­6, 16­20; III­Leg­3­5, 0­4, 13­20, 17­28; IV­Leg­3­5, 3­8, 20­27, 20­26.</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFB96FEB3FB7D" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">
<emphasis id="B90BEAC4FFA8FFB7FF7BFB96FDDBFBA3" bold="true" box="[246,537,1040,1066]" pageId="7" pageNumber="241">Measurements, female</emphasis>
(n=7). Idiosoma L 1600­2070; dorsal plate L 535­570, W 410­465; genital flaps L 275­315, W 135­150; genital acetabula (ac.1ac. 3) D 50­60, 35­40, 60­75; capitulum L 400­460, rostrum L 110­125, chelicera total L 525­590, cheliceral stylet L 175­200; pedipalpal segments (P15) L: 90­100, 175­190, 110­125, 235­255, 37­50; legs segments L: ILeg­16: 110­150, 150­165, 175­215, 260­ 300, 310­350, 310­365; IILeg­16: 125­150, 150­180, 210­250, 360­425, 410­475, 410­475; IIILeg­16: 125­140, 160­190, 225­265, 375­450, 425­500, 425­475; IVLeg­16: 235­265, 250­265, 350­400, 500­590, 500­565, 435­500.
</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BFB7AFE18FA68" blockId="7.[151,1444,197,1505]" pageId="7" pageNumber="241">
<emphasis id="B90BEAC4FFA8FFB7FF7BFB7AFDC1FA9F" bold="true" box="[246,515,1276,1302]" pageId="7" pageNumber="241">Measurements, male</emphasis>
(n=8). Idiosoma L 1400­1570; dorsal plate L 500­560, W 400­420; genital flaps L 260­315, W 120­130; genital acetabula (ac.1ac.3) D 37­43, 30­35, 50­63; capitulum L 360­400, rostrum L 100­125; chelicera: basal segment L 350­365, cheliceral stylet L 135­165; pedipalpal segments (P15) L: 80­90, 160­175, 110­120, 210­230, 30­38; legs segments L: ILeg­16: 100­125, 125­140, 160­ 190, 260­290, 325­350, 375­400; IILeg­16: 100­125, 135­140, 210­240, 350­400, 435­465, 475­500; III Leg­16: 125­140, 150­190, 225­250, 360­400, 435­465, 485­515; IVLeg­16: 210­240, 225­250, 325­375, 485­550, 475­525, 485­500
</paragraph>
</subSubSection>
<subSubSection id="C365655DFFA8FFB7FF1AF98FFD7CF81E" pageId="7" pageNumber="241" type="discussion">
<paragraph id="8BC036D6FFA8FFB7FF1AF98FFAB8F8DD" blockId="7.[151,1444,1545,1943]" pageId="7" pageNumber="241">
<emphasis id="B90BEAC4FFA8FFB7FF1AF98FFED5F9AA" bold="true" box="[151,279,1545,1571]" pageId="7" pageNumber="241">Remarks.</emphasis>
The water mite
<taxonomicName id="4C7F4D55FFA8FFB7FE62F98CFD38F9AA" authorityName="Thon" authorityYear="1899" box="[495,762,1546,1571]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFA8FFB7FE62F98CFD38F9AA" box="[495,762,1546,1571]" italics="true" pageId="7" pageNumber="241">Hydryphantes hellichi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C7F4D55FFA8FFB7FC3FF98CFB0DF9AD" authority="(Geer, 1778)" baseAuthorityName="Geer" baseAuthorityYear="1778" box="[946,1231,1546,1572]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="ruber">
<emphasis id="B90BEAC4FFA8FFB7FC3FF98CFBE7F9AA" box="[946,1061,1546,1571]" italics="true" pageId="7" pageNumber="241">H. ruber</emphasis>
(Geer, 1778)
</taxonomicName>
. However, clear differences can be found in the morphology of the larva, deutonymph and adults of
<taxonomicName id="4C7F4D55FFA8FFB7FB13F9AAFADCF9CC" authorityName="Thon" authorityYear="1899" box="[1182,1310,1580,1605]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFA8FFB7FB13F9AAFADCF9CC" box="[1182,1310,1580,1605]" italics="true" pageId="7" pageNumber="241">H. hellichi</emphasis>
</taxonomicName>
compared with the latter species.
<taxonomicName id="4C7F4D55FFA8FFB7FE3FF9C8FD78F9EE" authorityName="Thon" authorityYear="1899" box="[434,698,1614,1639]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFA8FFB7FE3FF9C8FD78F9EE" box="[434,698,1614,1639]" italics="true" pageId="7" pageNumber="241">Hydryphantes hellichi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C7F4D55FFA8FFB7FCD2F9C8FC05F9EE" baseAuthorityName="Geer" baseAuthorityYear="1778" box="[863,967,1614,1639]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="ruber">
<emphasis id="B90BEAC4FFA8FFB7FCD2F9C8FC05F9EE" box="[863,967,1614,1639]" italics="true" pageId="7" pageNumber="241">H. ruber</emphasis>
</taxonomicName>
by the following characters (character states of the larva, deutonymph and adults are given in parenthesis):
<emphasis id="B90BEAC4FFA8FFB7FC6DF9E9FBE3F900" bold="true" box="[992,1057,1647,1673]" pageId="7" pageNumber="241">larva</emphasis>
: the distance between bases of trichobothria
<emphasis id="B90BEAC4FFA8FFB7FEB7F917FE9DF923" box="[314,351,1681,1706]" italics="true" pageId="7" pageNumber="241">Oi</emphasis>
larger than their length,
<figureCitation id="13442A53FFA8FFB7FD06F917FD11F922" box="[651,723,1681,1707]" captionStart="Figures 12" captionStartId="1.[151,237,1862,1886]" captionTargetBox="[167,1428,1038,1854]" captionTargetId="figure-239@1.[151,1434,999,1856]" captionTargetPageId="1" captionText="Figures 12. Hydryphantes hellichi Thon, 1899, larva: 1­ dorsal view, 2­ ventral view. Scale bar: 50µm." figureDoi="http://doi.org/10.5281/zenodo.8031925" httpUri="https://zenodo.org/record/8031925/files/figure.png" pageId="7" pageNumber="241">Fig. 1</figureCitation>
(smaller than their length,
<figureCitation id="13442A53FFA8FFB7FBAEF917FBBFF922" box="[1059,1149,1681,1707]" captionStart="Figures 3035" captionStartId="8.[151,237,1722,1746]" captionTargetBox="[175,1413,209,1701]" captionTargetId="figure-13@8.[151,1436,587,1703]" captionTargetPageId="8" captionText="Figures 3035.Hydryphantes ruber (Geer, 1788): 30 dorsal platelets; 31 chelicera, dorsal view; 32 excretory pore plate; 33 I­Leg­46, simple setae are not shown; 34 frontal plate; 35 capitulum, lateral view; 3033 larva, 3435 female. Scale bars: 3031 = 50µm, 3233 = 20µm, 34 = 200µm, 35 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031937" httpUri="https://zenodo.org/record/8031937/files/figure.png" pageId="7" pageNumber="241">Fig. 30</figureCitation>
), the basal segments of chelicerae with wide strips,
<figureCitation id="13442A53FFA8FFB7FE6EF935FDEAF944" box="[483,552,1715,1741]" captionStart="Figures 39" captionStartId="2.[151,237,1561,1585]" captionTargetBox="[171,1391,360,1528]" captionTargetId="figure-70@2.[151,1436,570,1554]" captionTargetPageId="2" captionText="Figures 39.Hydryphantes hellichi Thon, 1899, larva: 36 excretory pore plate, 7 capitulum, 8 chelicerae, dorsal view, 9 pedipalp. Scale bars: 36, 9 = 50µm, 78 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031927" httpUri="https://zenodo.org/record/8031927/files/figure.png" pageId="7" pageNumber="241">Fig. 8</figureCitation>
(with narrow strips,
<figureCitation id="13442A53FFA8FFB7FCADF935FCB5F944" box="[800,887,1715,1741]" captionStart="Figures 3035" captionStartId="8.[151,237,1722,1746]" captionTargetBox="[175,1413,209,1701]" captionTargetId="figure-13@8.[151,1436,587,1703]" captionTargetPageId="8" captionText="Figures 3035.Hydryphantes ruber (Geer, 1788): 30 dorsal platelets; 31 chelicera, dorsal view; 32 excretory pore plate; 33 I­Leg­46, simple setae are not shown; 34 frontal plate; 35 capitulum, lateral view; 3033 larva, 3435 female. Scale bars: 3031 = 50µm, 3233 = 20µm, 34 = 200µm, 35 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031937" httpUri="https://zenodo.org/record/8031937/files/figure.png" pageId="7" pageNumber="241">Fig. 31</figureCitation>
), excretory pore plate wider than long, figures 3­6 (as long as wide,
<figureCitation id="13442A53FFA8FFB7FE26F952FDC5F967" box="[427,519,1748,1774]" captionStart="Figures 3035" captionStartId="8.[151,237,1722,1746]" captionTargetBox="[175,1413,209,1701]" captionTargetId="figure-13@8.[151,1436,587,1703]" captionTargetPageId="8" captionText="Figures 3035.Hydryphantes ruber (Geer, 1788): 30 dorsal platelets; 31 chelicera, dorsal view; 32 excretory pore plate; 33 I­Leg­46, simple setae are not shown; 34 frontal plate; 35 capitulum, lateral view; 3033 larva, 3435 female. Scale bars: 3031 = 50µm, 3233 = 20µm, 34 = 200µm, 35 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031937" httpUri="https://zenodo.org/record/8031937/files/figure.png" pageId="7" pageNumber="241">Fig. 32</figureCitation>
), ILeg­4 solenidion and eupathidium equal in length,
<figureCitation id="13442A53FFA8FFB7FB39F952FACEF967" box="[1204,1292,1748,1774]" captionStart="Figures 1014" captionStartId="3.[151,237,1637,1661]" captionTargetBox="[176,1423,208,1599]" captionTargetId="figure-13@3.[151,1436,590,1631]" captionText="Figures 1014. Hydryphantes hellichi Thon, 1899, larva: 10 ­ ILeg­46; 11 ­ IILeg­46; 12 ­ IIILeg­46; 13 ­ claws of leg I; 14 ­ claws of leg III. Simple setae on IIIILeg­46 are not shown." figureDoi="http://doi.org/10.5281/zenodo.8031929" httpUri="https://zenodo.org/record/8031929/files/figure.png" pageId="7" pageNumber="241">Fig. 10</figureCitation>
(solenidion longer than eupathidium,
<figureCitation id="13442A53FFA8FFB7FE40F970FDE7F899" box="[461,549,1782,1808]" captionStart="Figures 3035" captionStartId="8.[151,237,1722,1746]" captionTargetBox="[175,1413,209,1701]" captionTargetId="figure-13@8.[151,1436,587,1703]" captionTargetPageId="8" captionText="Figures 3035.Hydryphantes ruber (Geer, 1788): 30 dorsal platelets; 31 chelicera, dorsal view; 32 excretory pore plate; 33 I­Leg­46, simple setae are not shown; 34 frontal plate; 35 capitulum, lateral view; 3033 larva, 3435 female. Scale bars: 3031 = 50µm, 3233 = 20µm, 34 = 200µm, 35 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031937" httpUri="https://zenodo.org/record/8031937/files/figure.png" pageId="7" pageNumber="241">Fig. 33</figureCitation>
), ILeg­6 de=ds (ILeg­6 de &lt;ds);
<emphasis id="B90BEAC4FFA8FFB7FC6DF970FACCF899" bold="true" box="[992,1294,1782,1808]" pageId="7" pageNumber="241">deutonymph and adults</emphasis>
: the frontal plate elongate with rather long posterior projections,
<figureCitation id="13442A53FFA8FFB7FCA1F89EFC50F8BB" box="[812,914,1816,1842]" captionStart="Figures 1516" captionStartId="4.[151,237,1640,1664]" captionTargetBox="[158,1423,808,1624]" captionTargetId="figure-310@4.[151,1436,808,1634]" captionTargetPageId="4" captionText="Figures 1516. Hydryphantes hellichi Thon, 1899, deutonymph: 16 dorsal view, 17 ventral view. Scale bar: 50µm." figureDoi="http://doi.org/10.5281/zenodo.8031931" httpUri="https://zenodo.org/record/8031931/files/figure.png" pageId="7" pageNumber="241">Figs. 15</figureCitation>
,
<figureCitation id="13442A53FFA8FFB7FC29F89EFC06F8BB" box="[932,964,1816,1842]" captionStart="Figures 1724" captionStartId="5.[151,237,1960,1984]" captionTargetBox="[196,1410,200,1946]" captionTargetId="figure-13@5.[168,1418,591,1954]" captionTargetPageId="5" captionText="Figures 1724. Hydryphantes hellichi Thon, 1899, deutonymph: 17 frontal plate; 18 genital field; 19 fragment of integument; 20 capitulum; 21 chelicera; 22 pedipalp; 23 ­ IILeg­45; 24 claw. Scale bars: 17, 23 = 200µm, 18, 20­21 = 100µm, 19, 22, 24 = 50µm." figureDoi="http://doi.org/10.5281/zenodo.8031933" httpUri="https://zenodo.org/record/8031933/files/figure.png" pageId="7" pageNumber="241">17</figureCitation>
,
<figureCitation id="13442A53FFA8FFB7FC5BF89EFC31F8BB" box="[982,1011,1816,1842]" captionStart="Figures 2529" captionStartId="6.[151,237,1768,1792]" captionTargetBox="[154,1421,210,1746]" captionTargetId="figure-13@6.[151,1436,600,1749]" captionTargetPageId="6" captionText="Figures 2529. Hydryphantes hellichi Thon, 1899, adults: 25 ­ frontal plate; 26 coxal plates; 27 genital field; 28 capitulum; 29 pedipalp. Scale bars: 2526, 28 = 200µm, 27, 29 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031935" httpUri="https://zenodo.org/record/8031935/files/figure.png" pageId="7" pageNumber="241">25</figureCitation>
(subquadrate, with short posterior projections,
<figureCitation id="13442A53FFA8FFB7FEA5F8BCFEBDF8DD" box="[296,383,1850,1876]" captionStart="Figures 3035" captionStartId="8.[151,237,1722,1746]" captionTargetBox="[175,1413,209,1701]" captionTargetId="figure-13@8.[151,1436,587,1703]" captionTargetPageId="8" captionText="Figures 3035.Hydryphantes ruber (Geer, 1788): 30 dorsal platelets; 31 chelicera, dorsal view; 32 excretory pore plate; 33 I­Leg­46, simple setae are not shown; 34 frontal plate; 35 capitulum, lateral view; 3033 larva, 3435 female. Scale bars: 3031 = 50µm, 3233 = 20µm, 34 = 200µm, 35 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031937" httpUri="https://zenodo.org/record/8031937/files/figure.png" pageId="7" pageNumber="241">Fig. 34</figureCitation>
), the capitulum with long rostrum,
<figureCitation id="13442A53FFA8FFB7FC91F8BCFCBCF8DD" box="[796,894,1850,1876]" captionStart="Figures 1724" captionStartId="5.[151,237,1960,1984]" captionTargetBox="[196,1410,200,1946]" captionTargetId="figure-13@5.[168,1418,591,1954]" captionTargetPageId="5" captionText="Figures 1724. Hydryphantes hellichi Thon, 1899, deutonymph: 17 frontal plate; 18 genital field; 19 fragment of integument; 20 capitulum; 21 chelicera; 22 pedipalp; 23 ­ IILeg­45; 24 claw. Scale bars: 17, 23 = 200µm, 18, 20­21 = 100µm, 19, 22, 24 = 50µm." figureDoi="http://doi.org/10.5281/zenodo.8031933" httpUri="https://zenodo.org/record/8031933/files/figure.png" pageId="7" pageNumber="241">Figs. 20</figureCitation>
,
<figureCitation id="13442A53FFA8FFB7FC07F8BCFC65F8DD" box="[906,935,1850,1876]" captionStart="Figures 2529" captionStartId="6.[151,237,1768,1792]" captionTargetBox="[154,1421,210,1746]" captionTargetId="figure-13@6.[151,1436,600,1749]" captionTargetPageId="6" captionText="Figures 2529. Hydryphantes hellichi Thon, 1899, adults: 25 ­ frontal plate; 26 coxal plates; 27 genital field; 28 capitulum; 29 pedipalp. Scale bars: 2526, 28 = 200µm, 27, 29 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031935" httpUri="https://zenodo.org/record/8031935/files/figure.png" pageId="7" pageNumber="241">28</figureCitation>
(with relatively short rostrum,
<figureCitation id="13442A53FFA8FFB7FA98F8BCFAAEF8DD" box="[1301,1388,1850,1876]" captionStart="Figures 3035" captionStartId="8.[151,237,1722,1746]" captionTargetBox="[175,1413,209,1701]" captionTargetId="figure-13@8.[151,1436,587,1703]" captionTargetPageId="8" captionText="Figures 3035.Hydryphantes ruber (Geer, 1788): 30 dorsal platelets; 31 chelicera, dorsal view; 32 excretory pore plate; 33 I­Leg­46, simple setae are not shown; 34 frontal plate; 35 capitulum, lateral view; 3033 larva, 3435 female. Scale bars: 3031 = 50µm, 3233 = 20µm, 34 = 200µm, 35 = 100µm." figureDoi="http://doi.org/10.5281/zenodo.8031937" httpUri="https://zenodo.org/record/8031937/files/figure.png" pageId="7" pageNumber="241">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="8BC036D6FFA8FFB7FF7BF8DDFD7CF81E" blockId="7.[151,1444,1545,1943]" pageId="7" pageNumber="241">
Thus, the morphology of all active stages of
<taxonomicName id="4C7F4D55FFA8FFB7FC81F8DDFC4BF8FD" authorityName="Thon" authorityYear="1899" box="[780,905,1883,1908]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFA8FFB7FC81F8DDFC4BF8FD" box="[780,905,1883,1908]" italics="true" pageId="7" pageNumber="241">H. hellichi</emphasis>
</taxonomicName>
clearly differs from that of
<taxonomicName id="4C7F4D55FFA8FFB7FB5AF8DDFAFFF8FD" baseAuthorityName="Geer" baseAuthorityYear="1778" box="[1239,1341,1883,1908]" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="ruber">
<emphasis id="B90BEAC4FFA8FFB7FB5AF8DDFAFFF8FD" box="[1239,1341,1883,1908]" italics="true" pageId="7" pageNumber="241">H. ruber</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7F4D55FFA8FFB7FA0DF8DDFF2DF81F" authorityName="Thon" authorityYear="1899" class="Arachnida" family="Hydryphantidae" genus="Hydryphantes" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="241" phylum="Arthropoda" rank="species" species="hellichi">
<emphasis id="B90BEAC4FFA8FFB7FA0DF8DDFF2DF81F" italics="true" pageId="7" pageNumber="241">H. hellichi</emphasis>
</taxonomicName>
should be treated as a separate species.
</paragraph>
</subSubSection>
</treatment>
</document>