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<document id="DFF396E30B3B84C47B1A6C295EF1FA6D" ID-DOI="10.5281/zenodo.13244881" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244881" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970250919" checkinUser="felipe" docAuthor="Hendrich, Lars &amp; Balke, Michael" docDate="2004" docId="03D687B7A73AFFB03A91FCC0BCC34E6F" docLanguage="en" docName="RafflesBZool.52.1.97-145.pdf" docOrigin="Raffles Bulletin of Zoology 52 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Copelatus tenebrosus Regimbart 1880" docType="treatment" docVersion="2" lastPageNumber="118" masterDocId="FFEFFFCFA72FFFA53903FF87BF1E4871" masterDocTitle="Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #" masterLastPageNumber="145" masterPageNumber="97" pageNumber="118" updateTime="1722975545229" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="919CCF1AD38BA3D1251525CF824CA4EE">Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #</mods:title>
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<mods:namePart id="FF594EBD13E892165C3698083401B6F5">Hendrich, Lars</mods:namePart>
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<treatment id="03D687B7A73AFFB03A91FCC0BCC34E6F" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03A91FCC0BCC34E6F" httpUri="http://treatment.plazi.org/id/03D687B7A73AFFB03A91FCC0BCC34E6F" lastPageNumber="118" pageId="21" pageNumber="118">
<subSubSection id="C365652AA73AFFB03A91FCC0BA594B2E" box="[914,1351,839,863]" pageId="21" pageNumber="118" type="nomenclature">
<paragraph id="8BC036A1A73AFFB03A91FCC0BA594B2E" blockId="21.[914,1351,839,863]" box="[914,1351,839,863]" pageId="21" pageNumber="118">
<heading id="D08881CDA73AFFB03A91FCC0BA594B2E" bold="true" box="[914,1351,839,863]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="118" reason="2">
<taxonomicName id="4C7F4D22A73AFFB03A91FCC0BA594B2E" ID-CoL="Y3GY" authority="Regimbart, 1880" authorityName="Regimbart" authorityYear="1880" box="[914,1351,839,863]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B90BEAB3A73AFFB03A91FCC0BA594B2E" bold="true" box="[914,1351,839,863]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A91FCC0BB9E4B2E" bold="true" box="[914,1152,839,863]" italics="true" pageId="21" pageNumber="118">Copelatus tenebrosus</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB03D84FCC0BA594B2E" author="Regimbart, M" box="[1159,1351,839,863]" pageId="21" pageNumber="118" pagination="209 - 216" refId="ref40404" refString="Regimbart, M., 1880. The new Dytiscidae and Gyrinidae collected during the recent scientific Sumatraexpedition. Notes from the Leyden Museum, 2: 209 - 216." type="journal article" year="1880">Régimbart, 1880</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03A2EFC03BBD24B9E" pageId="21" pageNumber="118" type="reference_group">
<paragraph id="8BC036A1A73AFFB03A2EFC03BBD24B9E" blockId="21.[813,1452,900,1007]" pageId="21" pageNumber="118">
<treatmentCitationGroup id="AB6F118FA73AFFB03A2EFC03BBD24B9E" pageId="21" pageNumber="118">
<treatmentCitation id="0ADE10B0A73AFFB03A2EFC03BBCD4BE8" author="Regimbart, M" box="[813,1235,900,921]" page="210" pageId="21" pageNumber="118" year="1880">
<taxonomicName id="4C7F4D22A73AFFB03A2EFC03BBCD4BE8" ID-CoL="Y3GY" authority="Regimbart, 1880: 210" authorityName="Regimbart" authorityPageNumber="210" authorityYear="1880" box="[813,1235,900,921]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B90BEAB3A73AFFB03A2EFC03BCE54BE8" box="[813,1019,900,921]" italics="true" pageId="21" pageNumber="118">Copelatus tenebrosus</emphasis>
<bibRefCitation id="EFEE4B50A73AFFB03D03FC03BBCD4BE8" author="Regimbart, M" box="[1024,1235,900,921]" pageId="21" pageNumber="118" pagination="209 - 216" refId="ref40404" refString="Regimbart, M., 1880. The new Dytiscidae and Gyrinidae collected during the recent scientific Sumatraexpedition. Notes from the Leyden Museum, 2: 209 - 216." type="journal article" year="1880">Régimbart, 1880: 210</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03DDFFC03BC644BC7" author="Zimmermann, A" page="217" pageId="21" pageNumber="118" year="1919">
<bibRefCitation id="EFEE4B50A73AFFB03DDFFC03BC644BC7" author="Zimmermann, A" pageId="21" pageNumber="118" pagination="68 - 249" refId="ref42338" refString="Zimmermann, A., 1919 a. Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem. Archiv fur Naturgeschichte, 83 (A), 12, 68 - 249 (1917)." type="journal article" year="1919">Zimmermann, 1919a: 217</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03A85FC26BCF34BC7" author="Zimmermann, A" box="[902,1005,929,950]" page="143" pageId="21" pageNumber="118" year="1920">
<bibRefCitation id="EFEE4B50A73AFFB03A85FC26BCF34BC7" author="Zimmermann, A" box="[902,1005,929,950]" pageId="21" pageNumber="118" pagination="1 - 326" refId="ref42405" refString="Zimmermann, A., 1920. Dytiscidae. In: Junk, W. &amp; S. Schenkling (eds.), Coleopterorum Catatalogus, Berlin, 4, pars 71, 1 - 326." type="journal article" year="1920">1920: 143</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03AFAFC26BBA14BC7" author="Fernando, C" box="[1017,1215,929,950]" page="87" pageId="21" pageNumber="118" year="1964">
<bibRefCitation id="EFEE4B50A73AFFB03AFAFC26BBA14BC7" author="Fernando, C" box="[1017,1215,929,950]" pageId="21" pageNumber="118" pagination="19 - 31" refId="ref36873" refString="Fernando, C. H., 1964. Notes on aquatic insects colonizing an isolated pond in Mawai, Johore. Bulletin of the National Museum, Singapore, 30: 19 - 31." type="journal article" year="1964">Fernando, 1964: 87</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03DC8FC26BAB64BC7" author="Gueorguiev, V" box="[1227,1448,929,950]" page="23" pageId="21" pageNumber="118" year="1968">
<bibRefCitation id="EFEE4B50A73AFFB03DC8FC26BAB64BC7" author="Gueorguiev, V" box="[1227,1448,929,950]" pageId="21" pageNumber="118" pagination="5 - 45" refId="ref37343" refString="Gueorguiev, V., 1968. Essai de classification des Coleopteres Dytiscidae, Tribus Copelatini (Colymbetinae). Izvestija na Zoologitjeskija Institut s Musei Sofia, 28: 5 - 45." type="journal article" year="1968">Guéorguiev, 1968: 23</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03A50FC3ABB134BA3" author="Vazirani, T" box="[851,1037,957,978]" page="317" pageId="21" pageNumber="118" year="1970">
<bibRefCitation id="EFEE4B50A73AFFB03A50FC3ABB134BA3" author="Vazirani, T" box="[851,1037,957,978]" pageId="21" pageNumber="118" pagination="93 - 129" refId="ref41247" refString="Vazirani, T. G., 1970. Contribution to the study of aquatic beetles (Coleoptera) V. A review of Hydroporinae: Dytiscidae in part, from India. Oriental Insects, 4 (1): 93 - 129." type="journal article" year="1970">Vazirani, 1970: 317</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03D15FC3ABB744BA3" author="Vazirani, T" box="[1046,1130,957,978]" page="57" pageId="21" pageNumber="118" year="1977">
<bibRefCitation id="EFEE4B50A73AFFB03D15FC3ABB744BA3" author="Vazirani, T" box="[1046,1130,957,978]" pageId="21" pageNumber="118" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">1977: 57</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73AFFB03D71FC3ABA524BA3" author="Yano, K &amp; Chu, P" box="[1138,1356,957,978]" page="107" pageId="21" pageNumber="118" year="1983">
<bibRefCitation id="EFEE4B50A73AFFB03D71FC3ABA524BA3" author="Yano, K &amp; Chu, P" box="[1138,1356,957,978]" pageId="21" pageNumber="118" pagination="103 - 118" refId="ref42131" refString="Yano, K., Y. I. Chu, P. W. Resma, &amp; M. Sato, 1983 b. Faunal and biological studies on the insects of paddy fields in Asia. XII. Aquatic Coleoptera from Taiwan and the Philippines. Chinese Journal of Entomology, 3 (2): 103 - 118." type="journal article" year="1983">Yano et al., 1983b: 107</bibRefCitation>
</treatmentCitation>
; Hendrich &amp; Balke, 1995: 44;
<treatmentCitation id="0ADE10B0A73AFFB03D14FC5DBBD64B9E" author="Nilsson, A" box="[1047,1224,986,1007]" page="71" pageId="21" pageNumber="118" year="2001">
<bibRefCitation id="EFEE4B50A73AFFB03D14FC5DBBD64B9E" author="Nilsson, A" box="[1047,1224,986,1007]" pageId="21" pageNumber="118" refId="ref39046" refString="Nilsson, A. N., 2001. Dytiscidae (Coleoptera). In: World Catalogue of Insects Volume 3. Apollo Books, Stenstrup, Denmark, 395 pp." type="book" year="2001">Nilsson, 2001: 71</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03A2EFB93BABC4C13" pageId="21" pageNumber="118" type="materials_examined">
<paragraph id="8BC036A1A73AFFB03A2EFB93BABC4C13" blockId="21.[813,1452,1044,1122]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A2EFB93BCE04C58" bold="true" box="[813,1022,1044,1065]" italics="true" pageId="21" pageNumber="118">Material examined. –</emphasis>
<specimenCount id="9D79FD28A73AFFB03D00FB93BB664C58" box="[1027,1144,1044,1065]" count="3" pageId="21" pageNumber="118" type="generic">3 specimens</specimenCount>
–
<materialsCitation id="3B173CFCA73AFFB03D8DFB93BBBF4C34" collectingDate="1972-01-10" collectionCode="CHF" collectorName="Graf &amp; Sungai Buloh" country="Singapore" location="Singapore" pageId="21" pageNumber="118" specimenCount="1">
<specimenCount id="9D79FD28A73AFFB03D8DFB93BBA04C58" box="[1166,1214,1044,1065]" count="1" pageId="21" pageNumber="118" type="generic">1 ex.</specimenCount>
,
<collectingCountry id="F3687631A73AFFB03DC5FB93BA344C58" box="[1222,1322,1044,1065]" name="Singapore" pageId="21" pageNumber="118">Singapore</collectingCountry>
,
<date id="FFC11061A73AFFB03C31FB93BAB64C58" box="[1330,1448,1044,1065]" pageId="21" pageNumber="118" value="1972-01-10">
<collectingDate id="EF85E989A73AFFB03C31FB93BAB64C58" box="[1330,1448,1044,1065]" pageId="21" pageNumber="118" value="1972-01-10">10 Jan.1972</collectingDate>
</date>
, T3751, coll.
<collectorName id="268A5377A73AFFB03AA1FBB6BCD34C37" box="[930,973,1073,1094]" pageId="21" pageNumber="118">Gräf</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03ADAFBB6BB124C37" box="[985,1036,1073,1094]" pageId="21" pageNumber="118">CHF</collectionCode>
).
<emphasis id="B90BEAB3A73AFFB03D19FBB7BBBF4C34" bold="true" box="[1050,1185,1072,1093]" pageId="21" pageNumber="118">
<collectorName id="268A5377A73AFFB03D19FBB7BBBF4C34" box="[1050,1185,1072,1093]" pageId="21" pageNumber="118">Sungai Buloh</collectorName>
</emphasis>
</materialsCitation>
-
<materialsCitation id="3B173CFCA73AFFB03DB2FBB6BA804C13" collectingDate="1990-12-04" collectionCode="ZRC" collectorName="C. M. Yang &amp; K. L. Yeo" country="Singapore" location="Nature Park" pageId="21" pageNumber="118" specimenCount="2">
<specimenCount id="9D79FD28A73AFFB03DB2FBB6BBFE4C37" box="[1201,1248,1073,1094]" count="2" pageId="21" pageNumber="118" type="generic">2 ex.</specimenCount>
,
<location id="8EA0607AA73AFFB03DEBFBB6BA454C37" LSID="urn:lsid:plazi:treatment:03D687B7A73AFFB03A91FCC0BCC34E6F:8EA0607AA73AFFB03DEBFBB6BA454C37" box="[1256,1371,1073,1094]" country="Singapore" name="Nature Park" pageId="21" pageNumber="118">Nature Park</location>
, shallow pool, No.1,
<date id="FFC11061A73AFFB03AA0FBCABB094C13" box="[931,1047,1101,1122]" pageId="21" pageNumber="118" value="1990-12-04">
<collectingDate id="EF85E989A73AFFB03AA0FBCABB094C13" box="[931,1047,1101,1122]" pageId="21" pageNumber="118" value="1990-12-04">4 Dec.1990</collectingDate>
</date>
, coll.
<collectorName id="268A5377A73AFFB03D57FBCABBD74C13" box="[1108,1225,1101,1122]" pageId="21" pageNumber="118">C. M. Yang</collectorName>
&amp;
<collectorName id="268A5377A73AFFB03DE9FBCABA534C13" box="[1258,1357,1101,1122]" pageId="21" pageNumber="118">K. L. Yeo</collectorName>
(
<collectionCode id="ED6EAE64A73AFFB03C60FBCABA864C13" box="[1379,1432,1101,1122]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="118">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03A2EFB00BA7F4C8E" pageId="21" pageNumber="118" type="distribution">
<paragraph id="8BC036A1A73AFFB03A2EFB00BA7F4C8E" blockId="21.[813,1451,1159,1279]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A2EFB00BCFD4CEE" bold="true" box="[813,995,1159,1183]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A2EFB00BCDC4CEE" bold="true" box="[813,962,1159,1183]" italics="true" pageId="21" pageNumber="118">Distribution</emphasis>
. –
</emphasis>
Northern Australia, India, Thailand, Peninsular Malaysia, Indonesia [Sumatra, Java, Bali], Laos, Myanmar, Vietnam (
<bibRefCitation id="EFEE4B50A73AFFB03D17FB40BBEC4CAF" author="Zimmermann, A" box="[1044,1266,1223,1246]" pageId="21" pageNumber="118" pagination="1 - 326" refId="ref42405" refString="Zimmermann, A., 1920. Dytiscidae. In: Junk, W. &amp; S. Schenkling (eds.), Coleopterorum Catatalogus, Berlin, 4, pars 71, 1 - 326." type="journal article" year="1920">Zimmermann, 1920</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB03C02FB40BAB94CAF" author="Vazirani, T" box="[1281,1447,1223,1246]" pageId="21" pageNumber="118" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73AFFB03A2EFB60BCAD4C8F" author="Watts, C" box="[813,947,1255,1278]" pageId="21" pageNumber="118" pagination="1 - 166" refId="ref41518" refString="Watts, C. H. S., 1978. A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology, Supplements Series, 57: 1 - 166." type="journal article" year="1978">Watts, 1978</bibRefCitation>
; Hendrich &amp; Balke, 1995).
<emphasis id="B90BEAB3A73AFFB03DE0FB60BA7F4C8E" bold="true" box="[1251,1377,1255,1279]" pageId="21" pageNumber="118">
<collectingCountry id="F3687631A73AFFB03DE0FB60BA424C8E" box="[1251,1372,1255,1279]" name="Singapore" pageId="21" pageNumber="118">Singapore</collectingCountry>
!
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73AFFB03A2EFAA0BCC34E6F" pageId="21" pageNumber="118" type="biology_ecology">
<paragraph id="8BC036A1A73AFFB03A2EFAA0BCC34E6F" blockId="21.[813,1452,1319,1566]" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A2EFAA0BB8E4D4F" box="[813,1168,1319,1343]" italics="true" pageId="21" pageNumber="118">
<emphasis id="B90BEAB3A73AFFB03A2EFAA0BCBF4D4E" bold="true" box="[813,929,1319,1343]" italics="true" pageId="21" pageNumber="118">Ecology. –</emphasis>
<taxonomicName id="4C7F4D22A73AFFB03AA4FAA0BB8E4D4F" authorityName="Regimbart" authorityYear="1880" box="[935,1168,1319,1342]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="tenebrosus">Copelatus tenebrosus</taxonomicName>
</emphasis>
inhabits different kinds of standing waters in open, exposed areas, such as pools, ponds and irrigation ditches. The species is often attracted to light and is the most common species of the genus can be found in deforested and cultivated areas in Southeast Asia. In
<collectingCountry id="F3687631A73AFFB03A2EFA40BC874DAF" box="[813,921,1479,1502]" name="Indonesia" pageId="21" pageNumber="118">Indonesia</collectingCountry>
,
<collectingCountry id="F3687631A73AFFB03AA7FA40BB1A4DAF" box="[932,1028,1479,1502]" name="Vietnam" pageId="21" pageNumber="118">Vietnam</collectingCountry>
,
<collectingCountry id="F3687631A73AFFB03D0CFA40BB714DAF" box="[1039,1135,1479,1502]" name="Thailand" pageId="21" pageNumber="118">Thailand</collectingCountry>
and in the
<collectingCountry id="F3687631A73AFFB03DE9FA40BA7D4DAF" box="[1258,1379,1479,1502]" name="Philippines" pageId="21" pageNumber="118">Philippines</collectingCountry>
it was also recorded from paddy fields (
<bibRefCitation id="EFEE4B50A73AFFB03D83FA60BA5F4D8F" author="Yano, K &amp; Chu, P" box="[1152,1345,1511,1534]" pageId="21" pageNumber="118" pagination="103 - 118" refId="ref42131" refString="Yano, K., Y. I. Chu, P. W. Resma, &amp; M. Sato, 1983 b. Faunal and biological studies on the insects of paddy fields in Asia. XII. Aquatic Coleoptera from Taiwan and the Philippines. Chinese Journal of Entomology, 3 (2): 103 - 118." type="journal article" year="1983">Yano et al., 1983b</bibRefCitation>
; Hendrich &amp; Balke, 1995).
</paragraph>
</subSubSection>
</treatment>
</document>
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