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<document id="305FE8B6ECC81640B0C8206251B7A9A7" ID-CLB-Dataset="7679" ID-DOI="10.11646/zootaxa.4942.1.2" ID-GBIF-Dataset="25d73144-c90c-45c3-a9a2-2454f87a9aef" ID-ISSN="1175-5326" ID-Zenodo-Dep="4596097" ID-ZooBank="79353662-7653-4F41-8B39-40E6E4B2E005" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615448142785" checkinUser="plazi" docAuthor="Haddad, Charles R., Jin, Chi, Platnick, Norman I. &amp; Booysen, Ruan" docDate="2021" docId="03D67155FFBEFFB4FF25FAE4A51A2BE9" docLanguage="en" docName="zootaxa.4942.1.2.pdf" docOrigin="Zootaxa 4942 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Capobula neethlingi Haddad &amp; Jin &amp; Platnick &amp; Booysen 2021, spec. nov." docType="treatment" docVersion="7" lastPageNumber="59" masterDocId="FFEF092DFFAFFFA6FFB2FFBAA0202918" masterDocTitle="Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae)" masterLastPageNumber="71" masterPageNumber="41" pageNumber="58" updateTime="1698933034801" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C5E9FC6CBE4E379D650761EC1B86ED00">Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="90477B3AB9356752FDE71385F820CE83">Haddad, Charles R.</mods:namePart>
<mods:nameIdentifier id="8662511D68850960B0B82308D4A020B9" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
<mods:affiliation id="C73DB13C851033F727385AA90A5A14F5">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa. &amp; haddadcr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760; &amp; booysenr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 0841 - 9143</mods:affiliation>
<mods:nameIdentifier id="70076AFE4FE93589BACDF606F756C148" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
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<mods:namePart id="03724B65FC803F811DC413F2491B7FF1">Jin, Chi</mods:namePart>
<mods:nameIdentifier id="18060855B24EE320F2ACCD3A2A0D193B" type="ORCID">0000-0002-4694-4870</mods:nameIdentifier>
<mods:affiliation id="2CD356E5613E5860B3909A1B7FD983C6">College of Landscape and Ecological Engineering, Hebei University of Engineering, Handan, Hebei 056038, P. R. China. &amp; jinchi _ spider @ 163. com; https: // orcid. org / 0000 - 0002 - 4694 - 4870</mods:affiliation>
</mods:name>
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<mods:role id="F9A011CDC34D30230792731E5D24BC0A">
<mods:roleTerm id="39F72EAA89A287DDD9FA37DC9DCEA295">Author</mods:roleTerm>
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<mods:namePart id="CFC1BCB8EF8F3D2DD52777C9DE197CF0">Platnick, Norman I.</mods:namePart>
<mods:nameIdentifier id="8126C5DBA2EF9875237C6637C74E59F6" type="ORCID">0000-0003-4868-7529</mods:nameIdentifier>
<mods:affiliation id="E70E98E7CFDC2458CA58454B78D9176C">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 th Street, New York, NY 10024, U. S. A. &amp; https: // orcid. org / 0000 - 0003 - 4868 - 7529; deceased</mods:affiliation>
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<mods:roleTerm id="EB46DF79CFCB7B406D7F366256665819">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A4B98B09CBB73A8FC296F9206A559C1F">Booysen, Ruan</mods:namePart>
<mods:nameIdentifier id="2BAC8779315EFDC9966D810775EB4257" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
<mods:affiliation id="6333EA01C911F8405DB961EF8B4A3FE9">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa. &amp; haddadcr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760; &amp; booysenr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 0841 - 9143</mods:affiliation>
<mods:nameIdentifier id="E2F2501C2D4BD03DEC1031DA95BE9CAB" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
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<mods:title id="1B00B5DD7417EECB95CA27DBAF2735B7">Zootaxa</mods:title>
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<mods:part id="96DA8EEC1580CA32C4E2FCA400642135">
<mods:date id="06CD849E27FA3805E0833632B9D76850">2021</mods:date>
<mods:detail id="78D60256C10AD1B7EFADEEAF3D945988" type="pubDate">
<mods:number id="8F79C3E3CB77F0B458166349803C0F7A">2021-03-11</mods:number>
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<mods:number id="2F215EB71478DA933B938073D789C120">4942</mods:number>
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<mods:number id="D51E1C28CCE696C295511E96FD9B5013">1</mods:number>
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<mods:classification id="F5F632385CB0DE4FFFFB90A0BB207436">journal article</mods:classification>
<mods:identifier id="BA896612328D53E4AE307B3E94C84E9B" type="CLB-Dataset">7679</mods:identifier>
<mods:identifier id="D26DBD7D4E6F0E2D2883B81764A0E36A" type="DOI">10.11646/zootaxa.4942.1.2</mods:identifier>
<mods:identifier id="917EEDF0B39305D6C2579896B8E4FAA6" type="GBIF-Dataset">25d73144-c90c-45c3-a9a2-2454f87a9aef</mods:identifier>
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<treatment id="03D67155FFBEFFB4FF25FAE4A51A2BE9" ID-DOI="http://doi.org/10.5281/zenodo.4618483" ID-GBIF-Taxon="180588634" ID-Zenodo-Dep="4618483" LSID="urn:lsid:plazi:treatment:03D67155FFBEFFB4FF25FAE4A51A2BE9" httpUri="http://treatment.plazi.org/id/03D67155FFBEFFB4FF25FAE4A51A2BE9" lastPageId="18" lastPageNumber="59" pageId="17" pageNumber="58">
<subSubSection id="C36593C8FFBEFFB7FF25FAE4A22D2C61" box="[151,525,1374,1401]" pageId="17" pageNumber="58" type="nomenclature">
<paragraph id="8BC0C043FFBEFFB7FF25FAE4A22D2C61" blockId="17.[151,525,1374,1437]" box="[151,525,1374,1401]" pageId="17" pageNumber="58">
<heading id="D088772FFFBEFFB7FF25FAE4A22D2C61" bold="true" box="[151,525,1374,1401]" fontSize="11" level="1" pageId="17" pageNumber="58" reason="1">
<emphasis id="B90B1C51FFBEFFB7FF25FAE4A22D2C61" bold="true" box="[151,525,1374,1401]" pageId="17" pageNumber="58">
<taxonomicName id="4C7FBBC0FFBEFFB7FF25FAE4A1AE2C61" ID-CoL="98Z27" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[151,398,1374,1401]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="neethlingi" status="sp. nov.">
<emphasis id="B90B1C51FFBEFFB7FF25FAE4A1AE2C61" bold="true" box="[151,398,1374,1401]" italics="true" pageId="17" pageNumber="58">Capobula neethlingi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238A12AFFBEFFB7FE27FAE5A22D2C61" box="[405,525,1375,1401]" pageId="17" pageNumber="58" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBEFFB7FF25FA39A1102C85" box="[151,304,1411,1437]" pageId="17" pageNumber="58" type="description">
<paragraph id="8BC0C043FFBEFFB7FF25FA39A1102C85" blockId="17.[151,525,1374,1437]" box="[151,304,1411,1437]" pageId="17" pageNumber="58">
<figureCitation id="1344DCC6FFBEFFB7FF25FA39A0FE2C85" box="[151,222,1411,1437]" captionStart="FIGURES 3–10" captionStartId="5.[151,264,1869,1893]" captionTargetBox="[151,1436,946,1846]" captionTargetId="figure-265@5.[151,1436,946,1846]" captionTargetPageId="5" captionText="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." figureDoi="http://doi.org/10.5281/zenodo.4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="17" pageNumber="58">Figs 9</figureCitation>
,
<figureCitation id="1344DCC6FFBEFFB7FF58FA39A1102C85" box="[234,304,1411,1437]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="17" pageNumber="58">71, 72</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBEFFB7FF25FA71A1322F81" pageId="17" pageNumber="58" type="materials_examined">
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<emphasis id="B90B1C51FFBEFFB7FF25FA71A1642CFC" bold="true" box="[151,324,1482,1509]" pageId="17" pageNumber="58">Type material.</emphasis>
<materialsCitation id="3B17CA1EFFBEFFB7FEFBFA71A48C2F11" ID-GBIF-Occurrence="3053986353" collectingDate="2012-12-07" collectionCode="NCA" collectorName="J. Neethling" country="South Africa" elevation="149" latitude="-33.969967" location="Saasveld Pass" longLatPrecision="1" longitude="22.529633" municipality="George" pageId="17" pageNumber="58" specimenCode="NCA 2019" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="holotype">
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<typeStatus id="54C47EE1FFBEFFB7FEFBFA71A1962CFD" box="[329,438,1483,1509]" pageId="17" pageNumber="58" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D790BCAFFBEFFB7FE09FA71A1F22CFD" box="[443,466,1483,1509]" pageId="17" pageNumber="58" type="female">♀</specimenCount>
:
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<collectingCountry id="F36880D3FFBEFFB7FE6CFA70A2952CFC" box="[478,693,1482,1508]" name="South Africa" pageId="17" pageNumber="58">SOUTH AFRICA</collectingCountry>
:
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<collectingRegion id="49BB0EA1FFBEFFB7FD72FA71A3792CFD" box="[704,857,1483,1509]" country="South Africa" name="Western Cape" pageId="17" pageNumber="58">Western Cape</collectingRegion>
</emphasis>
:
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,
<location id="8EA09698FFBEFFB7FC77FA71A4432CFD" LSID="urn:lsid:plazi:treatment:03D67155FFBEFFB4FF25FAE4A51A2BE9:8EA09698FFBEFFB7FC77FA71A4432CFD" box="[965,1123,1483,1509]" country="South Africa" latitude="-33.969967" longLatPrecision="1" longitude="22.529633" municipality="George" name="Saasveld Pass" pageId="17" pageNumber="58" stateProvince="Western Cape">Saasveld Pass</location>
,
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,
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,
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a.s.l.
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,
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<collectingDate id="EF851F6BFFBEFFB7FE9EFA55A18A2F11" box="[300,426,1519,1545]" pageId="17" pageNumber="58" value="2012-12-07">7.XII.2012</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBEFFB7FE5BFA55A2512F11" box="[489,625,1519,1545]" pageId="17" pageNumber="58">J. Neethling</collectorName>
(leaf litter, indigenous forest) (
<specimenCode id="DBD96838FFBEFFB7FC65FA55A4762F11" box="[983,1110,1519,1545]" collectionCode="NCA" pageId="17" pageNumber="58">NCA 2019</specimenCode>
/1002a)
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.
<materialsCitation id="3B17CA1EFFBEFFB7FB0AFA55A5BC2F4A" ID-GBIF-Occurrence="3053986366" collectingDate="2001-01-15" collectingDateMax="2015-10-08" collectingDateMin="2001-01-15" collectionCode="NCA" collectorName="Z. Mbo" country="South Africa" county="Laingsburg" elevation="735" latitude="-33.35" location="Swartberg Nature Reserve" longLatPrecision="1205" longitude="21.683332" municipality="Anysberg Nature Reserve" pageId="17" pageNumber="58" specimenCode="NCA 2019, NCA 2016, NCA 2002" specimenCount="3" specimenCount-female="3" stateProvince="Western Cape" typeStatus="paratype">
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<typeStatus id="54C47EE1FFBEFFB7FB0AFA55A5102F11" box="[1208,1328,1519,1545]" pageId="17" pageNumber="58" type="paratype">Paratypes</typeStatus>
</emphasis>
: together with
<typeStatus id="54C47EE1FFBEFFB7FF63F9A9A1162F35" box="[209,310,1555,1581]" pageId="17" pageNumber="58" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D790BCAFFBEFFB7FEF0F9A9A1442F35" box="[322,356,1555,1581]" pageId="17" pageNumber="58" type="female">1♀</specimenCount>
(
<specimenCode id="DBD96838FFBEFFB7FEC6F9A9A1D22F36" box="[372,498,1555,1582]" collectionCode="NCA" pageId="17" pageNumber="58">NCA 2019</specimenCode>
/1002b).
<emphasis id="B90B1C51FFBEFFB7FDE7F9A8A3132F34" bold="true" box="[597,819,1554,1580]" pageId="17" pageNumber="58">
<collectingCountry id="F36880D3FFBEFFB7FDE7F9A8A30E2F34" box="[597,814,1554,1580]" name="South Africa" pageId="17" pageNumber="58">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="B90B1C51FFBEFFB7FC89F9A9A3F72F35" box="[827,983,1555,1581]" italics="true" pageId="17" pageNumber="58">
<collectingRegion id="49BB0EA1FFBEFFB7FC89F9A9A3F72F35" box="[827,983,1555,1581]" country="South Africa" name="Western Cape" pageId="17" pageNumber="58">Western Cape</collectingRegion>
</emphasis>
:
<collectingCounty id="62A1B8CFFFBEFFB7FC55F9A9A4492F35" box="[999,1129,1555,1581]" pageId="17" pageNumber="58">Laingsburg</collectingCounty>
,
<collectingMunicipality id="6BA45A39FFBEFFB7FBC1F9A9A5B82F35" box="[1139,1432,1555,1581]" pageId="17" pageNumber="58">Anysberg Nature Reserve</collectingMunicipality>
,
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,
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,
<elevation id="00522770FFBEFFB7FE62F98CA2752F49" box="[464,597,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="17" pageNumber="58" unit="m" value="735.0">
<quantity id="4C876DA6FFBEFFB7FE62F98CA2382F49" box="[464,536,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="17" pageNumber="58" unit="m" value="735.0">735 m</quantity>
a.s.l.
</elevation>
,
<date id="FFC1E683FFBEFFB7FDD0F98CA32D2F49" box="[610,781,1590,1617]" pageId="17" pageNumber="58" value="2015-09-08" valueMax="2015-10-08" valueMin="2015-09-08">
<collectingDate id="EF851F6BFFBEFFB7FDD0F98CA32D2F49" box="[610,781,1590,1617]" pageId="17" pageNumber="58" value="2015-09-08" valueMax="2015-10-08" valueMin="2015-09-08">8.IX–8.X.2015</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBEFFB7FCF9F98DA3BF2F49" box="[843,927,1591,1617]" pageId="17" pageNumber="58">Z. Mbo</collectorName>
(
<collectingMethod id="523EB854FFBEFFB7FC1DF98DA4142F49" box="[943,1076,1591,1617]" pageId="17" pageNumber="58">pitfall traps</collectingMethod>
, karoo veld) (
<specimenCode id="DBD96838FFBEFFB7FB62F98DA56D2F4A" box="[1232,1357,1591,1618]" collectionCode="NCA" pageId="17" pageNumber="58">NCA 2016</specimenCode>
/2464);
</materialsCitation>
<materialsCitation id="3B17CA1EFFBEFFB7FF25F9E1A12E2F81" ID-GBIF-Occurrence="3053986318" collectingDate="2001-01-15" collectingDateMax="2015-10-08" collectingDateMin="2001-01-15" collectionCode="NCA" collectorName="Z. van der Walt" country="South Africa" county="Laingsburg" elevation="735" latitude="-33.35" location="Swartberg Nature Reserve" longLatPrecision="1205" longitude="21.683332" municipality="Anysberg Nature Reserve" pageId="17" pageNumber="58" specimenCode="NCA 2019, NCA 2016, NCA 2002" specimenCount="3" specimenCount-female="3" stateProvince="Western Cape" typeStatus="paratype">
<location id="8EA09698FFBEFFB7FF25F9E1A1E02F6D" LSID="urn:lsid:plazi:treatment:03D67155FFBEFFB4FF25FAE4A51A2BE9:8EA09698FFBEFFB7FF25F9E1A1E02F6D" box="[151,448,1627,1653]" country="South Africa" county="Laingsburg" latitude="-33.35" longLatPrecision="1205" longitude="21.683332" municipality="Anysberg Nature Reserve" name="Swartberg Nature Reserve" pageId="17" pageNumber="58" stateProvince="Western Cape">Swartberg Nature Reserve</location>
,
<collectorName id="268AA595FFBEFFB7FE79F9E1A2452F6D" box="[459,613,1627,1653]" pageId="17" pageNumber="58">Gamkaskloof</collectorName>
,
<geoCoordinate id="EE4BA684FFBEFFB7FDDDF9E1A2ED2F6D" box="[623,717,1627,1653]" degrees="33" direction="south" minutes="21" orientation="latitude" pageId="17" pageNumber="58" precision="925" value="-33.35">33°21’S</geoCoordinate>
,
<geoCoordinate id="EE4BA684FFBEFFB7FD6AF9E1A3182F6D" box="[728,824,1627,1653]" degrees="21" direction="east" minutes="41" orientation="longitude" pageId="17" pageNumber="58" precision="925" value="21.683332">21°41’E</geoCoordinate>
,
<date id="FFC1E683FFBEFFB7FCF6F9E0A3912F6D" box="[836,945,1626,1653]" pageId="17" pageNumber="58" value="2001-01-15">
<collectingDate id="EF851F6BFFBEFFB7FCF6F9E0A3912F6D" box="[836,945,1626,1653]" pageId="17" pageNumber="58" value="2001-01-15">15.I.2001</collectingDate>
</date>
, leg.
<collectorName id="268AA595FFBEFFB7FC5EF9E1A4B72F6D" box="[1004,1175,1627,1653]" pageId="17" pageNumber="58">Z. van der Walt</collectorName>
(on ground),
<specimenCount id="9D790BCAFFBEFFB7FA82F9E1A5732F6D" box="[1328,1363,1627,1653]" pageId="17" pageNumber="58" type="female">2♀</specimenCount>
(
<specimenCode id="DBD96838FFBEFFB7FAD3F9E1A0F32F81" collectionCode="NCA" pageId="17" pageNumber="58">NCA 2002</specimenCode>
/198)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBEFFB7FF75F918A1682E55" pageId="17" pageNumber="58" type="diagnosis">
<paragraph id="8BC0C043FFBEFFB7FF75F918A1682E55" blockId="17.[151,1437,1482,2013]" pageId="17" pageNumber="58">
<emphasis id="B90B1C51FFBEFFB7FF75F918A1622FA4" bold="true" box="[199,322,1698,1724]" pageId="17" pageNumber="58">Diagnosis.</emphasis>
Females of
<taxonomicName id="4C7FBBC0FFBEFFB7FE7BF919A2792FA5" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[457,601,1699,1725]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="neethlingi" status="sp. nov.">
<emphasis id="B90B1C51FFBEFFB7FE7BF919A2792FA5" box="[457,601,1699,1725]" italics="true" pageId="17" pageNumber="58">C. neethlingi</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFBEFFB7FDD2F919A2F12FA5" bold="true" box="[608,721,1699,1725]" pageId="17" pageNumber="58">
<taxonomicNameLabel id="A238A12AFFBEFFB7FDD2F919A2F12FA5" box="[608,721,1699,1725]" pageId="17" pageNumber="58" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
have a similar epigyne to that of
<taxonomicName id="4C7FBBC0FFBEFFB7FBF1F919A4E62FA4" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[1091,1222,1699,1725]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="montana" status="sp. nov.">
<emphasis id="B90B1C51FFBEFFB7FBF1F919A4E62FA4" box="[1091,1222,1699,1725]" italics="true" pageId="17" pageNumber="58">C. montana</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFBEFFB7FB7EF919A51E2FA5" bold="true" box="[1228,1342,1699,1725]" pageId="17" pageNumber="58">
<taxonomicNameLabel id="A238A12AFFBEFFB7FB7EF919A51E2FA5" box="[1228,1342,1699,1725]" pageId="17" pageNumber="58" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished by the V- rather than J-shaped ridges containing the copulatory openings, the slightly separated copulatory ducts (touching medially in
<taxonomicName id="4C7FBBC0FFBEFFB7FDFEF951A2FC2E1D" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[588,732,1771,1797]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="neethlingi" status="sp. nov.">
<emphasis id="B90B1C51FFBEFFB7FDFEF951A2FC2E1D" box="[588,732,1771,1797]" italics="true" pageId="17" pageNumber="58">C. neethlingi</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFBEFFB7FD50F951A3732E1D" bold="true" box="[738,851,1771,1797]" pageId="17" pageNumber="58">
<taxonomicNameLabel id="A238A12AFFBEFFB7FD50F951A3732E1D" box="[738,851,1771,1797]" pageId="17" pageNumber="58" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
), and the bursae that are separated by approximately half their diameter, while almost touching in
<taxonomicName id="4C7FBBC0FFBEFFB7FD38F8B5A33A2E31" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[650,794,1807,1833]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="neethlingi" status="sp. nov.">
<emphasis id="B90B1C51FFBEFFB7FD38F8B5A33A2E31" box="[650,794,1807,1833]" italics="true" pageId="17" pageNumber="58">C. neethlingi</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFBEFFB7FC92F8B5A3B12E31" bold="true" box="[800,913,1807,1833]" pageId="17" pageNumber="58">
<taxonomicNameLabel id="A238A12AFFBEFFB7FC92F8B5A3B12E31" box="[800,913,1807,1833]" pageId="17" pageNumber="58" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(compare
<figureCitation id="1344DCC6FFBEFFB7FBB5F8B5A48B2E31" box="[1031,1195,1807,1833]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="17" pageNumber="58">Figs 71 and 72</figureCitation>
with
<figureCitation id="1344DCC6FFBEFFB7FB5BF8B5A5B02E32" box="[1257,1424,1806,1834]" captionStart="FIGURES 65–70" captionStartId="16.[151,264,1820,1844]" captionTargetBox="[193,1395,1034,1795]" captionTargetId="figure-375@16.[189,1398,1029,1797]" captionTargetPageId="16" captionText="FIGURES 65–70. Capobula montana spec. nov., genitalic morphology of female (65, 66) and male (67–70) paratypes from Platberg Nature Reserve: 65 Epigyne, ventral; 66 Vulva, dorsal; 67–69 Left palp (67 prolateral, 68 retrolateral, 69 retrolateral); 70 Palpal femur, patella and tibia, retro-dorsal. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4596123" httpUri="https://zenodo.org/record/4596123/files/figure.png" pageId="17" pageNumber="58">Figs 65 and 66</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBEFFB4FF75F8EDA1E32BD5" lastPageId="18" lastPageNumber="59" pageId="17" pageNumber="58" type="etymology">
<paragraph id="8BC0C043FFBEFFB7FF75F8EDA0EF2E8D" blockId="17.[151,1437,1482,2013]" pageId="17" pageNumber="58">
<emphasis id="B90B1C51FFBEFFB7FF75F8EDA16E2E69" bold="true" box="[199,334,1879,1905]" pageId="17" pageNumber="58">Etymology.</emphasis>
Named for arachnologist Jan Andries Neethling, who collected the
<typeStatus id="54C47EE1FFBEFFB7FBD5F8EDA4ED2E69" box="[1127,1229,1879,1905]" pageId="17" pageNumber="58" type="holotype">holotype</typeStatus>
; name in genitive case.
</paragraph>
<paragraph id="8BC0C043FFBEFFB4FF75F825A20229CD" blockId="17.[151,1437,1482,2013]" lastBlockId="18.[151,1437,150,753]" lastPageId="18" lastPageNumber="59" pageId="17" pageNumber="58">
<emphasis id="B90B1C51FFBEFFB7FF75F825A2532EA0" bold="true" box="[199,627,1950,1977]" pageId="17" pageNumber="58">
Female (
<typeStatus id="54C47EE1FFBEFFB7FE9DF824A1B92EA0" box="[303,409,1950,1976]" pageId="17" pageNumber="58" type="holotype">holotype</typeStatus>
, NCA 2019/1002).
</emphasis>
Measurements: CL 0.84, CW 0.64, AL 1.02, AW 0.87, TL 1.95 (1.80– 1.98), PERW 0.30, MOQAW 0.12, MOQPW 0.15, MOQL 0.16. Length of leg segments: I 0.60 + 0.24 + 0.52 + 0.45 + 0.25 = 2.06; II 0.54 + 0.23 + 0.41 + 0.38 + 0.24 = 1.80; III 0.46 + 0.21 + 0.33 + 0.38 + 0.23 = 1.61; IV 0.59 + 0.24 + 0.49 + 0.56 + 0.27 = 2.15.
</paragraph>
<paragraph id="8BC0C043FFBDFFB4FF75FF65A39028B5" blockId="18.[151,1437,150,753]" pageId="18" pageNumber="59">Colour: carapace deep orange-brown, with black mottling at centre, pits and lateral margins with black edges; chelicerae deep yellow-brown, with faint black mottling; endites and labium yellow-brown, cream at distal ends; sternum bright orange, pits slightly darker, lateral margins orange-brown; palps pale yellow-brown; legs with femora and tibiae I and II pale orange-brown, yellow-brown distally; patellae, metatarsi and tarsi I and II, and legs III and IV yellow-brown; abdomen dark grey dorsally and laterally, dorsally with eight fine cream chevrons in posterior half to spinnerets; venter slightly paler mottled grey; spinnerets cream.</paragraph>
<paragraph id="8BC0C043FFBDFFB4FF75FE0DA26028ED" blockId="18.[151,1437,150,753]" pageId="18" pageNumber="59">Leg spination: femora and patellae: spineless; tibiae: I plv 6 rlv 6, II plv 6 rlv 5; metatarsi: I plv 4 rlv 4, II plv 4 rlv 4; tarsi: I plv 3 rlv 3, II plv 3 rlv 2.</paragraph>
<paragraph id="8BC0C043FFBDFFB4FF75FE45A1802B9D" blockId="18.[151,1437,150,753]" pageId="18" pageNumber="59">
Epigyne with small copulatory openings in J-shaped epigynal ridges (
<figureCitation id="1344DCC6FFBDFFB4FC6AFE45A40B2B01" box="[984,1067,511,537]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="18" pageNumber="59">Fig. 71</figureCitation>
); copulatory ducts initially curving medially, then posteriorly, looping dorsally and anterolaterally before entering teardrop-shaped primary spermathecae along their interior margin; bursae subtriangular, apices converging mesally, similar in size to primary spermathecae (
<figureCitation id="1344DCC6FFBDFFB4FE8DFDD1A1B32B9D" box="[319,403,619,645]" captionStart="FIGURES 71–74" captionStartId="19.[151,264,1279,1303]" captionTargetBox="[196,1395,184,1245]" captionTargetId="figure-18@19.[189,1398,181,1255]" captionTargetPageId="19" captionText="FIGURES 71–74. Capobula neethlingi spec. nov. (71, 72) and C. ukhahlamba spec. nov. (73, 74), genitalic morphology of paratype females from Anysberg Nature Reserve and Cathedral Peak, respectively: 71, 73 Epigyne, ventral; 72, 74 Vulva, dorsal. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4596125" httpUri="https://zenodo.org/record/4596125/files/figure.png" pageId="18" pageNumber="59">Fig. 72</figureCitation>
).
</paragraph>
<paragraph id="8BC0C043FFBDFFB4FF75FD34A1E32BD5" blockId="18.[151,1437,150,753]" pageId="18" pageNumber="59">
<emphasis id="B90B1C51FFBDFFB4FF75FD34A19A2BB0" bold="true" box="[199,442,654,681]" pageId="18" pageNumber="59">Habitat and biology.</emphasis>
This species was recorded in two very contrasting biotopes,
<emphasis id="B90B1C51FFBDFFB4FBD3FD35A4A02BB0" box="[1121,1152,655,680]" italics="true" pageId="18" pageNumber="59">viz</emphasis>
. moist Afromontane Forest and xeric Nama Karoo.
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFBDFFB4FF75FD6CA51A2BE9" box="[199,1338,726,753]" pageId="18" pageNumber="59" type="distribution">
<paragraph id="8BC0C043FFBDFFB4FF75FD6CA51A2BE9" blockId="18.[151,1437,150,753]" box="[199,1338,726,753]" pageId="18" pageNumber="59">
<emphasis id="B90B1C51FFBDFFB4FF75FD6CA1412BE8" bold="true" box="[199,353,726,752]" pageId="18" pageNumber="59">Distribution.</emphasis>
Only known from three localities in the
<collectingRegion id="49BB0EA1FFBDFFB4FC9AFD6DA4102BE9" box="[808,1072,727,753]" country="South Africa" name="Western Cape" pageId="18" pageNumber="59">Western Cape Province</collectingRegion>
,
<collectingCountry id="F36880D3FFBDFFB4FB8EFD6DA4EB2BE9" box="[1084,1227,727,753]" name="South Africa" pageId="18" pageNumber="59">South Africa</collectingCountry>
(
<figureCitation id="1344DCC6FFBDFFB4FB6BFD6DA50E2BE9" box="[1241,1326,726,753]" captionStart="FIGURE 75" captionStartId="20.[151,250,1541,1565]" captionTargetBox="[189,1398,561,1517]" captionTargetId="figure-129@20.[189,1398,561,1517]" captionTargetPageId="20" captionText="FIGURE 75. Distribution of Capobula capensis spec. nov. (circles), C. infima (Simon, 1896) comb. nov. (triangles), C. montana spec. nov. (squares), C. neethlingi spec. nov. (stars) and C. ukhahlamba spec. nov. (inverted triangles) in southern Africa." figureDoi="http://doi.org/10.5281/zenodo.4596127" httpUri="https://zenodo.org/record/4596127/files/figure.png" pageId="18" pageNumber="59">Fig. 75</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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