348 lines
60 KiB
XML
348 lines
60 KiB
XML
<document id="AA5631E1B96E2F603ECBB67EA7CF2DB3" ID-CLB-Dataset="53457" ID-DOI="10.11646/zootaxa.5293.3.6" ID-GBIF-Dataset="f5f379bf-cbea-409d-9910-95cc6187acf6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7961475" ID-ZooBank="8B6480D9-3A65-4319-BC4C-089F7AC6946F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1684826307319" checkinUser="plazi" docAuthor="Telnov, Dmitry & Gusakov, Alexey A." docDate="2023" docId="03D63A302539FFCA10EFF9D2FDBEF812" docLanguage="en" docName="zootaxa.5293.3.7.pdf" docOrigin="Zootaxa 5293 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holcopyge Champion 1890, syn. nov." docType="treatment" docVersion="5" lastPageNumber="545" masterDocId="FFEF42482538FFCE1078FFE9FFABFF87" masterDocTitle="Rediscovery of Tomosomus Motschulsky, 1855 (Coleoptera: Anthicidae) with redescription of the genus, a new genus-rank synonym, and a key to species" masterLastPageNumber="556" masterPageNumber="541" pageNumber="542" updateTime="1685386713976" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="548CE93104BB8B8D4B912A72B51DA916">Rediscovery of Tomosomus Motschulsky, 1855 (Coleoptera: Anthicidae) with redescription of the genus, a new genus-rank synonym, and a key to species</mods:title>
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<mods:namePart id="19FDE9E09C8F9E1A10F7B8AC2C7167BE">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="A3FF8AD71B595632E3D817A93F341C2E">Department of Life Sciences, Natural History Museum, SW 7 5 BD, London, United Kingdom & Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401, & Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004, Rīga, Latvia</mods:affiliation>
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<mods:namePart id="C3E429800F00A6314B86D3B92B705268">Gusakov, Alexey A.</mods:namePart>
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<mods:affiliation id="97FC6EC6984AA403474FA448AAF7748D">Zoological Museum, Moscow Lomonosov State University, 2 Bol’shaya Nikitskaya Str., 125009, Moscow, Russia https: // orcid. org / 0000 - 0001 - 7709 - 7515</mods:affiliation>
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<mods:title id="16DBFFE4E09856885B891502A6CCBD76">Zootaxa</mods:title>
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<mods:date id="E4DEA7A08174FE4DB26FD42190745D19">2023</mods:date>
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<subSubSection id="C365D8AD2539FFCF10EFF9D2FED3F915" pageId="1" pageNumber="542" type="nomenclature">
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<paragraph id="8BC08B262539FFCF10EFF9D2FDD3F9D2" blockId="1.[151,632,1594,1621]" box="[151,632,1594,1621]" pageId="1" pageNumber="542">
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<heading id="D0883C4A2539FFCF10EFF9D2FDD3F9D2" bold="true" box="[151,632,1594,1621]" fontSize="11" level="1" pageId="1" pageNumber="542" reason="1">
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<emphasis id="B90B57342539FFCF10EFF9D2FDD3F9D2" bold="true" box="[151,632,1594,1621]" pageId="1" pageNumber="542">
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=
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<taxonomicName id="4C7FF0A52539FFCF10D7F9D3FDAEF9D3" authority="Champion, 1890" authorityName="Champion" authorityYear="1890" box="[175,517,1594,1621]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus" status="syn. nov.">
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<emphasis id="B90B57342539FFCF10D7F9D3FE9BF9D2" bold="true" box="[175,304,1594,1621]" italics="true" pageId="1" pageNumber="542">Holcopyge</emphasis>
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<bibRefCitation id="EFEEF6D72539FFCF114FF9D3FDAEF9D3" author="Champion, G. C." box="[311,517,1594,1620]" pageId="1" pageNumber="542" pagination="292 - 293" refId="ref6084" refString="Champion, G. C. (1890) On a new genus of Anthicidae from tropical South America. The Entomologists Monthly Magazine, 26, 292 - 293." type="journal article" year="1890">Champion, 1890</bibRefCitation>
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</taxonomicName>
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<taxonomicNameLabel id="A238EA4F2539FFCF1274F9D2FDD3F9D2" box="[524,632,1595,1621]" pageId="1" pageNumber="542">syn. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC08B262539FFCF10EFF993FED3F915" blockId="1.[151,931,1658,1713]" box="[151,376,1658,1682]" pageId="1" pageNumber="542">
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<bibRefCitation id="EFEEF6D72539FFCF10EFF993FEDFF915" author="Champion, G. C." box="[151,372,1658,1682]" pageId="1" pageNumber="542" pagination="292 - 293" refId="ref6084" refString="Champion, G. C. (1890) On a new genus of Anthicidae from tropical South America. The Entomologists Monthly Magazine, 26, 292 - 293." type="journal article" year="1890">Champion (1890: 92)</bibRefCitation>
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.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C365D8AD2539FFCF10EFF970FC08F937" box="[151,931,1688,1713]" pageId="1" pageNumber="542" type="materials_examined">
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<paragraph id="8BC08B262539FFCF10EFF970FC08F937" blockId="1.[151,931,1658,1713]" box="[151,931,1688,1713]" pageId="1" pageNumber="542">
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<typeStatus id="54C435842539FFCF10EFF970FF62F936" box="[151,201,1689,1713]" pageId="1" pageNumber="542">Type</typeStatus>
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species:
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<taxonomicName id="4C7FF0A52539FFCF115EF971FD66F936" authority="Champion, 1890" authorityName="Champion" authorityYear="1890" box="[294,717,1688,1713]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="pallidicornis">
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<emphasis id="B90B57342539FFCF115EF971FDB7F937" box="[294,540,1688,1712]" italics="true" pageId="1" pageNumber="542">Holcopyge pallidicornis</emphasis>
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<bibRefCitation id="EFEEF6D72539FFCF125AF971FD66F936" author="Champion, G. C." box="[546,717,1688,1713]" pageId="1" pageNumber="542" pagination="292 - 293" refId="ref6084" refString="Champion, G. C. (1890) On a new genus of Anthicidae from tropical South America. The Entomologists Monthly Magazine, 26, 292 - 293." type="journal article" year="1890">Champion, 1890</bibRefCitation>
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</taxonomicName>
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, present designation.
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</paragraph>
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<subSubSection id="C365D8AD2539FFCF10EFF936FC98F856" pageId="1" pageNumber="542" type="reference_group">
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<emphasis id="B90B57342539FFCF10EFF936FE82F97E" box="[151,297,1759,1785]" italics="true" pageId="1" pageNumber="542">
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<taxonomicName id="4C7FF0A52539FFCF10EFF936FEA7F97E" authorityName="Champion" authorityYear="1890" box="[151,268,1759,1785]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">Holcopyge</taxonomicName>
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—
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</emphasis>
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<treatmentCitation id="0ADEAD372539FFCF1151F936FD81F97E" author="Champion, G. C." box="[297,554,1759,1785]" page="292" pageId="1" pageNumber="542" year="1890">
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<bibRefCitation id="EFEEF6D72539FFCF1151F936FD81F97E" author="Champion, G. C." box="[297,554,1759,1785]" pageId="1" pageNumber="542" pagination="292 - 293" refId="ref6084" refString="Champion, G. C. (1890) On a new genus of Anthicidae from tropical South America. The Entomologists Monthly Magazine, 26, 292 - 293." type="journal article" year="1890">Champion (1890: 292)</bibRefCitation>
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</treatmentCitation>
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||
original description, without subfamily placement, compared with
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||
<taxonomicName id="4C7FF0A52539FFCF1559F936FE06F89A" authority="LaFerte-Senectere, 1849" authorityName="LaFerte-Senectere" authorityYear="1849" class="Insecta" family="Anthicidae" genus="Tomoderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">
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<emphasis id="B90B57342539FFCF1559F936FA37F97E" box="[1313,1436,1759,1785]" italics="true" pageId="1" pageNumber="542">Tomoderus</emphasis>
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LaFerté-Sénectère, 1849
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||
</taxonomicName>
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||
(Tomoderinae Bonadona, 1961);
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||
<treatmentCitation id="0ADEAD372539FFCF134CF8EAFC42F89A" author="Pic, M." box="[820,1001,1795,1821]" page="213" pageId="1" pageNumber="542" year="1897">
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<bibRefCitation id="EFEEF6D72539FFCF134CF8EAFC42F89A" author="Pic, M." box="[820,1001,1795,1821]" pageId="1" pageNumber="542" pagination="212 - 224" refId="ref6661" refString="Pic, M. (1897) Premier supplement a ma liste des Anthicides (in Ann. Belg. 1894, pp. 43). Annales de la Societe Entomologique de Belgique, 41, 212 - 224." type="journal article" year="1897">Pic (1897: 213)</bibRefCitation>
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</treatmentCitation>
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note;
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||
<treatmentCitation id="0ADEAD372539FFCF144DF8EAFB71F89A" author="Pic, M." box="[1077,1242,1795,1821]" page="26" pageId="1" pageNumber="542" year="1911">
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<bibRefCitation id="EFEEF6D72539FFCF144DF8EAFB71F89A" author="Pic, M." box="[1077,1242,1795,1821]" pageId="1" pageNumber="542" pagination="102" refId="ref6702" refString="Pic, M. (1911) Anthicidae. Pars 36. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 102." type="book chapter" year="1911">Pic (1911: 26)</bibRefCitation>
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</treatmentCitation>
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||
checklist;
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||
<treatmentCitation id="0ADEAD372539FFCF1521F8EAFEB5F8C6" author="Lucas, R." page="330" pageId="1" pageNumber="542" year="1920">
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<bibRefCitation id="EFEEF6D72539FFCF1521F8EAFEB5F8C6" author="Lucas, R." pageId="1" pageNumber="542" refId="ref6481" refString="Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische Verlag, Berlin, 696 pp." type="book" year="1920">Lucas (1920: 330)</bibRefCitation>
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</treatmentCitation>
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||
checklist;
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||
<treatmentCitation id="0ADEAD372539FFCF11E7F8CEFD36F8C6" author="Heberdey, R. F." box="[415,669,1831,1857]" page="691" pageId="1" pageNumber="542" year="1938">
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<bibRefCitation id="EFEEF6D72539FFCF11E7F8CEFD36F8C6" author="Heberdey, R. F." box="[415,669,1831,1857]" pageId="1" pageNumber="542" pagination="667 - 734" refId="ref6389" refString="Heberdey, R. F. (1938) Beitrage zum Bau des Subelytralraumes und zur Atmung der Coleopteren. Zeitschrift fur Morphologie und Okologie der Tiere, 33 (4), 667 - 734. https: // doi. org / 10.1007 / BF 00407573" type="journal article" year="1938">Heberdey (1938: 691)</bibRefCitation>
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</treatmentCitation>
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||
functional morphology;
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||
<treatmentCitation id="0ADEAD372539FFCF13C6F8CEFB2EF8C6" author="Werner, F. G." box="[958,1157,1830,1857]" page="3" pageId="1" pageNumber="542" year="1965">
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<bibRefCitation id="EFEEF6D72539FFCF13C6F8CEFB2EF8C6" author="Werner, F. G." box="[958,1157,1830,1857]" pageId="1" pageNumber="542" pagination="1 - 15" refId="ref6863" refString="Werner, F. G. (1965) A key to the described species of New World Anthicidae of the genus Acanthinus LaFerte, with new synonymy (Coleoptera). Papeis Avulsos do Deptartamento de Zoologia Secretario da Agricultura, 18 (Article 1), 1 - 15." type="journal article" year="1965">Werner (1965: 3)</bibRefCitation>
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||
</treatmentCitation>
|
||
key to South American
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||
<taxonomicName id="4C7FF0A52539FFCF10EFF8A2FEB9F8E2" authorityName="Latreille" authorityYear="1819" box="[151,274,1867,1893]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
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genera;
|
||
<treatmentCitation id="0ADEAD372539FFCF110FF8A2FDE1F8E2" author="Werner, F. G." box="[375,586,1867,1893]" page="11" pageId="1" pageNumber="542" year="1966">
|
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<bibRefCitation id="EFEEF6D72539FFCF110FF8A2FDE1F8E2" author="Werner, F. G." box="[375,586,1867,1893]" pageId="1" pageNumber="542" pagination="9 - 23" refId="ref6915" refString="Werner, F. G. (1966) A preliminary account of the Anthicidae of Venezuela. Revista de la Facultad de Agronomia, III (4), 9 - 23." type="journal article" year="1966">Werner (1966: 11)</bibRefCitation>
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||
</treatmentCitation>
|
||
key to Venezuelan
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||
<taxonomicName id="4C7FF0A52539FFCF1356F8A2FC02F8E2" authorityName="Latreille" authorityYear="1819" box="[814,937,1867,1893]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
|
||
genera;
|
||
<treatmentCitation id="0ADEAD372539FFCF1477F8A2FAABF8E2" author="Uhmann, G." box="[1039,1280,1867,1893]" page="171" pageId="1" pageNumber="542" year="1976">
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||
<bibRefCitation id="EFEEF6D72539FFCF1477F8A2FAABF8E2" author="Uhmann, G." box="[1039,1280,1867,1893]" pageId="1" pageNumber="542" pagination="166 - 182" refId="ref6778" refString="Uhmann, G. (1976) Die Gattungen der Anthicidae und ihre systematische Anordnung. Beitrag zur Kenntnis der Anthicidae (Coleoptera). Entomologische Blatter, 72 (3), 166 - 182." type="journal article" year="1976">Uhmann (1976: 171)</bibRefCitation>
|
||
</treatmentCitation>
|
||
key to world
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||
<taxonomicName id="4C7FF0A52539FFCF10EFF886FEB9F80E" authorityName="Latreille" authorityYear="1819" box="[151,274,1903,1929]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
|
||
genera, erroneously arranged within
|
||
<taxonomicName id="4C7FF0A52539FFCF12C4F886FC82F80E" authorityName="Latreille" authorityYear="1819" box="[700,809,1903,1929]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="tribe" tribe="Anthicini">Anthicini</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0ADEAD372539FFCF134FF886FBBAF80E" author="Uhmann, G." box="[823,1041,1903,1929]" page="76" pageId="1" pageNumber="542" year="1978">
|
||
<bibRefCitation id="EFEEF6D72539FFCF134FF886FBBAF80E" author="Uhmann, G." box="[823,1041,1903,1929]" pageId="1" pageNumber="542" pagination="75 - 80" refId="ref6815" refString="Uhmann, G. (1978) Die Gattungen der Anthicidae und ihre systematische Anordnung. Teil 2. Die systematische Anordnung. 3. Beitrag zur Kenntnis der Anthicidae (Coleoptera). Entomologische Blatter, 74 (1 - 2), 75 - 80." type="journal article" year="1978">Uhmann (1978: 76</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 79) checklist of world
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||
<taxonomicName id="4C7FF0A52539FFCF1559F886FA37F80E" authorityName="Latreille" authorityYear="1819" box="[1313,1436,1903,1929]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
|
||
genera, erroneously attributed to
|
||
<taxonomicName id="4C7FF0A52539FFCF1186F87AFDC0F82A" authorityName="Latreille" authorityYear="1819" box="[510,619,1939,1965]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="tribe" tribe="Anthicini">Anthicini</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0ADEAD372539FFCF120BF87AFCF4F82A" author="Chandler, D. S." box="[627,863,1938,1965]" page="557" pageId="1" pageNumber="542" year="2002">
|
||
<bibRefCitation id="EFEEF6D72539FFCF120BF87AFCF4F82A" author="Chandler, D. S." box="[627,863,1938,1965]" pageId="1" pageNumber="542" pagination="1819" refId="ref6151" refString="Chandler, D. S. (2002) 117. Anthicidae Latreille 1819. In: Arnett, R. H., Jr., Thomas, M. C., Skelley, P. E. & Frank, J. H. (Eds.), American Beetles. Vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press, Boca Raton, London, New York, Washington, D. C., pp. 549 - 558." type="book chapter" year="2002">Chandler (2002: 557)</bibRefCitation>
|
||
</treatmentCitation>
|
||
comment on a generic composition of Tomoderinae;
|
||
<treatmentCitation id="0ADEAD372539FFCF10EFF85EFE23F856" author="Chandler, D. S." box="[151,392,1975,2001]" page="731" pageId="1" pageNumber="542" year="2010">
|
||
<bibRefCitation id="EFEEF6D72539FFCF10EFF85EFE23F856" author="Chandler, D. S." box="[151,392,1975,2001]" pageId="1" pageNumber="542" pagination="729 - 741" refId="ref6240" refString="Chandler, D. S. (2010) 11.26. Anthicidae Latreille, 1819. In: Leschen, R. A. B., Beutel, R. G. & Lawrence, J. F. (Eds.), Coleoptera, Beetles. Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). Handbook of Zoology. Arthropoda: Insecta. De Gruyter, Berlin & New York, pp. 729 - 741. https: // doi. org / 10.1515 / 9783110911213.729" type="book chapter" year="2010">Chandler (2010: 731)</bibRefCitation>
|
||
</treatmentCitation>
|
||
checklist of world
|
||
<taxonomicName id="4C7FF0A52539FFCF1227F85EFD71F856" authorityName="Latreille" authorityYear="1819" box="[607,730,1975,2001]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
|
||
genera.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365D8AD2539FFCA10BFF832FDD5FE0E" lastPageId="4" lastPageNumber="545" pageId="1" pageNumber="542" type="materials_examined">
|
||
<paragraph id="8BC08B262539FFCF10BFF832FCEEF872" blockId="1.[151,1436,1759,2037]" box="[199,837,2010,2037]" pageId="1" pageNumber="542">
|
||
<emphasis id="B90B57342539FFCF10BFF832FE43F873" bold="true" box="[199,488,2010,2037]" pageId="1" pageNumber="542">
|
||
<typeStatus id="54C435842539FFCF10BFF832FEA9F872" box="[199,258,2011,2037]" pageId="1" pageNumber="542">Type</typeStatus>
|
||
material examined
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="134497A32539FFCF118FF832FDF5F872" box="[503,606,2010,2037]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="8.[151,250,1424,1448]" captionStartId-1="9.[151,250,1520,1544]" captionStartId-2="10.[151,250,1912,1936]" captionStartId-3="11.[151,250,1289,1313]" captionStartId-4="12.[151,250,1831,1855]" captionTargetBox-0="[158,1422,274,1400]" captionTargetBox-1="[309,1277,181,1496]" captionTargetBox-2="[157,1430,197,1882]" captionTargetBox-3="[167,1409,181,1265]" captionTargetBox-4="[152,1431,189,1808]" captionTargetId-0="figure-63@8.[151,1436,260,1400]" captionTargetId-1="figure-17@9.[309,1277,181,1496]" captionTargetId-2="figure-27@10.[151,1436,181,1888]" captionTargetId-3="figure-17@11.[167,1419,181,1265]" captionTargetId-4="figure-27@12.[151,1436,181,1808]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 5. Tomosomus pallidicornis (Champion, 1890), lectotype ♁. A—Habitus, dorsal view; B—ditto, lateral view." captionText-1="FIGURE 6. Tomosomus pallidicornis (Champion, 1890), paralectotype ♁, habitus, dorsal view." captionText-2="FIGURE 7. Tomosomus pallidicornis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—Tergite VII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus, lateral view [not to scale]." captionText-3="FIGURE 8. Tomosomus meridionalis (Champion, 1890), lectotype ♁. A—Habitus, dorsal view; B—ditto, lateral view." captionText-4="FIGURE 9. Tomosomus meridionalis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—TergiteVII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternites VIII & IX; E—Aedeagus, ventral view [note—aedeagus tip is somewhat damaged]; F—ditto, endophallic armature (enlarged) [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.7961489" figureDoi-1="http://doi.org/10.5281/zenodo.7961493" figureDoi-2="http://doi.org/10.5281/zenodo.7961497" figureDoi-3="http://doi.org/10.5281/zenodo.7961499" figureDoi-4="http://doi.org/10.5281/zenodo.7961501" httpUri-0="https://zenodo.org/record/7961489/files/figure.png" httpUri-1="https://zenodo.org/record/7961493/files/figure.png" httpUri-2="https://zenodo.org/record/7961497/files/figure.png" httpUri-3="https://zenodo.org/record/7961499/files/figure.png" httpUri-4="https://zenodo.org/record/7961501/files/figure.png" pageId="1" pageNumber="542">Figs 5–9</figureCitation>
|
||
). See redescriptions.
|
||
</paragraph>
|
||
<caption id="DF00DBAE253AFFCC10EFF87EFBB7F854" ID-DOI="http://doi.org/10.5281/zenodo.7961479" ID-Zenodo-Dep="7961479" httpUri="https://zenodo.org/record/7961479/files/figure.png" pageId="2" pageNumber="543" startId="2.[151,250,1943,1967]" targetBox="[173,1407,187,1914]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BC08B26253AFFCC10EFF87EFBB7F854" blockId="2.[151,1436,1943,2006]" pageId="2" pageNumber="543">
|
||
<emphasis id="B90B5734253AFFCC10EFF87EFEB8F828" bold="true" box="[151,275,1943,1967]" pageId="2" pageNumber="543">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C7FF0A5253AFFCC1161F871FD4AF828" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[281,737,1943,1967]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="subtestaceus">
|
||
<emphasis id="B90B5734253AFFCC1161F871FDBDF828" box="[281,534,1943,1967]" italics="true" pageId="2" pageNumber="543">Tomosomus subtestaceus</emphasis>
|
||
<bibRefCitation id="EFEEF6D7253AFFCC1265F87EFD4AF828" author="Motschulsky, V." box="[541,737,1943,1967]" pageId="2" pageNumber="543" pagination="8 - 25" refId="ref6524" refString="Motschulsky, V. (1855) Voyages, Lettre de M. de Motschulsky a M. Menetries. № 2. Etudes entomologiques, [1855], 8 - 25." type="journal article" year="1855">Motschulsky, 1855</bibRefCitation>
|
||
</taxonomicName>
|
||
. A—Paralectotypes ♀♀, habitus, laterodorsal view; B—Lectotype ♁ (upper specimen) and paralectotypes ♁♁, habitus, dorsal view; C—Labels [not to scale].
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00DBAE253BFFCD10EFF92AFAC0F887" ID-DOI="http://doi.org/10.5281/zenodo.7961481" ID-Zenodo-Dep="7961481" httpUri="https://zenodo.org/record/7961481/files/figure.png" pageId="3" pageNumber="544" startId="3.[151,250,1731,1755]" targetBox="[157,1430,187,1701]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BC08B26253BFFCD10EFF92AFAC0F887" blockId="3.[151,1436,1731,1792]" pageId="3" pageNumber="544">
|
||
<emphasis id="B90B5734253BFFCD10EFF92AFEBFF95B" bold="true" box="[151,276,1731,1756]" pageId="3" pageNumber="544">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7FF0A5253BFFCD1164F92DFD4DF95B" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[284,742,1732,1756]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="subtestaceus">
|
||
<emphasis id="B90B5734253BFFCD1164F92DFDB1F95B" box="[284,538,1732,1756]" italics="true" pageId="3" pageNumber="544">Tomosomus subtestaceus</emphasis>
|
||
<bibRefCitation id="EFEEF6D7253BFFCD1259F92DFD4DF95B" author="Motschulsky, V." box="[545,742,1732,1756]" pageId="3" pageNumber="544" pagination="8 - 25" refId="ref6524" refString="Motschulsky, V. (1855) Voyages, Lettre de M. de Motschulsky a M. Menetries. № 2. Etudes entomologiques, [1855], 8 - 25." type="journal article" year="1855">Motschulsky, 1855</bibRefCitation>
|
||
</taxonomicName>
|
||
. A—Lectotype ♁, habitus, dorsal view; B—ditto, lateral view; C— Apex of elytra, tergite VII and aedeagus, dorsal view [not to scale]. Images courtesy Kirill V. Makarov (Moscow, Russia).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC08B26253BFFCD10BFF8A2FB6AF872" blockId="3.[151,1437,1867,2038]" pageId="3" pageNumber="544">
|
||
<emphasis id="B90B5734253BFFCD10BFF8A2FEDCF8E2" bold="true" box="[199,375,1867,1893]" pageId="3" pageNumber="544">Nomenclature.</emphasis>
|
||
<taxonomicName id="4C7FF0A5253BFFCD1107F8A2FE53F8E2" authorityName="Champion" authorityYear="1890" box="[383,504,1867,1893]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253BFFCD1107F8A2FE53F8E2" box="[383,504,1867,1893]" italics="true" pageId="3" pageNumber="544">Holcopyge</emphasis>
|
||
</taxonomicName>
|
||
was established by
|
||
<bibRefCitation id="EFEEF6D7253BFFCD12A3F8A2FC09F8E2" author="Champion, G. C." box="[731,930,1867,1893]" pageId="3" pageNumber="544" pagination="292 - 293" refId="ref6084" refString="Champion, G. C. (1890) On a new genus of Anthicidae from tropical South America. The Entomologists Monthly Magazine, 26, 292 - 293." type="journal article" year="1890">Champion (1890)</bibRefCitation>
|
||
for two species described in the same paper,
|
||
<taxonomicName id="4C7FF0A5253BFFCD10EFF899FEE3F80E" authorityName="Telnov & Gusakov" authorityYear="2023" baseAuthorityName="Champion" baseAuthorityYear="1890" box="[151,328,1903,1929]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="meridionalis">
|
||
<emphasis id="B90B5734253BFFCD10EFF899FEE3F80E" box="[151,328,1903,1929]" italics="true" pageId="3" pageNumber="544">H. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7FF0A5253BFFCD1107F899FD9AF80E" authorityName="Champion" authorityYear="1890" box="[383,561,1903,1929]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="pallidicornis">
|
||
<emphasis id="B90B5734253BFFCD1107F899FD9AF80E" box="[383,561,1903,1929]" italics="true" pageId="3" pageNumber="544">H. pallidicornis</emphasis>
|
||
</taxonomicName>
|
||
. No
|
||
<typeStatus id="54C43584253BFFCD1212F899FD31F80D" box="[618,666,1904,1930]" pageId="3" pageNumber="544">type</typeStatus>
|
||
species was designated for
|
||
<taxonomicName id="4C7FF0A5253BFFCD13ADF886FBE5F80E" authorityName="Champion" authorityYear="1890" box="[981,1102,1903,1929]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253BFFCD13ADF886FBE5F80E" box="[981,1102,1903,1929]" italics="true" pageId="3" pageNumber="544">Holcopyge</emphasis>
|
||
</taxonomicName>
|
||
, so herewith we designate
|
||
<taxonomicName id="4C7FF0A5253BFFCD15F9F899FE8DF82A" authorityName="Champion" authorityYear="1890" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="species" species="pallidicornis">
|
||
<emphasis id="B90B5734253BFFCD15F9F899FE8DF82A" italics="true" pageId="3" pageNumber="544">H. pallidicornis</emphasis>
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="54C43584253BFFCD1100F87DFE03F829" box="[376,424,1940,1966]" pageId="3" pageNumber="544">type</typeStatus>
|
||
of its genus.
|
||
<taxonomicName id="4C7FF0A5253BFFCD123BF87AFD17F82A" authorityName="Champion" authorityYear="1890" box="[579,700,1939,1965]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253BFFCD123BF87AFD17F82A" box="[579,700,1939,1965]" italics="true" pageId="3" pageNumber="544">Holcopyge</emphasis>
|
||
</taxonomicName>
|
||
was initially placed in
|
||
<taxonomicName id="4C7FF0A5253BFFCD13BDF87AFBEBF82A" authorityName="Latreille" authorityYear="1819" box="[965,1088,1939,1965]" class="Insecta" family="Anthicidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="family">Anthicidae</taxonomicName>
|
||
but without family placement, pointing out morphological similarities with
|
||
<taxonomicName id="4C7FF0A5253BFFCD12F3F85EFB8BF856" authority="LaFerte-Senectere, 1849" authorityName="LaFerte-Senectere" authorityYear="1849" box="[651,1056,1975,2001]" class="Insecta" family="Anthicidae" genus="Tomoderus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="544" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253BFFCD12F3F85EFCADF856" box="[651,774,1975,2001]" italics="true" pageId="3" pageNumber="544">Tomoderus</emphasis>
|
||
LaFerté-Sénectère, 1849
|
||
</taxonomicName>
|
||
of Tomoderinae Bonadona, 1961. Formally placed in Tomoderinae by
|
||
<bibRefCitation id="EFEEF6D7253BFFCD1257F832FD4BF872" author="Chandler, D. S." box="[559,736,2011,2037]" pageId="3" pageNumber="544" pagination="1819" refId="ref6151" refString="Chandler, D. S. (2002) 117. Anthicidae Latreille 1819. In: Arnett, R. H., Jr., Thomas, M. C., Skelley, P. E. & Frank, J. H. (Eds.), American Beetles. Vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press, Boca Raton, London, New York, Washington, D. C., pp. 549 - 558." type="book chapter" year="2002">Chandler (2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEEF6D7253BFFCD1293F832FC83F872" author="Chandler, D. S." box="[747,808,2011,2037]" pageId="3" pageNumber="544" pagination="729 - 741" refId="ref6240" refString="Chandler, D. S. (2010) 11.26. Anthicidae Latreille, 1819. In: Leschen, R. A. B., Beutel, R. G. & Lawrence, J. F. (Eds.), Coleoptera, Beetles. Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). Handbook of Zoology. Arthropoda: Insecta. De Gruyter, Berlin & New York, pp. 729 - 741. https: // doi. org / 10.1515 / 9783110911213.729" type="book chapter" year="2010">2010</bibRefCitation>
|
||
). This genus has never been revised.
|
||
</paragraph>
|
||
<paragraph id="8BC08B26253CFFCA10BFFF7EFBE3FE9A" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="545">
|
||
Study of the type species of both
|
||
<taxonomicName id="4C7FF0A5253CFFCA124CFF7EFD1CFF37" authorityName="Motschulsky" authorityYear="1855" box="[564,695,151,176]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253CFFCA124CFF7EFD1CFF37" box="[564,695,151,176]" italics="true" pageId="4" pageNumber="545">Tomosomus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7FF0A5253CFFCA1293FF7EFCCFFF36" authorityName="Champion" authorityYear="1890" box="[747,868,151,177]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253CFFCA1293FF7EFCCFFF36" box="[747,868,151,177]" italics="true" pageId="4" pageNumber="545">Holcopyge</emphasis>
|
||
</taxonomicName>
|
||
suggests both genera are congeneric. A new genus-rank synonym is therefore proposed.
|
||
<taxonomicName id="4C7FF0A5253CFFCA124AFF52FCE3FF52" authorityName="Motschulsky" authorityYear="1855" box="[562,840,187,213]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="species" species="subtestaceus">
|
||
<emphasis id="B90B5734253CFFCA124AFF52FCE3FF52" box="[562,840,187,213]" italics="true" pageId="4" pageNumber="545">Tomosomus subtestaceus</emphasis>
|
||
</taxonomicName>
|
||
was described from an unstated number of specimens (
|
||
<bibRefCitation id="EFEEF6D7253CFFCA10E7FF36FEDFFF7E" author="Motschulsky, V." box="[159,372,222,249]" pageId="4" pageNumber="545" pagination="8 - 25" refId="ref6524" refString="Motschulsky, V. (1855) Voyages, Lettre de M. de Motschulsky a M. Menetries. № 2. Etudes entomologiques, [1855], 8 - 25." type="journal article" year="1855">Motschulsky 1855</bibRefCitation>
|
||
). A series of
|
||
<specimenCount id="9D7940AF253CFFCA127CFF36FD33FF7E" box="[516,664,223,249]" pageId="4" pageNumber="545" type="generic" typeStatus="syntypes">five syntypes</specimenCount>
|
||
of this taxon is stored at
|
||
<collectionCode id="ED6E13E3253CFFCA13C9FF36FBA7FF7E" box="[945,1036,223,249]" pageId="4" pageNumber="545">MZUM</collectionCode>
|
||
.
|
||
<taxonomicName id="4C7FF0A5253CFFCA146EFF36FA8EFF7E" authorityName="Champion" authorityYear="1890" box="[1046,1317,223,249]" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="species" species="pallidicornis">
|
||
<emphasis id="B90B5734253CFFCA146EFF36FA8EFF7E" box="[1046,1317,223,249]" italics="true" pageId="4" pageNumber="545">Holcopyge pallidicornis</emphasis>
|
||
</taxonomicName>
|
||
(proposed as type of its genus herein, see above) was described from
|
||
<specimenCount id="9D7940AF253CFFCA1351FEEAFC61FE9A" box="[809,970,259,285]" pageId="4" pageNumber="545" type="generic" typeStatus="syntypes">three syntypes</specimenCount>
|
||
at
|
||
<collectionCode id="ED6E13E3253CFFCA1394FEEAFBEFFE9A" box="[1004,1092,259,285]" pageId="4" pageNumber="545">BMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC08B26253CFFCA10BFFECEFDD5FE0E" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="545">
|
||
All three species yet arranged to
|
||
<taxonomicName id="4C7FF0A5253CFFCA1249FECEFD1FFEC7" authorityName="Motschulsky" authorityYear="1855" box="[561,692,295,320]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253CFFCA1249FECEFD1FFEC7" box="[561,692,295,320]" italics="true" pageId="4" pageNumber="545">Tomosomus</emphasis>
|
||
</taxonomicName>
|
||
appear valid (also consider the key below). Two new combinations are proposed for the former
|
||
<taxonomicName id="4C7FF0A5253CFFCA119BFEA2FD17FEE2" box="[483,700,331,357]" pageId="4" pageNumber="545">
|
||
<emphasis id="B90B5734253CFFCA119BFEA2FDF7FEE2" box="[483,604,331,357]" italics="true" pageId="4" pageNumber="545">Holcopyge</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C7FF0A5253CFFCA12B2FEA2FB6AFEE2" authority="(Champion, 1890)" authorityName="Telnov & Gusakov" authorityYear="2023" baseAuthorityName="Champion" baseAuthorityYear="1890" box="[714,1217,331,357]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="species" species="meridionalis" status="comb. nov.">
|
||
<emphasis id="B90B5734253CFFCA12B2FEA2FC4DFEE2" box="[714,998,331,357]" italics="true" pageId="4" pageNumber="545">Tomosomus meridionalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEEF6D7253CFFCA1380FEA2FB12FEE2" author="Champion, G. C." box="[1016,1209,331,357]" pageId="4" pageNumber="545" pagination="292 - 293" refId="ref6084" refString="Champion, G. C. (1890) On a new genus of Anthicidae from tropical South America. The Entomologists Monthly Magazine, 26, 292 - 293." type="journal article" year="1890">Champion, 1890</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A238EA4F253CFFCA14B3FEA2FAE0FEE2" box="[1227,1355,331,357]" pageId="4" pageNumber="545" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7FF0A5253CFFCA15F0FEA2FE50FE0E" authority="(Champion, 1890)" authorityName="Telnov & Gusakov" authorityYear="2023" baseAuthorityName="Champion" baseAuthorityYear="1890" class="Insecta" family="Anthicidae" genus="Holcopyge" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="species" species="pallidicornis" status="comb. nov.">
|
||
<emphasis id="B90B5734253CFFCA15F0FEA2FE8DFE0E" italics="true" pageId="4" pageNumber="545">T. pallidicornis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEEF6D7253CFFCA114DFE86FE59FE0E" author="Champion, G. C." box="[309,498,367,393]" pageId="4" pageNumber="545" pagination="292 - 293" refId="ref6084" refString="Champion, G. C. (1890) On a new genus of Anthicidae from tropical South America. The Entomologists Monthly Magazine, 26, 292 - 293." type="journal article" year="1890">Champion, 1890</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A238EA4F253CFFCA127AFE86FDD5FE0E" box="[514,638,367,393]" pageId="4" pageNumber="545" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365D8AD253CFFCA10BFFE7BFAD7F882" pageId="4" pageNumber="545" type="description">
|
||
<paragraph id="8BC08B26253CFFCA10BFFE7BFAD7F882" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="545">
|
||
<emphasis id="B90B5734253CFFCA10BFFE7BFDADFE2A" bold="true" box="[199,518,402,429]" pageId="4" pageNumber="545">Redescription of the genus.</emphasis>
|
||
In general appearance strongly resembles
|
||
<taxonomicName id="4C7FF0A5253CFFCA13A1FE7AFBFFFE2A" authorityName="LaFerte-Senectere" authorityYear="1849" box="[985,1108,403,429]" class="Insecta" family="Anthicidae" genus="Tomoderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253CFFCA13A1FE7AFBFFFE2A" box="[985,1108,403,429]" italics="true" pageId="4" pageNumber="545">Tomoderus</emphasis>
|
||
</taxonomicName>
|
||
. Dorsum and venter glossy to moderately glossy. Head prognathous, subtriangular, slightly convex in dorsal aspect. Frontoclypeal suture present, straight. Antennal insertion visible in dorsal view, placed at anterolateral margin of frons at some distance from compound eye. Compound eye moderate, not emarginate, irregularly elliptical, facets not numerous, rather strongly convex. Interfacetal setae not observed. Head base broadly rounded to slightly concave. Antenna slightly widened in apical third, constituted of eleven moniliform antennomeres. Second antennomere not shortened. Penultimate antennomere slightly transverse. Mandible apex bidentate. Maxillary palpus quadrimerous. Terminal maxillary palpomere small, securiform to strongly triangular. Head dorsal setae inconspicuous, appressed. Few long, tactile setae along lateral and basal margin, also on vertex. Cranial “neck” narrow, completely concealed by occiput and anterior margin of pronotum. Pronotum distinctly bilobate, with broad and deep postmedian lateral constriction. Anterior lobe distinctly wider and larger than posterior. Anterior rim very narrow both dorsally and laterally. Antebasal sulcus not indicated both dorsally and laterally. Anterior lobe dorsally longitudinally sulcate or not. Lateral constriction starts from a pit in a lateral wall of pronotum just above transition to hypomeron. Dorsal setae of pronotum inconspicuous, appressed to suberect. Lateral margin of anterior lobe with four long, erect tactile setae, of posterior lobe—with one similar seta. Scutellar shield small, triangular, apically rounded. Elytra ovoid to elongate ovoid, slightly convex in dorsal aspect. Humerus obsolete (all known species apterous). Sutural stria present at least in basal two thirds of elytron, narrow. Posterior sutural angle of elytron either acute, not rounded. Epipleuron nearly complete. Elytral setae long, suberect to erect. Profemur distinctly clavate, meso- and metathoracic femurs slightly clavate or not. Tibiae subequal in length to corresponding femora. Tibial terminal spurs not present. Tibia denser setose compare to corresponding femur. Penultimate antennomere not or indistinctly bilobate. Pretarsal claw simple. Procoxa conical, contiguous, basally separated by low intercoxal ridge. Procoxal cavity (
|
||
<figureCitation id="134497A3253CFFCA1542FB8AFA38FBFA" box="[1338,1427,1123,1149]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[189,1420,201,1903]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961485" httpUri="https://zenodo.org/record/7961485/files/figure.png" pageId="4" pageNumber="545">Fig. 3C</figureCitation>
|
||
) closed internally, open externally, closed behind by moderately long postcoxal plate. Procoxal rests well-developed. Mesoventrite (
|
||
<figureCitation id="134497A3253CFFCA1143FB42FE3AFB42" box="[315,401,1195,1221]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[189,1420,201,1903]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961485" httpUri="https://zenodo.org/record/7961485/files/figure.png" pageId="4" pageNumber="545">Fig. 3C</figureCitation>
|
||
) short, broadly triangular, broadly rounded at anterior margin, lateral edges slightly arched. Mesanepisterna broadly separated by anterior margin on mesoventrite, broadly subtriangular. Mesoventral intercoxal process long and narrow, projecting along almost whole length of mesocoxa, apically rounded, meeting anterior process of metasternum. Mesepimeron elongate triangular. Mesocoxa subovate, mesocoxal cavity closed laterally by mesepimeron. Metaventrite (
|
||
<figureCitation id="134497A3253CFFCA1184FAD2FDFAFAD2" box="[508,593,1339,1365]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[189,1420,201,1903]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961485" httpUri="https://zenodo.org/record/7961485/files/figure.png" pageId="4" pageNumber="545">Fig. 3C</figureCitation>
|
||
) remarkably short and narrow, anterior intercoxal process moderately long, subacute angulate. Metaventral bridge between meso- and metathoracic coxal cavities very narrow. Metaventrite strongly, triangularly expanding laterad to meso- and metacoxal cavities.Metathoracic discrimen not present.Posterior margin of metaventrite broadly emarginate medially to accommodate broad, apically rounded anterior intercoxal process of morphological sternite III (
|
||
<figureCitation id="134497A3253CFFCA1243FA22FD38FA62" box="[571,659,1483,1509]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[189,1420,201,1903]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961485" httpUri="https://zenodo.org/record/7961485/files/figure.png" pageId="4" pageNumber="545">Fig. 3C</figureCitation>
|
||
). Metacoxa strongly transverse. Five free abdominal ventrites. Tergite VII rounded at posterior margin, dorsally with broad, conspicuous median longitudinal furrow lateral margins of which slightly deflected and irregularly microdenticulate (
|
||
<figureCitation id="134497A3253CFFCA1347F9FAFC0AF9AA" box="[831,929,1555,1581]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="4" pageNumber="545">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA13C9F9FAFC5DF9AA" box="[945,1014,1555,1581]" captionStart="FIGURE 7" captionStartId="10.[151,250,1912,1936]" captionTargetBox="[157,1430,197,1882]" captionTargetId="figure-27@10.[151,1436,181,1888]" captionTargetPageId="10" captionText="FIGURE 7. Tomosomus pallidicornis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—Tergite VII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus, lateral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961497" httpUri="https://zenodo.org/record/7961497/files/figure.png" pageId="4" pageNumber="545">7A–B</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA147FF9FAFBE5F9AA" box="[1031,1102,1555,1581]" captionStart="FIGURE 9" captionStartId="12.[151,250,1831,1855]" captionTargetBox="[152,1431,189,1808]" captionTargetId="figure-27@12.[151,1436,181,1808]" captionTargetPageId="12" captionText="FIGURE 9. Tomosomus meridionalis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—TergiteVII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternites VIII & IX; E—Aedeagus, ventral view [note—aedeagus tip is somewhat damaged]; F—ditto, endophallic armature (enlarged) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961501" httpUri="https://zenodo.org/record/7961501/files/figure.png" pageId="4" pageNumber="545">9A–B</figureCitation>
|
||
). Morphological sternite VII rounded to subtruncate at posterior margin (
|
||
<figureCitation id="134497A3253CFFCA12FDF9DEFD4BF9D6" box="[645,736,1591,1617]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="4" pageNumber="545">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA1295F9DEFCA4F9D6" box="[749,783,1591,1617]" captionStart="FIGURE 7" captionStartId="10.[151,250,1912,1936]" captionTargetBox="[157,1430,197,1882]" captionTargetId="figure-27@10.[151,1436,181,1888]" captionTargetPageId="10" captionText="FIGURE 7. Tomosomus pallidicornis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—Tergite VII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus, lateral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961497" httpUri="https://zenodo.org/record/7961497/files/figure.png" pageId="4" pageNumber="545">7C</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA1364F9DEFCEBF9D6" box="[796,832,1591,1617]" captionStart="FIGURE 9" captionStartId="12.[151,250,1831,1855]" captionTargetBox="[152,1431,189,1808]" captionTargetId="figure-27@12.[151,1436,181,1808]" captionTargetPageId="12" captionText="FIGURE 9. Tomosomus meridionalis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—TergiteVII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternites VIII & IX; E—Aedeagus, ventral view [note—aedeagus tip is somewhat damaged]; F—ditto, endophallic armature (enlarged) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961501" httpUri="https://zenodo.org/record/7961501/files/figure.png" pageId="4" pageNumber="545">9C</figureCitation>
|
||
). Tergite VIII variably emarginate at posterior margin (
|
||
<figureCitation id="134497A3253CFFCA10E7F9B2FF50F9F2" box="[159,251,1627,1653]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="4" pageNumber="545">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA117EF9B2FE81F9F2" box="[262,298,1627,1653]" captionStart="FIGURE 7" captionStartId="10.[151,250,1912,1936]" captionTargetBox="[157,1430,197,1882]" captionTargetId="figure-27@10.[151,1436,181,1888]" captionTargetPageId="10" captionText="FIGURE 7. Tomosomus pallidicornis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—Tergite VII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus, lateral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961497" httpUri="https://zenodo.org/record/7961497/files/figure.png" pageId="4" pageNumber="545">7D</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA114EF9B2FEF7F9F2" box="[310,348,1627,1653]" captionStart="FIGURE 9" captionStartId="12.[151,250,1831,1855]" captionTargetBox="[152,1431,189,1808]" captionTargetId="figure-27@12.[151,1436,181,1808]" captionTargetPageId="12" captionText="FIGURE 9. Tomosomus meridionalis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—TergiteVII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternites VIII & IX; E—Aedeagus, ventral view [note—aedeagus tip is somewhat damaged]; F—ditto, endophallic armature (enlarged) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961501" httpUri="https://zenodo.org/record/7961501/files/figure.png" pageId="4" pageNumber="545">9D</figureCitation>
|
||
). Morphological sternite IX complex, with free or fused arms and accessory pair of free arms and median, Y- or O-shaped sclerite (
|
||
<figureCitation id="134497A3253CFFCA1274F996FDC1F91E" box="[524,618,1663,1689]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="4" pageNumber="545">Figs 4D</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA120EF996FD33F91E" box="[630,664,1663,1689]" captionStart="FIGURE 7" captionStartId="10.[151,250,1912,1936]" captionTargetBox="[157,1430,197,1882]" captionTargetId="figure-27@10.[151,1436,181,1888]" captionTargetPageId="10" captionText="FIGURE 7. Tomosomus pallidicornis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—Tergite VII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus, lateral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961497" httpUri="https://zenodo.org/record/7961497/files/figure.png" pageId="4" pageNumber="545">7E</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA12DCF996FD61F91E" box="[676,714,1663,1689]" captionStart="FIGURE 9" captionStartId="12.[151,250,1831,1855]" captionTargetBox="[152,1431,189,1808]" captionTargetId="figure-27@12.[151,1436,181,1808]" captionTargetPageId="12" captionText="FIGURE 9. Tomosomus meridionalis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—TergiteVII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternites VIII & IX; E—Aedeagus, ventral view [note—aedeagus tip is somewhat damaged]; F—ditto, endophallic armature (enlarged) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961501" httpUri="https://zenodo.org/record/7961501/files/figure.png" pageId="4" pageNumber="545">9D</figureCitation>
|
||
). Aedeagus irregularly shaped, penis encapsulated to subtubular tegmen (
|
||
<figureCitation id="134497A3253CFFCA1081F94AFEDBF93A" box="[249,368,1699,1725]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="4" pageNumber="545">Figs 4E–F</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA1103F94AFE37F93A" box="[379,412,1699,1725]" captionStart="FIGURE 7" captionStartId="10.[151,250,1912,1936]" captionTargetBox="[157,1430,197,1882]" captionTargetId="figure-27@10.[151,1436,181,1888]" captionTargetPageId="10" captionText="FIGURE 7. Tomosomus pallidicornis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—Tergite VII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternite VIII, ventral view; E—Morphological sternite IX; F—Aedeagus, lateral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961497" httpUri="https://zenodo.org/record/7961497/files/figure.png" pageId="4" pageNumber="545">7E</figureCitation>
|
||
,
|
||
<figureCitation id="134497A3253CFFCA11D1F94AFE67F93A" box="[425,460,1699,1725]" captionStart="FIGURE 9" captionStartId="12.[151,250,1831,1855]" captionTargetBox="[152,1431,189,1808]" captionTargetId="figure-27@12.[151,1436,181,1808]" captionTargetPageId="12" captionText="FIGURE 9. Tomosomus meridionalis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—TergiteVII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternites VIII & IX; E—Aedeagus, ventral view [note—aedeagus tip is somewhat damaged]; F—ditto, endophallic armature (enlarged) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961501" httpUri="https://zenodo.org/record/7961501/files/figure.png" pageId="4" pageNumber="545">9E</figureCitation>
|
||
). Phallobase fused to tegmen. Endophallus with (
|
||
<figureCitation id="134497A3253CFFCA1380F94AFBE7F93A" box="[1016,1100,1699,1725]" captionStart="FIGURE 9" captionStartId="12.[151,250,1831,1855]" captionTargetBox="[152,1431,189,1808]" captionTargetId="figure-27@12.[151,1436,181,1808]" captionTargetPageId="12" captionText="FIGURE 9. Tomosomus meridionalis (Champion, 1890), lectotype ♁, terminalia and genitalia.A—TergiteVII, dorsal view; B— Part of a median furrow of tergite VII, dorsal view (enlarged); C—Morphological sternite VII, ventral view; D—Morphological sternites VIII & IX; E—Aedeagus, ventral view [note—aedeagus tip is somewhat damaged]; F—ditto, endophallic armature (enlarged) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961501" httpUri="https://zenodo.org/record/7961501/files/figure.png" pageId="4" pageNumber="545">Fig. 9F</figureCitation>
|
||
) or without armature. Female tergite and morphological sternite VII rounded to subtruncate at posterior margin. Ovipositor with coxites close together in midventral region, in part divided ventrally. Styli small, borne on incompletely bisegmented coxites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365D8AD253CFFCA10BFF8E7FDBEF812" pageId="4" pageNumber="545" type="diagnosis">
|
||
<paragraph id="8BC08B26253CFFCA10BFF8E7FDBEF812" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="545">
|
||
<emphasis id="B90B5734253CFFCA10BFF8E7FE79F8AF" bold="true" box="[199,466,1806,1832]" pageId="4" pageNumber="545">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C7FF0A5253CFFCA11A5F8E6FDCBF8AF" authorityName="Motschulsky" authorityYear="1855" box="[477,608,1807,1832]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="545" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90B5734253CFFCA11A5F8E6FDCBF8AF" box="[477,608,1807,1832]" italics="true" pageId="4" pageNumber="545">Tomosomus</emphasis>
|
||
</taxonomicName>
|
||
is different from all tomoderine genera primarily in the longitudinally furrowed tergite VII in the combination with the remarkably short, narrow metaventrite, the asymmetrical, subtubular aedeagus not entirely enclosing the penis, the apically emarginate morphological sternite VIII, and the peculiarly shaped morphological sternite IX.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |