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<document ID-DOI="10.11646/zootaxa.5278.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7905995" ID-ZooBank="697AC989-141D-4A89-8F02-5A4644E303EA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683528451479" checkinUser="plazi" docAuthor="Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak &amp; Toda, Masanori J." docDate="2023" docId="03C687D9FFC2FFF4A5C6FDCB8415E2A5" docLanguage="en" docName="zootaxa.5278.2.1.pdf" docOrigin="Zootaxa 5278 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colocasiomyia zeylanica" docType="treatment" docVersion="2" lastPageNumber="221" masterDocId="FFFFFFA1FFD6FFE0A551FFBF817EE53C" masterDocTitle="Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species" masterLastPageNumber="238" masterPageNumber="201" pageNumber="221" updateTime="1683528945942" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species</mods:title>
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<mods:namePart>Zhang, Guang</mods:namePart>
<mods:affiliation>Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology (Institute of Biodiversity), Yunnan University, Kunming, 650091 China &amp; Guizhou Provincial Museum, Guiyang, 550081 China</mods:affiliation>
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<mods:affiliation>Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology (Institute of Biodiversity), Yunnan University, Kunming, 650091 China &amp; gaojj @ ynu. edu. cn; https: // orcid. org / 0000 - 0003 - 4636 - 2130</mods:affiliation>
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<mods:namePart>Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart>Yafuso, Masako</mods:namePart>
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<mods:affiliation>Uema 1 - 2 - 21, Naha, Okinawa 902 - 0073, Japan. yafusom @ juno. ocn. ne. jp; https: // orcid. org / 0000 - 0001 - 9737 - 7792</mods:affiliation>
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<mods:namePart>Suwito, Awit</mods:namePart>
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<mods:affiliation>Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. sungaicola @ gmail. com; https: // orcid. org / 0000 - 0003 - 2600 - 2488</mods:affiliation>
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<mods:namePart>Meleng, Paulus Ak</mods:namePart>
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<mods:affiliation>Research Development and Innovation Division, Forest Department Sarawak, KM 10, Jalan Datuk Amar Kalong Ningkan, 93250, Kuching, Sarawak, Malaysia. paulusm @ sarawak. gov. my; https: // orcid. org / 0000 - 0003 - 3777 - 0597</mods:affiliation>
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<mods:namePart>Toda, Masanori J.</mods:namePart>
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<taxonomicName ID-CoL="X6V9" baseAuthorityName="Okada" baseAuthorityYear="1986" box="[151,448,628,655]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="zeylanica">
<emphasis bold="true" box="[151,448,628,655]" italics="true" pageId="20" pageNumber="221">Colocasiomyia zeylanica</emphasis>
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species group
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<paragraph blockId="20.[151,1075,697,721]" box="[151,1075,697,721]" pageId="20" pageNumber="221">
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<taxonomicName ID-CoL="X6V9" baseAuthorityName="Okada" baseAuthorityYear="1986" box="[151,406,697,721]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="zeylanica">
<emphasis box="[151,406,697,721]" italics="true" pageId="20" pageNumber="221">Colocasiomyia zeylanica</emphasis>
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species group:
<treatmentCitation author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[563,814,697,721]" page="694" pageId="20" pageNumber="221" year="2006">
<bibRefCitation author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[563,814,697,721]" pageId="20" pageNumber="221" pagination="684 - 702" refId="ref21931" refString="Sultana, F., Hu, Y. G., Toda, M. J., Takenaka, K. &amp; Yafuso, M. (2006) Phylogeny and classification of Colocasiomyia (Diptera, Drosophilidae), and its evolution of pollination mutualism with aroid plants. Systematic Entomology, 31, 684 - 702. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00344. x" type="journal article" year="2006">
Sultana
<emphasis box="[646,697,697,721]" italics="true" pageId="20" pageNumber="221">et al.</emphasis>
, 2006: 694
</bibRefCitation>
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;
<treatmentCitation author="Fartyal, R. S. &amp; Gao, J. J. &amp; Toda, M. J. &amp; Hu, Y. G. &amp; Takano, K. T. &amp; Suwito, A. &amp; Katoh, T. &amp; Takigahira, T. &amp; Yin, J. T." box="[825,1071,697,721]" page="769" pageId="20" pageNumber="221" year="2013">
<bibRefCitation author="Fartyal, R. S. &amp; Gao, J. J. &amp; Toda, M. J. &amp; Hu, Y. G. &amp; Takano, K. T. &amp; Suwito, A. &amp; Katoh, T. &amp; Takigahira, T. &amp; Yin, J. T." box="[825,1071,697,721]" pageId="20" pageNumber="221" pagination="763 - 782" refId="ref20189" refString="Fartyal, R. S., Gao, J. J., Toda, M. J., Hu, Y. G., Takano, K. T., Suwito, A., Katoh, T., Takigahira, T. &amp; Yin, J. T. (2013) Colocasiomyia (Diptera: Drosophilidae) revised phylogenetically, with a new species group having peculiar lifecycles on monsteroid (Araceae) host plants. Systematic Entomology, 38, 763 - 782. https: // doi. org / 10.1111 / syen. 12027" type="journal article" year="2013">
Fartyal
<emphasis box="[903,954,697,721]" italics="true" pageId="20" pageNumber="221">et al.</emphasis>
, 2013: 769
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.
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</paragraph>
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<paragraph blockId="20.[151,1437,767,1945]" pageId="20" pageNumber="221">
<emphasis bold="true" box="[151,266,767,793]" pageId="20" pageNumber="221">Diagnosis</emphasis>
(modified from
<bibRefCitation author="Fartyal, R. S. &amp; Gao, J. J. &amp; Toda, M. J. &amp; Hu, Y. G. &amp; Takano, K. T. &amp; Suwito, A. &amp; Katoh, T. &amp; Takigahira, T. &amp; Yin, J. T." box="[454,665,767,793]" pageId="20" pageNumber="221" pagination="763 - 782" refId="ref20189" refString="Fartyal, R. S., Gao, J. J., Toda, M. J., Hu, Y. G., Takano, K. T., Suwito, A., Katoh, T., Takigahira, T. &amp; Yin, J. T. (2013) Colocasiomyia (Diptera: Drosophilidae) revised phylogenetically, with a new species group having peculiar lifecycles on monsteroid (Araceae) host plants. Systematic Entomology, 38, 763 - 782. https: // doi. org / 10.1111 / syen. 12027" type="journal article" year="2013">
Fartyal
<emphasis box="[541,597,767,793]" italics="true" pageId="20" pageNumber="221">et al.</emphasis>
2013
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). Pedicel with uniformly short, stout setae only, lacking prominently long one(s) (ch.2-1;
<figureCitation box="[378,444,803,829]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,181,1406]" captionTargetId="figure-20@5.[151,1436,181,1406]" captionTargetPageId="5" captionText="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906005" httpUri="https://zenodo.org/record/7906005/files/figure.png" pageId="20" pageNumber="221">Fig. 3</figureCitation>
, “
<figureCitation box="[468,535,803,829]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="20" pageNumber="221">Fig. 6</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[581,749,803,829]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Two peg-like setae present only on fore tarsomere II (ch.36- 0, ch.38-0;
<figureCitation box="[287,407,839,865]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="20" pageNumber="221">Fig. 9A–E</figureCitation>
<figureCitation box="[407,415,858,872]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="left" box="[407,415,858,872]" fontSize="6" pageId="20" pageNumber="221">1</subScript>
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, “
<figureCitation box="[444,534,839,865]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">Fig. 1C</figureCitation>
” in
<bibRefCitation author="Okada, T." box="[590,733,839,865]" pageId="20" pageNumber="221" pagination="53 - 59" refId="ref21490" refString="Okada, T. (1986) Taximetrical analyses of costal chaetotaxy of the genus Drosophilella Duda, with description of a new species from Sri Lanka (Diptera, Drosophilidae). Proceedings of the Japanese Society of Systematic Zoology, 34, 53 - 59." type="journal article" year="1986">Okada 1986</bibRefCitation>
, “
<figureCitation box="[759,831,839,865]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="20" pageNumber="221">Fig. 9</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[886,1058,839,865]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Hypandrium forming like tube surrounding phallus (ch.66-2; Fig. 11). Gonocoxites absent (ch.69-4; Fig. 11).
</paragraph>
<paragraph blockId="20.[151,1437,767,1945]" pageId="20" pageNumber="221">
<emphasis bold="true" box="[199,425,911,937]" pageId="20" pageNumber="221">Shared characters.</emphasis>
Distance between antennal scapes wider than half of scape diameter (ch.1-1;
<figureCitation box="[1304,1429,911,944]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="20" pageNumber="221">
Fig. 2A–E
<subScript attach="left" box="[1421,1429,930,944]" fontSize="6" pageId="20" pageNumber="221">2</subScript>
</figureCitation>
, “
<figureCitation box="[162,254,947,973]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">Fig. 1A</figureCitation>
” in
<bibRefCitation author="Okada, T." box="[309,453,947,973]" pageId="20" pageNumber="221" pagination="53 - 59" refId="ref21490" refString="Okada, T. (1986) Taximetrical analyses of costal chaetotaxy of the genus Drosophilella Duda, with description of a new species from Sri Lanka (Diptera, Drosophilidae). Proceedings of the Japanese Society of Systematic Zoology, 34, 53 - 59." type="journal article" year="1986">Okada 1986</bibRefCitation>
, “
<figureCitation box="[479,550,947,973]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="20" pageNumber="221">Fig. 6</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[606,778,947,973]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Antennal, first flagellomere grayish yellow, with small invaginated organ but no hollow organ (ch.3-0;
<figureCitation box="[680,746,983,1009]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,181,1406]" captionTargetId="figure-20@5.[151,1436,181,1406]" captionTargetPageId="5" captionText="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906005" httpUri="https://zenodo.org/record/7906005/files/figure.png" pageId="20" pageNumber="221">Fig. 3</figureCitation>
, “
<figureCitation box="[769,836,983,1009]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="20" pageNumber="221">Fig. 6</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[881,1048,983,1009]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
); arista entirely densely pubescent, without branches (ch.5-1;
<figureCitation box="[447,514,1019,1045]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,181,1406]" captionTargetId="figure-20@5.[151,1436,181,1406]" captionTargetPageId="5" captionText="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906005" httpUri="https://zenodo.org/record/7906005/files/figure.png" pageId="20" pageNumber="221">Fig. 3</figureCitation>
, “
<figureCitation box="[539,661,1019,1045]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">Fig. 1A, B</figureCitation>
” in
<bibRefCitation author="Okada, T." box="[710,850,1019,1045]" pageId="20" pageNumber="221" pagination="53 - 59" refId="ref21490" refString="Okada, T. (1986) Taximetrical analyses of costal chaetotaxy of the genus Drosophilella Duda, with description of a new species from Sri Lanka (Diptera, Drosophilidae). Proceedings of the Japanese Society of Systematic Zoology, 34, 53 - 59." type="journal article" year="1986">Okada 1986</bibRefCitation>
, “
<figureCitation box="[872,941,1019,1045]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="20" pageNumber="221">Fig. 6</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[989,1158,1019,1045]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Eye dark red (
<figureCitation box="[1339,1423,1019,1045]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[222,1364,181,1919]" captionTargetId="figure-20@19.[222,1364,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Habitus (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906035" httpUri="https://zenodo.org/record/7906035/files/figure.png" pageId="20" pageNumber="221">Fig. 14</figureCitation>
). Frons blackish brown; frontal vittae mat. Supracervical setae distally more or less curved, longer than inner, occipital setae (ch.11-0, ch.12-0;
<figureCitation box="[423,492,1091,1117]" captionStart="FIGURE 4" captionStartId="6.[151,250,1863,1887]" captionTargetBox="[162,1420,181,1839]" captionTargetId="figure-19@6.[157,1429,181,1839]" captionTargetPageId="6" captionText="FIGURE 4. Head parts (caudal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: dl a, dorsolateral tentorial apodeme; dm a, dorsomedial tentorial apodeme; i ocp s, inner occipital seta; o vt s, outer vertical seta; spc s, supracervical seta. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906007" httpUri="https://zenodo.org/record/7906007/files/figure.png" pageId="20" pageNumber="221">Fig. 4</figureCitation>
). Palpus pale grayish yellow, flat or convex in ventrodistal portion (ch.16-0;
<figureCitation box="[1363,1431,1091,1118]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="20" pageNumber="221">Fig. 5</figureCitation>
, “
<figureCitation box="[162,249,1127,1153]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">Fig. 1B</figureCitation>
” in
<bibRefCitation author="Okada, T." box="[301,441,1127,1153]" pageId="20" pageNumber="221" pagination="53 - 59" refId="ref21490" refString="Okada, T. (1986) Taximetrical analyses of costal chaetotaxy of the genus Drosophilella Duda, with description of a new species from Sri Lanka (Diptera, Drosophilidae). Proceedings of the Japanese Society of Systematic Zoology, 34, 53 - 59." type="journal article" year="1986">Okada 1986</bibRefCitation>
, “
<figureCitation box="[465,534,1127,1153]" captionStart="FIGURE 7" captionStartId="9.[151,250,1213,1237]" captionTargetBox="[151,1435,309,1190]" captionTargetId="figure-91@9.[151,1436,309,1190]" captionTargetPageId="9" captionText="FIGURE 7. Thoraxes (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906015" httpUri="https://zenodo.org/record/7906015/files/figure.png" pageId="20" pageNumber="221">Fig. 7</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[584,754,1127,1153]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). All cibarial sensilla very small or minute; anterior sensilla irregularly arranged in square or triangle (ch.18-1, ch.19-1;
<figureCitation box="[850,969,1163,1190]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="20" pageNumber="221">Fig. 5A–E</figureCitation>
<figureCitation box="[970,978,1182,1196]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="right" box="[970,978,1182,1196]" fontSize="6" pageId="20" pageNumber="221">2</subScript>
</figureCitation>
, “
<figureCitation box="[1007,1078,1163,1189]" captionStart="FIGURE 7" captionStartId="9.[151,250,1213,1237]" captionTargetBox="[151,1435,309,1190]" captionTargetId="figure-91@9.[151,1436,309,1190]" captionTargetPageId="9" captionText="FIGURE 7. Thoraxes (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906015" httpUri="https://zenodo.org/record/7906015/files/figure.png" pageId="20" pageNumber="221">Fig. 7</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[1133,1304,1163,1189]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Proboscis without a pair of lateral, spherical bumps covered with numerous, short, stout setae (ch.22-0;
<figureCitation box="[1229,1299,1199,1226]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="20" pageNumber="221">Fig. 5</figureCitation>
, “
<figureCitation box="[1324,1394,1199,1225]" captionStart="FIGURE 7" captionStartId="9.[151,250,1213,1237]" captionTargetBox="[151,1435,309,1190]" captionTargetId="figure-91@9.[151,1436,309,1190]" captionTargetPageId="9" captionText="FIGURE 7. Thoraxes (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906015" httpUri="https://zenodo.org/record/7906015/files/figure.png" pageId="20" pageNumber="221">Fig. 7</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[151,323,1235,1261]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Additional, presutural, dorsocentral setae absent (
<figureCitation box="[915,986,1235,1261]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="20" pageNumber="221">Fig. 6</figureCitation>
, “
<figureCitation box="[1012,1104,1235,1261]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">Fig. 1A</figureCitation>
” in
<bibRefCitation author="Okada, T." box="[1157,1301,1235,1261]" pageId="20" pageNumber="221" pagination="53 - 59" refId="ref21490" refString="Okada, T. (1986) Taximetrical analyses of costal chaetotaxy of the genus Drosophilella Duda, with description of a new species from Sri Lanka (Diptera, Drosophilidae). Proceedings of the Japanese Society of Systematic Zoology, 34, 53 - 59." type="journal article" year="1986">Okada 1986</bibRefCitation>
). Basal and apical, scutellar setae nearly parallel (
<figureCitation box="[591,663,1271,1297]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="20" pageNumber="221">Fig. 6</figureCitation>
). Halter grayish brown. Wing hyaline; veins yellow (
<figureCitation box="[1288,1360,1271,1297]" captionStart="FIGURE 8" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[190,1397,181,1919]" captionTargetId="figure-19@10.[190,1397,181,1919]" captionTargetPageId="10" captionText="FIGURE 8. Right wings of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, anterodistal portion (scale bars: 0.1 mm); 2, the whole (scale bars: 0.5 mm). Abbreviations: dm-m, discal-medial crossvein; M4, 4th branch of media." figureDoi="http://doi.org/10.5281/zenodo.7906019" httpUri="https://zenodo.org/record/7906019/files/figure.png" pageId="20" pageNumber="221">Fig. 8</figureCitation>
). Fore tarsomere II slightly elongated below (ch.34-1;
<figureCitation box="[689,806,1307,1333]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="20" pageNumber="221">Fig. 9A–E</figureCitation>
<figureCitation box="[807,815,1326,1340]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="right" box="[807,815,1326,1340]" fontSize="6" pageId="20" pageNumber="221">1</subScript>
</figureCitation>
, “
<figureCitation box="[840,926,1307,1333]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">Fig. 1C</figureCitation>
” in
<bibRefCitation author="Okada, T." box="[976,1116,1307,1333]" pageId="20" pageNumber="221" pagination="53 - 59" refId="ref21490" refString="Okada, T. (1986) Taximetrical analyses of costal chaetotaxy of the genus Drosophilella Duda, with description of a new species from Sri Lanka (Diptera, Drosophilidae). Proceedings of the Japanese Society of Systematic Zoology, 34, 53 - 59." type="journal article" year="1986">Okada 1986</bibRefCitation>
, “
<figureCitation box="[1138,1207,1307,1333]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="20" pageNumber="221">Fig. 9</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[1255,1424,1307,1333]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Preapical setae absent on all tibiae (ch.41-1;
<figureCitation box="[643,709,1343,1369]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="20" pageNumber="221">Fig. 9</figureCitation>
, “
<figureCitation box="[731,817,1343,1369]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">Fig. 1A</figureCitation>
” in
<bibRefCitation author="Okada, T." box="[862,1001,1343,1369]" pageId="20" pageNumber="221" pagination="53 - 59" refId="ref21490" refString="Okada, T. (1986) Taximetrical analyses of costal chaetotaxy of the genus Drosophilella Duda, with description of a new species from Sri Lanka (Diptera, Drosophilidae). Proceedings of the Japanese Society of Systematic Zoology, 34, 53 - 59." type="journal article" year="1986">Okada 1986</bibRefCitation>
). Dorsomedian, terminal, down-curved seta on tarsomere Vs not longer than claw (ch.42-0;
<figureCitation box="[750,819,1379,1405]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="20" pageNumber="221">Fig. 9</figureCitation>
). Mid tibia apicoventrally with 2 (1 long and 1 small) stout setae (ch.39-0;
<figureCitation box="[390,509,1415,1441]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="20" pageNumber="221">Fig. 9A–E</figureCitation>
<figureCitation box="[509,517,1434,1448]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="left" box="[509,517,1434,1448]" fontSize="6" pageId="20" pageNumber="221">2</subScript>
</figureCitation>
). Spiracles simple, small (ch.44-0). Male sternite VI pubescent entirely, without process (ch.48-0, ch.50-0;
<figureCitation box="[453,585,1451,1477]" captionStart="FIGURE 10" captionStartId="12.[151,250,1971,1995]" captionTargetBox="[155,1436,597,1948]" captionTargetId="figure-179@12.[151,1436,597,1948]" captionTargetPageId="12" captionText="FIGURE 10. Abdominal sternites of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female.Abbreviation: s, sternite. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906023" httpUri="https://zenodo.org/record/7906023/files/figure.png" pageId="20" pageNumber="221">Fig. 10A–E</figureCitation>
<figureCitation box="[585,593,1470,1484]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="left" box="[585,593,1470,1484]" fontSize="6" pageId="20" pageNumber="221">1</subScript>
</figureCitation>
). Epandrium anteroventrally much elongated, not fused to cercus, without peg-like, stout seta at posteroventral apex; apodeme narrow lobe on anteromedial to -dorsal margin or absent (ch.55- 1, ch.57-0, ch.58-0, ch.62-0; Figs 11-1A–C
<subScript attach="right" box="[631,651,1542,1556]" fontSize="6" pageId="20" pageNumber="221">1,2</subScript>
, 11-2D, E). Hypandrial setae absent (ch.67-1; Figs 11-1A–C
<subScript attach="left" box="[1325,1345,1542,1556]" fontSize="6" pageId="20" pageNumber="221">3,4</subScript>
, 11-2D, E). Postgonites fused to aedeagal sheath, forming together phallal-sheath tube distally more or less separated into a pair of lateral lobes (ch.71-1, ch.72-1, ch.73-0; Figs 11-1A–C
<subScript attach="right" box="[835,855,1614,1628]" fontSize="6" pageId="20" pageNumber="221">3,4</subScript>
, 11-2D, E). Phallapodeme mostly rod-like (ch.77-0; Figs 11-1A–C
<subScript attach="left" box="[311,331,1650,1664]" fontSize="6" pageId="20" pageNumber="221">3,4</subScript>
, 11-2D, E; “
<figureCitation box="[476,551,1631,1657]" captionStart="FIGURE 10" captionStartId="12.[151,250,1971,1995]" captionTargetBox="[155,1436,597,1948]" captionTargetId="figure-179@12.[151,1436,597,1948]" captionTargetPageId="12" captionText="FIGURE 10. Abdominal sternites of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female.Abbreviation: s, sternite. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906023" httpUri="https://zenodo.org/record/7906023/files/figure.png" pageId="20" pageNumber="221">Fig.10</figureCitation>
” in
<bibRefCitation author="Grimaldi, D. A." box="[600,769,1631,1657]" pageId="20" pageNumber="221" pagination="417 - 426" refId="ref20653" refString="Grimaldi, D. A. (1991) Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new record, and a new species from Nepal. Entomologica Scandinavica, 22, 417 - 426. https: // doi. org / 10.1163 / 187631291 X 00219" type="journal article" year="1991">Grimaldi 1991</bibRefCitation>
). Phallal guide absent (ch.78-0; Figs 11-1A–C
<subScript attach="left" box="[1294,1302,1650,1664]" fontSize="6" pageId="20" pageNumber="221">4</subScript>
, 11-2D, E). Epiproct nearly entirely pubescent (ch.79-0;
<figureCitation box="[658,790,1667,1693]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="20" pageNumber="221">Fig. 12A–E</figureCitation>
<figureCitation box="[789,797,1686,1700]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="left" box="[789,797,1686,1700]" fontSize="6" pageId="20" pageNumber="221">1</subScript>
</figureCitation>
). Epigynium pubescent only on dorsal portion (ch.82-0;
<figureCitation box="[151,284,1703,1729]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="20" pageNumber="221">Fig. 12A–E</figureCitation>
<figureCitation box="[283,291,1722,1736]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="left" box="[283,291,1722,1736]" fontSize="6" pageId="20" pageNumber="221">1</subScript>
</figureCitation>
). Hypogynial, anteroventral bridge absent (ch.85-1;
<figureCitation box="[890,1023,1703,1729]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="20" pageNumber="221">Fig. 12A–E</figureCitation>
<figureCitation box="[1022,1030,1722,1736]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="20" pageNumber="221">
<subScript attach="left" box="[1022,1030,1722,1736]" fontSize="6" pageId="20" pageNumber="221">2</subScript>
</figureCitation>
); valves separated from each other, distally differentiated into more or less narrow process from basal, broad portion, with only trichoid sensilla (ch.86- 0, ch.87-1, ch.88-1;
<figureCitation box="[377,460,1775,1801]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="20" pageNumber="221">Fig. 12</figureCitation>
).
</paragraph>
<paragraph blockId="20.[151,1437,767,1945]" box="[199,1203,1811,1838]" pageId="20" pageNumber="221">
<emphasis bold="true" box="[199,370,1811,1837]" pageId="20" pageNumber="221">Included taxa.</emphasis>
<taxonomicName baseAuthorityName="Okada" baseAuthorityYear="1986" box="[377,658,1811,1837]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="zeylanica">
<emphasis box="[377,658,1811,1837]" italics="true" pageId="20" pageNumber="221">Colocasiomyia zeylanica</emphasis>
</taxonomicName>
species subgroup; and
<taxonomicName authorityName="Zhang &amp; Gao &amp; Takano &amp; Yafuso &amp; Suwito &amp; Meleng &amp; Toda" authorityYear="2023" box="[919,1084,1811,1838]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="oligochaeta">
<emphasis box="[919,938,1812,1838]" italics="true" pageId="20" pageNumber="221">C</emphasis>
.
<emphasis box="[952,1084,1811,1837]" italics="true" pageId="20" pageNumber="221">oligochaeta</emphasis>
</taxonomicName>
subgroup.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="221" type="key">
<paragraph blockId="20.[151,1437,767,1945]" pageId="20" pageNumber="221">
<emphasis bold="true" box="[199,253,1847,1873]" pageId="20" pageNumber="221">Key.</emphasis>
The digital, multiple-entry key “
<taxonomicName baseAuthorityName="Okada" baseAuthorityYear="1986" box="[627,909,1847,1873]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="zeylanica">
<emphasis box="[627,909,1847,1873]" italics="true" pageId="20" pageNumber="221">Colocasiomyia zeylanica</emphasis>
</taxonomicName>
species group” to all species of this group has been constructed partly based on the character matrix of Appendix 2, and is available from the following website: https://evolgen.biol.se.tmu.ac.jp/Classification/index.jsp (Biological Classification and Identification System).
</paragraph>
</subSubSection>
</treatment>
</document>
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