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<document id="10B7C45FB0DFD01ABBC0341F4F3B0592" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F065FFFFFF42FAC8FB15F9A4" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Coprophanaeoides) bluti Genier &amp; Arnaud, new species" docType="treatment" docVersion="10" lastPageNumber="9" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="8" updateTime="1698234910999" updateUser="plazi">
<mods:mods id="2478CDF798C1215BEFFD5418C802C8AF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A5C3449A3E5474937A5BA129D28568C3">
<mods:title id="BAA80A2B41D9DDBFD2C463C4A411E8D6">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
</mods:titleInfo>
<mods:name id="5C66DACD4E6A622F1ECC1AD3103FD4D7" type="personal">
<mods:role id="FF9DCCB8F22071D76DB7BBC0DCCC7BED">
<mods:roleTerm id="3E5AFD40BBA9BF938F072988B5A46110">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5CA89BD712B1B279D00B0ED9E32E5C0E">François Génier</mods:namePart>
</mods:name>
<mods:name id="01E00C2AC3C40C9B4223135B5C272B93" type="personal">
<mods:role id="EA0D289EE39DCDDEE8C278A8EB8A06E9">
<mods:roleTerm id="A3F1C6C28104CBC953518794EA611479">Author</mods:roleTerm>
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<mods:namePart id="A98F7F9397ACF164B48ADAC37C768847">Patrick Arnaud</mods:namePart>
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<mods:typeOfResource id="42546591D9AD9C977DE77FBC563B93FC">text</mods:typeOfResource>
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<mods:title id="3D9C682520D4270B01B8D22E6624B585">Zootaxa</mods:title>
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<mods:part id="922DBB71301B7DD11D1B9A2E0841BB8E">
<mods:date id="FA342FE1F77EACA675D121493F90E6C7">2016</mods:date>
<mods:detail id="4F2AECE8780B4B350889ED828EF46467" type="volume">
<mods:number id="DF52548C9C27F789D82ED5B5C7A3C8C9">4099</mods:number>
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<mods:detail id="E26CC0E3D8277F04A29D6F8E710A9FCE" type="issue">
<mods:number id="0039A544DEC552757A38060250BEE58E">1</mods:number>
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<mods:classification id="7C6347CFF7A1894C37B0C3F9194BC765">journal volume</mods:classification>
<mods:identifier id="B4988817613991ED2465C31395D54B97" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
<mods:identifier id="9EFE2FF82AF38FA0131FFACE9FF64408" type="GBIF-Dataset">5d7eab55-66e2-43ed-b764-bac0c6e5b6d6</mods:identifier>
<mods:identifier id="E05B1BF91F4150B05A27A065CC443EDE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E27C20C0A8E6B9A0C6DE642B0157FDF4" type="Zenodo-Dep">268274</mods:identifier>
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<treatment id="03C68782F065FFFFFF42FAC8FB15F9A4" ID-DOI="http://doi.org/10.5281/zenodo.5780252" ID-GBIF-Taxon="127631974" ID-Zenodo-Dep="5780252" LSID="urn:lsid:plazi:treatment:03C68782F065FFFFFF42FAC8FB15F9A4" httpUri="http://treatment.plazi.org/id/03C68782F065FFFFFF42FAC8FB15F9A4" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="C375651FF065FFF0FF42FAC8FBD6FAEF" box="[151,1072,1317,1343]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BD03694F065FFF0FF42FAC8FBD6FAEF" blockId="7.[151,1072,1317,1376]" box="[151,1072,1317,1343]" pageId="7" pageNumber="8">
<heading id="D09881F8F065FFF0FF42FAC8FBD6FAEF" bold="true" box="[151,1072,1317,1343]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B91BEA86F065FFF0FF42FAC8FBD6FAEF" bold="true" box="[151,1072,1317,1343]" pageId="7" pageNumber="8">
1.
<taxonomicName id="4C6F4D17F065FFF0FF61FAC8FC75FAEF" ID-CoL="34R29" authority="Genier &amp; Arnaud" authorityName="Genier &amp; Arnaud" box="[180,915,1317,1343]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bluti" status="sp. nov." subGenus="Coprophanaeoides">
<emphasis id="B91BEA86F065FFF0FF61FAC8FD4DFAEF" bold="true" box="[180,683,1317,1343]" italics="true" pageId="7" pageNumber="8">Dendropaemon (Coprophanaeoides) bluti</emphasis>
Génier &amp; Arnaud
</taxonomicName>
,
<taxonomicNameLabel id="A22857FDF065FFF0FC75FAC8FBD6FAEF" box="[928,1072,1317,1343]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF065FFF0FF42FAAAFE82FAB0" box="[151,356,1351,1376]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BD03694F065FFF0FF42FAAAFE82FAB0" blockId="7.[151,1072,1317,1376]" box="[151,356,1351,1376]" pageId="7" pageNumber="8">
(
<figureCitation id="13542A11F065FFF0FF4AFAAAFF0AFAB0" box="[159,236,1351,1376]" captionStart="FIGURES 1 – 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 – 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="7" pageNumber="8">Figs. 1</figureCitation>
,
<figureCitation id="13542A11F065FFF0FF2FFAA5FEC0FAB0" box="[250,294,1352,1376]" captionStart="FIGURES 110 – 117" captionStartId="112.[151,264,1933,1956]" captionTargetBox="[219,1362,202,1901]" captionTargetId="figure@112.[212,1375,193,1913]" captionTargetPageId="112" captionText="FIGURES 110 – 117. Metatarsus, ventral view. 110. D. bluti; 111. D. carinifer; 112. D. aenigmaticus; 113. D. amazonicus; 114. D. angustulus; 115. D. viridis; 116. D. viridipennis; 117. D. convexus." httpUri="https://zenodo.org/record/268282/files/figure.png" pageId="7" pageNumber="8">110</figureCitation>
,
<figureCitation id="13542A11F065FFF0FEE4FAAAFEBAFAB0" box="[305,348,1351,1376]" captionStart="FIGURE 156" captionStartId="116.[151,250,1237,1259]" captionTargetBox="[150,1436,193,1215]" captionTargetId="figure@116.[150,1436,193,1215]" captionTargetPageId="116" captionText="FIGURE 156. Known distribution of D. bluti, D. carinifer, D. compressipennis, D. cribrosus, D. furtadoi, D. hirticollis, D. inflatus, D. pauliani, D. pilosissimus, D. renatii." httpUri="https://zenodo.org/record/268286/files/figure.png" pageId="7" pageNumber="8">156</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF065FFF0FF42FA62FCF1FA78" box="[151,791,1423,1448]" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BD03694F065FFF0FF42FA62FCF1FA78" blockId="7.[151,1436,1423,2025]" box="[151,791,1423,1448]" pageId="7" pageNumber="8">
<emphasis id="B91BEA86F065FFF0FF42FA62FECBFA78" bold="true" box="[151,301,1423,1448]" pageId="7" pageNumber="8">
<typeStatus id="54D48836F065FFF0FF42FA62FF34FA78" box="[151,210,1423,1448]" pageId="7" pageNumber="8">Type</typeStatus>
locality
</emphasis>
. Culiseu, Quellgebiet des Xingu,
<collectingCountry id="F3787604F065FFF0FD7CFA62FCF4FA78" box="[681,786,1423,1448]" name="Brazil" pageId="7" pageNumber="8">Brasilien</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF065FFF0FF12FA59FF01F9AF" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BD03694F065FFF0FF12FA59FF01F9AF" blockId="7.[151,1436,1423,2025]" pageId="7" pageNumber="8">
<emphasis id="B91BEA86F065FFF0FF12FA59FEDCFA1D" bold="true" box="[199,314,1460,1485]" pageId="7" pageNumber="8">Diagnosis</emphasis>
. The acutely notched external side of each clypeal tooth will place this species in
<taxonomicName id="4C6F4D17F065FFF0FF42FA3AFE8DFA20" box="[151,363,1495,1520]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F065FFF0FF42FA3AFE8DFA20" box="[151,363,1495,1520]" italics="true" pageId="7" pageNumber="8">Coprophanaeoides</emphasis>
</taxonomicName>
. Within
<taxonomicName id="4C6F4D17F065FFF0FE0BFA3AFD54FA20" box="[478,690,1495,1520]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F065FFF0FE0BFA3AFD54FA20" box="[478,690,1495,1520]" italics="true" pageId="7" pageNumber="8">Coprophanaeoides</emphasis>
</taxonomicName>
, the completely glabrous pronotum will separate it from
<emphasis id="B91BEA86F065FFF0FA54FA34FB78F9C5" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C6F4D17F065FFF0FA54FA34FF1FF9C4" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="carinifer">D. carinifer</taxonomicName>
,
<taxonomicName id="4C6F4D17F065FFF0FED3FA10FE01F9C4" box="[262,487,1533,1556]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="compressipennis">D. compressipennis</taxonomicName>
,
<taxonomicName id="4C6F4D17F065FFF0FE20FA10FD62F9C5" box="[501,644,1532,1557]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cribrosus">D. cribrosus</taxonomicName>
,
<taxonomicName id="4C6F4D17F065FFF0FD47FA10FCF2F9C5" box="[658,788,1532,1557]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="furtadoi">D. furtadoi</taxonomicName>
,
<taxonomicName id="4C6F4D17F065FFF0FCF7FA10FC55F9C5" box="[802,947,1532,1557]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hirticollis">D. hirticollis</taxonomicName>
and
<taxonomicName id="4C6F4D17F065FFF0FC25FA10FB78F9C5" box="[1008,1182,1532,1557]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pilosissimus">D. pilosissimus</taxonomicName>
</emphasis>
. The distinct pronotal posterior edge will separate it from.
<taxonomicName id="4C6F4D17F065FFF0FDFEF9CCFD08F9E8" box="[555,750,1569,1592]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="inemarginatus">
<emphasis id="B91BEA86F065FFF0FDFEF9CCFD08F9E8" box="[555,750,1569,1592]" italics="true" pageId="7" pageNumber="8">D. inemarginatus</emphasis>
</taxonomicName>
. Its smaller size and well defines elytral striae will separate it from
<taxonomicName id="4C6F4D17F065FFF0FF03F9A8FEA9F98D" box="[214,335,1604,1629]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="B91BEA86F065FFF0FF03F9A8FEA9F98D" box="[214,335,1604,1629]" italics="true" pageId="7" pageNumber="8">D. inflatus</emphasis>
</taxonomicName>
and finally the smaller eyes and less produced anterior pronotal angles will separate it from
<taxonomicName id="4C6F4D17F065FFF0FA54F9A8FF06F950" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F065FFF0FA54F9A8FF06F950" italics="true" pageId="7" pageNumber="8">D. renatii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF065FFFFFF12F961FC16FA10" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8BD03694F065FFFFFF12F961FD3DFADD" blockId="7.[151,1436,1423,2025]" lastBlockId="8.[151,1437,151,1653]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B91BEA86F065FFF0FF12F961FEBFF975" bold="true" box="[199,345,1676,1701]" pageId="7" pageNumber="8">Description.</emphasis>
Female
<typeStatus id="54D48836F065FFF0FE68F961FDF9F975" box="[445,543,1676,1701]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F065FFF0FDE5F961FD92F974" box="[560,628,1676,1701]" captionStart="FIGURES 1 – 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 – 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
).
<emphasis id="B91BEA86F065FFF0FD5BF961FD2DF975" bold="true" box="[654,715,1676,1701]" pageId="7" pageNumber="8">Body</emphasis>
. Body moderately large, length
<quantity id="4C979B71F065FFF0FBEAF961FB41F974" box="[1087,1191,1676,1701]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="8" unit="mm" value="12.5">12.5 mm</quantity>
, maximum width 7.0 mm; body subrectangular in dorsal view; dorsum narrowly flat.
<emphasis id="B91BEA86F065FFF0FCA8F942FC58F918" bold="true" box="[893,958,1711,1736]" pageId="7" pageNumber="8">Color</emphasis>
. Dorsal surface black, glossy, with green metallic sheen; head with green metallic sheen on genae and frons; pronotum with green metallic sheen except disc and irregular area laterally; elytra with uniform green metallic sheen; ventrum with faint greenish metallic sheen; pygidium with green metallic sheen; legs with coppery and greenish metallic sheen on femora and tibiae.
<emphasis id="B91BEA86F065FFF0FA80F8F1FA73F8E5" bold="true" box="[1365,1429,1820,1845]" pageId="7" pageNumber="8">Head</emphasis>
. Clypeus broadly arcuate, anterior portion upturned; clypeal teeth ogival; clypeal median emargination narrowly vshaped, clypeal edge acutely notched on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin ill-defined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with ill-defined rugulae and minute tubercles laterally, smooth internally, lacking distinct transverse carina, simply convex; clypeofrontal carina low, more than 6 times wider than high, straight in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes small in dorsal view, interocular ratio 4.7.
<emphasis id="B91BEA86F06AFFFFFCA2FF51FC16FF05" bold="true" box="[887,1008,188,213]" pageId="8" pageNumber="9">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.5; disc of pronotum smooth basally, with transverse rugulae anteriorly, with an illdefined shallow longitudinal depression on posterior half; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a transverse anteriorly arcuate carina; anterior angles surface finely granulate, similar to lateral margin along posterior edge of anterior margin; lateral fossae oval, bordered laterally by a blunt carina and anteriorly by a blunt tubercle; lateral portions strongly explanate; pronotal basal fossae ill-defined, slightly concave; posterior margin well-defined, lacking crenulation and setae.
<emphasis id="B91BEA86F06AFFFFFC18FE79FBFFFE7D" bold="true" box="[973,1049,404,429]" pageId="8" pageNumber="9">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.1; elytral base distinctly marginate; elytral striae 1–4 moderately wide basally and fine apically, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures fine and well-defined, adjacent strial edge encroaching on interval, stria 1 weakly impressed apically, going straight to elytral apical margin; interstriae slightly convex, minutely punctate throughout, surface glossy.
<emphasis id="B91BEA86F06AFFFFFF42FDAAFEB2FDB0" bold="true" box="[151,340,583,608]" pageId="8" pageNumber="9">Thoracic sterna</emphasis>
. Proepisternal carina absent; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, y-shaped.
<emphasis id="B91BEA86F06AFFFFFE2BFD81FDD0FD55" bold="true" box="[510,566,620,645]" pageId="8" pageNumber="9">Legs</emphasis>
. Profemur posterior surface slightly but distinctly convex and glabrous internally, posterointernal margin rather thin, uneven, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with rugose irregular punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge straight in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, three-segmented, first segment short, approximately as long as wide at apex. Metafemur broadly oval in anterior view, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with an irregular fine sulcus medially. Metatibia rather slender, gradually widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy, metatibial posterior surface flat between longitudinal row of setae and lateral edge, glossy between punctures. Metatarsus three-segmented (
<figureCitation id="13542A11F06AFFFFFC8CFB89FC2FFBAC" box="[857,969,1124,1149]" captionStart="FIGURES 110 – 117" captionStartId="112.[151,264,1933,1956]" captionTargetBox="[219,1362,202,1901]" captionTargetId="figure@112.[212,1375,193,1913]" captionTargetPageId="112" captionText="FIGURES 110 – 117. Metatarsus, ventral view. 110. D. bluti; 111. D. carinifer; 112. D. aenigmaticus; 113. D. amazonicus; 114. D. angustulus; 115. D. viridis; 116. D. viridipennis; 117. D. convexus." httpUri="https://zenodo.org/record/268282/files/figure.png" pageId="8" pageNumber="9">Fig. 110</figureCitation>
), first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
<emphasis id="B91BEA86F06AFFFFFEF6FB41FDF3FB15" bold="true" box="[291,533,1196,1221]" pageId="8" pageNumber="9">Abdominal sternites</emphasis>
. Sternites 3–6 longitudinally flat; sternites 4–6 with 1–3 unaligned rows of setae laterally, glabrous medially; sternite 7 approximately longitudinally flat medially, subequal in length to segment 6 along midline; pygidium minutely punctate on disc.
</paragraph>
<paragraph id="8BD03694F06AFFFFFF12FAFAFD2EFAFF" blockId="8.[151,1437,151,1653]" box="[199,712,1303,1328]" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF12FAFAFE90FAE0" bold="true" box="[199,374,1303,1328]" pageId="8" pageNumber="9">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF06AFFFFFE5DFAF5FE0AFAE0" box="[392,492,1303,1328]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
). Length:
<quantity id="4C979B71F06AFFFFFD88FAFAFD22FAFF" box="[605,708,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="9" unit="mm" value="12.5">12.5 mm</quantity>
.
</paragraph>
<paragraph id="8BD03694F06AFFFFFF12FAD1FC16FA10" blockId="8.[151,1437,151,1653]" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF12FAD1FE4DFA85" bold="true" box="[199,427,1340,1365]" pageId="8" pageNumber="9">
Primary
<typeStatus id="54D48836F06AFFFFFEEDFAD1FE8DFA85" box="[312,363,1340,1365]" pageId="8" pageNumber="9">type</typeStatus>
data
</emphasis>
.
<typeStatus id="54D48836F06AFFFFFE6BFAD1FDC0FA85" box="[446,550,1340,1365]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
female (ZMHB): [♀]; [Campos] handwritten, green card; [1.-
<date id="FFD11054F06AFFFFFB22FAD1FA94FA84" box="[1271,1394,1340,1364]" pageId="8" pageNumber="9" value="1887-10-09">10.9.1887</date>
/ 2. Xingu-Expd.] handwritten; [
<collectingCountry id="F3787604F06AFFFFFE07FAB2FDDFFAA8" box="[466,569,1375,1400]" name="Brazil" pageId="8" pageNumber="9">Brasilien</collectingCountry>
/ Quellgebiet d. Xingu,/ Culiseu/Ehrenreiche S.] green card; [
<taxonomicName id="4C6F4D17F06AFFFFFB31FAB2FA75FAA8" box="[1252,1427,1375,1400]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F06AFFFFFF42FA69FEDEFA4D" authority="Olsuf." authorityName="Olsuf." box="[151,312,1412,1437]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="renatii">renatii Olsuf.</taxonomicName>
♀/ Xingu, Blut det. 1938] handwritten; [WORLD/ SCARAB./ DATABASE/ WSD00016732];
<typeStatus id="54D48836F06AFFFFFF42FA45FECCFA10" box="[151,298,1448,1472]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C6F4D17F06AFFFFFEECFA4AFE00FA10" box="[313,486,1447,1472]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F06AFFFFFEECFA4AFE00FA10" box="[313,486,1447,1472]" italics="true" pageId="8" pageNumber="9">Dendropaemon</emphasis>
</taxonomicName>
/
<taxonomicName id="4C6F4D17F06AFFFFFE20FA4AFDCEFA10" box="[501,552,1447,1472]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bluti" status="sp. nov.">
<emphasis id="B91BEA86F06AFFFFFE20FA4AFDCEFA10" box="[501,552,1447,1472]" italics="true" pageId="8" pageNumber="9">bluti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857FDF06AFFFFFDE5FA45FD84FA10" box="[560,610,1448,1472]" pageId="8" pageNumber="9" rank="species">n.sp.</taxonomicNameLabel>
/ Génier &amp; Arnaud, 2014] red card.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06AFFFFFF12FA21FD61FA35" box="[199,647,1484,1509]" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BD03694F06AFFFFFF12FA21FD61FA35" blockId="8.[151,1437,151,1653]" box="[199,647,1484,1509]" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF12FA21FE4FFA35" bold="true" box="[199,425,1484,1509]" pageId="8" pageNumber="9">Material examined</emphasis>
. Primary
<typeStatus id="54D48836F06AFFFFFDCDFA20FDAEFA35" box="[536,584,1485,1509]" pageId="8" pageNumber="9">type</typeStatus>
only.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06AFFFFFF12FA02FDE4F9FD" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BD03694F06AFFFFFF12FA02FB47F9D8" blockId="8.[151,1437,151,1653]" box="[199,1185,1519,1544]" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF12FA02FEA1F9D8" bold="true" box="[199,327,1519,1544]" pageId="8" pageNumber="9">Etymology</emphasis>
. A patronym in honor of Heinrich Blut who published on the genus in 1939.
</paragraph>
<paragraph id="8BD03694F06AFFFFFF12F9F9FDE4F9FD" blockId="8.[151,1437,151,1653]" box="[199,514,1556,1581]" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF12F9F9FE9BF9FD" bold="true" box="[199,381,1556,1581]" pageId="8" pageNumber="9">Natural history</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06AFFFFFF12F9DAFB15F9A4" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BD03694F06AFFFFFF12F9DAFD6AF980" blockId="8.[151,1437,151,1653]" box="[199,652,1591,1616]" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF12F9DAFED2F980" bold="true" box="[199,308,1591,1616]" pageId="8" pageNumber="9">Remarks</emphasis>
.
<collectingCountry id="F3787604F06AFFFFFE97F9DAFE9AF980" box="[322,380,1591,1616]" name="Malta" pageId="8" pageNumber="9">Male</collectingCountry>
and variation unknown.
</paragraph>
<paragraph id="8BD03694F06AFFFFFF12F9B1FB15F9A4" blockId="8.[151,1437,151,1653]" box="[199,1267,1628,1653]" pageId="8" pageNumber="9">
Blut misidentified this species as
<taxonomicName id="4C6F4D17F06AFFFFFDEBF9B0FD4DF9A4" box="[574,683,1629,1652]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F06AFFFFFDEBF9B0FD4DF9A4" box="[574,683,1629,1652]" italics="true" pageId="8" pageNumber="9">D. renatii</emphasis>
</taxonomicName>
and redescribed
<taxonomicName id="4C6F4D17F06AFFFFFCBCF9B0FC33F9A4" box="[873,981,1629,1652]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F06AFFFFFCBCF9B0FC33F9A4" box="[873,981,1629,1652]" italics="true" pageId="8" pageNumber="9">D. renatii</emphasis>
</taxonomicName>
as
<taxonomicName id="4C6F4D17F06AFFFFFC2FF9B0FB0AF9A5" box="[1018,1260,1628,1653]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="refulgens" subSpecies="olsufieffi">
<emphasis id="B91BEA86F06AFFFFFC2FF9B0FB0AF9A5" box="[1018,1260,1628,1653]" italics="true" pageId="8" pageNumber="9">D. refulgens olsufieffi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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