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<document id="A51333017E6D7D7C35C15E5F15D791F0" ID-CLB-Dataset="45831" ID-DOI="10.5281/zenodo.279398" ID-GBIF-Dataset="34ce6298-fc68-42cb-8075-40fb85ddf824" ID-ISSN="1175-5326" ID-Zenodo-Dep="279398" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460417465786" checkinUser="plazi" docAuthor="Pilato, Giovanni, Kiosya, Yevgen, Lisi, Oscar, Inshina, Valentina &amp; Biserov, Vladimir" docDate="2011" docId="03C35E455800FFF4FF431AC12421FEE4" docLanguage="en" docName="zt03123p031.pdf" docOrigin="Zootaxa 3123" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hypsibius pallidoides Pilato, Kiosya, Lisi, Inshina &amp; Biserov, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FFFA263D580CFFFAFFD41B482036FFF8" masterDocTitle="Annotated list of Tardigrada records from Ukraine with the description of three new species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="13" updateTime="1698329079364" updateUser="ExternalLinkService">
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<mods:title id="67B8FC03326912CA0DDD2354DD697412">Annotated list of Tardigrada records from Ukraine with the description of three new species</mods:title>
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<mods:namePart id="5C40B7B94CA3B43D128931610A9156D4">Pilato, Giovanni</mods:namePart>
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<mods:namePart id="CAE0764AB98076205A1F82F21BEFA76B">Lisi, Oscar</mods:namePart>
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<mods:namePart id="8D1534D9A7FA00E8801D1BDCB99FCBC1">Inshina, Valentina</mods:namePart>
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<mods:namePart id="2303F302A275FBBBF60B60F50026DD8E">Biserov, Vladimir</mods:namePart>
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<treatment id="03C35E455800FFF4FF431AC12421FEE4" ID-DOI="http://doi.org/10.5281/zenodo.5616582" ID-GBIF-Taxon="119425565" ID-Zenodo-Dep="5616582" LSID="urn:lsid:plazi:treatment:03C35E455800FFF4FF431AC12421FEE4" httpUri="http://treatment.plazi.org/id/03C35E455800FFF4FF431AC12421FEE4" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="C370BCD85800FFF6FF431AC120DDFE3C" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BD5EF535800FFF6FF431AC121CBFE5A" blockId="12.[151,509,392,452]" box="[151,509,392,419]" pageId="12" pageNumber="13">
<heading id="D09D583F5800FFF6FF431AC121CBFE5A" bold="true" box="[151,509,392,419]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B91E33415800FFF6FF431AC121CBFE5A" bold="true" box="[151,509,392,419]" pageId="12" pageNumber="13">
<taxonomicName id="4C6A94D05800FFF6FF431AC121AEFE5B" box="[151,408,393,419]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415800FFF6FF431AC121AEFE5B" bold="true" box="[151,408,393,419]" italics="true" pageId="12" pageNumber="13">Hypsibius pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5800FFF6FE741AC021CBFE5A" box="[416,509,392,418]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD5EF535800FFF6FF431AE420DDFE3C" blockId="12.[151,509,392,452]" box="[151,235,428,452]" pageId="12" pageNumber="13">
(
<figureCitation id="1351F3D65800FFF6FF4B1AE420D5FE3C" box="[159,227,428,452]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C370BCD85800FFF7FF431ABB2119FE17" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BD5EF535800FFF6FF431ABB2253FDAC" blockId="12.[151,1436,499,596]" pageId="12" pageNumber="13">
<emphasis id="B91E33415800FFF6FF431ABB21B9FDF4" bold="true" box="[151,399,499,524]" pageId="12" pageNumber="13">Material examined.</emphasis>
Kherson Oblast, Ivano-Rybalchansky district of Chernomorsky biosphere reserve (46°27ʹ25ʺN, 32°8ʹ56ʺE), collector D.A. Korolesova, moss on wood,
<date id="FFD4C9935800FFF6FC6F19502403FDD7" box="[955,1077,535,559]" pageId="12" pageNumber="13" value="2008-06">June 2008</date>
:
<typeStatus id="54D151F15800FFF6FB96195F2491FDD7" box="[1090,1191,535,559]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, seven
<typeStatus id="54D151F15800FFF6FB2B1950255CFDC8" box="[1279,1386,536,560]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
and additional
<specimenCount id="9D6C24DA5800FFF6FEDA19742190FDAC" box="[270,422,572,596]" pageId="12" pageNumber="13" type="generic">27 specimens</specimenCount>
and nine exuvia.
</paragraph>
<caption id="DF15BFDB5800FFF6FF431C022228F845" httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="12" pageNumber="13" targetBox="[164,1420,603,1851]" targetPageId="12">
<paragraph id="8BD5EF535800FFF6FF431C022228F845" blockId="12.[151,1437,1866,1981]" pageId="12" pageNumber="13">
<emphasis id="B91E33415800FFF6FF431C022125F898" bold="true" box="[151,275,1866,1889]" pageId="12" pageNumber="13">FIGURE 7.</emphasis>
AD,
<taxonomicName id="4C6A94D05800FFF6FE8D1C032200F899" box="[345,566,1867,1889]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415800FFF6FE8D1C032200F899" box="[345,566,1867,1889]" italics="true" pageId="12" pageNumber="13">Hypsibius pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5800FFF6FDE81C0322B8F899" box="[572,654,1867,1889]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B91E33415800FFF6FDE81C0322B1F898" bold="true" box="[572,647,1867,1888]" pageId="12" pageNumber="13">sp. nov</emphasis>
.
</taxonomicNameLabel>
: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E,
<taxonomicName id="4C6A94D05800FFF6FC601CC02442F866" box="[948,1140,1928,1950]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="pallidus">
<emphasis id="B91E33415800FFF6FC601CC02442F866" box="[948,1140,1928,1950]" italics="true" pageId="12" pageNumber="13">Hypsibius pallidus</emphasis>
</taxonomicName>
: Bucco-pharyngal apparatus (slide No.1251). (Scale bars = 10 µm).
</paragraph>
</caption>
<paragraph id="8BD5EF535801FFF7FF131BD0216CFEE2" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
<emphasis id="B91E33415801FFF7FF131BD021BEFF48" bold="true" box="[199,392,151,176]" pageId="13" pageNumber="14">
<typeStatus id="54D151F15801FFF7FF131BD02134FF48" box="[199,258,152,176]" pageId="13" pageNumber="14">Type</typeStatus>
repository.
</emphasis>
<typeStatus id="54D151F15801FFF7FE5A1BDF21CFFF57" box="[398,505,151,175]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
, seven
<typeStatus id="54D151F15801FFF7FD9E1BD02283FF48" box="[586,693,152,176]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
and three exuvia (slide No.5430) are deposited in the collection of Binda &amp; Pilato (Museum of the Department of Animal Biology “Marcello La Greca”, University of Catania,
<collectingCountry id="F37DAFC35801FFF7FA8C1BF325A6FF2B" box="[1368,1424,187,211]" name="Italy" pageId="13" pageNumber="14">Italy</collectingCountry>
). Other specimens are deposited in the Kiosya's collection (School of Biology,
<collectingCountry id="F37DAFC35801FFF7FBD61BA82403FF0F" box="[1026,1077,224,247]" name="Vietnam" pageId="13" pageNumber="14">V.N.</collectingCountry>
Karazin Kharkiv National University,
<collectingCountry id="F37DAFC35801FFF7FF3B1A4A2178FEE2" box="[239,334,258,282]" name="Ukraine" pageId="13" pageNumber="14">Ukraine</collectingCountry>
).
</paragraph>
<paragraph id="8BD5EF535801FFF7FF131A6D2119FE17" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
<emphasis id="B91E33415801FFF7FF131A6D2197FEC6" bold="true" box="[199,417,293,318]" pageId="13" pageNumber="14">Specific diagnosis.</emphasis>
Eye spots present; cuticle smooth; buccal armature without teeth; buccal tube very narrow; pharyngeal bulb with two macroplacoids and a very small, dot-shaped, septulum; claws similar to
<taxonomicName id="4C6A94D05801FFF7FB3C1A012088FE7D" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidus">
<emphasis id="B91E33415801FFF7FB3C1A012088FE7D" italics="true" pageId="13" pageNumber="14">Hypsibius pallidus</emphasis>
</taxonomicName>
(i.e. the main branches of the external claws were inserted on the secondary branches well above the claw base); accessory points present; lunules absent; no cuticular bar on the first three pairs of legs; the cuticular bar between the claws on the hind legs seemed to be absent, but this character needs to be confirmed. Smooth eggs laid in the exuvia.
</paragraph>
</subSubSection>
<subSubSection id="C370BCD85801FFF7FF131AB2230DFBFB" pageId="13" pageNumber="14" type="description">
<paragraph id="8BD5EF535801FFF7FF131AB222BDFD1F" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
<emphasis id="B91E33415801FFF7FF131AB22223FDEB" bold="true" box="[199,533,506,531]" pageId="13" pageNumber="14">
Description of the
<typeStatus id="54D151F15801FFF7FE731AB32226FDEB" box="[423,528,507,531]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
.
</emphasis>
Body length 252 µm; colorless, cuticle smooth without cuticular pores or dots. Eye spots present. Bucco-pharyngeal apparatus of
<taxonomicName id="4C6A94D05801FFF7FD2119562353FDCE" box="[757,869,542,566]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="genus">
<emphasis id="B91E33415801FFF7FD2119562353FDCE" box="[757,869,542,566]" italics="true" pageId="13" pageNumber="14">Hypsibius</emphasis>
</taxonomicName>
<typeStatus id="54D151F15801FFF7FCA719572395FDCF" box="[883,931,543,567]" pageId="13" pageNumber="14">type</typeStatus>
(
<figureCitation id="1351F3D65801FFF7FC6D19562476FDCE" box="[953,1088,542,566]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Fig. 7A, B</figureCitation>
). Mouth subterminal without peribuccal lamellae; small buccal cavity without rings of teeth. Narrow, rigid buccal tube, without ventral lamina, 22.3 µm long and 1.5 µm wide externally (
<emphasis id="B91E33415801FFF7FD5D192E22A8FD85" box="[649,670,614,637]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
= 6.7); stylet supports inserted on the buccal tube at 55.5 % of its length (
<emphasis id="B91E33415801FFF7FF3819C12137FD58" box="[236,257,649,672]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
= 55.5). Pharyngeal bulb (20.0 µm × 18.2 µm) with apophyses, two macroplacoids and a very small, dotshaped septulum (
<figureCitation id="1351F3D65801FFF7FEBE19E421DAFD3C" box="[362,492,684,708]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Fig. 7A, B</figureCitation>
, arrow). First macroplacoid 3.8 µm long (
<emphasis id="B91E33415801FFF7FC3419E523C0FD3C" box="[992,1014,685,708]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
=17.0), second 2.6 µm (
<emphasis id="B91E33415801FFF7FACB19E52503FD3C" box="[1311,1333,685,708]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
= 11.7); macroplacoid row length 6.9 µm (
<emphasis id="B91E33415801FFF7FDC21999221AFD10" box="[534,556,721,744]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
= 30.9).
</paragraph>
<paragraph id="8BD5EF535801FFF7FF1319BB20FDFC18" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
Claws of the
<taxonomicName id="4C6A94D05801FFF7FE8E19BB21FCFCF3" box="[346,458,755,779]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="genus">
<emphasis id="B91E33415801FFF7FE8E19BB21FCFCF3" box="[346,458,755,779]" italics="true" pageId="13" pageNumber="14">Hypsibius</emphasis>
</taxonomicName>
<typeStatus id="54D151F15801FFF7FE0519BC2232FCF4" box="[465,516,756,780]" pageId="13" pageNumber="14">type</typeStatus>
, well developed; the main branches of the external claws were inserted on the secondary branches well above the claw base (
<figureCitation id="1351F3D65801FFF7FDA9185E22C0FCD6" box="[637,758,790,814]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Fig. 7C, D</figureCitation>
) as in
<taxonomicName id="4C6A94D05801FFF7FCEB185E2426FCD6" box="[831,1040,790,814]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidus">
<emphasis id="B91E33415801FFF7FCEB185E2426FCD6" box="[831,1040,790,814]" italics="true" pageId="13" pageNumber="14">Hypsibius pallidus</emphasis>
</taxonomicName>
. Main brancheses of all claws with very thin accessory points. External and internal claws on the second and third pairs of legs 12.7 µm (
<emphasis id="B91E33415801FFF7FAF71873250FFCAA" box="[1315,1337,827,850]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
= 57.0) and 7.6 µm long (
<emphasis id="B91E33415801FFF7FE8A18162142FC8D" box="[350,372,862,885]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
= 34.1) respectively; claw orientation on the hind legs did not allow correct measurements, but in
<tableCitation id="C6E8DAE85801FFF7FF6018C82131FC60" box="[180,263,896,920]" captionStart="TABLE 4" captionStartId="14.[151,235,329,350]" captionTargetBox="[151,1436,409,1567]" captionTargetPageId="14" captionText="TABLE 4. Measurements of some structures of the holotype and two paratypes of Hypsibius pallidoides sp. nov., a specimen of H. pallidus and the holotype of H. septulatus." httpUri="http://table.plazi.org/id/DF15BFDB5802FFF4FF431A0122B6FE86" pageId="13" pageNumber="14" tableUuid="DF15BFDB5802FFF4FF431A0122B6FE86">Table 4</tableCitation>
the measurements of two
<typeStatus id="54D151F15801FFF7FDE418C922ADFC61" box="[560,667,897,921]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
are shown. Lunules absent; a cuticular bar between the claws of the hind legs seemed to be absent, but this character needs to be confirmed; no cuticular bar on the first three pairs of legs.
</paragraph>
<paragraph id="8BD5EF535801FFF7FF1318A2230DFBFB" blockId="13.[151,1436,151,2020]" box="[199,827,1002,1028]" pageId="13" pageNumber="14">
<emphasis id="B91E33415801FFF7FF1318A221E4FBFB" bold="true" box="[199,466,1002,1027]" pageId="13" pageNumber="14">Decription of the eggs.</emphasis>
Smooth eggs laid in the exuvia.
</paragraph>
</subSubSection>
<subSubSection id="C370BCD85801FFF7FF131F472242FBB2" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BD5EF535801FFF7FF131F472242FBB2" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
<emphasis id="B91E33415801FFF7FF131F47210DFBDF" bold="true" box="[199,315,1039,1063]" pageId="13" pageNumber="14">Remarks.</emphasis>
The
<typeStatus id="54D151F15801FFF7FEA21F4721D7FBDF" box="[374,481,1039,1063]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
were similar to the
<typeStatus id="54D151F15801FFF7FD101F462310FBDE" box="[708,806,1038,1062]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
in qualitative and metric characters. The measurements of three specimens are reported in
<tableCitation id="C6E8DAE85801FFF7FDCE1F7A2246FBB2" box="[538,624,1074,1098]" captionStart="TABLE 4" captionStartId="14.[151,235,329,350]" captionTargetBox="[151,1436,409,1567]" captionTargetPageId="14" captionText="TABLE 4. Measurements of some structures of the holotype and two paratypes of Hypsibius pallidoides sp. nov., a specimen of H. pallidus and the holotype of H. septulatus." httpUri="http://table.plazi.org/id/DF15BFDB5802FFF4FF431A0122B6FE86" pageId="13" pageNumber="14" tableUuid="DF15BFDB5802FFF4FF431A0122B6FE86">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C370BCD85801FFF4FF131F1E2421FEE4" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BD5EF535801FFF7FF131F1E20F3FB69" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
<emphasis id="B91E33415801FFF7FF131F1E2178FB96" bold="true" box="[199,334,1110,1134]" pageId="13" pageNumber="14">Etymology.</emphasis>
The specific name
<taxonomicName id="4C6A94D05801FFF7FDFB1F1E229AFB96" box="[559,684,1110,1134]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides">
<emphasis id="B91E33415801FFF7FDFB1F1E229AFB96" box="[559,684,1110,1134]" italics="true" pageId="13" pageNumber="14">pallidoides</emphasis>
</taxonomicName>
indicates that this new species closely resembles
<taxonomicName id="4C6A94D05801FFF7FB321F1E2088FB69" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidus">
<emphasis id="B91E33415801FFF7FB321F1E2088FB69" italics="true" pageId="13" pageNumber="14">Hypsibius pallidus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD5EF535801FFF7FF131FD42206FA9E" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
<emphasis id="B91E33415801FFF7FF131FD421E5FB4D" bold="true" box="[199,467,1180,1205]" pageId="13" pageNumber="14">Differential diagnosis.</emphasis>
<taxonomicName id="4C6A94D05801FFF7FE341FD522EFFB4D" box="[480,729,1181,1205]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FE341FD522EFFB4D" box="[480,729,1181,1205]" italics="true" pageId="13" pageNumber="14">Hypsibius pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5801FFF7FD321FD52373FB4C" box="[742,837,1181,1205]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B91E33415801FFF7FD321FD52308FB4D" bold="true" box="[742,830,1181,1205]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
was very similar to
<taxonomicName id="4C6A94D05801FFF7FB921FD624FEFB4D" box="[1094,1224,1181,1205]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidus">
<emphasis id="B91E33415801FFF7FB921FD624FEFB4D" box="[1094,1224,1181,1205]" italics="true" pageId="13" pageNumber="14">H. pallidus</emphasis>
</taxonomicName>
, which lacks any microplacoid or septulum, and had higher values of
<emphasis id="B91E33415801FFF7FD0B1F8922C3FB20" box="[735,757,1217,1240]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
index for the macroplacoids (
<tableCitation id="C6E8DAE85801FFF7FB971F8824ADFB20" box="[1091,1179,1216,1240]" captionStart="TABLE 4" captionStartId="14.[151,235,329,350]" captionTargetBox="[151,1436,409,1567]" captionTargetPageId="14" captionText="TABLE 4. Measurements of some structures of the holotype and two paratypes of Hypsibius pallidoides sp. nov., a specimen of H. pallidus and the holotype of H. septulatus." httpUri="http://table.plazi.org/id/DF15BFDB5802FFF4FF431A0122B6FE86" pageId="13" pageNumber="14" tableUuid="DF15BFDB5802FFF4FF431A0122B6FE86">Table 4</tableCitation>
;
<figureCitation id="1351F3D65801FFF7FB731F88255BFB20" box="[1191,1389,1216,1240]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Figs. 7AB and 7</figureCitation>
E). Five species of
<taxonomicName id="4C6A94D05801FFF7FE9D1FAB218FFB03" box="[329,441,1251,1275]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="genus">
<emphasis id="B91E33415801FFF7FE9D1FAB218FFB03" box="[329,441,1251,1275]" italics="true" pageId="13" pageNumber="14">Hypsibius</emphasis>
</taxonomicName>
were known to have smooth cuticle, two macroplacoids and septulum:
<taxonomicName id="4C6A94D05801FFF7FB331FAC2547FB03" box="[1255,1393,1251,1275]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="dujardini">
<emphasis id="B91E33415801FFF7FB331FAC2547FB03" box="[1255,1393,1251,1275]" italics="true" pageId="13" pageNumber="14">H. dujardini</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A94D05801FFF7FA551FAC212CFAE7" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="iskandarovi">
<emphasis id="B91E33415801FFF7FA551FAC212CFAE7" italics="true" pageId="13" pageNumber="14">H. iskandarovi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A94D05801FFF7FEFC1E40218AFAE7" box="[296,444,1287,1311]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="septulatus">
<emphasis id="B91E33415801FFF7FEFC1E40218AFAE7" box="[296,444,1287,1311]" italics="true" pageId="13" pageNumber="14">H. septulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A94D05801FFF7FE1D1E402437FAE7" authority="Miller, McInnes &amp; Bergstrom, 2005" authorityName="Miller, McInnes &amp; Bergstrom" authorityYear="2005" box="[457,1025,1287,1312]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="heardensis">
<emphasis id="B91E33415801FFF7FE1D1E402255FAE7" box="[457,611,1287,1311]" italics="true" pageId="13" pageNumber="14">H. heardensis</emphasis>
<bibRefCitation id="EFFB92A25801FFF7FDBF1E4F2437FAE7" author="Miller" box="[619,1025,1287,1312]" pageId="13" pageNumber="14" refString="Miller, W. R., McInnes, S. J. &amp; Bergstrom, D. M. (2005) Tardigrades of the Australian Antarctic: Hypsibius heardensis (Eutardigrada: Hypsibiidae: dujardini group) a new species from sub-Antarctic Heard Island. Zootaxa, 1022, 57 - 64." type="journal article" year="2005">Miller, McInnes &amp; Bergstrom, 2005</bibRefCitation>
</taxonomicName>
, and
<emphasis id="B91E33415801FFF7FBEF1E4025ADFAE7" box="[1083,1435,1287,1311]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C6A94D05801FFF7FBEF1E4024C5FAE7" box="[1083,1267,1287,1311]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="seychellensis">H. seychellensis</taxonomicName>
.
<taxonomicName id="4C6A94D05801FFF7FB291E4025ADFAE7" box="[1277,1435,1287,1311]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">H. pallidoides</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A22D8E3A5801FFF7FF431E6320C4FABA" box="[151,242,1323,1347]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B91E33415801FFF7FF431E6320DDFABB" bold="true" box="[151,235,1323,1347]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
differed from all in having external claws of
<taxonomicName id="4C6A94D05801FFF7FD2B1E6223E7FABA" box="[767,977,1322,1346]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidus">
<emphasis id="B91E33415801FFF7FD2B1E6223E7FABA" box="[767,977,1322,1346]" italics="true" pageId="13" pageNumber="14">Hypsibius pallidus</emphasis>
</taxonomicName>
-
<typeStatus id="54D151F15801FFF7FC0D1E63243DFABB" box="[985,1035,1323,1347]" pageId="13" pageNumber="14">type</typeStatus>
and in having smaller, dot-shaped, septulum. Other differences include:
</paragraph>
<paragraph id="8BD5EF535801FFF7FF111E392550F9EF" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
the lack of eye spots and presence of a pseudoseptulum in
<taxonomicName id="4C6A94D05801FFF7FC9B1E3A23C5FA71" box="[847,1011,1393,1417]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="iskandarovi">
<emphasis id="B91E33415801FFF7FC9B1E3A23C5FA71" box="[847,1011,1393,1417]" italics="true" pageId="13" pageNumber="14">H. iskandarovi</emphasis>
</taxonomicName>
(
<figureCitation id="1351F3D65801FFF7FBD61E3924BCFA71" box="[1026,1162,1393,1418]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Figs. 7 A, B</figureCitation>
and
<figureCitation id="1351F3D65801FFF7FB6B1E3924DAFA71" box="[1215,1260,1393,1417]" captionStart="FIGURE 8. A, B" captionStartId="15.[151,250,1134,1157]" captionTargetBox="[264,1308,189,1109]" captionTargetId="figure@15.[258,1317,183,1117]" captionTargetPageId="15" captionText="FIGURE 8. A, B, Hypsibius iskandarovi (paratype, slide No. 4334). A, Bucco-pharyngeal apparatus; septulum and pseudoseptulum are visible (arrow). B, Claws of the fourth pair of legs; the lunules (arrow) are visible. C, D, Hypsibius septulatus (holotype, slide No. 4780). C, Bucco-pharyngeal apparatus. D, Claws of the third pair of legs; the cuticular bar is visible (arrow). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279406/files/figure.png" pageId="13" pageNumber="14">8 A</figureCitation>
); the stylet supports inserted on the buccal tube in a more cephalic position (
<emphasis id="B91E33415801FFF7FC841EDE2350FA55" box="[848,870,1430,1453]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
=
<quantity id="4C9242B65801FFF7FC511EDC2425FA54" box="[901,1043,1428,1452]" metricMagnitude="0" metricUnit="m" metricValue="1.39319" metricValueMax="1.4097" metricValueMin="1.37668" pageId="13" pageNumber="14" unit="in" value="54.85" valueMax="55.5" valueMin="54.2">54.255.5 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FBCF1EDE248FFA55" box="[1051,1209,1429,1453]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FBCF1EDE2419FA55" box="[1051,1071,1430,1453]" italics="true" pageId="13" pageNumber="14">H</emphasis>
.
<emphasis id="B91E33415801FFF7FBE91EDD248FFA55" box="[1085,1209,1429,1453]" italics="true" pageId="13" pageNumber="14">pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5801FFF7FB161EDD252CFA54" box="[1218,1306,1429,1453]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B91E33415801FFF7FB161EDD2523FA55" bold="true" box="[1218,1301,1429,1453]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
, more than
<quantity id="4C9242B65801FFF7FF431EF020E7FA28" box="[151,209,1464,1488]" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" pageId="13" pageNumber="14" unit="in" value="58.0">58 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FF0D1EF12149FA28" box="[217,383,1464,1488]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="iskandarovi">
<emphasis id="B91E33415801FFF7FF0D1EF12149FA28" box="[217,383,1464,1488]" italics="true" pageId="13" pageNumber="14">H. iskandarovi</emphasis>
</taxonomicName>
); shorter claws (
<emphasis id="B91E33415801FFF7FDEB1EF12262FA28" box="[575,596,1465,1488]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
index relative to the posterior claws of the hind legs:
<quantity id="4C9242B65801FFF7FB6F1EF0257CFA28" box="[1211,1354,1464,1488]" metricMagnitude="0" metricUnit="m" metricValue="1.78689" metricValueMax="1.8313399999999997" metricValueMin="1.7424399999999998" pageId="13" pageNumber="14" unit="in" value="70.35" valueMax="72.1" valueMin="68.6">68.672.1 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FA851EF1217FFA0C" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FA851EF1217FFA0C" italics="true" pageId="13" pageNumber="14">Hypsibius pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5801FFF7FE841E94219EFA0B" box="[336,424,1500,1524]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B91E33415801FFF7FE841E942194FA0C" bold="true" box="[336,418,1500,1524]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<quantity id="4C9242B65801FFF7FE631E932275FA0B" box="[439,579,1499,1523]" metricMagnitude="0" metricUnit="m" metricValue="2.16789" metricValueMax="2.42062" metricValueMin="1.91516" pageId="13" pageNumber="14" unit="in" value="85.35" valueMax="95.3" valueMin="75.4">75.495.3 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FD9E1E9522D9FA0C" box="[586,751,1500,1524]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="iskandarovi">
<emphasis id="B91E33415801FFF7FD9E1E9522D9FA0C" box="[586,751,1500,1524]" italics="true" pageId="13" pageNumber="14">H. iskandarovi</emphasis>
</taxonomicName>
according to
<bibRefCitation id="EFFB92A25801FFF7FC5C1E94240CFA0B" author="Tumanov" box="[904,1082,1499,1523]" pageId="13" pageNumber="14" refString="Tumanov, D. V. (1997) Hypsibius iskandarovi sp. n., a new species of Tardigrada from fresh waters of North-West Russia (Tardigrada: Hypsibiidae). Zoosystematica Rossica, 5, 2, 219 - 220." type="journal article" year="1997">Tumanov, 1997</bibRefCitation>
); presence of lunules (
<figureCitation id="1351F3D65801FFF7FAE11E93209DF9EF" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Figs. 7C, D</figureCitation>
and
<figureCitation id="1351F3D65801FFF7FF351EB72135F9EF" box="[225,259,1535,1559]" captionStart="FIGURE 8. A, B" captionStartId="15.[151,250,1134,1157]" captionTargetBox="[264,1308,189,1109]" captionTargetId="figure@15.[258,1317,183,1117]" captionTargetPageId="15" captionText="FIGURE 8. A, B, Hypsibius iskandarovi (paratype, slide No. 4334). A, Bucco-pharyngeal apparatus; septulum and pseudoseptulum are visible (arrow). B, Claws of the fourth pair of legs; the lunules (arrow) are visible. C, D, Hypsibius septulatus (holotype, slide No. 4780). C, Bucco-pharyngeal apparatus. D, Claws of the third pair of legs; the cuticular bar is visible (arrow). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279406/files/figure.png" pageId="13" pageNumber="14">8B</figureCitation>
) and cuticular bar on the first three pairs of legs for
<taxonomicName id="4C6A94D05801FFF7FC981D4823C7F9EF" box="[844,1009,1535,1559]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="iskandarovi">
<emphasis id="B91E33415801FFF7FC981D4823C7F9EF" box="[844,1009,1535,1559]" italics="true" pageId="13" pageNumber="14">H. iskandarovi</emphasis>
</taxonomicName>
, all absent from the new species.
</paragraph>
<paragraph id="8BD5EF535801FFF7FF111D6A2430F95D" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
Cuticular undulations in
<taxonomicName id="4C6A94D05801FFF7FE0F1D6C2259F9C3" box="[475,623,1571,1595]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="septulatus">
<emphasis id="B91E33415801FFF7FE0F1D6C2259F9C3" box="[475,623,1571,1595]" italics="true" pageId="13" pageNumber="14">H. septulatus</emphasis>
</taxonomicName>
were lacking in
<taxonomicName id="4C6A94D05801FFF7FCFE1D6C23FFF9C3" box="[810,969,1571,1595]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FCFE1D6C2308F9C3" box="[810,830,1572,1595]" italics="true" pageId="13" pageNumber="14">H</emphasis>
.
<emphasis id="B91E33415801FFF7FC981D6B23FFF9C3" box="[844,969,1571,1595]" italics="true" pageId="13" pageNumber="14">pallidoides</emphasis>
</taxonomicName>
<emphasis id="B91E33415801FFF7FC041D6B2415F9C3" bold="true" box="[976,1059,1571,1595]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A22D8E3A5801FFF7FC041D6B2415F9C3" box="[976,1059,1571,1595]" pageId="13" pageNumber="14" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
, the buccal tube was slightly narrower and the stylet supports inserted on the buccal tube in a more cephalic position (
<emphasis id="B91E33415801FFF7FBB91D0F24B5F9A6" box="[1133,1155,1607,1630]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
=
<quantity id="4C9242B65801FFF7FB701D0E2505F9A6" box="[1188,1331,1606,1630]" metricMagnitude="0" metricUnit="m" metricValue="1.39319" metricValueMax="1.4097" metricValueMin="1.37668" pageId="13" pageNumber="14" unit="in" value="54.85" valueMax="55.5" valueMin="54.2">54.255.5 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FAEF1D0F20D6F979" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FAEF1D0F2579F9A6" box="[1339,1359,1607,1630]" italics="true" pageId="13" pageNumber="14">H</emphasis>
.
<emphasis id="B91E33415801FFF7FA8B1D0E20D6F979" italics="true" pageId="13" pageNumber="14">pallidoides</emphasis>
</taxonomicName>
<emphasis id="B91E33415801FFF7FF3C1D22210AF97A" bold="true" box="[232,316,1642,1666]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A22D8E3A5801FFF7FF3C1D22210AF97A" box="[232,316,1642,1666]" pageId="13" pageNumber="14" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
,
<quantity id="4C9242B65801FFF7FE9F1D2121ACF979" box="[331,410,1641,1665]" metricMagnitude="0" metricUnit="m" metricValue="1.6383" pageId="13" pageNumber="14" unit="in" value="64.5">64.5 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FE761D222201F979" box="[418,567,1641,1665]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="septulatus">
<emphasis id="B91E33415801FFF7FE761D222201F979" box="[418,567,1641,1665]" italics="true" pageId="13" pageNumber="14">H. septulatus</emphasis>
</taxonomicName>
) (
<tableCitation id="C6E8DAE85801FFF7FD841D21229EF979" box="[592,680,1641,1665]" captionStart="TABLE 4" captionStartId="14.[151,235,329,350]" captionTargetBox="[151,1436,409,1567]" captionTargetPageId="14" captionText="TABLE 4. Measurements of some structures of the holotype and two paratypes of Hypsibius pallidoides sp. nov., a specimen of H. pallidus and the holotype of H. septulatus." httpUri="http://table.plazi.org/id/DF15BFDB5802FFF4FF431A0122B6FE86" pageId="13" pageNumber="14" tableUuid="DF15BFDB5802FFF4FF431A0122B6FE86">Table 4</tableCitation>
;
<figureCitation id="1351F3D65801FFF7FD611D21230FF979" box="[693,825,1641,1666]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Figs. 7A, B</figureCitation>
and
<figureCitation id="1351F3D65801FFF7FCA51D212349F979" box="[881,895,1641,1665]" captionStart="FIGURE 8. A, B" captionStartId="15.[151,250,1134,1157]" captionTargetBox="[264,1308,189,1109]" captionTargetId="figure@15.[258,1317,183,1117]" captionTargetPageId="15" captionText="FIGURE 8. A, B, Hypsibius iskandarovi (paratype, slide No. 4334). A, Bucco-pharyngeal apparatus; septulum and pseudoseptulum are visible (arrow). B, Claws of the fourth pair of legs; the lunules (arrow) are visible. C, D, Hypsibius septulatus (holotype, slide No. 4780). C, Bucco-pharyngeal apparatus. D, Claws of the third pair of legs; the cuticular bar is visible (arrow). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279406/files/figure.png" pageId="13" pageNumber="14">8</figureCitation>
C); septulum much smaller, claws were slightly shorter (
<tableCitation id="C6E8DAE85801FFF7FF271DC5217FF95D" box="[243,329,1677,1701]" captionStart="TABLE 4" captionStartId="14.[151,235,329,350]" captionTargetBox="[151,1436,409,1567]" captionTargetPageId="14" captionText="TABLE 4. Measurements of some structures of the holotype and two paratypes of Hypsibius pallidoides sp. nov., a specimen of H. pallidus and the holotype of H. septulatus." httpUri="http://table.plazi.org/id/DF15BFDB5802FFF4FF431A0122B6FE86" pageId="13" pageNumber="14" tableUuid="DF15BFDB5802FFF4FF431A0122B6FE86">Table 4</tableCitation>
) and absence of cuticular bar on the legs (
<figureCitation id="1351F3D65801FFF7FCF51DC523AAF95D" box="[801,924,1677,1701]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Figs 7C, D</figureCitation>
and
<figureCitation id="1351F3D65801FFF7FC071DC523D7F95D" box="[979,993,1677,1701]" captionStart="FIGURE 8. A, B" captionStartId="15.[151,250,1134,1157]" captionTargetBox="[264,1308,189,1109]" captionTargetId="figure@15.[258,1317,183,1117]" captionTargetPageId="15" captionText="FIGURE 8. A, B, Hypsibius iskandarovi (paratype, slide No. 4334). A, Bucco-pharyngeal apparatus; septulum and pseudoseptulum are visible (arrow). B, Claws of the fourth pair of legs; the lunules (arrow) are visible. C, D, Hypsibius septulatus (holotype, slide No. 4780). C, Bucco-pharyngeal apparatus. D, Claws of the third pair of legs; the cuticular bar is visible (arrow). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279406/files/figure.png" pageId="13" pageNumber="14">8</figureCitation>
D).
</paragraph>
<paragraph id="8BD5EF535801FFF7FF111DF92214F914" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
<taxonomicName id="4C6A94D05801FFF7FF111DF92169F930" box="[197,351,1712,1736]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="heardensis">
<emphasis id="B91E33415801FFF7FF111DF92169F930" box="[197,351,1712,1736]" italics="true" pageId="13" pageNumber="14">H. heardensis</emphasis>
</taxonomicName>
lacked eye spots, present in the new species, and possessed cuticular bar on the legs, which were absent from
<taxonomicName id="4C6A94D05801FFF7FEF71D9D21F7F914" box="[291,449,1748,1772]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FEF71D9D21F7F914" box="[291,449,1748,1772]" italics="true" pageId="13" pageNumber="14">H. pallidoides</emphasis>
</taxonomicName>
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<emphasis id="B91E33415801FFF7FE1C1D9C222DF914" bold="true" box="[456,539,1748,1772]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BD5EF535801FFF7FF111DBF2170F8AE" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
The stylet supports of
<taxonomicName id="4C6A94D05801FFF7FE6B1DB0226BF8F7" box="[447,605,1783,1807]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FE6B1DB0226BF8F7" box="[447,605,1783,1807]" italics="true" pageId="13" pageNumber="14">H. pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5801FFF7FDB11DB02288F8F7" box="[613,702,1784,1808]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B91E33415801FFF7FDB11DB02281F8E8" bold="true" box="[613,695,1784,1808]" pageId="13" pageNumber="14">sp. nov</emphasis>
.
</taxonomicNameLabel>
were inserted on the buccal tube in a more cephalic position than for
<taxonomicName id="4C6A94D05801FFF7FF6A1C542144F8CB" box="[190,370,1819,1843]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="seychellensis">
<emphasis id="B91E33415801FFF7FF6A1C542144F8CB" box="[190,370,1819,1843]" italics="true" pageId="13" pageNumber="14">H. seychellensis</emphasis>
</taxonomicName>
(
<emphasis id="B91E33415801FFF7FE561C5421AEF8CB" box="[386,408,1820,1843]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
=
<quantity id="4C9242B65801FFF7FE601C532277F8CB" box="[436,577,1819,1843]" metricMagnitude="0" metricUnit="m" metricValue="1.39319" metricValueMax="1.4097" metricValueMin="1.37668" pageId="13" pageNumber="14" unit="in" value="54.85" valueMax="55.5" valueMin="54.2">54.255.5 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FD931C5422D3F8CB" box="[583,741,1819,1843]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FD931C54226DF8CB" box="[583,603,1820,1843]" italics="true" pageId="13" pageNumber="14">H</emphasis>
.
<emphasis id="B91E33415801FFF7FDBD1C5322D3F8CB" box="[617,741,1819,1843]" italics="true" pageId="13" pageNumber="14">pallidoides</emphasis>
</taxonomicName>
<emphasis id="B91E33415801FFF7FD381C532373F8CB" bold="true" box="[748,837,1819,1843]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A22D8E3A5801FFF7FD381C532373F8CB" box="[748,837,1819,1843]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<quantity id="4C9242B65801FFF7FC871C5323D6F8CB" box="[851,992,1819,1843]" metricMagnitude="0" metricUnit="m" metricValue="1.6002" metricValueMax="1.61798" metricValueMin="1.58242" pageId="13" pageNumber="14" unit="in" value="63.0" valueMax="63.7" valueMin="62.3">62.363.7 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FC321C5424AFF8CB" box="[998,1177,1819,1843]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="seychellensis">
<emphasis id="B91E33415801FFF7FC321C5424AFF8CB" box="[998,1177,1819,1843]" italics="true" pageId="13" pageNumber="14">H. seychellensis</emphasis>
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) and the placoids were slightly shorter.
</paragraph>
<paragraph id="8BD5EF535801FFF7FF111C2925BDF81C" blockId="13.[151,1436,151,2020]" pageId="13" pageNumber="14">
The buccal tube of
<taxonomicName id="4C6A94D05801FFF7FE751C2B221DF882" box="[417,555,1890,1914]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="dujardini">
<emphasis id="B91E33415801FFF7FE751C2B221DF882" box="[417,555,1890,1914]" italics="true" pageId="13" pageNumber="14">H. dujardini</emphasis>
</taxonomicName>
gradually increased in width towards the posterior end, while the buccal tube of
<taxonomicName id="4C6A94D05801FFF7FF621CCE2160F865" box="[182,342,1925,1949]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FF621CCE20FCF865" box="[182,202,1926,1949]" italics="true" pageId="13" pageNumber="14">H</emphasis>
.
<emphasis id="B91E33415801FFF7FF0E1CCD2160F865" box="[218,342,1925,1949]" italics="true" pageId="13" pageNumber="14">pallidoides</emphasis>
</taxonomicName>
<emphasis id="B91E33415801FFF7FE8B1CCD2182F865" bold="true" box="[351,436,1925,1949]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A22D8E3A5801FFF7FE8B1CCD2182F865" box="[351,436,1925,1949]" pageId="13" pageNumber="14" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
had a uniform width (
<figureCitation id="1351F3D65801FFF7FD611CCD232EF866" box="[693,792,1925,1950]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="13" pageNumber="14">Figs. 7A</figureCitation>
and
<figureCitation id="1351F3D65801FFF7FC851CCD234EF865" box="[849,888,1925,1949]" captionStart="FIGURE 9. A, B" captionStartId="16.[151,250,1102,1125]" captionTargetBox="[258,1316,195,1055]" captionTargetId="figure@16.[258,1317,195,1055]" captionTargetPageId="16" captionText="FIGURE 9. A, B, Hypsibius dujardini (slide No. 2728). A, Bucco-pharyngeal apparatus. B, Claws of the second pair of legs. C, D, Hypibius allisoni (slide No. 4338). C, Bucco-pharyngeal apparatus (arrow indicates the small microplacoid). D, Claws of the third pair of legs. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279407/files/figure.png" pageId="13" pageNumber="14">9A</figureCitation>
); the stylet supports were inserted on the buccal tube in a more cephalic position (
<emphasis id="B91E33415801FFF7FDDA1CE22212F839" box="[526,548,1962,1985]" italics="true" pageId="13" pageNumber="14">pt</emphasis>
=
<quantity id="4C9242B65801FFF7FD961CE022F9F838" box="[578,719,1960,1984]" metricMagnitude="0" metricUnit="m" metricValue="1.39319" metricValueMax="1.4097" metricValueMin="1.37668" pageId="13" pageNumber="14" unit="in" value="54.85" valueMax="55.5" valueMin="54.2">54.255.5 in</quantity>
<taxonomicName id="4C6A94D05801FFF7FD021CE22343F839" box="[726,885,1961,1985]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415801FFF7FD021CE222DCF839" box="[726,746,1962,1985]" italics="true" pageId="13" pageNumber="14">H</emphasis>
.
<emphasis id="B91E33415801FFF7FD2C1CE12343F839" box="[760,885,1961,1985]" italics="true" pageId="13" pageNumber="14">pallidoides</emphasis>
</taxonomicName>
<emphasis id="B91E33415801FFF7FCA81CE123F8F839" bold="true" box="[892,974,1961,1985]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A22D8E3A5801FFF7FCA81CE123F8F839" box="[892,974,1961,1985]" pageId="13" pageNumber="14" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
, about
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<taxonomicName id="4C6A94D05801FFF7FBB61CE224DDF839" box="[1122,1259,1961,1985]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="dujardini">
<emphasis id="B91E33415801FFF7FBB61CE224DDF839" box="[1122,1259,1961,1985]" italics="true" pageId="13" pageNumber="14">H. dujardini</emphasis>
</taxonomicName>
); and the cuticular bar between the claws on the hind legs was present in
<taxonomicName id="4C6A94D05801FFF7FCFD1C852384F81C" box="[809,946,1996,2020]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="14" phylum="Tardigrada" rank="species" species="dujardini">
<emphasis id="B91E33415801FFF7FCFD1C852384F81C" box="[809,946,1996,2020]" italics="true" pageId="13" pageNumber="14">H. dujardini</emphasis>
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, but apparently absent in the new species.
</paragraph>
<paragraph id="8BD5EF535802FFF4FF111BD02421FEE4" blockId="14.[151,1436,151,284]" pageId="14" pageNumber="15">
<taxonomicName id="4C6A94D05802FFF4FF111BD021B9FF48" box="[197,399,152,176]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="15" phylum="Tardigrada" rank="species" species="allisoni">
<emphasis id="B91E33415802FFF4FF111BD021B9FF48" box="[197,399,152,176]" italics="true" pageId="14" pageNumber="15">Hypsibius allisoni</emphasis>
</taxonomicName>
had a very small microplacoid (
<figureCitation id="1351F3D65802FFF4FD231BDF230CFF57" box="[759,826,151,175]" captionStart="FIGURE 9. A, B" captionStartId="16.[151,250,1102,1125]" captionTargetBox="[258,1316,195,1055]" captionTargetId="figure@16.[258,1317,195,1055]" captionTargetPageId="16" captionText="FIGURE 9. A, B, Hypsibius dujardini (slide No. 2728). A, Bucco-pharyngeal apparatus. B, Claws of the second pair of legs. C, D, Hypibius allisoni (slide No. 4338). C, Bucco-pharyngeal apparatus (arrow indicates the small microplacoid). D, Claws of the third pair of legs. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279407/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
C), not the septulum present in
<taxonomicName id="4C6A94D05802FFF4FB431BD12503FF48" box="[1175,1333,152,176]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="15" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415802FFF4FB431BD12503FF48" box="[1175,1333,152,176]" italics="true" pageId="14" pageNumber="15">H. pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5802FFF4FAE81BD025A2FF57" box="[1340,1428,152,176]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B91E33415802FFF4FAE81BD025B8FF48" bold="true" box="[1340,1422,152,176]" pageId="14" pageNumber="15">sp. nov</emphasis>
.
</taxonomicNameLabel>
, though it can be difficult to distinguish the difference between a microplacoid and a septulum. We therefore stress that the claw shape of
<taxonomicName id="4C6A94D05802FFF4FE4C1BA822BBFF00" box="[408,653,224,248]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="15" phylum="Tardigrada" rank="species" species="pallidoides" status="sp. nov.">
<emphasis id="B91E33415802FFF4FE4C1BA822BBFF00" box="[408,653,224,248]" italics="true" pageId="14" pageNumber="15">Hypsibius pallidoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D8E3A5802FFF4FD401BA822D9FF0F" box="[660,751,224,248]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B91E33415802FFF4FD401BA822DEFF00" bold="true" box="[660,744,224,248]" pageId="14" pageNumber="15">sp. nov</emphasis>
.
</taxonomicNameLabel>
was very different from
<taxonomicName id="4C6A94D05802FFF4FBDF1BA924B4FF00" box="[1035,1154,224,248]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="15" phylum="Tardigrada" rank="species" species="allisoni">
<emphasis id="B91E33415802FFF4FBDF1BA924B4FF00" box="[1035,1154,224,248]" italics="true" pageId="14" pageNumber="15">H. allisoni</emphasis>
</taxonomicName>
, as the main branch was inserted on the secondary branch in a more distal position (
<figureCitation id="1351F3D65802FFF4FCF81A4C2398FEE4" box="[812,942,260,284]" captionStart="FIGURE 7. A D" captionStartId="12.[151,250,1866,1889]" captionTargetBox="[164,1420,603,1851]" captionTargetId="figure@12.[164,1424,603,1851]" captionTargetPageId="12" captionText="FIGURE 7. A D, Hypsibius pallidoides sp. nov.: A, Bucco-pharyngeal apparatus of the holotype (slide No. 5430) (the arrow indicates the very small septulum). B, Bucco-pharyngeal apparatus of a paratype (slide No. 5430) (the arrow indicates the very small septulum). C, D, Claws of the second pair of legs of the holotype. E, Hypsibius pallidus: Bucco-pharyngal apparatus (slide No. 1251). (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279405/files/figure.png" pageId="14" pageNumber="15">Figs. 7C, D</figureCitation>
and
<figureCitation id="1351F3D65802FFF4FC311A4C23C2FEE4" box="[997,1012,260,284]" captionStart="FIGURE 9. A, B" captionStartId="16.[151,250,1102,1125]" captionTargetBox="[258,1316,195,1055]" captionTargetId="figure@16.[258,1317,195,1055]" captionTargetPageId="16" captionText="FIGURE 9. A, B, Hypsibius dujardini (slide No. 2728). A, Bucco-pharyngeal apparatus. B, Claws of the second pair of legs. C, D, Hypibius allisoni (slide No. 4338). C, Bucco-pharyngeal apparatus (arrow indicates the small microplacoid). D, Claws of the third pair of legs. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279407/files/figure.png" pageId="14" pageNumber="15">9</figureCitation>
D).
</paragraph>
</subSubSection>
</treatment>
</document>