489 lines
79 KiB
XML
489 lines
79 KiB
XML
<document id="44A8C963CEAF1309A939F3D0332A7424" ID-CLB-Dataset="46809" ID-DOI="10.11646/zootaxa.4078.1.16" ID-GBIF-Dataset="9382c912-1e52-41c8-8499-1c515191c8ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="261493" ID-ZooBank="0102DC85-29D8-4B7A-BD24-984C115189AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1460361037286" checkinUser="plazi" docAuthor="Barão, Kim Ribeiro, Garbelotto, Thereza De Almeida, Campos, Luiz Alexandre & Grazia, Jocelia" docDate="2016" docId="03BF87AB1B2AFFCA9AC499CB33F3FE5D" docLanguage="en" docName="zootaxa.4078.1.16.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae)" docType="treatment" docVersion="6" lastPageNumber="171" masterDocId="FF86FFD31B2BFFC99A539C4A366FFFE9" masterDocTitle="Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae)" masterLastPageNumber="186" masterPageNumber="168" pageNumber="169" updateTime="1698294510363" updateUser="ExternalLinkService">
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<mods:title id="0FDA6E726C9AC5AAFB1B1F44F425E40C">Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae)</mods:title>
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<mods:namePart id="3983CA72180A77F4857279FFA43AE46A">Barão, Kim Ribeiro</mods:namePart>
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<mods:affiliation id="32B55CEEFB86823B6F80FC7DBE5DE2AA">Programa de Pós-Graduação em Biologia Animal & Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
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<mods:nameIdentifier id="9EC26A29E608972C94E3093E1A80A44D" type="email">kbarao@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="FBF6202DD95AF90CCD0525F53A2556D7">Garbelotto, Thereza De Almeida</mods:namePart>
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<mods:affiliation id="4B59E3485074839A263A3583D2D85266">Programa de Pós-Graduação em Biologia Animal & Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
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<mods:nameIdentifier id="2B66B83F95C7AED3682B8DCB8849018E" type="email">therezagarbelotto@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="96FAF519A83EA59BC6A5EB1E045D0C9F">Author</mods:roleTerm>
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<mods:namePart id="959240402F948DEF31A6FC0BFDEAF45C">Campos, Luiz Alexandre</mods:namePart>
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<mods:affiliation id="BC027EF45A76953510BA89E629927EDE">Programa de Pós-Graduação em Biologia Animal & Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
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<mods:nameIdentifier id="B9F44A9A3215D5CAC5703A1D2C6FAAF3" type="email">luiz.campos@ufrgs.br</mods:nameIdentifier>
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<mods:namePart id="D9D5E1FA479A20730792219C0015811C">Grazia, Jocelia</mods:namePart>
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<mods:affiliation id="20367697C9D45F8AAA0714661D759234">Programa de Pós-Graduação em Biologia Animal & Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
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<mods:nameIdentifier id="80F99A7FC56C9F9AC4B8BD2A78BCDF99" type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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<mods:title id="19048BE6334F96904FC8F0786C37E716">Zootaxa</mods:title>
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<mods:date id="3318E72329F7E40C3B6E7CDCFF447C02">2016</mods:date>
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<mods:number id="05324C52AE5D1DE55B33F7F55646F12F">2016-02-09</mods:number>
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<mods:number id="7359AA7BF73FDE18D6E9CDACA40C1AEF">4078</mods:number>
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<mods:identifier id="95F9243C1E26CA27F9B872E621DAA161" type="DOI">10.11646/zootaxa.4078.1.16</mods:identifier>
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<treatment id="03BF87AB1B2AFFCA9AC499CB33F3FE5D" ID-DOI="http://doi.org/10.5281/zenodo.6081299" ID-Zenodo-Dep="6081299" LSID="urn:lsid:plazi:treatment:03BF87AB1B2AFFCA9AC499CB33F3FE5D" httpUri="http://treatment.plazi.org/id/03BF87AB1B2AFFCA9AC499CB33F3FE5D" lastPageId="3" lastPageNumber="171" pageId="1" pageNumber="169">
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<heading id="D0E181D11B2AFFC89AC499CB37C3FA72" bold="true" box="[151,428,1409,1435]" fontSize="11" level="1" pageId="1" pageNumber="169" reason="1">
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<emphasis id="B962EAAF1B2AFFC89AC499CB37C3FA72" bold="true" box="[151,428,1409,1435]" pageId="1" pageNumber="169">
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<emphasis id="B962EAAF1B2AFFC89AC499CB376EFA72" bold="true" box="[151,257,1409,1435]" italics="true" pageId="1" pageNumber="169">Braunus</emphasis>
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<bibRefCitation id="EF874B4C1B2AFFC89B5B99CB37C3FA72" author="Distant" box="[264,428,1409,1435]" pageId="1" pageNumber="169" refString="Distant, W. L. (1899) Rhyncotal notes - III. Heteroptera: Discocephalinae and Pentatominae (part.). The Annals and Magazine of Natural History, including Zoology, Botany and Geology (seventh series), 4 (14), 421 - 445." type="journal article" year="1899">Distant, 1899</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30C65361B2AFFC89AC499833471FA15" pageId="1" pageNumber="169" type="reference_group">
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<paragraph id="8BA936BD1B2AFFC89AC499833471FA15" blockId="1.[151,705,1480,1533]" pageId="1" pageNumber="169">
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<treatmentCitationGroup id="AB0611931B2AFFC89AC499833471FA15" pageId="1" pageNumber="169">
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<treatmentCitation id="0AB710AC1B2AFFC89AC49983378AFA37" box="[151,485,1480,1502]" pageId="1" pageNumber="169">
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<emphasis id="B962EAAF1B2AFFC89AC499833682FA37" box="[151,237,1481,1502]" italics="true" pageId="1" pageNumber="169">Braunus</emphasis>
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<bibRefCitation id="EF874B4C1B2AFFC89AA099823710FA37" author="Distant" box="[243,383,1480,1502]" pageId="1" pageNumber="169" refString="Distant, W. L. (1899) Rhyncotal notes - III. Heteroptera: Discocephalinae and Pentatominae (part.). The Annals and Magazine of Natural History, including Zoology, Botany and Geology (seventh series), 4 (14), 421 - 445." type="journal article" year="1899">Distant, 1899</bibRefCitation>
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: 422–423
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</treatmentCitation>
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;
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<treatmentCitation id="0AB710AC1B2AFFC89BBC998234D3FA37" box="[495,700,1480,1502]" pageId="1" pageNumber="169">
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<bibRefCitation id="EF874B4C1B2AFFC89BBC998234E6FA37" author="Kirkaldy" box="[495,649,1480,1502]" pageId="1" pageNumber="169" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I. Cimicidae. Felix L. Dames, Berlin, xl + 392 pp." type="book" year="1909">Kirkaldy, 1909</bibRefCitation>
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: 220
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</treatmentCitation>
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.
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<treatmentCitation id="0AB710AC1B2AFFC89AC499A237D8FA15" box="[151,439,1510,1533]" pageId="1" pageNumber="169">
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<taxonomicName id="4C164D3E1B2AFFC89AC499A237D8FA15" authority="McDonald, 1982: 327" authorityName="McDonald" authorityPageNumber="327" authorityYear="1982" box="[151,439,1510,1533]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="169" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EAAF1B2AFFC89AC499A236A0FA14" box="[151,207,1512,1533]" italics="true" pageId="1" pageNumber="169">Lojus</emphasis>
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||
<bibRefCitation id="EF874B4C1B2AFFC89A8699AC37EBFA15" author="McDonald" box="[213,388,1510,1532]" pageId="1" pageNumber="169" refString="McDonald, F. J. D. (1982) Lojus asperus, a new genus and species (Hemiptera: Pentatomidae). Annals of the Entomological Society of America, 75, 327 - 329. http: // dx. doi. org / 10.1093 / aesa / 75.3.327" type="journal article" year="1982">McDonald, 1982</bibRefCitation>
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: 327
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</taxonomicName>
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.
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||
<emphasis id="B962EAAF1B2AFFC89B9299AD3471FA15" bold="true" box="[449,542,1511,1532]" pageId="1" pageNumber="169">syn. nov.</emphasis>
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</paragraph>
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<subSubSection id="C30C65361B2AFFC89AC49A633788F98F" pageId="1" pageNumber="169" type="type_taxon">
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<paragraph id="8BA936BD1B2AFFC89AC49A633788F98F" blockId="1.[151,1437,1577,2034]" pageId="1" pageNumber="169">
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<emphasis id="B962EAAF1B2AFFC89AC49A633758F9AB" bold="true" box="[151,311,1577,1602]" pageId="1" pageNumber="169">
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<typeStatus id="54AD881F1B2AFFC89AC49A6336BDF9AB" box="[151,210,1577,1602]" pageId="1" pageNumber="169">Type</typeStatus>
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species.
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</emphasis>
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||
<emphasis id="B962EAAF1B2AFFC89B109A6037CDF9A8" box="[323,418,1578,1601]" italics="true" pageId="1" pageNumber="169">Braunus</emphasis>
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||
:
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<emphasis id="B962EAAF1B2AFFC89BE69A633442F9AB" box="[437,557,1577,1602]" italics="true" pageId="1" pageNumber="169">Coriplatus</emphasis>
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<taxonomicName id="4C164D3E1B2AFFC8986A9A60350CF9AB" authority="Walker, 1867" authorityName="Walker" authorityYear="1867" box="[569,867,1577,1602]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="169" phylum="Arthropoda" rank="species" species="sciocorinus">
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<emphasis id="B962EAAF1B2AFFC8986A9A6034D5F9A8" box="[569,698,1578,1601]" italics="true" pageId="1" pageNumber="169">sciocorinus</emphasis>
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<bibRefCitation id="EF874B4C1B2AFFC898959A63350CF9AB" author="Walker" box="[710,867,1577,1602]" pageId="1" pageNumber="169" refString="Walker, F. L. S. (1867) Catalogue of the specimens of Heteropterous-Hemiptera in the collection of the British Museum - Part I: Scutata. E. Newman, London, 240 pp." type="book" year="1867">Walker, 1867</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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<taxonomicName id="4C164D3E1B2AFFC89E4D9A603233F9A8" box="[1054,1116,1578,1601]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="169" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EAAF1B2AFFC89E4D9A603233F9A8" box="[1054,1116,1578,1601]" italics="true" pageId="1" pageNumber="169">Lojus</emphasis>
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</taxonomicName>
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:
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<taxonomicName id="4C164D3E1B2AFFC89E239A6036BDF98F" authority="McDonald, 1982" authorityName="McDonald" authorityYear="1982" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="169" phylum="Arthropoda" rank="species" species="asperus">
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<emphasis id="B962EAAF1B2AFFC89E239A60337EF9A8" box="[1136,1297,1578,1601]" italics="true" pageId="1" pageNumber="169">Lojus asperus</emphasis>
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<bibRefCitation id="EF874B4C1B2AFFC89F4E9A6336BDF98F" author="McDonald" pageId="1" pageNumber="169" refString="McDonald, F. J. D. (1982) Lojus asperus, a new genus and species (Hemiptera: Pentatomidae). Annals of the Entomological Society of America, 75, 327 - 329. http: // dx. doi. org / 10.1093 / aesa / 75.3.327" type="journal article" year="1982">McDonald, 1982</bibRefCitation>
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||
</taxonomicName>
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||
, by original designation.
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||
</paragraph>
|
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</subSubSection>
|
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<subSubSection id="C30C65361B2AFFC89A949A3B3709F8D7" pageId="1" pageNumber="169" type="diagnosis">
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<paragraph id="8BA936BD1B2AFFC89A949A3B3709F8D7" blockId="1.[151,1437,1577,2034]" pageId="1" pageNumber="169">
|
||
<emphasis id="B962EAAF1B2AFFC89A949A3B372EF963" bold="true" box="[199,321,1649,1674]" pageId="1" pageNumber="169">Diagnosis.</emphasis>
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Mandibular plates widely triangular apically, at least as long as clypeus. Costal margin of each forewing with a basal process. Peritreme of the metathoracic gland obsolete; mesial margin of each ostiole bordered by a globoid tubercle. Abdominal sternite III projected over abdominal sternite IV ventro-medially. Male genitalia: dorsal rim of pygophore concave with 1+1 trapezoidal expansions laterad to segment X; segment X rectangular. Female genitalia: sutural margin of each gonocoxite 8 parallel basally, usually juxtaposed, and divergent apically.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30C65361B2AFFCB9A949B033214FCB4" lastPageId="2" lastPageNumber="170" pageId="1" pageNumber="169" type="description">
|
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<paragraph id="8BA936BD1B2AFFCB9A949B033789FDFE" blockId="1.[151,1437,1577,2034]" lastBlockId="2.[151,1437,151,1184]" lastPageId="2" lastPageNumber="170" pageId="1" pageNumber="169">
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||
<emphasis id="B962EAAF1B2AFFC89A949B033736F88B" bold="true" box="[199,345,1865,1890]" pageId="1" pageNumber="169">Description.</emphasis>
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||
Brown to dark-brown or mottled brown and ivory; legs dark-brown or yellowish with brown, circular spots; dorsal and ventral surfaces of connexivum light brown medially. Integument strongly corrugated dorsally. Head longer than wide; mandibular plates equal to or longer than clypeus, but usually not meeting anterior to clypeus, widely triangular apically; anterior margin of each bucculae triangular, each posterior margin tapering towards base of head; ocelli from rudimentary to well-developed; antennae five-segmented; labium arising before anterior margin of eyes, attaining from hind coxae to abdominal sternite III. Anterior angles of pronotum obtuse or acutely projected; anterolateral margins smooth or with spine-like tubercles; humeral angles strongly projected laterad or anteriad, each with a single process or quadrate with two or more processes marginally; posterolateral margin each with or without a median tubercle and indentation; posterior margin straight to strongly concave; cicatrices swollen (
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<figureCitation id="132D2A381B29FFCB9B239D6D378BFEA9" box="[368,484,295,320]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 1–11</figureCitation>
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). Scutellum foveate in basal angles. Costal margin of each forewing with an obtuse process basally; radial vein calloused or not apically; membranal veins coarse and reticulated; length of corium and membrane variable (
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<figureCitation id="132D2A381B29FFCB9BD89D25347CFE61" box="[395,531,367,392]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,627,650]" captionTargetBox="[151,1436,195,603]" captionTargetId="figure@4.[151,1436,195,603]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Braunus sciocorinus (Walker, 1867), general morphology. 12 – 13. habitus; 12. dorsal view; 13. ventral view; 14. male forewing; 15. female forewing; 16. male hindwing; 17. external scent efferent system, inset showing ostiolar area. Abreviattions: ev = evaporatorium; ms = mesopleura; mt = metapleura; o = ostiole. Scale bars: 11 – 12 = 2 mm; 13 – 16 = 0.5 mm." httpUri="https://zenodo.org/record/261496/files/figure.png" pageId="2" pageNumber="170">Figs 14–15</figureCitation>
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,
|
||
<figureCitation id="132D2A381B29FFCB98759D25341FFE61" box="[550,624,367,392]" captionStart="FIGURES 54 – 59" captionStartId="10.[151,264,748,771]" captionTargetBox="[151,1435,267,725]" captionTargetId="figure@10.[151,1436,267,725]" captionTargetPageId="10" captionText="FIGURES 54 – 59. Braunus ocellatus (Thomas, 2001) comb. nov.. 54 – 55. habitus; 54. dorsal view; 55. ventral view; 56. male forewing; 57. female forewing; 58. male hindwing; 59. external scent efferent system. Abreviattions: ev = evaporatorium; ms = mesopleura; mt = metapleura; o = ostiole. Scale bars: 47 – 48 = 2 mm; 49 – 52 = 0.5 mm." httpUri="https://zenodo.org/record/261502/files/figure.png" pageId="2" pageNumber="170">56–57</figureCitation>
|
||
). Peritreme of metasternal gland obsolete; mesial margin of ostiole bordered by a rounded process; evaporatorium concolorous with pleural color pattern, reaching halfway to lateral metapleural margin and to the posterior margin of mesopleura (
|
||
<figureCitation id="132D2A381B29FFCB99099DFD35DFFE39" box="[858,944,439,464]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,627,650]" captionTargetBox="[151,1436,195,603]" captionTargetId="figure@4.[151,1436,195,603]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Braunus sciocorinus (Walker, 1867), general morphology. 12 – 13. habitus; 12. dorsal view; 13. ventral view; 14. male forewing; 15. female forewing; 16. male hindwing; 17. external scent efferent system, inset showing ostiolar area. Abreviattions: ev = evaporatorium; ms = mesopleura; mt = metapleura; o = ostiole. Scale bars: 11 – 12 = 2 mm; 13 – 16 = 0.5 mm." httpUri="https://zenodo.org/record/261496/files/figure.png" pageId="2" pageNumber="170">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB99EF9DFD35B5FE39" box="[956,986,439,464]" captionStart="FIGURES 48 – 53" captionStartId="9.[151,264,731,754]" captionTargetBox="[151,1436,195,707]" captionTargetId="figure@9.[151,1436,195,707]" captionTargetPageId="9" captionText="FIGURES 48 – 53. Braunus prionotus Barão & Garbelotto sp. nov. 48 – 49. habitus; 48. dorsal view; 49. ventral view; 50. head, dorsal view; 51. external scent efferent system; 52 – 53. genital plates. Abbreviations: ev = evaporatorium; gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; la 8 = laterotergites 8; la 9 = laterotergites 9; ms = mesopleura; mt = metapleura; o = ostiole; X = abdominal segment X. Scale bars: 48 – 49 = 2 mm; 50 – 53 = 1 mm." httpUri="https://zenodo.org/record/261501/files/figure.png" pageId="2" pageNumber="170">51</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB99B59DFD3269FE39" box="[998,1030,439,464]" captionStart="FIGURES 54 – 59" captionStartId="10.[151,264,748,771]" captionTargetBox="[151,1435,267,725]" captionTargetId="figure@10.[151,1436,267,725]" captionTargetPageId="10" captionText="FIGURES 54 – 59. Braunus ocellatus (Thomas, 2001) comb. nov.. 54 – 55. habitus; 54. dorsal view; 55. ventral view; 56. male forewing; 57. female forewing; 58. male hindwing; 59. external scent efferent system. Abreviattions: ev = evaporatorium; ms = mesopleura; mt = metapleura; o = ostiole. Scale bars: 47 – 48 = 2 mm; 49 – 52 = 0.5 mm." httpUri="https://zenodo.org/record/261502/files/figure.png" pageId="2" pageNumber="170">59</figureCitation>
|
||
). Posterolateral angles of abdominal sternites each projected posterolaterally, obtuse apically. Median portion of abdominal sternite III projected onto base of abdominal sternite IV.
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B29FFCB9A949E6E3203FD24" blockId="2.[151,1437,151,1184]" pageId="2" pageNumber="170">
|
||
<emphasis id="B962EAAF1B29FFCB9A949E6E37EDFDD4" bold="true" box="[199,386,548,573]" pageId="2" pageNumber="170">Male genitalia.</emphasis>
|
||
Pygophore sub-quadrate (
|
||
<figureCitation id="132D2A381B29FFCB98949E6E353DFDD5" box="[711,850,548,573]" captionStart="FIGURES 18 – 31" captionStartId="5.[151,264,1815,1838]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@5.[151,1436,195,1791]" captionTargetPageId="5" captionText="FIGURES 18 – 31. Braunus sciocorinus (Walker, 1867), male genitalia. 18 – 23, pygophore; 18 – 19. dorsal view; 20 – 21. posterior view; 22 – 23. ventral view; 24. right paramere, lateral view; 25. abdominal segment X, dorsal view; 26 – 31. phallus; 26 – 27. dorsal view; 28 – 29. lateral view; 30 – 31. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 17 – 22 = 0.5; 23 – 30 = 0.25; inset of Fig. 17 = 0.2 mm." httpUri="https://zenodo.org/record/261497/files/figure.png" pageId="2" pageNumber="170">Figs 18–23</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB99349E6E35DFFDD5" box="[871,944,548,572]" captionStart="FIGURES 36 – 43" captionStartId="7.[151,264,1639,1662]" captionTargetBox="[151,1436,555,1615]" captionTargetId="figure@7.[151,1436,555,1616]" captionTargetPageId="7" captionText="FIGURES 36 – 43. Braunus machadoi Barão & Garbelotto sp. nov. 36 – 37. habitus; 36. dorsal view; 37. ventral view; 38 – 43. pygophore; 38 – 39. dorsal view; 40 – 41. posterior view; 42 – 43. ventral view. Abbreviations: dr = dorsal rim; pa = posterolateral angle; par = paramere; sl = mid depression of superior layer of ventral rim; vr = ventral rim; X = abdominal segment X. Scale bars: 35 – 36 = 2 mm; 37 – 42 = 0.5 mm." httpUri="https://zenodo.org/record/261499/files/figure.png" pageId="2" pageNumber="170">38–43</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB99959E6E327FFDD4" box="[966,1040,548,573]" captionStart="FIGURES 60 – 73" captionStartId="11.[151,264,1804,1827]" captionTargetBox="[151,1432,193,1782]" captionTargetId="figure@11.[151,1436,193,1783]" captionTargetPageId="11" captionText="FIGURES 60 – 73. Braunus ocellatus (Thomas, 2001) comb. nov., male genitalia. 60 – 65. pygophore; 60 – 61. dorsal view; 62 – 63. posterior view; 64 – 65. ventral view; 66. right paramere, lateral view; 67. abdominal segment X, dorsal view; 68 – 73. phallus; 68 – 69. dorsal view; 70 – 71. lateral view; 72 – 73. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 52 – 58 = 0.5 mm; 59 – 60 = 0.25 mm." httpUri="https://zenodo.org/record/261503/files/figure.png" pageId="2" pageNumber="170">60–65</figureCitation>
|
||
). Dorsal rim concave with 1+1 trapezoidal projections laterally to segment X. Segment X rectangular (
|
||
<figureCitation id="132D2A381B29FFCB99E49E0D3260FD89" box="[951,1039,583,608]" captionStart="FIGURES 18 – 31" captionStartId="5.[151,264,1815,1838]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@5.[151,1436,195,1791]" captionTargetPageId="5" captionText="FIGURES 18 – 31. Braunus sciocorinus (Walker, 1867), male genitalia. 18 – 23, pygophore; 18 – 19. dorsal view; 20 – 21. posterior view; 22 – 23. ventral view; 24. right paramere, lateral view; 25. abdominal segment X, dorsal view; 26 – 31. phallus; 26 – 27. dorsal view; 28 – 29. lateral view; 30 – 31. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 17 – 22 = 0.5; 23 – 30 = 0.25; inset of Fig. 17 = 0.2 mm." httpUri="https://zenodo.org/record/261497/files/figure.png" pageId="2" pageNumber="170">Figs 25</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB9E489E0D3254FD89" box="[1051,1083,583,608]" captionStart="FIGURES 60 – 73" captionStartId="11.[151,264,1804,1827]" captionTargetBox="[151,1432,193,1782]" captionTargetId="figure@11.[151,1436,193,1783]" captionTargetPageId="11" captionText="FIGURES 60 – 73. Braunus ocellatus (Thomas, 2001) comb. nov., male genitalia. 60 – 65. pygophore; 60 – 61. dorsal view; 62 – 63. posterior view; 64 – 65. ventral view; 66. right paramere, lateral view; 67. abdominal segment X, dorsal view; 68 – 73. phallus; 68 – 69. dorsal view; 70 – 71. lateral view; 72 – 73. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 52 – 58 = 0.5 mm; 59 – 60 = 0.25 mm." httpUri="https://zenodo.org/record/261503/files/figure.png" pageId="2" pageNumber="170">67</figureCitation>
|
||
). Parameres F-shaped (
|
||
<figureCitation id="132D2A381B29FFCB9F139E0D33F8FD89" box="[1344,1431,583,608]" captionStart="FIGURES 18 – 31" captionStartId="5.[151,264,1815,1838]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@5.[151,1436,195,1791]" captionTargetPageId="5" captionText="FIGURES 18 – 31. Braunus sciocorinus (Walker, 1867), male genitalia. 18 – 23, pygophore; 18 – 19. dorsal view; 20 – 21. posterior view; 22 – 23. ventral view; 24. right paramere, lateral view; 25. abdominal segment X, dorsal view; 26 – 31. phallus; 26 – 27. dorsal view; 28 – 29. lateral view; 30 – 31. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 17 – 22 = 0.5; 23 – 30 = 0.25; inset of Fig. 17 = 0.2 mm." httpUri="https://zenodo.org/record/261497/files/figure.png" pageId="2" pageNumber="170">Figs 24</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB9AC49E2636D9FD6C" box="[151,182,620,645]" captionStart="FIGURES 60 – 73" captionStartId="11.[151,264,1804,1827]" captionTargetBox="[151,1432,193,1782]" captionTargetId="figure@11.[151,1436,193,1783]" captionTargetPageId="11" captionText="FIGURES 60 – 73. Braunus ocellatus (Thomas, 2001) comb. nov., male genitalia. 60 – 65. pygophore; 60 – 61. dorsal view; 62 – 63. posterior view; 64 – 65. ventral view; 66. right paramere, lateral view; 67. abdominal segment X, dorsal view; 68 – 73. phallus; 68 – 69. dorsal view; 70 – 71. lateral view; 72 – 73. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 52 – 58 = 0.5 mm; 59 – 60 = 0.25 mm." httpUri="https://zenodo.org/record/261503/files/figure.png" pageId="2" pageNumber="170">66</figureCitation>
|
||
), placed laterad to segment X. Inferior layer of ventral rim uniformly excavated or emarginated; superior layer of ventral rim depressed medially. Apical margin of phallotheca with a membranous thecal shield (
|
||
<figureCitation id="132D2A381B29FFCB9F409EC533F7FD41" box="[1299,1432,655,680]" captionStart="FIGURES 18 – 31" captionStartId="5.[151,264,1815,1838]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@5.[151,1436,195,1791]" captionTargetPageId="5" captionText="FIGURES 18 – 31. Braunus sciocorinus (Walker, 1867), male genitalia. 18 – 23, pygophore; 18 – 19. dorsal view; 20 – 21. posterior view; 22 – 23. ventral view; 24. right paramere, lateral view; 25. abdominal segment X, dorsal view; 26 – 31. phallus; 26 – 27. dorsal view; 28 – 29. lateral view; 30 – 31. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 17 – 22 = 0.5; 23 – 30 = 0.25; inset of Fig. 17 = 0.2 mm." httpUri="https://zenodo.org/record/261497/files/figure.png" pageId="2" pageNumber="170">Figs 26–31</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB9AC49EFE368FFD24" box="[151,224,692,717]" captionStart="FIGURES 60 – 73" captionStartId="11.[151,264,1804,1827]" captionTargetBox="[151,1432,193,1782]" captionTargetId="figure@11.[151,1436,193,1783]" captionTargetPageId="11" captionText="FIGURES 60 – 73. Braunus ocellatus (Thomas, 2001) comb. nov., male genitalia. 60 – 65. pygophore; 60 – 61. dorsal view; 62 – 63. posterior view; 64 – 65. ventral view; 66. right paramere, lateral view; 67. abdominal segment X, dorsal view; 68 – 73. phallus; 68 – 69. dorsal view; 70 – 71. lateral view; 72 – 73. ventral view. Abbreviations: dr = dorsal rim; ds = ductus seminis distalis; lcp = laterodorsal conjunctival process; pa = posterolateral angle; par = paramere; ph = phallotheca; sl = mid depression of superior layer of ventral rim; ts = thecal shield; v = vesica; vcp = ventral conjunctival process; vr = ventral rim; X = abdominal segment X. Scale bars: 52 – 58 = 0.5 mm; 59 – 60 = 0.25 mm." httpUri="https://zenodo.org/record/261503/files/figure.png" pageId="2" pageNumber="170">68–73</figureCitation>
|
||
;
|
||
<emphasis id="B962EAAF1B29FFCB9ABF9EFF3690FD25" box="[236,255,693,716]" italics="true" pageId="2" pageNumber="170">ts</emphasis>
|
||
). Ventral and laterodorsal conjunctival processes present, ventral hatchet-like.
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B29FFCB9A949E9D3214FCB4" blockId="2.[151,1437,151,1184]" pageId="2" pageNumber="170">
|
||
<emphasis id="B962EAAF1B29FFCB9A949E9D37E2FD19" bold="true" box="[199,397,727,752]" pageId="2" pageNumber="170">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="132D2A381B29FFCB9BCF9E9D3471FD19" box="[412,542,727,752]" captionStart="FIGURES 32 – 35" captionStartId="6.[151,264,851,874]" captionTargetBox="[151,1436,193,830]" captionTargetId="figure@6.[151,1436,193,830]" captionTargetPageId="6" captionText="FIGURES 32 – 35. Braunus sciocorinus (Walker, 1867), female genitalia. 32 – 33. genital plates, ventroposterior view; 34 – 35. female receptaculum seminis and ausenwand, ventral view. Abbreviations: ch = chitinellipsen; cs = capsula seminalis; gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; g 9 = gonapophyses 9; la 8 = laterotergites 8; la 9 = laterotergites 9; lf = lateral folding of gonapophyses 9; md = median duct of vesicular area; pi = pars intermedialis; tvi = thickning of vaginal intima; X = abdominal segment X. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261498/files/figure.png" pageId="2" pageNumber="170">Figs 32–33</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB98799E9D341BFD19" box="[554,628,727,752]" captionStart="FIGURES 44 – 47" captionStartId="8.[151,264,734,757]" captionTargetBox="[151,1436,193,712]" captionTargetId="figure@8.[151,1436,193,713]" captionTargetPageId="8" captionText="FIGURES 44 – 47. Braunus gibbus Garbelotto & Barão sp. nov. 44 – 45. habitus; 44. dorsal view; 45. ventral view; 46 – 47. genital plates. Abbreviations: gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; la 8 = laterotergites 8; la 9 = laterotergites 9; X = abdominal segment X. Scale bars: 43 – 44 = 2 mm; 45 – 46 = 0.5 mm." httpUri="https://zenodo.org/record/261500/files/figure.png" pageId="2" pageNumber="170">46–47</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB982C9E9D34A7FD19" box="[639,712,727,752]" captionStart="FIGURES 48 – 53" captionStartId="9.[151,264,731,754]" captionTargetBox="[151,1436,195,707]" captionTargetId="figure@9.[151,1436,195,707]" captionTargetPageId="9" captionText="FIGURES 48 – 53. Braunus prionotus Barão & Garbelotto sp. nov. 48 – 49. habitus; 48. dorsal view; 49. ventral view; 50. head, dorsal view; 51. external scent efferent system; 52 – 53. genital plates. Abbreviations: ev = evaporatorium; gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; la 8 = laterotergites 8; la 9 = laterotergites 9; ms = mesopleura; mt = metapleura; o = ostiole; X = abdominal segment X. Scale bars: 48 – 49 = 2 mm; 50 – 53 = 1 mm." httpUri="https://zenodo.org/record/261501/files/figure.png" pageId="2" pageNumber="170">52–53</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB98879E9D3573FD19" box="[724,796,727,752]" captionStart="FIGURES 74 – 77" captionStartId="12.[151,264,838,861]" captionTargetBox="[151,1436,193,817]" captionTargetId="figure@12.[151,1436,193,818]" captionTargetPageId="12" captionText="FIGURES 74 – 77. Braunus ocellatus (Thomas, 2001) comb. nov., female genitalia. 74 – 75. genital plates, ventroposterior view; 76 – 77. female receptaculum seminis and ausenwand, ventral view. Abbreviations: cs = capsula seminalis; gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; g 9 = gonapophyses 9; la 8 = laterotergites 8; la 9 = laterotergites 9; lf = lateral folding of gonapophyses 9; md = median duct of vesicular area; pi = pars intermedialis; rs = ring sclerites; tvi = thickning of vaginal intima; X = abdominal segment X. Scale bars: 67 – 68 = 0.5 mm; 69 – 70 = 0.25 mm." httpUri="https://zenodo.org/record/261504/files/figure.png" pageId="2" pageNumber="170">74–75</figureCitation>
|
||
).
|
||
<emphasis id="B962EAAF1B29FFCB99619E9D352EFD19" bold="true" box="[818,833,727,752]" pageId="2" pageNumber="170">S</emphasis>
|
||
utural margins of gonocoxites 8 each parallel basally, usually juxtaposed, and divergent apically. Gonapophyses 8 visible, usually carinated medially. Spiracle of laterotergites 8 visible, placed posterior to lateral angle of each gonocoxite 8. Gonocoxites 9 rectangular or trapezoidal. Laterotergites 9 basally depressed, rounded apically. Segment X rectangular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C65361B29FFCB9A949F2D3741FC4D" pageId="2" pageNumber="170" type="distribution">
|
||
<paragraph id="8BA936BD1B29FFCB9A949F2D3741FC4D" blockId="2.[151,1437,151,1184]" pageId="2" pageNumber="170">
|
||
<emphasis id="B962EAAF1B29FFCB9A949F2D370FFC69" bold="true" box="[199,352,871,896]" pageId="2" pageNumber="170">Distribution.</emphasis>
|
||
Central America: Costa Rica. South America: Venezuela, Colombia, Ecuador, Peru and Bolivia (
|
||
<figureCitation id="132D2A381B29FFCB9ACC9FC6374DFC4D" box="[159,290,908,933]" captionStart="FIGURES 78 – 82" captionStartId="13.[151,264,1110,1133]" captionTargetBox="[151,1436,591,1086]" captionTargetId="figure@13.[151,1436,591,1086]" captionTargetPageId="13" captionText="FIGURES 78 – 82. Distribution of species of Braunus Distant, 1899 and localities of some collection events. 78. map of localities; 79. a collection locality of B. sciocorinus (Walker, 1867) and B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, Yanayacu Biological Station; photo by D. Forero); 80. type locality of B. machadoi Barão & Garbelotto sp. nov. (PERU, Cuzco, Wayqecha Research Center; photo by C. Weirauch); 81 – 82. a collection locality of B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, 1.3 km SW of Baeza; photos by J. Skuhrovec)." httpUri="https://zenodo.org/record/261505/files/figure.png" pageId="2" pageNumber="170">Figs 78–82</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C65361B29FFCB9A949FE53514FB49" pageId="2" pageNumber="170" type="discussion">
|
||
<paragraph id="8BA936BD1B29FFCB9A949FE53514FB49" blockId="2.[151,1437,151,1184]" pageId="2" pageNumber="170">
|
||
<emphasis id="B962EAAF1B29FFCB9A949FE53754FC21" bold="true" box="[199,315,943,968]" pageId="2" pageNumber="170">Remarks.</emphasis>
|
||
<bibRefCitation id="EF874B4C1B29FFCB9B109FE53461FC21" author="McDonald" box="[323,526,943,968]" pageId="2" pageNumber="170" refString="McDonald, F. J. D. (1982) Lojus asperus, a new genus and species (Hemiptera: Pentatomidae). Annals of the Entomological Society of America, 75, 327 - 329. http: // dx. doi. org / 10.1093 / aesa / 75.3.327" type="journal article" year="1982">McDonald (1982)</bibRefCitation>
|
||
related
|
||
<emphasis id="B962EAAF1B29FFCB983F9FFB34A4FC21" box="[620,715,945,968]" italics="true" pageId="2" pageNumber="170">Braunus</emphasis>
|
||
[sic
|
||
<taxonomicName id="4C164D3E1B29FFCB99569FFB352CFC21" box="[773,835,945,968]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB99569FFB352CFC21" box="[773,835,945,968]" italics="true" pageId="2" pageNumber="170">Lojus</emphasis>
|
||
</taxonomicName>
|
||
] to
|
||
<taxonomicName id="4C164D3E1B29FFCB99209FE53215FC21" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" box="[883,1146,943,968]" class="Insecta" family="Pentatomidae" genus="Berecynthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB99209FE53593FC21" box="[883,1020,943,968]" italics="true" pageId="2" pageNumber="170">Berecynthus</emphasis>
|
||
Stål, 1862
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D3E1B29FFCB9EDB9FE53308FC21" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" box="[1160,1383,943,968]" class="Insecta" family="Pentatomidae" genus="Padaeus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9EDB9FE53287FC21" box="[1160,1256,943,968]" italics="true" pageId="2" pageNumber="170">Padaeus</emphasis>
|
||
Stål, 1862
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C164D3E1B29FFCB9AC49F9F37E8FC05" authority="Spinola, 1837" authorityName="Spinola" authorityYear="1837" box="[151,391,980,1005]" class="Insecta" family="Pentatomidae" genus="Proxys" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9AC49F9F368BFC05" box="[151,228,981,1004]" italics="true" pageId="2" pageNumber="170">Proxys</emphasis>
|
||
Spinola, 1837
|
||
</taxonomicName>
|
||
based on the shape of metasternal gland (MG) peritreme and projections on the anterolateral margins of pronotum. However, while the MG peritreme is obsolete in
|
||
<emphasis id="B962EAAF1B29FFCB99E39FB33260FBF9" box="[944,1039,1017,1040]" italics="true" pageId="2" pageNumber="170">Braunus</emphasis>
|
||
, it is fully developed, spout-shaped on
|
||
<taxonomicName id="4C164D3E1B29FFCB9AE89856372BFBDC" box="[187,324,1052,1077]" class="Insecta" family="Pentatomidae" genus="Berecynthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9AE89856372BFBDC" box="[187,324,1052,1077]" italics="true" pageId="2" pageNumber="170">Berecynthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D3E1B29FFCB9B07985637DBFBDC" box="[340,436,1052,1077]" class="Insecta" family="Pentatomidae" genus="Padaeus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9B07985637DBFBDC" box="[340,436,1052,1077]" italics="true" pageId="2" pageNumber="170">Padaeus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D3E1B29FFCB9BBE98573456FBDD" box="[493,569,1053,1076]" class="Insecta" family="Pentatomidae" genus="Proxys" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9BBE98573456FBDD" box="[493,569,1053,1076]" italics="true" pageId="2" pageNumber="170">Proxys</emphasis>
|
||
</taxonomicName>
|
||
. Similarly, the anterolateral margins of the pronotum are armed with fingerlike process in
|
||
<emphasis id="B962EAAF1B29FFCB9B09980B37D6FBB1" box="[346,441,1089,1112]" italics="true" pageId="2" pageNumber="170">Braunus</emphasis>
|
||
and
|
||
<taxonomicName id="4C164D3E1B29FFCB9851987534E4FBB1" box="[514,651,1087,1112]" class="Insecta" family="Pentatomidae" genus="Berecynthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9851987534E4FBB1" box="[514,651,1087,1112]" italics="true" pageId="2" pageNumber="170">Berecynthus</emphasis>
|
||
</taxonomicName>
|
||
, whereas smooth in
|
||
<taxonomicName id="4C164D3E1B29FFCB99C598753599FBB1" box="[918,1014,1087,1112]" class="Insecta" family="Pentatomidae" genus="Padaeus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB99C598753599FBB1" box="[918,1014,1087,1112]" italics="true" pageId="2" pageNumber="170">Padaeus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D3E1B29FFCB9E6D980B32E5FBB1" box="[1086,1162,1089,1112]" class="Insecta" family="Pentatomidae" genus="Proxys" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9E6D980B32E5FBB1" box="[1086,1162,1089,1112]" italics="true" pageId="2" pageNumber="170">Proxys</emphasis>
|
||
</taxonomicName>
|
||
. The corrugate dorsal appearance, brachyptery, obsolete MG peritreme, and genitalia structure of
|
||
<emphasis id="B962EAAF1B29FFCB9E52982F320FFB95" box="[1025,1120,1125,1148]" italics="true" pageId="2" pageNumber="170">Braunus</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C164D3E1B29FFCB9EAC982F37D7FB49" authority="Grazia & Barcellos, 2004" authorityName="Grazia & Barcellos" authorityYear="2004" class="Insecta" family="Pentatomidae" genus="Pentatomiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAAF1B29FFCB9EAC982F33F3FB95" box="[1279,1436,1125,1148]" italics="true" pageId="2" pageNumber="170">Pentatomiana</emphasis>
|
||
Grazia & Barcellos, 2004
|
||
</taxonomicName>
|
||
, as noted by Grazia & Barcellos (2004).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
<subSection id="E2992F561B29FFCA9AC498BE33F3FE5D" lastPageId="3" lastPageNumber="171" pageId="2" pageNumber="170" type="key">
|
||
<paragraph id="8BA936BD1B29FFCB9AC498BE34EBFAE7" blockId="2.[151,644,1268,1294]" box="[151,644,1268,1294]" pageId="2" pageNumber="170">
|
||
<heading id="D0E181D11B29FFCB9AC498BE34EBFAE7" bold="true" box="[151,644,1268,1294]" fontSize="11" level="1" pageId="2" pageNumber="170" reason="1">
|
||
<emphasis id="B962EAAF1B29FFCB9AC498BE34EBFAE7" bold="true" box="[151,644,1268,1294]" pageId="2" pageNumber="170">
|
||
Key to species of
|
||
<emphasis id="B962EAAF1B29FFCB9B3E98BE37B8FAE7" bold="true" box="[365,471,1268,1294]" italics="true" pageId="2" pageNumber="170">Braunus</emphasis>
|
||
<bibRefCitation id="EF874B4C1B29FFCB9B8C98BE34EBFAE7" author="Distant" box="[479,644,1268,1294]" pageId="2" pageNumber="170" refString="Distant, W. L. (1899) Rhyncotal notes - III. Heteroptera: Discocephalinae and Pentatominae (part.). The Annals and Magazine of Natural History, including Zoology, Botany and Geology (seventh series), 4 (14), 421 - 445." type="journal article" year="1899">Distant, 1899</bibRefCitation>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<subSubSection id="C30C65361B29FFCA9AC4997033F3FE5D" lastPageId="3" lastPageNumber="171" pageId="2" pageNumber="170" type="key">
|
||
<key id="FF87618C1B29FFCA9AC4997033F3FE5D" lastPageId="3" lastPageNumber="171" pageId="2" pageNumber="170">
|
||
<keyStep id="30E22E181B29FFCB9AC4997033F4FA4E" pageId="2" pageNumber="170">
|
||
<paragraph id="8BA936BD1B29FFCB9AC4997033F3FA84" blockId="2.[151,1436,1338,1447]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC4997033F3FA84" pageId="2" pageNumber="170">
|
||
1. Lateral margin of each mandibular plate denticulate (
|
||
<figureCitation id="132D2A381B29FFCB98889970354CFAA6" box="[731,803,1338,1359]" captionStart="FIGURES 48 – 53" captionStartId="9.[151,264,731,754]" captionTargetBox="[151,1436,195,707]" captionTargetId="figure@9.[151,1436,195,707]" captionTargetPageId="9" captionText="FIGURES 48 – 53. Braunus prionotus Barão & Garbelotto sp. nov. 48 – 49. habitus; 48. dorsal view; 49. ventral view; 50. head, dorsal view; 51. external scent efferent system; 52 – 53. genital plates. Abbreviations: ev = evaporatorium; gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; la 8 = laterotergites 8; la 9 = laterotergites 9; ms = mesopleura; mt = metapleura; o = ostiole; X = abdominal segment X. Scale bars: 48 – 49 = 2 mm; 50 – 53 = 1 mm." httpUri="https://zenodo.org/record/261501/files/figure.png" pageId="2" pageNumber="170">Fig. 50</figureCitation>
|
||
); humeral angles of pronotum projected anteriad, each with multiple processes (
|
||
<figureCitation id="132D2A381B29FFCB9B3F991D37C4FA85" box="[364,427,1367,1388]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A381B29FFCB9BE6991D37BFFA85" box="[437,464,1367,1388]" captionStart="FIGURES 48 – 53" captionStartId="9.[151,264,731,754]" captionTargetBox="[151,1436,195,707]" captionTargetId="figure@9.[151,1436,195,707]" captionTargetPageId="9" captionText="FIGURES 48 – 53. Braunus prionotus Barão & Garbelotto sp. nov. 48 – 49. habitus; 48. dorsal view; 49. ventral view; 50. head, dorsal view; 51. external scent efferent system; 52 – 53. genital plates. Abbreviations: ev = evaporatorium; gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; la 8 = laterotergites 8; la 9 = laterotergites 9; ms = mesopleura; mt = metapleura; o = ostiole; X = abdominal segment X. Scale bars: 48 – 49 = 2 mm; 50 – 53 = 1 mm." httpUri="https://zenodo.org/record/261501/files/figure.png" pageId="2" pageNumber="170">48</figureCitation>
|
||
); apex of scutellum explanate (
|
||
<figureCitation id="132D2A381B29FFCB98A8991D352EFA85" box="[763,833,1367,1388]" captionStart="FIGURES 48 – 53" captionStartId="9.[151,264,731,754]" captionTargetBox="[151,1436,195,707]" captionTargetId="figure@9.[151,1436,195,707]" captionTargetPageId="9" captionText="FIGURES 48 – 53. Braunus prionotus Barão & Garbelotto sp. nov. 48 – 49. habitus; 48. dorsal view; 49. ventral view; 50. head, dorsal view; 51. external scent efferent system; 52 – 53. genital plates. Abbreviations: ev = evaporatorium; gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; la 8 = laterotergites 8; la 9 = laterotergites 9; ms = mesopleura; mt = metapleura; o = ostiole; X = abdominal segment X. Scale bars: 48 – 49 = 2 mm; 50 – 53 = 1 mm." httpUri="https://zenodo.org/record/261501/files/figure.png" pageId="2" pageNumber="170">Fig. 48</figureCitation>
|
||
)................
|
||
<taxonomicName id="4C164D3E1B29FFCB9E5D99123327FA85" authority="Barao & Garbelotto" authorityName="Barao & Garbelotto" box="[1038,1352,1367,1389]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="species" species="prionotus" status="sp. nov.">
|
||
<emphasis id="B962EAAF1B29FFCB9E5D991232EAFA84" box="[1038,1157,1368,1389]" italics="true" pageId="2" pageNumber="170">B. prionotus</emphasis>
|
||
Barão & Garbelotto
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25157D41B29FFCB9F1C991D33F3FA84" box="[1359,1436,1367,1389]" pageId="2" pageNumber="170" rank="species">
|
||
<emphasis id="B962EAAF1B29FFCB9F1C991D33F3FA84" bold="true" box="[1359,1436,1367,1389]" pageId="2" pageNumber="170">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B29FFCB9AC4993F33F4FA4E" blockId="2.[151,1436,1338,1447]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC4993F33F4FA4E" pageId="2" pageNumber="170">
|
||
1’. Lateral margin of each mandibular plate smooth; humeral angles of pronotum projected laterad, each with 1–4 processes (
|
||
<figureCitation id="132D2A381B29FFCB9F22993F372CFA4E" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 1–5, 7–11</figureCitation>
|
||
); apex of scutellum smooth..................................................................... 2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E22E181B29FFCB9AC499E533F4F9F5" pageId="2" pageNumber="170">
|
||
<paragraph id="8BA936BD1B29FFCB9AC499E533F3FA0B" blockId="2.[151,1436,1455,1564]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC499E533F3FA0B" pageId="2" pageNumber="170">
|
||
2(1). Anterolateral margins of pronotum smooth, except for a weak tubercle at each anterior angle; humeral angles of pronotum each with single digitoid process at apex (
|
||
<figureCitation id="132D2A381B29FFCB981299873416FA0B" box="[577,633,1485,1506]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Fig. 8</figureCitation>
|
||
).......................................
|
||
<taxonomicName id="4C164D3E1B29FFCB9E0299873347FA0B" authority="Thomas" authorityName="Thomas" box="[1105,1320,1484,1506]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="species" species="ateuchus" status="comb. nov.">
|
||
<emphasis id="B962EAAF1B29FFCB9E02998732ABFA0B" box="[1105,1220,1484,1506]" italics="true" pageId="2" pageNumber="170">B. ateuchus</emphasis>
|
||
(Thomas)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25157D41B29FFCB9F7D998633F3FA0B" box="[1326,1436,1484,1506]" pageId="2" pageNumber="170">
|
||
<emphasis id="B962EAAF1B29FFCB9F7D998633F3FA0B" bold="true" box="[1326,1436,1484,1506]" pageId="2" pageNumber="170">comb. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B29FFCB9AC499A033F4F9F5" blockId="2.[151,1436,1455,1564]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC499A033F4F9F5" pageId="2" pageNumber="170">
|
||
2’. Anterolateral margins of pronotum each with spine-like tubercles; humeral angles of pronotum each with two or more processes apically (
|
||
<figureCitation id="132D2A381B29FFCB9B289A4D344BF9F5" box="[379,548,1543,1564]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 1–5, 7, 9–11</figureCitation>
|
||
)........................................................................ 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E22E181B29FFCB9AC49A6F33F4F97B" pageId="2" pageNumber="170">
|
||
<paragraph id="8BA936BD1B29FFCB9AC49A6F33F4F9BE" blockId="2.[151,1436,1573,1682]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC49A6F33F4F9BE" pageId="2" pageNumber="170">
|
||
3(2’). Anterior angles of pronotum each with a spine-like tubercle surpassing the lateral margin of each compound eye in dorsal view (
|
||
<figureCitation id="132D2A381B29FFCB9AB49A08373BF9BE" box="[231,340,1602,1623]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 1–4, 7</figureCitation>
|
||
)......................................................................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B29FFCB9AC49A1533F4F97B" blockId="2.[151,1436,1573,1682]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC49A1533F4F97B" pageId="2" pageNumber="170">
|
||
3’. Anterior angles of pronotum each with a spine-like tubercle not surpassing the lateral margin of each compound eye in dorsal view (
|
||
<figureCitation id="132D2A381B29FFCB9B4F9A3737FBF97B" box="[284,404,1661,1682]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 5, 9–11</figureCitation>
|
||
).................................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E22E181B29FFCB9AC49AD033F3F8EE" pageId="2" pageNumber="170">
|
||
<paragraph id="8BA936BD1B29FFCB9AC49AD033F4F925" blockId="2.[151,1436,1690,1799]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC49AD033F4F925" pageId="2" pageNumber="170">
|
||
4(3). Head uniformly dark brown dorsally; posterolateral margins of pronotum sinuated, each with median indentation (
|
||
<figureCitation id="132D2A381B29FFCB9F649AD033E1F946" box="[1335,1422,1690,1711]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 1–3</figureCitation>
|
||
); internal margin of each pronotal cicatrix with white spot..........................
|
||
<taxonomicName id="4C164D3E1B29FFCB9E659AF23375F925" authority="Walker" authorityName="Walker" box="[1078,1306,1719,1741]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="species" species="sciocorinus">
|
||
<emphasis id="B962EAAF1B29FFCB9E659AF232AFF924" box="[1078,1216,1720,1741]" italics="true" pageId="2" pageNumber="170">B. sciocorinus</emphasis>
|
||
(Walker)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132D2A381B29FFCB9F749AFD33FBF925" box="[1319,1428,1719,1740]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,627,650]" captionTargetBox="[151,1436,195,603]" captionTargetId="figure@4.[151,1436,195,603]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Braunus sciocorinus (Walker, 1867), general morphology. 12 – 13. habitus; 12. dorsal view; 13. ventral view; 14. male forewing; 15. female forewing; 16. male hindwing; 17. external scent efferent system, inset showing ostiolar area. Abreviattions: ev = evaporatorium; ms = mesopleura; mt = metapleura; o = ostiole. Scale bars: 11 – 12 = 2 mm; 13 – 16 = 0.5 mm." httpUri="https://zenodo.org/record/261496/files/figure.png" pageId="2" pageNumber="170">Figs 12–13</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B29FFCB9AC49A9F33F3F8EE" blockId="2.[151,1436,1690,1799]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC49A9F33F3F8EE" pageId="2" pageNumber="170">
|
||
4’. Head uniformly castaneous dorsally; posterolateral margins of pronotum sinuated, each with tubercle and indentation medially (
|
||
<figureCitation id="132D2A381B29FFCB9AB49AB83751F8EE" box="[231,318,1778,1799]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Figs 4, 7</figureCitation>
|
||
); internal margin of each pronotal cicatrix concolorous with remainder of pronotum......................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E22E181B29FFCB9AC49B4533F3F851" pageId="2" pageNumber="170">
|
||
<paragraph id="8BA936BD1B29FFCB9AC49B4533F3F8B6" blockId="2.[151,1436,1807,1976]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC49B4533F3F8B6" pageId="2" pageNumber="170">
|
||
5(4’). Outline of each mandibular plate black dorsally; anterolateral margins of pronotum each with stout tubercles; each humeral angle trilobed apically; posterior margin of pronotum slightly concave (
|
||
<figureCitation id="132D2A381B29FFCB99CB9B6735BBF8AB" box="[920,980,1837,1858]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Fig. 4</figureCitation>
|
||
); abdomen dark brown ventrally with 1+1 yellowish bands mediolaterally..................................
|
||
<taxonomicName id="4C164D3E1B29FFCB99D99B0032A7F8B6" authority="Barao & Garbelotto" authorityName="Barao & Garbelotto" box="[906,1224,1865,1887]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
|
||
<emphasis id="B962EAAF1B29FFCB99D99B00326DF8B6" box="[906,1026,1865,1887]" italics="true" pageId="2" pageNumber="170">B. machadoi</emphasis>
|
||
Barão & Garbelotto
|
||
</taxonomicName>
|
||
<emphasis id="B962EAAF1B29FFCB9E9D9B003375F889" bold="true" box="[1230,1306,1866,1888]" pageId="2" pageNumber="170">
|
||
<taxonomicNameLabel id="A25157D41B29FFCB9E9D9B003375F889" box="[1230,1306,1866,1888]" pageId="2" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="132D2A381B29FFCB9F749B0033FAF8B6" box="[1319,1429,1866,1887]" captionStart="FIGURES 36 – 43" captionStartId="7.[151,264,1639,1662]" captionTargetBox="[151,1436,555,1615]" captionTargetId="figure@7.[151,1436,555,1616]" captionTargetPageId="7" captionText="FIGURES 36 – 43. Braunus machadoi Barão & Garbelotto sp. nov. 36 – 37. habitus; 36. dorsal view; 37. ventral view; 38 – 43. pygophore; 38 – 39. dorsal view; 40 – 41. posterior view; 42 – 43. ventral view. Abbreviations: dr = dorsal rim; pa = posterolateral angle; par = paramere; sl = mid depression of superior layer of ventral rim; vr = ventral rim; X = abdominal segment X. Scale bars: 35 – 36 = 2 mm; 37 – 42 = 0.5 mm." httpUri="https://zenodo.org/record/261499/files/figure.png" pageId="2" pageNumber="170">Figs 36–37</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B29FFCB9AC49B2D33F3F851" blockId="2.[151,1436,1807,1976]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC49B2D33F3F851" pageId="2" pageNumber="170">
|
||
5’. Outline of each mandibular plate concolorous with remainder of head; anterolateral margins of pronotum each with small tubercles; each humeral angle singled lobed apically; posterior margin of pronotum straight (
|
||
<figureCitation id="132D2A381B29FFCB9E329BCF32F2F873" box="[1121,1181,1925,1946]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Fig. 7</figureCitation>
|
||
); abdomen uniformly reddish-brown ventrally.......................................................
|
||
<taxonomicName id="4C164D3E1B29FFCB9E169BE93348F85E" authority="(McDonald)" baseAuthorityName="McDonald" box="[1093,1319,1954,1976]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="species" species="asperus">
|
||
<emphasis id="B962EAAF1B29FFCB9E169BE932C4F851" box="[1093,1195,1955,1976]" italics="true" pageId="2" pageNumber="170">B. asperus</emphasis>
|
||
(McDonald)
|
||
</taxonomicName>
|
||
<emphasis id="B962EAAF1B29FFCB9F7D9BE833F3F851" bold="true" box="[1326,1436,1954,1976]" pageId="2" pageNumber="170">comb. nov.</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E22E181B29FFCA9AC49BF533F4FF23" lastPageId="3" lastPageNumber="171" pageId="2" pageNumber="170">
|
||
<paragraph id="8BA936BD1B29FFCB9AC49BF533F3F81B" blockId="2.[151,1436,1983,2034]" pageId="2" pageNumber="170">
|
||
<keyLead id="30E795881B29FFCB9AC49BF533F3F81B" pageId="2" pageNumber="170">
|
||
6(3’). Tubercles on each anterolateral margin of pronotum obtuse; humeral angles of pronotum each with two processes apically, one anteriorly and one posteriorly (
|
||
<figureCitation id="132D2A381B29FFCB98599B97343FF81B" box="[522,592,2013,2034]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="2" pageNumber="170">Fig. 10</figureCitation>
|
||
).......................................
|
||
<taxonomicName id="4C164D3E1B29FFCB9E609B973347F81B" authority="Thomas" authorityName="Thomas" box="[1075,1320,2012,2034]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="170" phylum="Arthropoda" rank="species" species="brettlizorum" status="comb. nov.">
|
||
<emphasis id="B962EAAF1B29FFCB9E609B9732ACF81B" box="[1075,1219,2012,2034]" italics="true" pageId="2" pageNumber="170">B. brettlizorum</emphasis>
|
||
(Thomas)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25157D41B29FFCB9F7D9B9633F3F81B" box="[1326,1436,2012,2034]" pageId="2" pageNumber="170">
|
||
<emphasis id="B962EAAF1B29FFCB9F7D9B9633F3F81B" bold="true" box="[1326,1436,2012,2034]" pageId="2" pageNumber="170">comb. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B28FFCA9AC49CDD33F4FF23" blockId="3.[151,1436,151,202]" pageId="3" pageNumber="171">
|
||
<keyLead id="30E795881B28FFCA9AC49CDD33F4FF23" pageId="3" pageNumber="171">
|
||
6’. Tubercles on each anterolateral margin of pronotum acute; humeral angles of pronotum each with three processes apically, two anterior and one posterior (
|
||
<figureCitation id="132D2A381B28FFCA9BB79CFF3432FF23" box="[484,605,181,202]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="3" pageNumber="171">Figs 5, 9, 11</figureCitation>
|
||
)................................................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E22E181B28FFCA9AC49C9833F4FED6" pageId="3" pageNumber="171">
|
||
<paragraph id="8BA936BD1B28FFCA9AC49C9833F3FEEC" blockId="3.[151,1436,210,320]" pageId="3" pageNumber="171">
|
||
<keyLead id="30E795881B28FFCA9AC49C9833F3FEEC" pageId="3" pageNumber="171">
|
||
7(6’). Ocelli absent; apex of each hemelytral radial vein not calloused; inferior layer of ventral rim of pygophore smooth........................................................................
|
||
<taxonomicName id="4C164D3E1B28FFCA99899CBA3348FEED" authority="Brailovsky & Rolston" authorityName="Brailovsky & Rolston" box="[986,1319,239,261]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="bordoni" status="comb. nov.">
|
||
<emphasis id="B962EAAF1B28FFCA99899CBA3250FEEC" box="[986,1087,239,261]" italics="true" pageId="3" pageNumber="171">B. bordoni</emphasis>
|
||
(Brailovsky & Rolston)
|
||
</taxonomicName>
|
||
<emphasis id="B962EAAF1B28FFCA9F7D9CA533F3FEEC" bold="true" box="[1326,1436,239,261]" pageId="3" pageNumber="171">
|
||
<taxonomicNameLabel id="A25157D41B28FFCA9F7D9CA533F3FEEC" box="[1326,1436,239,261]" pageId="3" pageNumber="171">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B28FFCA9AC49D4733F4FED6" blockId="3.[151,1436,210,320]" pageId="3" pageNumber="171">
|
||
<keyLead id="30E795881B28FFCA9AC49D4733F4FED6" pageId="3" pageNumber="171">
|
||
7’. Ocelli present; apex of each hemelytral radial vein calloused; inferior layer of ventral rim of pygophore projected medially (except
|
||
<taxonomicName id="4C164D3E1B28FFCA9B7E9D603423FED6" authority="Garbelotto & Barao" authorityName="Garbelotto & Barao" box="[301,588,297,319]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov.">
|
||
<emphasis id="B962EAAF1B28FFCA9B7E9D6037E7FED6" box="[301,392,297,319]" italics="true" pageId="3" pageNumber="171">B. gibbus</emphasis>
|
||
Garbelotto & Barão
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25157D41B28FFCA98019D6034F0FEA9" box="[594,671,298,320]" pageId="3" pageNumber="171" rank="species">
|
||
<emphasis id="B962EAAF1B28FFCA98019D6034F0FEA9" bold="true" box="[594,671,298,320]" pageId="3" pageNumber="171">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male unknown)................................................. 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E22E181B28FFCA9AC49D0D33F3FE5D" pageId="3" pageNumber="171">
|
||
<paragraph id="8BA936BD1B28FFCA9AC49D0D33F3FE93" blockId="3.[151,1436,327,437]" pageId="3" pageNumber="171">
|
||
<keyLead id="30E795881B28FFCA9AC49D0D33F3FE93" pageId="3" pageNumber="171">
|
||
8(7’). Dark brown dorsally; abdomen uniformly dark brown ventrally; anterior lobe of pronotum as long as posterior lobe of pronotum (
|
||
<figureCitation id="132D2A381B28FFCA9B409D2F3738FE93" box="[275,343,357,378]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="3" pageNumber="171">Fig. 11</figureCitation>
|
||
)....................................................
|
||
<taxonomicName id="4C164D3E1B28FFCA99829D2F32C8FE93" authority="Thomas" authorityName="Thomas" box="[977,1191,356,378]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="ocellatus" status="comb. nov.">
|
||
<emphasis id="B962EAAF1B28FFCA99829D2F322CFE93" box="[977,1091,356,378]" italics="true" pageId="3" pageNumber="171">B. ocellatus</emphasis>
|
||
(Thomas)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25157D41B28FFCA9EFF9D2E3375FE93" box="[1196,1306,356,378]" pageId="3" pageNumber="171">
|
||
<emphasis id="B962EAAF1B28FFCA9EFF9D2E3375FE93" bold="true" box="[1196,1306,356,378]" pageId="3" pageNumber="171">comb. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="132D2A381B28FFCA9F749D2F33FAFE93" box="[1319,1429,357,378]" captionStart="FIGURES 54 – 59" captionStartId="10.[151,264,748,771]" captionTargetBox="[151,1435,267,725]" captionTargetId="figure@10.[151,1436,267,725]" captionTargetPageId="10" captionText="FIGURES 54 – 59. Braunus ocellatus (Thomas, 2001) comb. nov.. 54 – 55. habitus; 54. dorsal view; 55. ventral view; 56. male forewing; 57. female forewing; 58. male hindwing; 59. external scent efferent system. Abreviattions: ev = evaporatorium; ms = mesopleura; mt = metapleura; o = ostiole. Scale bars: 47 – 48 = 2 mm; 49 – 52 = 0.5 mm." httpUri="https://zenodo.org/record/261502/files/figure.png" pageId="3" pageNumber="171">Figs 54–55</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA936BD1B28FFCA9AC49DC833F3FE5D" blockId="3.[151,1436,327,437]" pageId="3" pageNumber="171">
|
||
<keyLead id="30E795881B28FFCA9AC49DC833F3FE5D" pageId="3" pageNumber="171">
|
||
8’. Light brown to yellowish dorsally; abdomen dark brown ventrally with 1+1 yellowish bands mediolaterally; anterior lobe of pronotum shorter than posterior lobe of pronotum (
|
||
<figureCitation id="132D2A381B28FFCA98999DD5356CFE5D" box="[714,771,415,436]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão & Garbelotto sp. nov.; 5. B. gibbus Garbelotto & Barão sp. nov.; 6. B. prionotus Barão & Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski & Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="3" pageNumber="171">Fig. 5</figureCitation>
|
||
)............
|
||
<taxonomicName id="4C164D3E1B28FFCA99FA9DEA32A7FE5D" authority="Garbelotto & Barao" authorityName="Garbelotto & Barao" box="[937,1224,415,437]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="171" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov.">
|
||
<emphasis id="B962EAAF1B28FFCA99FA9DEA326BFE5C" box="[937,1028,415,437]" italics="true" pageId="3" pageNumber="171">B. gibbus</emphasis>
|
||
Garbelotto & Barão
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25157D41B28FFCA9E9D9DD53375FE5C" box="[1230,1306,415,437]" pageId="3" pageNumber="171" rank="species">
|
||
<emphasis id="B962EAAF1B28FFCA9E9D9DD53375FE5C" bold="true" box="[1230,1306,415,437]" pageId="3" pageNumber="171">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="132D2A381B28FFCA9F749DD533FAFE5D" box="[1319,1429,415,436]" captionStart="FIGURES 44 – 47" captionStartId="8.[151,264,734,757]" captionTargetBox="[151,1436,193,712]" captionTargetId="figure@8.[151,1436,193,713]" captionTargetPageId="8" captionText="FIGURES 44 – 47. Braunus gibbus Garbelotto & Barão sp. nov. 44 – 45. habitus; 44. dorsal view; 45. ventral view; 46 – 47. genital plates. Abbreviations: gc 8 = gonocoxites 8; gc 9 = gonocoxites 9; g 8 = gonapophyses 8; la 8 = laterotergites 8; la 9 = laterotergites 9; X = abdominal segment X. Scale bars: 43 – 44 = 2 mm; 45 – 46 = 0.5 mm." httpUri="https://zenodo.org/record/261500/files/figure.png" pageId="3" pageNumber="171">Figs 44–45</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |