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<mods:title id="4B444ACF1A7DA913A19970853A5BD319">Description of a second deep-water species in the alpheid shrimp genus Bannereus Bruce, 1988 (Malacostraca: Caridea)</mods:title>
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<mods:namePart id="738DA0A759A722E46F246636E107E5C4">Anker, Arthur</mods:namePart>
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<treatment id="03BF3D13437CFFA6FF59FEB6FB1B0B7E" ID-DOI="http://doi.org/10.5281/zenodo.4323822" ID-GBIF-Taxon="176086567" ID-Zenodo-Dep="4323822" LSID="urn:lsid:plazi:treatment:03BF3D13437CFFA6FF59FEB6FB1B0B7E" httpUri="http://treatment.plazi.org/id/03BF3D13437CFFA6FF59FEB6FB1B0B7E" lastPageId="8" lastPageNumber="141" pageId="3" pageNumber="136">
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<paragraph id="8BA98C05437CFFADFF59FEB6FE5C0F70" blockId="3.[151,460,289,352]" box="[151,460,289,316]" pageId="3" pageNumber="136">
<heading id="D0E13B69437CFFADFF59FEB6FE5C0F70" bold="true" box="[151,460,289,316]" fontSize="11" level="1" pageId="3" pageNumber="136" reason="1">
<emphasis id="B9625017437CFFADFF59FEB6FE5C0F70" bold="true" box="[151,460,289,316]" pageId="3" pageNumber="136">
<taxonomicName id="4C16F786437CFFADFF59FEB6FEF70F70" ID-CoL="874K2" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[151,359,289,316]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B9625017437CFFADFF59FEB6FEF70F70" bold="true" box="[151,359,289,316]" italics="true" pageId="3" pageNumber="136">Bannereus chani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251ED6C437CFFADFEA0FEB6FE5C0F70" box="[366,460,290,316]" pageId="3" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BA98C05437CFFADFF59FED2FF690F2C" blockId="3.[151,460,289,352]" box="[151,249,326,352]" pageId="3" pageNumber="136">
<figureCitation id="132D9080437CFFADFF59FED2FF690F2C" box="[151,249,326,352]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1868,1892]" captionStartId-1="4.[151,250,1869,1893]" captionStartId-2="6.[151,250,1875,1899]" captionStartId-3="7.[151,250,1943,1967]" captionTargetBox-0="[176,1414,194,1837]" captionTargetBox-1="[156,1426,357,1837]" captionTargetBox-2="[218,1363,194,1846]" captionTargetBox-3="[184,1424,194,1909]" captionTargetId-0="figure-21@2.[164,1423,181,1844]" captionTargetId-1="figure-113@4.[151,1436,345,1845]" captionTargetId-2="figure-21@6.[210,1377,181,1851]" captionTargetId-3="figure-19@7.[154,1434,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." captionText-1="FIGURE 2. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral." captionText-2="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." captionText-3="FIGURE 4. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) pleon, third to sixth pleonites, lateral; (B) telson, dorsal; (C) right first pleopod, ventral; (D) right second pleopod, ventral; (E) right uropod, dorsal; (F) same, posterior portion of endopod and exopod showing setal sockets of stiff setae, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.4315590" figureDoi-1="http://doi.org/10.5281/zenodo.4315592" figureDoi-2="http://doi.org/10.5281/zenodo.4315594" figureDoi-3="http://doi.org/10.5281/zenodo.4315596" httpUri-0="https://zenodo.org/record/4315590/files/figure.png" httpUri-1="https://zenodo.org/record/4315592/files/figure.png" httpUri-2="https://zenodo.org/record/4315594/files/figure.png" httpUri-3="https://zenodo.org/record/4315596/files/figure.png" pageId="3" pageNumber="136">Figs 14</figureCitation>
</paragraph>
<paragraph id="8BA98C05437CFFADFF59FE1EFD390F8D" blockId="3.[151,1436,394,449]" pageId="3" pageNumber="136">
(?)
<taxonomicName id="4C16F786437CFFADFF72FE1FFE030FEE" authorityName="Bruce" authorityYear="1988" box="[188,403,394,418]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B9625017437CFFADFF72FE1FFE030FEE" box="[188,403,394,418]" italics="true" pageId="3" pageNumber="136">Bannereus anomalus</emphasis>
</taxonomicName>
<bibRefCitation id="EF87F1F4437CFFADFE61FE1FFDCC0FEE" author="Bruce, A. J." box="[431,604,394,418]" pageId="3" pageNumber="136" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988: 140</bibRefCitation>
(
<emphasis id="B9625017437CFFADFDA3FE1FFD210FEE" box="[621,689,395,418]" italics="true" pageId="3" pageNumber="136">partim</emphasis>
, male specimen), figs. 2DK, 3, 4HJ (as on figure, legend erroneous), 5BD, GJ, 6C [not
<taxonomicName id="4C16F786437CFFADFE57FE3EFD0F0F8D" authority="Bruce, 1988" authorityName="Bruce" authorityYear="1988" box="[409,671,425,449]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B9625017437CFFADFE57FE3EFD880F8D" box="[409,536,425,449]" italics="true" pageId="3" pageNumber="136">B. anomalus</emphasis>
<bibRefCitation id="EF87F1F4437CFFADFDD0FE3DFD0F0F8D" author="Bruce, A. J." box="[542,671,425,449]" pageId="3" pageNumber="136" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce, 1988</bibRefCitation>
</taxonomicName>
].
</paragraph>
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<subSubSection id="C30CDF8E437CFFADFF59FE7BFCF70C1D" pageId="3" pageNumber="136" type="materials_examined">
<paragraph id="8BA98C05437CFFADFF59FE7BFCF70C1D" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
<emphasis id="B9625017437CFFADFF59FE7BFEAF0C45" bold="true" box="[151,319,495,521]" pageId="3" pageNumber="136">Type material</emphasis>
.
<materialsCitation id="3B7E8658437CFFADFE83FE7BFCF40C1D" ID-GBIF-Occurrence="2995098301" collectingDate="2002-05-20" collectionCode="T, R" collectorName="Sta. DW &amp; T. Y. Chan &amp; R. von Cosel &amp; B. Richer de Forges" country="Taiwan" county="Richer de Forges" elevation="329" latitude="22.313334" location="Taiwan" longLatPrecision="125" longitude="121.49333" municipality="Campaign" pageId="3" pageNumber="136" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AD32A7437CFFADFE83FE7BFE220C45" box="[333,434,495,521]" pageId="3" pageNumber="136" type="holotype">holotype</typeStatus>
, ovigerous female (cl
<quantity id="4CEE21E0437CFFADFD63FE7BFC980C46" box="[685,776,495,522]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="136" unit="mm" value="6.3">6.3 mm</quantity>
), MNHN-IU-2017-11749,
<location id="8EC9DADE437CFFADFBF8FE7BFB1B0C45" LSID="urn:lsid:plazi:treatment:03BF3D13437CFFA6FF59FEB6FB1B0B7E:8EC9DADE437CFFADFBF8FE7BFB1B0C45" box="[1078,1163,495,521]" country="Taiwan" county="Richer de Forges" latitude="22.313334" longLatPrecision="125" longitude="121.49333" municipality="Campaign" name="Taiwan" pageId="3" pageNumber="136">Taiwan</location>
, off south-eastern coast,
<collectingMunicipality id="6BCD167F437CFFADFF59FD87FE9A0C61" box="[151,266,531,557]" pageId="3" pageNumber="136">Campaign</collectingMunicipality>
<collectingCountry id="F301CC95437CFFADFEDFFD87FEED0C61" box="[273,381,531,557]" name="Taiwan" pageId="3" pageNumber="136">TAIWAN</collectingCountry>
2002, N/O “Ocean Researcher”,
<collectorName id="26E3E9D3437CFFADFD3AFD87FCC90C62" box="[756,857,531,558]" pageId="3" pageNumber="136">Sta. DW</collectorName>
151,
<geoCoordinate id="EE22EAC2437CFFADFC43FD87FB920C61" box="[909,1026,531,557]" degrees="22" direction="north" minutes="18.8" orientation="latitude" pageId="3" pageNumber="136" precision="92" value="22.313334">22°18.8N</geoCoordinate>
<geoCoordinate id="EE22EAC2437CFFADFBC4FD87FB1D0C61" box="[1034,1165,531,557]" degrees="121" direction="east" minutes="29.6" orientation="longitude" pageId="3" pageNumber="136" precision="92" value="121.49333">121°29.6E</geoCoordinate>
, depth:
<quantity id="4CEE21E0437CFFADFB28FD87FAF70C62" box="[1254,1383,531,558]" metricMagnitude="2" metricUnit="m" metricValue="3.285" metricValueMax="3.56" metricValueMin="3.01" pageId="3" pageNumber="136" unit="m" value="328.5" valueMax="356.0" valueMin="301.0">
<elevation id="003B6B36437CFFADFB28FD87FAF70C62" box="[1254,1383,531,558]" metricMagnitude="2" metricUnit="m" metricValue="3.285" metricValueMax="3.56" metricValueMin="3.01" pageId="3" pageNumber="136" unit="m" value="328.5" valueMax="356.0" valueMin="301.0">301356 m</elevation>
</quantity>
, leg.
<collectorName id="26E3E9D3437CFFADFF59FDACFE9A0C1D" box="[151,266,567,594]" pageId="3" pageNumber="136">T.Y. Chan</collectorName>
,
<collectorName id="26E3E9D3437CFFADFEDBFDACFE370C1D" box="[277,423,567,594]" pageId="3" pageNumber="136">R. von Cosel</collectorName>
and
<collectorName id="26E3E9D3437CFFADFE13FDACFD500C1E" box="[477,704,567,594]" pageId="3" pageNumber="136">B. Richer de Forges</collectorName>
,
<date id="FFA8AAC5437CFFADFD04FDA3FCF40C1D" box="[714,868,567,594]" pageId="3" pageNumber="136" value="2002-05-20">
<collectingDate id="EFEC532D437CFFADFD04FDA3FCF40C1D" box="[714,868,567,594]" pageId="3" pageNumber="136" value="2002-05-20">20 May 2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CDF8E437CFFABFF09FDCFFE790AED" lastPageId="5" lastPageNumber="138" pageId="3" pageNumber="136" type="description">
<paragraph id="8BA98C05437CFFADFF09FDCFFCE40CF1" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
<emphasis id="B9625017437CFFADFF09FDCFFEC10C39" bold="true" box="[199,337,603,629]" pageId="3" pageNumber="136">Description</emphasis>
. Medium-sized alpheid shrimp. Carapace glabrous, somewhat swollen, not compressed laterally. Rostrum present as small median convexity of frontal margin; orbital teeth absent (
<figureCitation id="132D9080437CFFADFB92FDEBFB4B0CD6" box="[1116,1243,639,666]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1A, B</figureCitation>
). Pterygostomial angle broadly rounded (
<figureCitation id="132D9080437CFFADFE6CFD37FE690CF1" box="[418,505,675,701]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1B</figureCitation>
). Cardiac notch moderately deep.
</paragraph>
<paragraph id="8BA98C05437CFFADFF09FD53FAF80D3D" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
First to fourth pleonites with pleura rounded posteroventrally, second greatly expanded in females; fifth pleuron with posteroventral margin somewhat more angular; sixth pleonite with blunt posterolateral projection near telson base, without articulated plate (
<figureCitation id="132D9080437CFFADFE38FC9BFDC00D65" box="[502,592,783,809]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[184,1424,194,1909]" captionTargetId="figure-19@7.[154,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) pleon, third to sixth pleonites, lateral; (B) telson, dorsal; (C) right first pleopod, ventral; (D) right second pleopod, ventral; (E) right uropod, dorsal; (F) same, posterior portion of endopod and exopod showing setal sockets of stiff setae, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4315596" httpUri="https://zenodo.org/record/4315596/files/figure.png" pageId="3" pageNumber="136">Fig. 4A</figureCitation>
). Telson subrectangular, considerably tapering distally, nearly twice as long as proximal width, with 2 pairs of small cuspidate setae (one of them minute) both situated in posterior half of telson; posterior margin straight, with 2 pairs of spiniform setae, mesial almost twice as long as lateral (
<figureCitation id="132D9080437CFFADFACDFCC3FACB0D3D" box="[1283,1371,855,881]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[184,1424,194,1909]" captionTargetId="figure-19@7.[154,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) pleon, third to sixth pleonites, lateral; (B) telson, dorsal; (C) right first pleopod, ventral; (D) right second pleopod, ventral; (E) right uropod, dorsal; (F) same, posterior portion of endopod and exopod showing setal sockets of stiff setae, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4315596" httpUri="https://zenodo.org/record/4315596/files/figure.png" pageId="3" pageNumber="136">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437CFFADFF09FCE8FC350D92" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
Eyes completely concealed by frontal projection of carapace in dorsal view, partly visible in lateral view, relatively well-developed, with normally pigmented corneas, latter occupying most of distolateral area of short stout eyestalk; anteromesial margin rounded, without tubercle (
<figureCitation id="132D9080437CFFADFCD3FC57FC070D92" box="[797,919,963,990]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1A, B</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437CFFADFF09FC73FA0B0AF9" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
Antennular peduncle with dorsally visible portion of first article subequal to second article in length (
<figureCitation id="132D9080437CFFADFAFBFC73FA1E0A4D" box="[1333,1422,999,1025]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1A</figureCitation>
). First article with stylocerite stout, rounded distally, not reaching distal margin of first article; ventromesial carina with large blunt tooth (
<figureCitation id="132D9080437CFFADFE54FBBBFE480A05" box="[410,472,1071,1097]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1</figureCitation>
AC). Second article about 1.3 times as long as wide, ventral surface with small subacute tooth distally (
<figureCitation id="132D9080437CFFADFEB1FBC7FE6C0A21" box="[383,508,1107,1133]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1A, C</figureCitation>
). Third article shortest. Lateral flagellum with fused portion composed of 4 units, accessory ramus very stout, subdivided into at least 3 units (tip broken off); several groups of aesthetascs extending from last unit of fused portion to last unit of accessory ramus; main ramus very slender distal to bifurcation (
<figureCitation id="132D9080437CFFADFAF6FB0FFA1D0AF9" box="[1336,1421,1179,1205]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437CFFADFF09FB2BFA810B0A" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
Antenna with basicerite moderately robust, armed with small subacute distoventral tooth (
<figureCitation id="132D9080437CFFADFB7DFB2BFA990A95" box="[1203,1289,1215,1241]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1B</figureCitation>
). Scaphocerite very broad, ovate, reaching to mid-length of third article of antennular peduncle; lateral margin broadly convex; blade strongly convex anteriorly, overreaching greatly reduced, blunt distolateral tooth (
<figureCitation id="132D9080437CFFADFBADFA93FA8B0B6E" box="[1123,1307,1287,1314]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1A, B, D, E</figureCitation>
). Carpocerite fairly slender, cylindrical, overreaching both scaphocerite and antennular peduncle (
<figureCitation id="132D9080437CFFADFBA9FABFFA940B0A" box="[1127,1284,1323,1350]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1A, B, D</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437CFFADFF09FADBFDE20851" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
Mouthparts not dissected, seemingly typical for genus in external observation. Third maxilliped pediform, robust, coxa with small rounded lateral plate; exopod moderately developed, not reaching penultimate article; antepenultimate article about 4 times as long as greatest width, somewhat curved, ventromesial margin slightly rugose, with several slender spiniform setae; penultimate article subcylindrical, about 2.5 times as long as distal width, with clusters of stout setae on ventromesial and distodorsal surface; ultimate article slender, elongate, tapering distally, tip with stout spiniform setae (
<figureCitation id="132D9080437CFFADFE23F997FDBB0851" box="[493,555,1539,1565]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1</figureCitation>
FH).
</paragraph>
<paragraph id="8BA98C05437CFFADFF09F9B3FC32092E" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
Only left (major) cheliped present (preserved) in
<typeStatus id="54AD32A7437CFFADFCCDF9B3FCF5080D" box="[771,869,1575,1601]" pageId="3" pageNumber="136" type="holotype">holotype</typeStatus>
(
<figureCitation id="132D9080437CFFADFCB9F9B3FC2F080D" box="[887,959,1575,1601]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1893]" captionTargetBox="[156,1426,357,1837]" captionTargetId="figure-113@4.[151,1436,345,1845]" captionTargetPageId="4" captionText="FIGURE 2. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.4315592" httpUri="https://zenodo.org/record/4315592/files/figure.png" pageId="3" pageNumber="136">Fig. 2</figureCitation>
). Major cheliped robust, greatly enlarged relative to body. Ischium short, unarmed (
<figureCitation id="132D9080437CFFADFDA6F9DFFD74082A" box="[616,740,1611,1638]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1893]" captionTargetBox="[156,1426,357,1837]" captionTargetId="figure-113@4.[151,1436,345,1845]" captionTargetPageId="4" captionText="FIGURE 2. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.4315592" httpUri="https://zenodo.org/record/4315592/files/figure.png" pageId="3" pageNumber="136">Fig. 2A, D</figureCitation>
). Merus subtriangular in cross-section, widening distally; ventromesial surface adjacent to margin with numerous spiniform setae inserted in irregular row; distomesial margin unarmed; distodorsal margin blunt (
<figureCitation id="132D9080437CFFADFDE1F907FD4008E2" box="[559,720,1683,1710]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1893]" captionTargetBox="[156,1426,357,1837]" captionTargetId="figure-113@4.[151,1436,345,1845]" captionTargetPageId="4" captionText="FIGURE 2. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.4315592" httpUri="https://zenodo.org/record/4315592/files/figure.png" pageId="3" pageNumber="136">Fig. 2A, D, E</figureCitation>
). Carpus very short, much broader than long, cup-shaped (
<figureCitation id="132D9080437CFFADFAA1F907FF70089E" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1893]" captionTargetBox="[156,1426,357,1837]" captionTargetId="figure-113@4.[151,1436,345,1845]" captionTargetPageId="4" captionText="FIGURE 2. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.4315592" httpUri="https://zenodo.org/record/4315592/files/figure.png" pageId="3" pageNumber="136">Fig. 2A, D</figureCitation>
). Chela with palm smooth, without ridges or other sculpture, swollen, elongate-ovoid in shape (
<figureCitation id="132D9080437CFFADFADAF923FA1E089E" box="[1300,1422,1719,1746]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1893]" captionTargetBox="[156,1426,357,1837]" captionTargetId="figure-113@4.[151,1436,345,1845]" captionTargetPageId="4" captionText="FIGURE 2. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.4315592" httpUri="https://zenodo.org/record/4315592/files/figure.png" pageId="3" pageNumber="136">Fig. 2A, B</figureCitation>
). Fingers about half-length of palm, not twisted mesially or laterally; fingertips strongly curved and crossing distally; pollex slightly shorter than dactylus; cutting surface deeply excavated, channel-like; mesial cutting margin with 2 rounded teeth proximally; lateral cutting margin with single low bump proximally; dactylus with cutting surface distinctly bulging, fitting into excavation of pollex (
<figureCitation id="132D9080437CFFADFD12F8D3FC05092E" box="[732,917,1863,1890]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1893]" captionTargetBox="[156,1426,357,1837]" captionTargetId="figure-113@4.[151,1436,345,1845]" captionTargetPageId="4" captionText="FIGURE 2. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.4315592" httpUri="https://zenodo.org/record/4315592/files/figure.png" pageId="3" pageNumber="136">Fig. 2A, B, C, F</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437CFFADFF09F8FFFC2E09BD" blockId="3.[151,1437,495,2034]" pageId="3" pageNumber="136">
Second pereiopod relatively short, stout (
<figureCitation id="132D9080437CFFADFD41F8FFFD4B09C9" box="[655,731,1899,1925]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1I</figureCitation>
). Merus somewhat longer than ischium. Carpus slightly shorter than merus, subdivided into 5 units (subarticles), proximal longest, their ratio approximately equal to 3: 1: 0.8: 0.8: 2 (
<figureCitation id="132D9080437CFFADFF3EF827FEAD0981" box="[240,317,1971,1997]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1I</figureCitation>
). Chela slightly shorter than carpus, with fingers about 0.7 length of palm; both fingers with dense fan-like tufts of setae near bifid tips; cutting edges unarmed (
<figureCitation id="132D9080437CFFADFC8CF843FC1109BD" box="[834,897,2007,2033]" captionStart="FIGURE 1" captionStartId="2.[151,250,1868,1892]" captionTargetBox="[176,1414,194,1837]" captionTargetId="figure-21@2.[164,1423,181,1844]" captionTargetPageId="2" captionText="FIGURE 1. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) frontal margin and cephalic appendages, dorsal; (B) same, lateral; (C) left antennule, ventral surface, lateral; (D) left antenna, ventral; (E) same, scaphocerite, dorsal; (F) right third maxilliped, lateral; (G) same, penultimate and antepenultimate articles, mesial; (H) left third maxilliped, tip of ultimate article, dorsomesial view; (I) left second pereiopod, lateral; (J) same, chela, mesial; (K) same, tip of dactylus, ventral; (L) same, tip of pollex, ventromesial." figureDoi="http://doi.org/10.5281/zenodo.4315590" httpUri="https://zenodo.org/record/4315590/files/figure.png" pageId="3" pageNumber="136">Fig. 1</figureCitation>
IL).
</paragraph>
<paragraph id="8BA98C05437BFFABFF09FF03FC900F0D" blockId="4.[151,1436,151,321]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="138" pageId="4" pageNumber="137">
Third pereiopod relatively stout (
<figureCitation id="132D9080437BFFAAFDF2FF03FD070EFD" box="[572,663,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="4" pageNumber="137">Fig. 3A</figureCitation>
). Ischium with 1 or 2 stout cuspidate seta(e) on ventrolateral surface (
<figureCitation id="132D9080437BFFAAFF6EFF2FFE870E99" box="[160,279,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="4" pageNumber="137">Fig. 3A, C</figureCitation>
). Merus about twice as long as ischium, somewhat inflated, with both dorsal and ventral margins slightly convex, about 3.5 times as long as maximal width, with small spiniform seta at about mid-length of ventral margin (
<figureCitation id="132D9080437BFFAAFF6EFE97FE890F51" box="[160,281,259,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="4" pageNumber="137">Fig. 3A, C</figureCitation>
). Carpus much slenderer than merus, about 0.4 length of merus, unarmed (
<figureCitation id="132D9080437BFFAAFB96FE97FB220F51" box="[1112,1202,259,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="4" pageNumber="137">Fig. 3A</figureCitation>
). Propodus elongate; ventral margin armed with row of 15 short spiniform setae, sometimes inserted by pairs, including 1 pair of longer spiniform setae distally, adjacent to dactylar base (
<figureCitation id="132D9080437AFFABFD13FF03FCAA0EFD" box="[733,826,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3A</figureCitation>
). Dactylus stout, about 0.2 length of propodus, biunguiculate; terminal unguis much longer than secondary unguis; both unguis directed slightly ventral relative to main dactylar axis and more or less parallel to each other; ventral margin with 4 minute, apparently articulated spinules, distal-most spinule situated near secondary unguis, proximal-most spinule situated at about mid-length of dactylus; dorsal margin with tuft of thickened setae (
<figureCitation id="132D9080437AFFABFDB7FEB3FD620F0D" box="[633,754,295,321]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3A, B</figureCitation>
).
</paragraph>
<caption id="DF69DC8D437BFFAAFF59F8D9FF4E099D" ID-DOI="http://doi.org/10.5281/zenodo.4315592" ID-Zenodo-Dep="4315592" httpUri="https://zenodo.org/record/4315592/files/figure.png" pageId="4" pageNumber="137" startId="4.[151,250,1869,1893]" targetBox="[156,1426,357,1837]" targetPageId="4">
<paragraph id="8BA98C05437BFFAAFF59F8D9FF4E099D" blockId="4.[151,1437,1869,2001]" pageId="4" pageNumber="137">
<emphasis id="B9625017437BFFAAFF59F8D9FE9E0929" bold="true" box="[151,270,1869,1893]" pageId="4" pageNumber="137">FIGURE 2</emphasis>
.
<taxonomicName id="4C16F786437BFFAAFED5F8DAFE550929" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[283,453,1869,1893]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B9625017437BFFAAFED5F8DAFE550929" box="[283,453,1869,1893]" italics="true" pageId="4" pageNumber="137">Bannereus chani</emphasis>
</taxonomicName>
<emphasis id="B9625017437BFFAAFE03F8D9FD8E0929" bold="true" box="[461,542,1869,1893]" pageId="4" pageNumber="137">
<taxonomicNameLabel id="A251ED6C437BFFAAFE03F8D9FD8E0929" box="[461,542,1869,1893]" pageId="4" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left (major) first pereiopod, lateral; (B) same, chela, mesial; (C) same, proximal teeth on cutting edge pollex, mesial; (D) same, ischium, merus, carpus and proximal portion of propodus, mesial; (E) same, detail of spiniform setae on merus, mesial; (F) same, chela fingers, lateral.
</paragraph>
</caption>
<paragraph id="8BA98C05437AFFABFF09FEDFFEB60C2D" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="138">
Fourth pereiopod generally similar to third, much slenderer (
<figureCitation id="132D9080437AFFABFCA0FEDFFC590F29" box="[878,969,331,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3D</figureCitation>
). Ischium bearing 1 or 2 cuspidate seta(e) on ventrolateral surface (
<figureCitation id="132D9080437AFFABFE63FEFBFDB40FC5" box="[429,548,367,393]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3D, F</figureCitation>
). Merus about 3.2 times as long as wide, less swollen, with margins less convex than in third pereiopod, unarmed or armed with 1 small cuspidate seta slightly proximal to mid-length (
<figureCitation id="132D9080437AFFABFAF3FE07FF3A0F9D" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3D, F</figureCitation>
). Carpus half-length of propodus, unarmed (
<figureCitation id="132D9080437AFFABFD67FE23FC950F9D" box="[681,773,439,465]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3D</figureCitation>
). Propodus elongate, with ventral margin armed with row of 11 short spiniform setae, including 1 pair of longer spiniform setae distally, adjacent to dactylar base (
<figureCitation id="132D9080437AFFABFAFBFE4FFA000FB9" box="[1333,1424,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3D</figureCitation>
). Dactylus less stout than that of third pereiopod but otherwise similar, about 0.2 length of propodus, biunguiculate; ventral margin with 4 minute, apparently articulated spinules, distal-most spinule located near secondary unguis (
<figureCitation id="132D9080437AFFABFF6EFDD3FE890C2D" box="[160,281,583,609]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3D, E</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437AFFABFF09FDFFFC570D11" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="138">
Fifth pereiopod shortest and slenderest (
<figureCitation id="132D9080437AFFABFD46FDFFFD720CC9" box="[648,738,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3G</figureCitation>
). Ischium unarmed (
<figureCitation id="132D9080437AFFABFC06FDFFFBB20CC9" box="[968,1058,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3G</figureCitation>
). Merus about three times as long as wide, slightly less than twice as long as ischium, unarmed or armed with 1 small cuspidate seta between 0.3 and 0.4 of merus length (
<figureCitation id="132D9080437AFFABFE4BFD27FE670C81" box="[389,503,691,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3G, I</figureCitation>
). Carpus about 0.6 length of merus (
<figureCitation id="132D9080437AFFABFC57FD27FC640C81" box="[921,1012,691,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3G</figureCitation>
). Propodus with ventromesial margin armed with 9 spiniform setae, including 1 pair of longer spiniform setae distally, adjacent to dactylar base, and 5 6 rows of grooming serrulate setae on distolateral surface; distodorsal margin with or without 1 spiniform seta at articulation with dactylus (
<figureCitation id="132D9080437AFFABFE08FC8BFDD50D75" box="[454,581,799,825]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3G, H</figureCitation>
). Dactylus similar to that of fourth pereiopod in proportions, biunguiculate; ventral margin with 3 minute, probably articulated spinules (
<figureCitation id="132D9080437AFFABFCF0FCD7FC2B0D11" box="[830,955,835,861]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="5" pageNumber="138">Fig. 3G, H</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437AFFABFF09FCF3FEB10A79" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="138">
Female first pleopod with well-developed endopod, overreaching mid-length of exopod, furnished with setae distally (
<figureCitation id="132D9080437AFFABFF34FC1FFEC40DE9" box="[250,340,907,933]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[184,1424,194,1909]" captionTargetId="figure-19@7.[154,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) pleon, third to sixth pleonites, lateral; (B) telson, dorsal; (C) right first pleopod, ventral; (D) right second pleopod, ventral; (E) right uropod, dorsal; (F) same, posterior portion of endopod and exopod showing setal sockets of stiff setae, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4315596" httpUri="https://zenodo.org/record/4315596/files/figure.png" pageId="5" pageNumber="138">Fig. 4C</figureCitation>
). Female second pleopod with short, stubby appendix interna (
<figureCitation id="132D9080437AFFABFBD6FC1FFBE40DE9" box="[1048,1140,907,933]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[184,1424,194,1909]" captionTargetId="figure-19@7.[154,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) pleon, third to sixth pleonites, lateral; (B) telson, dorsal; (C) right first pleopod, ventral; (D) right second pleopod, ventral; (E) right uropod, dorsal; (F) same, posterior portion of endopod and exopod showing setal sockets of stiff setae, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4315596" httpUri="https://zenodo.org/record/4315596/files/figure.png" pageId="5" pageNumber="138">Fig. 4D</figureCitation>
). Uropod with lateral lobe of protopod blunt; exopod broad, truncate distally, with distolateral tooth poorly marked; adjacent spiniform seta well developed, reaching beyond distal margin of exopod; diaeresis not distinct; endopod broadly ovate, about same length as exopod; distal area of both exopod and endopod with fields of stiff setae, in addition to marginal setae (
<figureCitation id="132D9080437AFFABFF6EFB8FFE840A79" box="[160,276,1051,1077]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[184,1424,194,1909]" captionTargetId="figure-19@7.[154,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) pleon, third to sixth pleonites, lateral; (B) telson, dorsal; (C) right first pleopod, ventral; (D) right second pleopod, ventral; (E) right uropod, dorsal; (F) same, posterior portion of endopod and exopod showing setal sockets of stiff setae, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4315596" httpUri="https://zenodo.org/record/4315596/files/figure.png" pageId="5" pageNumber="138">Fig. 4E, F</figureCitation>
).
</paragraph>
<paragraph id="8BA98C05437AFFABFF09FBABFB410A31" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="138">Gill-exopod formula: 5 pleurobranchs (above bases of P15); 0 podobranchs; 1 arthrobranch (at base of Mxp3); 1 mastigobranch = strap-like epipod (on coxa of Mxp3); 0 setobranchs; 3 exopods (on Mxp13).</paragraph>
<paragraph id="8BA98C05437AFFABFF09FB12FE790AED" blockId="5.[151,1437,151,2013]" box="[199,489,1158,1185]" pageId="5" pageNumber="138">
<emphasis id="B9625017437AFFABFF09FB12FEF50AEC" bold="true" box="[199,357,1158,1184]" pageId="5" pageNumber="138">Colour in life</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30CDF8E437AFFABFF09FB3EFC7E0AA6" pageId="5" pageNumber="138" type="distribution">
<paragraph id="8BA98C05437AFFABFF09FB3EFC7E0AA6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="138">
<emphasis id="B9625017437AFFABFF09FB3EFEC90A88" bold="true" box="[199,345,1194,1220]" pageId="5" pageNumber="138">Distribution</emphasis>
. Western Pacific: presently known with certainty only from
<typeStatus id="54AD32A7437AFFABFC3DFB3FFBB30A89" box="[1011,1059,1195,1221]" pageId="5" pageNumber="138">type</typeStatus>
locality off south-eastern
<collectingCountry id="F301CC95437AFFABFA8CFB3FFA080A89" box="[1346,1432,1195,1221]" name="Taiwan" pageId="5" pageNumber="138">Taiwan</collectingCountry>
; possibly also in the Coral Sea off eastern
<collectingCountry id="F301CC95437AFFABFDA8FB5BFD5D0AA5" box="[614,717,1231,1257]" name="Australia" pageId="5" pageNumber="138">Australia</collectingCountry>
(
<bibRefCitation id="EF87F1F4437AFFABFD15FB5BFCF20AA5" author="Bruce, A. J." box="[731,866,1231,1257]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
; see below).
</paragraph>
</subSubSection>
<subSubSection id="C30CDF8E437AFFABFF09FB67FBC50B19" pageId="5" pageNumber="138" type="biology_ecology">
<paragraph id="8BA98C05437AFFABFF09FB67FBC50B19" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="138">
<emphasis id="B9625017437AFFABFF09FB67FEB60B41" bold="true" box="[199,294,1267,1293]" pageId="5" pageNumber="138">Ecology</emphasis>
. The
<typeStatus id="54AD32A7437AFFABFEA8FB67FE580B41" box="[358,456,1267,1293]" pageId="5" pageNumber="138" type="holotype">holotype</typeStatus>
was dredged from a depth of
<quantity id="4CEE21E0437AFFABFCD6FB66FC090B41" box="[792,921,1266,1293]" metricMagnitude="2" metricUnit="m" metricValue="3.285" metricValueMax="3.56" metricValueMin="3.01" pageId="5" pageNumber="138" unit="m" value="328.5" valueMax="356.0" valueMin="301.0">301356 m</quantity>
, without any further field notes, e.g. mentioning a possible association with hexactinellid sponges. The male from the Coral Sea reported by
<bibRefCitation id="EF87F1F4437AFFABFB04FA83FACE0B7D" author="Bruce, A. J." box="[1226,1374,1303,1329]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce (1988)</bibRefCitation>
as
<taxonomicName id="4C16F786437AFFABFA4AFA83FE940B19" authorityName="Bruce" authorityYear="1988" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B9625017437AFFABFA4AFA83FE940B19" italics="true" pageId="5" pageNumber="138">B. anomalus</emphasis>
</taxonomicName>
was found inside a hexactinellid sponge brought up from a depth of
<specimenCount id="9D10478C437AFFABFBC9FAAFFBC50B19" box="[1031,1109,1339,1365]" pageId="5" pageNumber="138" type="generic">
<quantity id="4CEE21E0437AFFABFBC9FAAFFBC10B19" box="[1031,1105,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="138" unit="m" value="360.0">360 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C30CDF8E437AFFABFF09FACBFA910BD1" pageId="5" pageNumber="138" type="etymology">
<paragraph id="8BA98C05437AFFABFF09FACBFA910BD1" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="138">
<emphasis id="B9625017437AFFABFF09FACBFED80B35" bold="true" box="[199,328,1375,1401]" pageId="5" pageNumber="138">Etymology</emphasis>
. The new species is named after our colleague Dr. Tin-Yam Chan (National
<collectingCountry id="F301CC95437AFFABFB50FACBFB600B35" box="[1182,1264,1375,1401]" name="Taiwan" pageId="5" pageNumber="138">Taiwan</collectingCountry>
Ocean University) for his immense contribution to decapod taxonomy, especially deep-water shrimps and lobsters.
</paragraph>
</subSubSection>
<subSubSection id="C30CDF8E437AFFA6FF09FA33FB1B0B7E" lastPageId="8" lastPageNumber="141" pageId="5" pageNumber="138" type="discussion">
<paragraph id="8BA98C05437AFFA6FF09FA33FF690F52" blockId="5.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,1330]" lastPageId="8" lastPageNumber="141" pageId="5" pageNumber="138">
<emphasis id="B9625017437AFFABFF09FA33FEA40B8D" bold="true" box="[199,308,1447,1473]" pageId="5" pageNumber="138">Remarks</emphasis>
.
<bibRefCitation id="EF87F1F4437AFFABFE8BFA33FE4C0B8D" author="Bruce, A. J." box="[325,476,1447,1473]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce (1988)</bibRefCitation>
noted several important differences between the female
<typeStatus id="54AD32A7437AFFABFBA0FA33FB400B8D" box="[1134,1232,1447,1473]" pageId="5" pageNumber="138" type="holotype">holotype</typeStatus>
and the non-type male of
<taxonomicName id="4C16F786437AFFABFF3DFA5FFE4D0BA9" authorityName="Bruce" authorityYear="1988" box="[243,477,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B9625017437AFFABFF3DFA5FFE4D0BA9" box="[243,477,1483,1509]" italics="true" pageId="5" pageNumber="138">Bannereus anomalus</emphasis>
</taxonomicName>
, which was the reason why the latter specimen was not given the
<typeStatus id="54AD32A7437AFFABFB71FA5FFA8F0BA9" box="[1215,1311,1483,1509]" pageId="5" pageNumber="138" type="paratype">paratype</typeStatus>
status. According to the author, “these differences appear sufficient to suggest that the male may represent a species distinct from the female, but on account of its damaged and incomplete nature, this cannot be stated with certainty”. Indeed, the differences between the female
<typeStatus id="54AD32A7437AFFABFDD0F9A3FD10081D" box="[542,640,1591,1617]" pageId="5" pageNumber="138" type="holotype">holotype</typeStatus>
and the non-type male appear to be rather significant. These differences include (i) the shape of the frontal margins of the carapace, rounded in the female
<emphasis id="B9625017437AFFABFBE5F9CFFBDA0838" box="[1067,1098,1627,1652]" italics="true" pageId="5" pageNumber="138">vs.</emphasis>
with a blunt rostral projection in the male (
<bibRefCitation id="EF87F1F4437AFFABFED1F9EBFE3508D5" author="Bruce, A. J." box="[287,421,1663,1689]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 2B, E); (ii) the absence of a strap-like epipod on the third maxilliped coxa in the female (not illustrated)
<emphasis id="B9625017437AFFABFE87F937FEF008F0" box="[329,352,1699,1724]" italics="true" pageId="5" pageNumber="138">vs</emphasis>
. its presence in the male (
<bibRefCitation id="EF87F1F4437AFFABFDB2F937FC9208F1" author="Bruce, A. J." box="[636,770,1699,1725]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 3J); (iii) the minor chela palm with bumps in the female
<emphasis id="B9625017437AFFABFF59F953FF3E08AC" box="[151,174,1735,1760]" italics="true" pageId="5" pageNumber="138">vs</emphasis>
. smooth in the single male chela (
<bibRefCitation id="EF87F1F4437AFFABFDFFF953FD2908AD" author="Bruce, A. J." box="[561,697,1735,1761]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 4G, H, mind a series of errors in the figure legend and in the text, e.g. confusion between the female and male minor cheliped or the cheliped accidentally called “minor second pereiopod”); (iv) the cheliped meri unarmed in the female (
<bibRefCitation id="EF87F1F4437AFFABFCE2F89BFC230965" author="Bruce, A. J." box="[812,947,1807,1833]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 4G)
<emphasis id="B9625017437AFFABFBD3F89BFBA40964" box="[1053,1076,1807,1832]" italics="true" pageId="5" pageNumber="138">vs</emphasis>
. armed with about 10 spiniform setae on the mesioventral margin of the single male cheliped (described as “ventrally spinose merus”, but not illustrated); (v) the fifth pereiopod merus and ischium unarmed in the female (not illustrated, but generally similar to those of the third pereiopod,
<bibRefCitation id="EF87F1F4437AFFABFE33F8EFFD1309D9" author="Bruce, A. J." box="[509,643,1915,1941]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 5E)
<emphasis id="B9625017437AFFABFD23F8EFFC9B09D8" box="[749,779,1915,1940]" italics="true" pageId="5" pageNumber="138">vs.</emphasis>
armed with ventral spiniform setae (“spines”) in the male (
<bibRefCitation id="EF87F1F4437AFFABFF51F80BFEB709F5" author="Bruce, A. J." box="[159,295,1951,1977]" pageId="5" pageNumber="138" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 5G, but see below); and (vi) the dactylus of the ambulatory (third to fifth) pereiopods stout, with single terminal unguis, with corpus armed with five to six small teeth on the ventral margin and one to two dorsal spinules at the base of the terminal unguis (
<bibRefCitation id="EF87F1F44377FFA6FDB1FF03FC970EFD" author="Bruce, A. J." box="[639,775,151,177]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 6A, B), in marked contrast to the noticeably slenderer dactylus of the male, with its terminal unguis bifurcated (with a stout ventral tooth), the corpus armed with only two teeth on the ventral margin and without dorsal spinules at the base of the terminal unguis (
<bibRefCitation id="EF87F1F44377FFA6FB1BFF4BFACF0EB5" author="Bruce, A. J." box="[1237,1375,223,249]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: figs. 5H, 6C).
</paragraph>
<caption id="DF69DC8D4379FFA8FF59F8C7FEBF09B7" ID-DOI="http://doi.org/10.5281/zenodo.4315594" ID-Zenodo-Dep="4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="6" pageNumber="139" startId="6.[151,250,1875,1899]" targetBox="[218,1363,194,1846]" targetPageId="6">
<paragraph id="8BA98C054379FFA8FF59F8C7FEBF09B7" blockId="6.[151,1437,1875,2043]" pageId="6" pageNumber="139">
<emphasis id="B96250174379FFA8FF59F8C7FE9E0927" bold="true" box="[151,270,1875,1899]" pageId="6" pageNumber="139">FIGURE 3</emphasis>
.
<taxonomicName id="4C16F7864379FFA8FED5F8C0FE550927" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[283,453,1875,1899]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174379FFA8FED5F8C0FE550927" box="[283,453,1875,1899]" italics="true" pageId="6" pageNumber="139">Bannereus chani</emphasis>
</taxonomicName>
<emphasis id="B96250174379FFA8FE02F8C7FD8D0927" bold="true" box="[460,541,1875,1899]" pageId="6" pageNumber="139">
<taxonomicNameLabel id="A251ED6C4379FFA8FE02F8C7FD8D0927" box="[460,541,1875,1899]" pageId="6" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial.
</paragraph>
</caption>
<caption id="DF69DC8D4378FFA9FF59F803FB0E09B4" ID-DOI="http://doi.org/10.5281/zenodo.4315596" ID-Zenodo-Dep="4315596" httpUri="https://zenodo.org/record/4315596/files/figure.png" pageId="7" pageNumber="140" startId="7.[151,250,1943,1967]" targetBox="[184,1424,194,1909]" targetPageId="7">
<paragraph id="8BA98C054378FFA9FF59F803FB0E09B4" blockId="7.[151,1436,1943,2040]" pageId="7" pageNumber="140">
<emphasis id="B96250174378FFA9FF59F803FE9F09E3" bold="true" box="[151,271,1943,1967]" pageId="7" pageNumber="140">FIGURE 4</emphasis>
.
<taxonomicName id="4C16F7864378FFA9FED3F80CFE5909FC" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[285,457,1944,1968]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="140" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174378FFA9FED3F80CFE5909FC" box="[285,457,1944,1968]" italics="true" pageId="7" pageNumber="140">Bannereus chani</emphasis>
</taxonomicName>
<emphasis id="B96250174378FFA9FE1CF80CFDB409FC" bold="true" box="[466,548,1944,1968]" pageId="7" pageNumber="140">
<taxonomicNameLabel id="A251ED6C4378FFA9FE1CF80CFDB409FC" box="[466,548,1944,1968]" pageId="7" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) pleon, third to sixth pleonites, lateral; (B) telson, dorsal; (C) right first pleopod, ventral; (D) right second pleopod, ventral; (E) right uropod, dorsal; (F) same, posterior portion of endopod and exopod showing setal sockets of stiff setae, dorsal.
</paragraph>
</caption>
<paragraph id="8BA98C054377FFA6FF09FEB3FB610C72" blockId="8.[151,1437,151,1330]" pageId="8" pageNumber="141">
The above-listed differences, especially those involving the chelipeds, ambulatory pereiopods and epipod, are significant, strongly supporting
<bibRefCitation id="EF87F1F44377FFA6FE30FEDFFD360F29" author="Bruce, A. J." box="[510,678,331,357]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruces (1988)</bibRefCitation>
suspicions that the male does not belong to the same species as the female
<typeStatus id="54AD32A74377FFA6FF22FEFBFEDE0FC5" box="[236,334,367,393]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C16F7864377FFA6FEB8FEFBFD940FC5" authorityName="Bruce" authorityYear="1988" box="[374,516,367,393]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FEB8FEFBFD940FC5" box="[374,516,367,393]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
. The main problem of
<bibRefCitation id="EF87F1F44377FFA6FCC2FEFBFC250FC5" author="Bruce, A. J." box="[780,949,367,393]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruces (1988)</bibRefCitation>
description lies in the fact that the accompanying illustrations were made from both specimens (see synonymy above). Therefore, all comparisons between
<taxonomicName id="4C16F7864377FFA6FF59FE23FF660F9D" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[151,246,439,465]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174377FFA6FF59FE23FF660F9D" box="[151,246,439,465]" italics="true" pageId="8" pageNumber="141">B. chani</emphasis>
</taxonomicName>
<emphasis id="B96250174377FFA6FF30FE23FECA0F9D" bold="true" box="[254,346,439,465]" pageId="8" pageNumber="141">
<taxonomicNameLabel id="A251ED6C4377FFA6FF30FE23FECA0F9D" box="[254,346,439,465]" pageId="8" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C16F7864377FFA6FE5DFE23FDB00F9D" authorityName="Bruce" authorityYear="1988" box="[403,544,439,465]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FE5DFE23FDB00F9D" box="[403,544,439,465]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
refer only to the female
<typeStatus id="54AD32A74377FFA6FCF0FE23FC300F9D" box="[830,928,439,465]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C16F7864377FFA6FC06FE23FBC60F9D" authorityName="Bruce" authorityYear="1988" box="[968,1110,439,465]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FC06FE23FBC60F9D" box="[968,1110,439,465]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF87F1F44377FFA6FBA8FE23FB7F0F9D" author="Bruce, A. J." box="[1126,1263,439,465]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: figs. 1, 2AC, 4AF, 5A, E, F, 6A, B) which was examined by the senior author in 2001, as well as a series of specimens from
<collectingCountry id="F301CC954377FFA6FAA9FE4FFE990C55" name="New Caledonia" pageId="8" pageNumber="141">New Caledonia</collectingCountry>
deposited in the MNNH. The latter include a complete female specimen collected relatively recently (in 2016), which corresponds particularly well to the
<typeStatus id="54AD32A74377FFA6FD08FDB7FCB80C71" box="[710,808,547,573]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
(see under comparative material above).
</paragraph>
<paragraph id="8BA98C054377FFA6FF09FDD3FC620A79" blockId="8.[151,1437,151,1330]" pageId="8" pageNumber="141">
The differences between
<taxonomicName id="4C16F7864377FFA6FE2DFDD3FDD10C2D" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[483,577,583,609]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174377FFA6FE2DFDD3FDD10C2D" box="[483,577,583,609]" italics="true" pageId="8" pageNumber="141">B. chani</emphasis>
</taxonomicName>
<emphasis id="B96250174377FFA6FD87FDD3FD340C2D" bold="true" box="[585,676,583,609]" pageId="8" pageNumber="141">
<taxonomicNameLabel id="A251ED6C4377FFA6FD87FDD3FD340C2D" box="[585,676,583,609]" pageId="8" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C16F7864377FFA6FD12FDD3FCF90C2D" authorityName="Bruce" authorityYear="1988" box="[732,873,583,609]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FD12FDD3FCF90C2D" box="[732,873,583,609]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
are essentially the same as between Bruces male specimen and the
<typeStatus id="54AD32A74377FFA6FEA5FDFFFE5D0CC9" box="[363,461,619,645]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
of the type species: the presence/absence of a blunt rostral projection [character (i) above]; the presence/absence of a strap-like epipod on the third maxilliped coxa [character (ii)]; the merus of at least one of the chelipeds armed with a row of spiniform setae or unarmed [character (iv)]; and obvious differences in the proportions of the dactyli of the third to fifth pereiopods [character (vi)]. The afore-mentioned character (iii) can be added to this series as well, because the palm of the major chela is smooth, i.e. without bumps, in
<taxonomicName id="4C16F7864377FFA6FB11FD6FFAAC0D59" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[1247,1340,763,789]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174377FFA6FB11FD6FFAAC0D59" box="[1247,1340,763,789]" italics="true" pageId="8" pageNumber="141">B. chani</emphasis>
</taxonomicName>
<emphasis id="B96250174377FFA6FA8DFD6FFA0C0D59" bold="true" box="[1347,1436,763,789]" pageId="8" pageNumber="141">
<taxonomicNameLabel id="A251ED6C4377FFA6FA8DFD6FFA0C0D59" box="[1347,1436,763,789]" pageId="8" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B96250174377FFA6FF59FC8BFF260D74" box="[151,182,799,824]" italics="true" pageId="8" pageNumber="141">vs.</emphasis>
with bumps in the
<typeStatus id="54AD32A74377FFA6FE5FFC8BFE630D75" box="[401,499,799,825]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C16F7864377FFA6FDD4FC8BFD370D75" authorityName="Bruce" authorityYear="1988" box="[538,679,799,825]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FDD4FC8BFD370D75" box="[538,679,799,825]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
; the only difference is that
<bibRefCitation id="EF87F1F44377FFA6FC11FC8BFBE40D75" author="Bruce, A. J." box="[991,1140,799,825]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce (1988)</bibRefCitation>
was comparing the minor chelae for this same feature. As to the character (v),
<bibRefCitation id="EF87F1F44377FFA6FD1BFCD7FCEB0D11" author="Bruce, A. J." box="[725,891,835,861]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruces (1988)</bibRefCitation>
tentative assignment of the detached walking leg illustrated in his figure 5G, H to the fifth pereiopod is not correct. In almost all alpheid shrimps, the fifth pereiopod has a grooming brush on the propodus, as illustrated for
<taxonomicName id="4C16F7864377FFA6FCD5FC1FFCE90DE9" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[795,889,907,933]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174377FFA6FCD5FC1FFCE90DE9" box="[795,889,907,933]" italics="true" pageId="8" pageNumber="141">B. chani</emphasis>
</taxonomicName>
<emphasis id="B96250174377FFA6FC4FFC1FFC4C0DE9" bold="true" box="[897,988,907,933]" pageId="8" pageNumber="141">
<taxonomicNameLabel id="A251ED6C4377FFA6FC4FFC1FFC4C0DE9" box="[897,988,907,933]" pageId="8" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132D90804377FFA6FC22FC1FFBD70DE9" box="[1004,1095,907,933]" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="8" pageNumber="141">Fig. 3G</figureCitation>
). The walking leg of the male specimen in
<bibRefCitation id="EF87F1F44377FFA6FEEDFC3BFE270D85" author="Bruce, A. J." box="[291,439,943,969]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce (1988)</bibRefCitation>
appears to be the fourth pereiopod, based on its general proportions and armature. Nevertheless, the
<typeStatus id="54AD32A74377FFA6FED4FC47FEEC0DA1" box="[282,380,979,1005]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C16F7864377FFA6FE6EFC47FE6D0DA1" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[416,509,979,1005]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174377FFA6FE6EFC47FE6D0DA1" box="[416,509,979,1005]" italics="true" pageId="8" pageNumber="141">B. chani</emphasis>
</taxonomicName>
<emphasis id="B96250174377FFA6FDCAFC47FDCD0DA1" bold="true" box="[516,605,979,1005]" pageId="8" pageNumber="141">
<taxonomicNameLabel id="A251ED6C4377FFA6FDCAFC47FDCD0DA1" box="[516,605,979,1005]" pageId="8" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
clearly differs from the
<typeStatus id="54AD32A74377FFA6FCA5FC47FC5D0DA1" box="[875,973,979,1005]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C16F7864377FFA6FC3CFC47FBEE0DA1" authorityName="Bruce" authorityYear="1988" box="[1010,1150,979,1005]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FC3CFC47FBEE0DA1" box="[1010,1150,979,1005]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
in the ischium and merus of the third pereiopod being armed with spiniform setae (one or two on the ischium and one on the merus,
<figureCitation id="132D90804377FFA6FA8EFC63FF3D0A79" captionStart="FIGURE 3" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[218,1363,194,1846]" captionTargetId="figure-21@6.[210,1377,181,1851]" captionTargetPageId="6" captionText="FIGURE 3. Bannereus chani sp. nov., holotype, ovigerous female (cl 6.3 mm), MNHN-IV-2017-11749: (A) left third pereiopod, lateral; (B) same, dactylus, lateral; (C) right third pereiopod, distal portion of basis to proximal portion of carpus, mesioventral; (D) right fourth pereiopod, lateral; (E) same, dactylus, lateral; (F) left fourth pereiopod, distal basis to proximal carpus, mesioventral; (G) right fifth pereiopod, lateral; (H) same, dactylus, lateral; (I) left fifth pereiopod, distal ischium to proximal carpus, mesial." figureDoi="http://doi.org/10.5281/zenodo.4315594" httpUri="https://zenodo.org/record/4315594/files/figure.png" pageId="8" pageNumber="141">Fig. 3A, C</figureCitation>
). In
<taxonomicName id="4C16F7864377FFA6FF2EFB8FFEFC0A79" authorityName="Bruce" authorityYear="1988" box="[224,364,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FF2EFB8FFEFC0A79" box="[224,364,1051,1077]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
, both of these articles are unarmed (
<bibRefCitation id="EF87F1F44377FFA6FCCDFB8FFC1A0A79" author="Bruce, A. J." box="[771,906,1051,1077]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
: fig. 5E).
</paragraph>
<paragraph id="8BA98C054377FFA6FF09FBABFB1B0B7E" blockId="8.[151,1437,151,1330]" pageId="8" pageNumber="141">
In summary, multiple lines of evidence suggest that
<bibRefCitation id="EF87F1F44377FFA6FCC9FBABFC3D0A15" author="Bruce, A. J." box="[775,941,1087,1113]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruces (1988)</bibRefCitation>
non-type male specimen may indeed belong to the same taxon as the species described herein as
<taxonomicName id="4C16F7864377FFA6FD14FBF7FCA60A31" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[730,822,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174377FFA6FD14FBF7FCA60A31" box="[730,822,1123,1149]" italics="true" pageId="8" pageNumber="141">B. chani</emphasis>
</taxonomicName>
<emphasis id="B96250174377FFA6FCF3FBF7FC060A31" bold="true" box="[829,918,1123,1149]" pageId="8" pageNumber="141">
<taxonomicNameLabel id="A251ED6C4377FFA6FCF3FBF7FC060A31" box="[829,918,1123,1149]" pageId="8" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which differs from
<taxonomicName id="4C16F7864377FFA6FBB7FBF7FA950A31" authorityName="Bruce" authorityYear="1988" box="[1145,1285,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FBB7FBF7FA950A31" box="[1145,1285,1123,1149]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
in at least six morphological details. If this hypothesis is eventually confirmed, it would imply that
<taxonomicName id="4C16F7864377FFA6FB95FB13FB280AED" authority="Anker &amp; Pachelle, 2020" authorityName="Anker &amp; Pachelle" authorityYear="2020" box="[1115,1208,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="B96250174377FFA6FB95FB13FB280AED" box="[1115,1208,1159,1185]" italics="true" pageId="8" pageNumber="141">B. chani</emphasis>
</taxonomicName>
<emphasis id="B96250174377FFA6FB0EFB13FA8A0AED" bold="true" box="[1216,1306,1159,1185]" pageId="8" pageNumber="141">
<taxonomicNameLabel id="A251ED6C4377FFA6FB0EFB13FA8A0AED" box="[1216,1306,1159,1185]" pageId="8" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a widely distributed western Pacific deep-water alpheid shrimp, ranging from
<collectingCountry id="F301CC954377FFA6FC68FB3FFC680A89" box="[934,1016,1195,1221]" name="Taiwan" pageId="8" pageNumber="141">Taiwan</collectingCountry>
to north-eastern
<collectingCountry id="F301CC954377FFA6FB74FB3FFAB40A89" box="[1210,1316,1195,1221]" name="Australia" pageId="8" pageNumber="141">Australia</collectingCountry>
. However, the general scarcity of the material of
<taxonomicName id="4C16F7864377FFA6FDF9FB5BFD3E0AA4" authorityName="Bruce" authorityYear="1988" box="[567,686,1231,1256]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B96250174377FFA6FDF9FB5BFD3E0AA4" box="[567,686,1231,1256]" italics="true" pageId="8" pageNumber="141">Bannereus</emphasis>
</taxonomicName>
precludes us from making more firm conclusions at this stage. With regard to
<taxonomicName id="4C16F7864377FFA6FECAFB67FE000B41" authorityName="Bruce" authorityYear="1988" box="[260,400,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Bannereus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B96250174377FFA6FECAFB67FE000B41" box="[260,400,1267,1293]" italics="true" pageId="8" pageNumber="141">B. anomalus</emphasis>
</taxonomicName>
, this species is presently known only from a few localities in the Coral Sea off
<collectingRegion id="49D242E74377FFA6FADFFB67FA080B41" box="[1297,1432,1267,1293]" country="Australia" name="Queensland" pageId="8" pageNumber="141">Queensland</collectingRegion>
,
<collectingCountry id="F301CC954377FFA6FF59FA83FF6E0B7D" box="[151,254,1303,1329]" name="Australia" pageId="8" pageNumber="141">Australia</collectingCountry>
(
<bibRefCitation id="EF87F1F44377FFA6FEC2FA83FE030B7D" author="Bruce, A. J." box="[268,403,1303,1329]" pageId="8" pageNumber="141" pagination="139 - 149" refId="ref5492" refString="Bruce, A. J. (1988) Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science, 42, 139 - 149." type="journal article" year="1988">Bruce 1988</bibRefCitation>
), and off
<collectingCountry id="F301CC954377FFA6FE30FA83FD3C0B7D" box="[510,684,1303,1329]" name="New Caledonia" pageId="8" pageNumber="141">New Caledonia</collectingCountry>
(present study, see Comparative material).
</paragraph>
</subSubSection>
</treatment>
</document>