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<document id="9D06D7D1DFF33E3100A4235529E6EC0E" ID-DOI="10.5852/ejt.2024.958.2665" ID-ISSN="2118-9773" ID-Zenodo-Dep="13837028" ID-ZooBank="425BCF68-144B-4861-BCED-CB570D8246D1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1727252768769" checkinUser="plazi" docAuthor="Constant, Jérôme &amp; Pham, Hong Thai" docDate="2024" docId="03B887FAFFA2FFFC2377FBF75DFE12FB" docLanguage="en" docName="EJT.2024.958.114-150.pdf" docOrigin="European Journal of Taxonomy 958" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2665/12349" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zanna limbourgi Constant 2024, sp. nov." docType="treatment" docUuid="CDC2CFD0-B1E8-4DA2-8114-5F4BD37D8782" docUuidSource="ZooBank" docVersion="3" lastPageNumber="32" masterDocId="FF81FF82FFBAFFE3213DFFAE5E6A1776" masterDocTitle="Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)" masterLastPageNumber="150" masterPageNumber="114" pageNumber="25" updateTime="1727378517280" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="96F35FE0D42628B5673472CBBA426B32">Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:name id="878FCE30D33AA4F5129855359D4E4917" type="personal">
<mods:role id="1A693D53EAC94EDF827ED02BD98B56B8">
<mods:roleTerm id="772CF82FF280C0AAA123CDCF1ECFC5BE">Author</mods:roleTerm>
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<mods:namePart id="2379F8DE86B9E8D24DD8655BC6D96320">Constant, Jérôme</mods:namePart>
<mods:nameIdentifier id="1A32670A75D94229BACBCB64DCC98964" type="LSID">6E6072A1-9415-4C8D-8E60-2504444DB290</mods:nameIdentifier>
<mods:affiliation id="D9FD90D151BAEEFEADC183D384E3695E">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier Street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<mods:nameIdentifier id="9D7E126B971C409F2D6B626832924C6E" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:name id="F0297BEE773461A7E951F5CF18F93431" type="personal">
<mods:role id="9AB287E2F1BA6EDAB681B08EC88BE524">
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<mods:namePart id="EF56851C04BBC38F21CC9B9220BB1C88">Pham, Hong Thai</mods:namePart>
<mods:nameIdentifier id="8C281B2ACB42CAE6D916C40570ADCE96" type="LSID">E34CB863-7E3B-4E8F-8738-B41C07D9F5F9</mods:nameIdentifier>
<mods:affiliation id="DD3BDC1D699AFEB081630528C28F9D7A">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.</mods:affiliation>
<mods:nameIdentifier id="F9E159828AFB428A91806761D0DB9B93" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="10D947C5E320950A7AB913CA403129BA">European Journal of Taxonomy</mods:title>
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<mods:part id="2097A374BC6A1360971D97089FE715D3">
<mods:date id="C3F21EF00CA0F0440FF7119F38BDEBEA">2024</mods:date>
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<mods:number id="93F0DD9ACDFD130DE7747D7D16E2ED65">2024-09-25</mods:number>
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<mods:detail id="EE77832354EFAAD68CF68C00787EC11C" type="volume">
<mods:number id="9F920713FB4E78C70C3072040B7A8A13">958</mods:number>
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<mods:start id="7E59D7AA8F8EA8F143482C3357C5AC45">114</mods:start>
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<mods:url id="35424FED93D92468125967BA5D9F227D">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2665/12349</mods:url>
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<mods:identifier id="35EA9EAA1C8ED4120A9642E4F10A2ED0" type="DOI">10.5852/ejt.2024.958.2665</mods:identifier>
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<treatment id="03B887FAFFA2FFFC2377FBF75DFE12FB" LSID="urn:lsid:zoobank.org:act:CDC2CFD0-B1E8-4DA2-8114-5F4BD37D8782" httpUri="http://treatment.plazi.org/id/03B887FAFFA2FFFC2377FBF75DFE12FB" lastPageId="31" lastPageNumber="32" pageId="24" pageNumber="25">
<subSubSection id="C30B6567FFA2FFFB2377FBF75D801305" box="[586,1002,1112,1139]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BAE36ECFFA2FFFB2377FBF75D801305" blockId="24.[358,1229,1112,1210]" box="[586,1002,1112,1139]" pageId="24" pageNumber="25">
<heading id="D0E68180FFA2FFFB2377FBF75D801305" box="[586,1002,1112,1139]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="4C114D6FFFA2FFFB2377FBF75DE21305" authority="Constant" authorityName="Constant" authorityYear="2024" box="[586,904,1112,1139]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB2377FBF75D721305" bold="true" box="[586,792,1112,1139]" italics="true" pageId="24" pageNumber="25">Zanna limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA2FFFB22B2FBF75D801305" box="[911,1002,1113,1139]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA2FFFB205BFBD25D8A13CF" pageId="24" pageNumber="25" type="description">
<paragraph id="8BAE36ECFFA2FFFB205BFBD25AA713E1" blockId="24.[358,1229,1112,1210]" box="[358,1229,1148,1175]" pageId="24" pageNumber="25">
<uri id="FF803AEEFFA2FFFB205BFBD25AA713E1" box="[358,1229,1148,1175]" pageId="24" pageNumber="25">
urn:lsid:zoobank.org:act:
<uuid id="FFB70C39FFA2FFFB23B5FBD25AA713E1" box="[648,1229,1148,1175]" pageId="24" pageNumber="25">CDC2CFD0-B1E8-4DA2-8114-5F4BD37D8782</uuid>
</uri>
</paragraph>
<paragraph id="8BAE36ECFFA2FFFB2369FB315D8A13CF" blockId="24.[358,1229,1112,1210]" box="[596,992,1183,1210]" pageId="24" pageNumber="25">
<figureCitation id="132A2A69FFA2FFFB2369FB315CF413CC" box="[596,670,1183,1210]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="24" pageNumber="25">Figs 5</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB2396FB315C9D13CC" box="[683,759,1183,1210]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="25.[189,230,1925,1951]" captionStartId-1="26.[197,236,1958,1984]" captionStartId-2="28.[185,226,1960,1986]" captionTargetBox-0="[240,1354,270,1868]" captionTargetBox-1="[230,1373,285,1917]" captionTargetBox-2="[213,1370,261,1921]" captionTargetId-0="figure-46@25.[230,1354,265,1868]" captionTargetId-1="figure-62@26.[226,1383,268,1920]" captionTargetId-2="figure-36@28.[213,1370,258,1921]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="28" captionText-0="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." captionText-1="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." captionText-2="Fig. 17. Zanna limbourgi Constant sp. nov., holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018. A. Habitus, lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi-0="http://doi.org/10.5281/zenodo.13837073" figureDoi-1="http://doi.org/10.5281/zenodo.13837076" figureDoi-2="http://doi.org/10.5281/zenodo.13837078" httpUri-0="https://zenodo.org/record/13837073/files/figure.png" httpUri-1="https://zenodo.org/record/13837076/files/figure.png" httpUri-2="https://zenodo.org/record/13837078/files/figure.png" pageId="24" pageNumber="25">15–17</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB223EFB315D1613CC" box="[771,892,1183,1210]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. A–D. Prothotax and basal half of head, lateral view. E–H. Head and thorax, dorsal view. I–L. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="24" pageNumber="25">18D, H, L</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB22B4FB315D8A13CF" box="[905,992,1183,1209]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. A–B. Z. bidoupana sp. nov. C–D. Z. chartieri Constant sp. nov. E–F. Z. kusamae sp. nov. G–H. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="24" pageNumber="25">19G–H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA2FFFB2180FB4E5D2F11B9" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BAE36ECFFA2FFFB2180FB4E5F5C138C" blockId="24.[189,310,1248,1274]" box="[189,310,1248,1274]" pageId="24" pageNumber="25">
<heading id="D0E68180FFA2FFFB2180FB4E5F5C138C" bold="true" box="[189,310,1248,1274]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B965EAFEFFA2FFFB2180FB4E5F5C138C" bold="true" box="[189,310,1248,1274]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA2FFFB2180FAA15D2F11B9" blockId="24.[189,1399,1295,1744]" pageId="24" pageNumber="25">
The species is closest to
<taxonomicName id="4C114D6FFFA2FFFB20CCFABE5CAE125F" authorityName="Constant &amp; Pham" authorityYear="2024" box="[497,708,1295,1321]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB20CCFABE5CAE125F" box="[497,708,1295,1321]" italics="true" pageId="24" pageNumber="25">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA2FFFB23F2FAA15D5A125F" box="[719,816,1295,1321]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C114D6FFFA2FFFB227DFABE5A56125F" authority="Constant" authorityName="Constant" authorityYear="2024" box="[832,1084,1295,1321]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB227DFABE5DA3125F" box="[832,969,1295,1321]" italics="true" pageId="24" pageNumber="25">Z. chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA2FFFB257AFAA15AC2125F" box="[1095,1192,1295,1321]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C114D6FFFA2FFFB25D0FABE5B1D125F" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1261,1399,1295,1321]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB25D0FABE5B1D125F" box="[1261,1399,1295,1321]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA2FFFB2180FA9D5F76123B" box="[189,284,1331,1357]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can be separated by the following characters: basal portion of head (
<figureCitation id="132A2A69FFA2FFFB243BFA9C5E9A1219" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">Fig. 16B, E–F</figureCitation>
) covered in moderately large, rather well-spaced, black pitting with ‘background’ of dense, very small black pitting (dense, irregular, rather large, coarse, sometimes coalescent black pitting in
<taxonomicName id="4C114D6FFFA2FFFB2180FA335F3312C0" authorityName="Constant &amp; Pham" authorityYear="2024" box="[189,345,1436,1462]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFA2FFFB2180FA335F3312C0" box="[189,345,1436,1462]" italics="true" pageId="24" pageNumber="25">Z. bidoupana</emphasis>
</taxonomicName>
–
<figureCitation id="132A2A69FFA2FFFB2045FA325C7E12C0" box="[376,532,1436,1462]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="24" pageNumber="25">Fig. 3B, E–F</figureCitation>
; moderately large, rather well-spaced, black pitting without ‘background’ of dense, very small black pitting in
<taxonomicName id="4C114D6FFFA2FFFB2342FA6E5D6212AF" authorityName="Constant &amp; Pham" authorityYear="2024" box="[639,776,1471,1497]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFA2FFFB2342FA6E5D6212AF" box="[639,776,1471,1497]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
–
<figureCitation id="132A2A69FFA2FFFB2216FA115DB712AF" box="[811,989,1471,1497]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="24" pageNumber="25">Fig. 12B, E–F</figureCitation>
); anal tube of male rather high in lateral view (
<figureCitation id="132A2A69FFA2FFFB2069FA4C5FAA128A" box="[340,448,1506,1532]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">Fig. 16A</figureCitation>
), about 1.65 times as long as high, with ventral angle in basal half (rather flattened dorsoventrally, 1.95–2.14 times as long as high in lateral view, with ventral angle at midlength in
<taxonomicName id="4C114D6FFFA2FFFB2180F9875F291134" authorityName="Constant" authorityYear="2024" box="[189,323,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFA2FFFB2180F9875F291134" box="[189,323,1576,1602]" italics="true" pageId="24" pageNumber="25">Z. chartieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFA2FFFB2040F9875C6E1134" authorityName="Constant &amp; Pham" authorityYear="2024" box="[381,516,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFA2FFFB2040F9875C6E1134" box="[381,516,1576,1602]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
–
<figureCitation id="132A2A69FFA2FFFB231CF9865CE81135" box="[545,642,1576,1603]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="24" pageNumber="25">Figs 7A</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB23B3F9875CAF1135" box="[654,709,1577,1603]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">13A</figureCitation>
); pygofer of male in lateral view (
<figureCitation id="132A2A69FFA2FFFB256AF9865AA81134" box="[1111,1218,1576,1602]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">Fig. 16A</figureCitation>
) with posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle (posterior margin projecting into a large posterior lobe angularly rounded in midheight in
<taxonomicName id="4C114D6FFFA2FFFB25E6F9DE5B0B11FF" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1243,1377,1647,1673]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFA2FFFB25E6F9DE5B0B11FF" box="[1243,1377,1647,1673]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
–
<figureCitation id="132A2A69FFA2FFFB2180F93C5F4211DA" box="[189,296,1682,1708]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">Fig. 13A</figureCitation>
; posterior margin moderately sinuate, forming a large, broadly rounded posterior lobe in dorsal half in
<taxonomicName id="4C114D6FFFA2FFFB202DF9185FC611B9" authorityName="Constant &amp; Pham" authorityYear="2024" box="[272,428,1717,1743]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFA2FFFB202DF9185FC611B9" box="[272,428,1717,1743]" italics="true" pageId="24" pageNumber="25">Z. bidoupana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFA2FFFB20DBF9185C1211B9" authorityName="Constant" authorityYear="2024" box="[486,632,1717,1743]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFA2FFFB20DBF9185C1211B9" box="[486,632,1717,1743]" italics="true" pageId="24" pageNumber="25">Z. limbourgi</emphasis>
</taxonomicName>
–
<figureCitation id="132A2A69FFA2FFFB23A9F91B5C9E11B9" box="[660,756,1717,1743]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="24" pageNumber="25">Figs 3A</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB223CF91B5D5D11B9" box="[769,823,1717,1743]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">16A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA2FFFB2180F9595FCA10F0" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BAE36ECFFA2FFFB2180F9595F2E1067" blockId="24.[189,324,1783,1809]" box="[189,324,1783,1809]" pageId="24" pageNumber="25">
<heading id="D0E68180FFA2FFFB2180F9595F2E1067" bold="true" box="[189,324,1783,1809]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B965EAFEFFA2FFFB2180F9595F2E1067" bold="true" box="[189,324,1783,1809]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA2FFFB2180F88B5FCA10F0" blockId="24.[189,1399,1829,1926]" pageId="24" pageNumber="25">
The species epithet is a patronym dedicated to my great (although still a bit fat, see
<bibRefCitation id="EF804B1DFFA2FFFB2582F88B5B041049" author="Constant J." box="[1215,1390,1829,1855]" pageId="24" pageNumber="25" pagination="41 - 54" refId="ref13027" refString="Constant J. 2008. Revision of the Eurybrachidae (XIII). The new Australian genus Chewobrachys (Hemiptera: Fulgoromorpha). Zootaxa 1898: 41 - 54." type="journal article" year="2008">Constant 2008</bibRefCitation>
) colleague Pol Limbourg (RBINS) who spotted the
<typeStatus id="54AA884EFFA2FFFB2212F8E75D081015" box="[815,866,1865,1891]" pageId="24" pageNumber="25">type</typeStatus>
specimens on a tree trunk in Phnom Aural Wildlife Sanctuary.
</paragraph>
</subSubSection>
<caption id="DF6E6664FFA3FFFA2180F82B5FB31090" ID-DOI="http://doi.org/10.5281/zenodo.13837073" ID-Zenodo-Dep="13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,230,1925,1951]" targetBox="[240,1354,270,1868]" targetPageId="25">
<paragraph id="8BAE36ECFFA3FFFA2180F82B5FB31090" blockId="25.[189,1399,1925,2022]" pageId="25" pageNumber="26">
<emphasis id="B965EAFEFFA3FFFA2180F82B5F7210E9" bold="true" box="[189,280,1925,1951]" pageId="25" pageNumber="26">Fig. 15.</emphasis>
<taxonomicName id="4C114D6FFFA3FFFA201DF8285C3910E9" authority="Constant" authorityName="Constant" authorityYear="2024" box="[288,595,1925,1951]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA3FFFA201DF8285F8E10E9" box="[288,484,1925,1951]" italics="true" pageId="25" pageNumber="26">Zanna limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA3FFFA2367F8285CDD10D6" box="[602,695,1926,1952]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS).
<emphasis id="B965EAFEFFA3FFFA22F1F82B5D8B10E9" bold="true" box="[972,993,1925,1951]" pageId="25" pageNumber="26">A</emphasis>
. Habitus dorsal view.
<emphasis id="B965EAFEFFA3FFFA25D8F82B5A9310E9" bold="true" box="[1253,1273,1925,1951]" pageId="25" pageNumber="26">B</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFA3FFFA2093F8065FA910B4" bold="true" box="[430,451,1960,1986]" pageId="25" pageNumber="26">C</emphasis>
. Habitus, ventral view.
<emphasis id="B965EAFEFFA3FFFA23E3F8075C9910B5" bold="true" box="[734,755,1961,1987]" pageId="25" pageNumber="26">D</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFA3FFFA253BF8075A7010B5" bold="true" box="[1030,1050,1961,1987]" pageId="25" pageNumber="26">E</emphasis>
. Head, ventral view.
<emphasis id="B965EAFEFFA3FFFA242AF8075B4310B5" bold="true" box="[1303,1321,1961,1987]" pageId="25" pageNumber="26">F</emphasis>
. Head and thorax, lateral view.
</paragraph>
</caption>
<caption id="DF6E6664FFA0FFF921F8F8085B151F70" ID-DOI="http://doi.org/10.5281/zenodo.13837076" ID-Zenodo-Dep="13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="26" pageNumber="27" startId="26.[197,236,1958,1984]" targetBox="[230,1373,285,1917]" targetPageId="26">
<paragraph id="8BAE36ECFFA0FFF921F8F8085B151F70" blockId="26.[197,1407,1958,2055]" pageId="26" pageNumber="27">
<emphasis id="B965EAFEFFA0FFF921F8F8085F7410B6" bold="true" box="[197,286,1958,1984]" pageId="26" pageNumber="27">Fig. 16.</emphasis>
<taxonomicName id="4C114D6FFFA0FFF92018F8095C2410B6" authority="Constant" authorityName="Constant" authorityYear="2024" box="[293,590,1958,1984]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA0FFF92018F8095F8910B6" box="[293,483,1958,1984]" italics="true" pageId="26" pageNumber="27">Zanna limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA0FFF92368F8095CC510B7" box="[597,687,1959,1985]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS), terminalia.
<emphasis id="B965EAFEFFA0FFF92506F8085A1810B6" bold="true" box="[1083,1138,1958,1984]" pageId="26" pageNumber="27">A–D</emphasis>
. Pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFA0FFF92001F8675F3B1095" bold="true" box="[316,337,1993,2019]" pageId="26" pageNumber="27">A</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFA0FFF92317F8675C541095" bold="true" box="[554,574,1993,2019]" pageId="26" pageNumber="27">B</emphasis>
. Caudal view.
<emphasis id="B965EAFEFFA0FFF923D4F8675C941095" bold="true" box="[745,766,1993,2019]" pageId="26" pageNumber="27">C</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFA0FFF92298F8675DD01095" bold="true" box="[933,954,1993,2019]" pageId="26" pageNumber="27">D</emphasis>
. Ventral view.
<emphasis id="B965EAFEFFA0FFF9255AF8675AF41095" bold="true" box="[1127,1182,1993,2019]" pageId="26" pageNumber="27">E–H</emphasis>
. Aedeagus.
<emphasis id="B965EAFEFFA0FFF92416F8675B541095" bold="true" box="[1323,1342,1993,2019]" pageId="26" pageNumber="27">E</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFA0FFF92072F8435F0B1F71" bold="true" box="[335,353,2029,2055]" pageId="26" pageNumber="27">F</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFA0FFF920C1F8425C791F70" bold="true" box="[508,531,2028,2054]" pageId="26" pageNumber="27">G</emphasis>
. Laterodorsal view.
<emphasis id="B965EAFEFFA0FFF923CCF8435D621F71" bold="true" box="[753,776,2029,2055]" pageId="26" pageNumber="27">H</emphasis>
. Ventral view.Abbreviations: see Materials and methods.
</paragraph>
</caption>
<subSubSection id="C30B6567FFA1FFF82180FEA75F7115B9" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BAE36ECFFA1FFF82180FEA75F061654" blockId="27.[189,364,264,291]" box="[189,364,264,291]" pageId="27" pageNumber="28">
<heading id="D0E68180FFA1FFF82180FEA75F061654" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B965EAFEFFA1FFF82180FEA75F061654" bold="true" box="[189,364,264,291]" pageId="27" pageNumber="28">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B793CB1FFA1FFF821D7FE995FE21577" collectingDate="2018-05-10" collectingDateMax="2018-05-14" collectingDateMin="2018-05-10" collectionCode="RBINS" collectorName="J. Constant &amp; P. Limbourg" country="Cambodia" elevation="300" latitude="11.983334" location="Srea Ken Village" longLatPrecision="1294" longitude="104.13333" municipality="Phnom Aural Wildlife Sanctuary" pageId="27" pageNumber="28" specimenCode="33.732" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Speu Province" typeStatus="holotype">
<paragraph id="8BAE36ECFFA1FFF821D7FE995F361627" blockId="27.[189,1399,311,513]" box="[234,348,311,337]" pageId="27" pageNumber="28">
<emphasis id="B965EAFEFFA1FFF821D7FE995F361627" bold="true" box="[234,348,311,337]" pageId="27" pageNumber="28">
<typeStatus id="54AA884EFFA1FFF821D7FE995F361627" box="[234,348,311,337]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FEF45FE21577" blockId="27.[189,1399,311,513]" pageId="27" pageNumber="28">
<collectingCountry id="F306767CFFA1FFF82180FEF45F311602" box="[189,347,346,372]" name="Cambodia" pageId="27" pageNumber="28">CAMBODIA</collectingCountry>
•
<specimenCount id="9D17FD65FFA1FFF82049FEF45FC81603" box="[372,418,346,373]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D5F80EFFA1FFF82092FEF55CBC1602" box="[431,726,346,373]" country="Cambodia" name="Kampong Speu" pageId="27" pageNumber="28">Kampong Speu Province</collectingRegion>
,
<collectingMunicipality id="6BCAAC96FFA1FFF823DFFEF45A081602" box="[738,1122,346,372]" pageId="27" pageNumber="28">Phnom Aural Wildlife Sanctuary</collectingMunicipality>
, near
<location id="8ECE6037FFA1FFF8259AFEF45B181602" LSID="urn:lsid:plazi:treatment:03B887FAFFA2FFFC2377FBF75DFE12FB:8ECE6037FFA1FFF8259AFEF45B181602" box="[1191,1394,346,373]" country="Cambodia" latitude="11.983334" longLatPrecision="1294" longitude="104.13333" municipality="Phnom Aural Wildlife Sanctuary" name="Srea Ken Village" pageId="27" pageNumber="28" stateProvince="Kampong Speu Province">Srea Ken Village</location>
;
<geoCoordinate id="EE25502BFFA1FFF82180FED25F4216EE" box="[189,296,380,408]" degrees="11" direction="north" minutes="59" orientation="latitude" pageId="27" pageNumber="28" precision="925" value="11.983334">11°59ʹ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFA1FFF82008FED25FC716EE" box="[309,429,380,408]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="27" pageNumber="28" precision="925" value="104.13333">104°08ʹ E</geoCoordinate>
; 10–14 May 2018;
<elevation id="003CD1DFFFA1FFF823AEFED05D3216E1" box="[659,856,381,408]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">
<quantity id="4CE99B09FFA1FFF823AEFED05D7216EE" box="[659,792,382,408]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</quantity>
a.s.l.
</elevation>
; GTI Project;
<collectorName id="26E4533AFFA1FFF8253FFED05AEE16EE" box="[1026,1156,382,408]" pageId="27" pageNumber="28">J. Constant</collectorName>
&amp;
<collectorName id="26E4533AFFA1FFF82596FED05B5616E1" box="[1195,1340,381,408]" pageId="27" pageNumber="28">P. Limbourg</collectorName>
leg.; I.G.:
<specimenCode id="DBB79E97FFA1FFF821C4FE0F5F2716CD" box="[249,333,417,443]" pageId="27" pageNumber="28">33.732</specimenCode>
; “Coll. I.R.Sc.N.B., Cambodia, Kampong Speu prov., Phnom Aural W.S., nr Srea Ken Vill.,
<date id="FFAF102CFFA1FFF82180FE6A5F3016A8" box="[189,346,452,478]" pageId="27" pageNumber="28" value="2018-05-10" valueMax="2018-05-14" valueMin="2018-05-10">
<collectingDate id="EFEBE9C4FFA1FFF82180FE6A5F3016A8" box="[189,346,452,478]" pageId="27" pageNumber="28" value="2018-05-10" valueMax="2018-05-14" valueMin="2018-05-10">10-14.v.2018</collectingDate>
</date>
,
<quantity id="4CE99B09FFA1FFF82055FE6A5F9B16A8" box="[360,497,452,478]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</quantity>
, 11°59ʹ N, 104°08ʹ E, GTI Project, Leg. J. Constant and P. Limbourg, I.G.: 33.732”;
<collectionCode id="ED00AE29FFA1FFF82017FE495FEE1577" box="[298,388,487,513]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="27" pageNumber="28" type="Museum">RBINS</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BAE36ECFFA1FFF821D7FD875F7115B9" blockId="27.[189,1399,553,719]" pageId="27" pageNumber="28">
<materialsCitation id="3B793CB1FFA1FFF821D7FD875DD31510" collectingDate="2018-05-10" collectingDateMax="2018-05-14" collectingDateMin="2018-05-10" collectionCode="RBINS" collectorName="J. Constant &amp; P. Limbourg" country="Cambodia" elevation="300" latitude="11.983334" location="CAMBODIA" longLatPrecision="1294" longitude="104.13333" municipality="Phnom Aural Wildlife Sanctuary" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Speu" typeStatus="paratype">
<emphasis id="B965EAFEFFA1FFF821D7FD875F0D1535" bold="true" box="[234,359,553,579]" pageId="27" pageNumber="28">
<typeStatus id="54AA884EFFA1FFF821D7FD875F0D1535" box="[234,359,553,579]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F306767CFFA1FFF82180FDE25F311510" box="[189,347,588,614]" name="Cambodia" pageId="27" pageNumber="28">CAMBODIA</collectingCountry>
•
<specimenCount id="9D17FD65FFA1FFF8204EFDE25FC81510" box="[371,418,588,614]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED00AE29FFA1FFF8225EFDE25DD31510" box="[867,953,588,614]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="27" pageNumber="28" type="Museum">RBINS</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B793CB1FFA1FFF822EFFDE25F7D15B9" collectingDate="2005-05-28" collectingMethod="light trap / Light Trap" collectionCode="RBINS" collectorName="I. Var &amp; P. Grootaert" country="Cambodia" latitude="13.708333" location="Kbal Spean" longLatPrecision="21" longitude="104.02556" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Siem Reap Province" typeStatus="paratype">
<specimenCount id="9D17FD65FFA1FFF822EFFDE25A681510" box="[978,1026,588,614]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D17FD65FFA1FFF82535FDE25A581510" box="[1032,1074,588,614]" count="1" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49D5F80EFFA1FFF8257DFDE25B581510" box="[1088,1330,588,614]" country="Cambodia" name="Siem Reap" pageId="27" pageNumber="28">Siem Reap Province</collectingRegion>
,
<location id="8ECE6037FFA1FFF82400FDE25F6215FF" LSID="urn:lsid:plazi:treatment:03B887FAFFA2FFFC2377FBF75DFE12FB:8ECE6037FFA1FFF82400FDE25F6215FF" country="Cambodia" latitude="13.708333" longLatPrecision="21" longitude="104.02556" name="Kbal Spean" pageId="27" pageNumber="28" stateProvince="Siem Reap Province">Kbal Spean</location>
; [
<geoCoordinate id="EE25502BFFA1FFF8201DFDC35FD615FF" box="[288,444,621,649]" degrees="13" direction="north" minutes="42" orientation="latitude" pageId="27" pageNumber="28" precision="15" seconds="30" value="13.708333">13°42ʹ30ʺ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFA1FFF820F0FDC35C1815FF" box="[461,626,621,649]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="27" pageNumber="28" precision="15" seconds="32" value="104.02556">104°01ʹ32ʺ E</geoCoordinate>
];
<collectingMethod id="52504EFBFFA1FFF823B1FDC15C9515FF" box="[652,767,623,649]" pageId="27" pageNumber="28">light trap</collectingMethod>
;
<date id="FFAF102CFFA1FFF8222DFDC15DD015FC" box="[784,954,623,650]" pageId="27" pageNumber="28" value="2005-05-28">28 May 2005</date>
;
<collectorName id="26E4533AFFA1FFF822F6FDC15A7A15FF" box="[971,1040,623,649]" pageId="27" pageNumber="28">I. Var</collectorName>
and
<collectorName id="26E4533AFFA1FFF8256EFDC15A8915FF" box="[1107,1251,623,649]" pageId="27" pageNumber="28">P. Grootaert</collectorName>
leg.; “Coll. I.R.Sc.N.B., Cambodia (Siem Reap prov), Kbal Spean,
<collectingMethod id="52504EFBFFA1FFF8226DFD3C5DBA15DB" box="[848,976,658,685]" pageId="27" pageNumber="28">Light Trap</collectingMethod>
,
<date id="FFAF102CFFA1FFF822E0FD3C5A3215DB" box="[989,1112,658,685]" pageId="27" pageNumber="28" value="2005-05-28">
<collectingDate id="EFEBE9C4FFA1FFF822E0FD3C5A3215DB" box="[989,1112,658,685]" pageId="27" pageNumber="28" value="2005-05-28">28.V.2005</collectingDate>
</date>
, Leg. Var &amp; Grootaert”;
<collectionCode id="ED00AE29FFA1FFF82180FD1B5F7D15B9" box="[189,279,693,719]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="27" pageNumber="28" type="Museum">RBINS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA1FFFC2180FD585FE31310" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28" type="description">
<paragraph id="8BAE36ECFFA1FFF82180FD585F241466" blockId="27.[189,334,758,784]" box="[189,334,758,784]" pageId="27" pageNumber="28">
<heading id="D0E68180FFA1FFF82180FD585F241466" bold="true" box="[189,334,758,784]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B965EAFEFFA1FFF82180FD585F241466" bold="true" box="[189,334,758,784]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FC8B5B641415" blockId="27.[189,1399,805,867]" pageId="27" pageNumber="28">
MEASUREMENTS AND RATIOS. LT:
<specimenCount id="9D17FD65FFA1FFF82374FC885C351436" box="[585,607,806,832]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
(n=2):
<quantity id="4CE99B09FFA1FFF823F9FC8B5D1E1436" box="[708,884,805,832]" metricMagnitude="-2" metricUnit="m" metricValue="4.515" metricValueMax="4.68" metricValueMin="4.35" pageId="27" pageNumber="28" unit="mm" value="45.15" valueMax="46.8" valueMin="43.5">43.5–46.8 mm</quantity>
. LTg/BTg =3.3; LH/BH =5.1; LF/BF= 5.3; BH/BPrH = 2.56; LH/LT =0.43; wingspan (extrapolated):
<specimenCount id="9D17FD65FFA1FFF8225FFCE75D161415" box="[866,892,841,867]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
:
<quantity id="4CE99B09FFA1FFF822BAFCE75D9A1415" box="[903,1008,841,867]" metricMagnitude="-2" metricUnit="m" metricValue="4.4799999999999995" pageId="27" pageNumber="28" unit="mm" value="44.8">44.8 mm</quantity>
(41.6–47),
<specimenCount id="9D17FD65FFA1FFF82545FCE75AFA1415" box="[1144,1168,841,867]" count="1" pageId="27" pageNumber="28" type="female">♀</specimenCount>
:
<quantity id="4CE99B09FFA1FFF825A3FCE65B601415" box="[1182,1290,840,867]" metricMagnitude="-2" metricUnit="m" metricValue="4.59" pageId="27" pageNumber="28" unit="mm" value="45.9">45.9 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FC245DFB1397" blockId="27.[189,1400,906,1249]" pageId="27" pageNumber="28">
HEAD (
<figureCitation id="132A2A69FFA1FFF82033FC245FD314D2" box="[270,441,906,933]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15B, E–F</figureCitation>
). Strongly elongate, representing 43% of total length, gently and evenly narrowing towards apex with apical 1/2 more or less parallel-sided in dorsal view; apex weakly widening, then obliquely cut in lateral view and with incomplete, rim-shaped orange carina (missing in ventral portion); pinkish brown densely covered in irregular very small black pitting, with some moderately larger pits mostly in basal half on sides and vertex and rarely coalescent; some slightly larger black pustules on side in distal portion, with always 2–3 along carina of vertex. Lateral carinae of frons indistinct in basal half, then nearly straight and slightly prominent towards apex, upcurved near apex and merging into a short dorsal carina reaching apex; lateral carinae of vertex indistinct in basal half, then rather weak and zigzagged, merging with upcurved part of lateral carinae of frons; median carina of vertex weakly marked in distal ⅓. Labium pale brown, reaching mesocoxae.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FAA65FB9121E" blockId="27.[189,1398,1288,1384]" pageId="27" pageNumber="28">
THORAX (
<figureCitation id="132A2A69FFA1FFF8200CFAA65F891254" box="[305,483,1288,1315]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15B, E–F</figureCitation>
). Pro- and mesonotum pinkish brown entirely covered in moderately dense, irregular, moderate black pitting with pits mostly smaller than space between them and rarely coalescent; median carina obsolete.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FA3E5AA11141" blockId="27.[188,1399,1424,1591]" pageId="27" pageNumber="28">
TEGMINA (
<figureCitation id="132A2A69FFA1FFF82007FA3E5F9912DC" box="[314,499,1424,1451]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15A, C–D</figureCitation>
). Strongly elongate and reticulate, pale pinkish brown, progressively paler, subhyaline towards apex, basal portion densely covered with minute dark red spots; all surface irregularly covered in small to moderately large black spots, mostly on veins, larger ones slightly protruding; costal and postclaval margins subparallel, weakly diverging from base towards apex and curved at level of apex of clavus; apical margin rounded. Veins orange in large basal portion. Clavus open.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180F9F05B2211ED" blockId="27.[189,1398,1629,1692]" pageId="27" pageNumber="28">VENATION. ScP+R forked in basal ⅓, MP forked more basally in basal 1/4; CuA forked near apex of clavus; PCu and A1 fused near apex of clavus; PCu+A1 fused with postclaval margin at nodal line.</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180F96D5A341076" blockId="27.[189,1398,1731,1793]" pageId="27" pageNumber="28">
HIND WINGS (
<figureCitation id="132A2A69FFA1FFF8205BF96D5F9611AB" box="[358,508,1731,1758]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15A, C</figureCitation>
). Subhyaline, milky white with veins slightly darker, tinged with reddish brown; vein PCu brown in basal half; postclaval margin emarginate at A1 vein.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180F8865FBF10DD" blockId="27.[189,1398,1831,1963]" pageId="27" pageNumber="28">
VENATION. ScP+R and MP forked beyond midlength; CuA forked slightly more basally then MP fork; PCu forked at basal ⅓; A1 forked rather close to base, A1 2 fused with A
<quantity id="4CE99B09FFA1FFF82530F8E55A501013" box="[1037,1082,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="27" pageNumber="28" unit="in" value="2.0">2 in</quantity>
distal portion and A1 2 +A2 forked slightly before reaching postclaval margin. Numerous crossveins in large portion along apical and postclaval margins.
</paragraph>
<caption id="DF6E6664FFA6FFFF2184F8065AD91093" ID-DOI="http://doi.org/10.5281/zenodo.13837078" ID-Zenodo-Dep="13837078" httpUri="https://zenodo.org/record/13837078/files/figure.png" pageId="28" pageNumber="29" startId="28.[185,226,1960,1986]" targetBox="[213,1370,261,1921]" targetPageId="28">
<paragraph id="8BAE36ECFFA6FFFF2184F8065AD91093" blockId="28.[185,1394,1960,2022]" pageId="28" pageNumber="29">
<emphasis id="B965EAFEFFA6FFFF2184F8065F7C10B4" bold="true" box="[185,278,1960,1986]" pageId="28" pageNumber="29">Fig. 17.</emphasis>
<taxonomicName id="4C114D6FFFA6FFFF2023F8075C3910B4" authority="Constant" authorityName="Constant" authorityYear="2024" box="[286,595,1960,1986]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA6FFFF2023F8075F8E10B4" box="[286,484,1960,1986]" italics="true" pageId="28" pageNumber="29">Zanna limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA6FFFF2366F8065CD210B4" box="[603,696,1960,1986]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018.
<emphasis id="B965EAFEFFA6FFFF20D1F8655C6B1093" bold="true" box="[492,513,1995,2021]" pageId="28" pageNumber="29">A</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFA6FFFF222DF8655D4E1093" bold="true" box="[784,804,1995,2021]" pageId="28" pageNumber="29">B</emphasis>
. Habitus, dorsal view.
<emphasis id="B965EAFEFFA6FFFF250CF8655A2C1093" bold="true" box="[1073,1094,1995,2021]" pageId="28" pageNumber="29">C</emphasis>
. Habitat.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFA7FFFC2180FEA75AB51630" blockId="29.[189,1399,265,467]" lastBlockId="31.[189,1399,265,327]" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
LEGS (
<figureCitation id="132A2A69FFA7FFFE2037FEA75FAB1655" box="[266,449,265,292]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="29" pageNumber="30">Fig. 15A, C–D</figureCitation>
). Pinkish stramineous, pilose, rather short and robust. Pro- and mesocoxae with some black markings; all femora with small to minute black spots and conspicuous black spots aligned in a ring in distal ⅓, rarely coalescent; pro- and mesotibiae with some small black spots (some arranged into irregular ring in basal ¼), and with apex brown; pro- and mesotarsi with segments darker apically; metatibiae weakly brown basally with all spines apically black, 4–5 lateral spines and 8 apical spines; first metatarsomere with 11–14 apical spines ventrally; second metatarsomere with 10 apical spines ventrally; first and second metatarsomere with a dense pad of microsetae ventrally; third metatarsomere elongate with a subapical blackish brown ring. Metatibiotarsal formula: (4–5) 8/11–14/10.
</paragraph>
<caption id="DF6E6664FFA7FFFE2182F8F65FA310AA" ID-DOI="http://doi.org/10.5281/zenodo.13837080" ID-Zenodo-Dep="13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="29" pageNumber="30" startId="29.[191,234,1880,1906]" targetBox="[185,1406,518,1841]" targetPageId="29">
<paragraph id="8BAE36ECFFA7FFFE2182F8F65FA310AA" blockId="29.[190,1401,1880,2013]" pageId="29" pageNumber="30">
<emphasis id="B965EAFEFFA7FFFE2182F8F65F711005" bold="true" box="[191,283,1880,1907]" pageId="29" pageNumber="30">Fig. 18.</emphasis>
<taxonomicName id="4C114D6FFFA7FFFE201CF8F45F061005" authorityName="Kirkaldy" authorityYear="1902" box="[289,364,1882,1907]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFA7FFFE201CF8F45F061005" box="[289,364,1882,1907]" italics="true" pageId="29" pageNumber="30">Zanna</emphasis>
</taxonomicName>
, new spp. from Cambodia and Vietnam, key characters.
<emphasis id="B965EAFEFFA7FFFE22C6F8F65A5E1004" bold="true" box="[1019,1076,1880,1906]" pageId="29" pageNumber="30">A–D</emphasis>
. Prothotax and basal half of head, lateral view.
<emphasis id="B965EAFEFFA7FFFE20A5F8D25FBB10E0" bold="true" box="[408,465,1916,1942]" pageId="29" pageNumber="30">E–H</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFA7FFFE2200F8D25D0010E0" bold="true" box="[829,874,1916,1942]" pageId="29" pageNumber="30">I–L</emphasis>
. Right tegmen.
<emphasis id="B965EAFEFFA7FFFE251CF8D25A1A10E0" bold="true" box="[1057,1136,1916,1942]" pageId="29" pageNumber="30">A, E, I</emphasis>
.
<taxonomicName id="4C114D6FFFA7FFFE2542F8D35B7010E0" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1151,1306,1916,1942]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFA7FFFE2542F8D35B7010E0" box="[1151,1306,1916,1942]" italics="true" pageId="29" pageNumber="30">Z. bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE241DF8D25B1310E0" box="[1312,1401,1916,1942]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA7FFFE2183F8315F7810CF" bold="true" box="[190,274,1951,1977]" pageId="29" pageNumber="30">B, F, J</emphasis>
.
<taxonomicName id="4C114D6FFFA7FFFE2019F80E5C4A10CF" authority="Constant" authorityName="Constant" authorityYear="2024" box="[292,544,1951,1978]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="34" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
Z.
<emphasis id="B965EAFEFFA7FFFE2075F8315FC510CF" box="[328,431,1951,1977]" italics="true" pageId="29" pageNumber="30">chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE2317F80E5CED10CC" box="[554,647,1952,1978]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA7FFFE23ACF8315C9410CF" bold="true" box="[657,766,1951,1977]" pageId="29" pageNumber="30">C, G, K.</emphasis>
<taxonomicName id="4C114D6FFFA7FFFE223AF80E5DFA10CF" authorityName="Constant &amp; Pham" authorityYear="2024" box="[775,912,1951,1977]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFA7FFFE223AF80E5DFA10CF" box="[775,912,1951,1977]" italics="true" pageId="29" pageNumber="30">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE22A4F80E5D9C10CC" box="[921,1014,1952,1978]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA7FFFE253DF8315A0810CF" bold="true" box="[1024,1122,1951,1977]" pageId="29" pageNumber="30">D, H, L</emphasis>
.
<taxonomicName id="4C114D6FFFA7FFFE254EF80E5B1310CF" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1139,1401,1951,1977]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA7FFFE254EF80E5B6D10CF" box="[1139,1287,1951,1977]" italics="true" pageId="29" pageNumber="30">Z. limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE2182F86D5F7710AB" box="[191,285,1987,2013]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, (not to scale).
</paragraph>
</caption>
<caption id="DF6E6664FFA4FFFD21FCF8CA5CDC109E" ID-DOI="http://doi.org/10.5281/zenodo.13837082" ID-Zenodo-Dep="13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="30" pageNumber="31" startId="30.[193,236,1892,1918]" targetBox="[193,1402,242,1853]" targetPageId="30">
<paragraph id="8BAE36ECFFA4FFFD21FCF8CA5CDC109E" blockId="30.[193,1403,1892,2024]" pageId="30" pageNumber="31">
<emphasis id="B965EAFEFFA4FFFD21FCF8CA5F761008" bold="true" box="[193,284,1892,1918]" pageId="30" pageNumber="31">Fig. 19.</emphasis>
<taxonomicName id="4C114D6FFFA4FFFD201DF8CB5F011008" authorityName="Kirkaldy" authorityYear="1902" box="[288,363,1893,1918]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFA4FFFD201DF8CB5F011008" box="[288,363,1893,1918]" italics="true" pageId="30" pageNumber="31">Zanna</emphasis>
</taxonomicName>
, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFA4FFFD21FCF8295F2410D7" bold="true" box="[193,334,1927,1953]" pageId="30" pageNumber="31">A, C, E, G</emphasis>
. Lateral view.
<emphasis id="B965EAFEFFA4FFFD2334F8295CFA10D7" bold="true" box="[521,656,1927,1953]" pageId="30" pageNumber="31">B, D, F, H</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFA4FFFD2278F8295D1710D7" bold="true" box="[837,893,1927,1953]" pageId="30" pageNumber="31">A–B</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD22ADF8265A5A10D7" authorityName="Constant &amp; Pham" authorityYear="2024" box="[912,1072,1927,1953]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD22ADF8265A5A10D7" box="[912,1072,1927,1953]" italics="true" pageId="30" pageNumber="31">Z. bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD2506F8265AF010D4" box="[1083,1178,1928,1954]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA4FFFD2598F8295AB410D7" bold="true" box="[1189,1246,1927,1953]" pageId="30" pageNumber="31">C–D</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD25CCF8265F4310B3" authority="Constant" authorityName="Constant" authorityYear="2024" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD25CCF8265B1110D7" box="[1265,1403,1927,1953]" italics="true" pageId="30" pageNumber="31">Z. chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD200FF8055FE410B3" box="[306,398,1963,1989]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA4FFFD20A5F8055FA110B3" bold="true" box="[408,459,1963,1989]" pageId="30" pageNumber="31">E–F</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD20E6F8055C0910B2" authorityName="Constant &amp; Pham" authorityYear="2024" box="[475,611,1962,1988]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD20E6F8055C0910B2" box="[475,611,1962,1988]" italics="true" pageId="30" pageNumber="31">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD2356F8055CAD10B3" box="[619,711,1963,1989]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA4FFFD23ECF8045D6610B2" bold="true" box="[721,780,1962,1988]" pageId="30" pageNumber="31">G–H</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD2221F8055A4A10B3" authority="Constant" authorityName="Constant" authorityYear="2024" box="[796,1056,1962,1989]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD2221F8055DDA10B2" box="[796,944,1962,1988]" italics="true" pageId="30" pageNumber="31">Z. limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD2514F8055AEC10B3" box="[1065,1158,1963,1989]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(scale bars =1 mm). Abbreviations: see Materials and methods.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFA5FFFC2180FEC05DCD1684" blockId="31.[189,1399,365,498]" pageId="31" pageNumber="32">
ABDOMEN (
<figureCitation id="132A2A69FFA5FFFC207DFEC05FA116FE" box="[320,459,365,392]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="31" pageNumber="32">Fig. 15A, C</figureCitation>
). Tergites smooth, segments blackish brown with more or less developed marking at posterior angles, and posterior margin, yellowish; sternites (except pregenital one) yellowish brown with irregular small black dots arranged in two groups in middle portion, and two groups on side of each segment, often with two black dots anteriorly to middle groups.
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180FDB75FE31310" blockId="31.[189,1399,536,1126]" pageId="31" pageNumber="32">
MALE TERMINALIA (
<figureCitation id="132A2A69FFA5FFFC2090FDB75C621544" box="[429,520,536,563]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16</figureCitation>
). Pygofer (
<emphasis id="B965EAFEFFA5FFFC23AFFDB75CD81544" box="[658,690,537,562]" italics="true" pageId="31" pageNumber="32">Py</emphasis>
–
<figureCitation id="132A2A69FFA5FFFC23ECFDB75D0E1544" box="[721,868,536,563]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16A–D</figureCitation>
) about 1.69 times as high as long in lateral view, robust with posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle; sides weakly excavate in dorsal portion (to observe in dorsal aspect,
<figureCitation id="132A2A69FFA5FFFC2183FD2C5F4615EA" box="[190,300,642,668]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16C</figureCitation>
). Anal tube (
<emphasis id="B965EAFEFFA5FFFC20FAFD2D5F8215EB" box="[455,488,643,669]" italics="true" pageId="31" pageNumber="32">An</emphasis>
–
<figureCitation id="132A2A69FFA5FFFC2338FD2C5CFF15EA" box="[517,661,642,668]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16A–C</figureCitation>
) robust, moderately flattened, 1.65 times as long as high and with ventral margin oblique in distal half and moderately angular before midlength in lateral view; in dorsal view, 1.08 times as long as wide, with sides sinuate, subparallel in basal portion, then roundly diverging towards apex, forming rounded posterior angles and with apical margin incurved; suboval in caudal view, flattened dorsally; epiproct and paraproct rather strongly surpassing anal tube, subtriangular and rounded posteriorly in dorsal aspect. Gonostyli (
<emphasis id="B965EAFEFFA5FFFC250DFC9D5A2F143B" box="[1072,1093,819,845]" italics="true" pageId="31" pageNumber="32">G</emphasis>
–
<figureCitation id="132A2A69FFA5FFFC2559FC9C5A9C143A" box="[1124,1270,818,844]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16A–D</figureCitation>
) elongate, 2.58 times as long as high; in lateral view, ventral margin broadly rounded and dorsal margin rather broadly rounded after lateral hook (
<emphasis id="B965EAFEFFA5FFFC2340FCD75CCE14E5" box="[637,676,889,915]" italics="true" pageId="31" pageNumber="32">lhg</emphasis>
), apical margin rather broadly rounded; lateral hook with basal portion projecting posterolaterad and apex strongly curved lateroventrad; in ventral view, lateral protrusion at about midlength, rather strongly, angularly marked and with posterior angle curved. Aedeagus (
<figureCitation id="132A2A69FFA5FFFC2094FC4C5C51148A" box="[425,571,994,1020]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. A–D. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. E–H. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16E–H</figureCitation>
) moderately elongate with lateral processes (
<emphasis id="B965EAFEFFA5FFFC2551FC4C5AF9148A" box="[1132,1171,994,1020]" italics="true" pageId="31" pageNumber="32">lpa</emphasis>
) forming multiple lobes; dorsal process (
<emphasis id="B965EAFEFFA5FFFC20E8FBAB5C681369" box="[469,514,1029,1055]" italics="true" pageId="31" pageNumber="32">dpa</emphasis>
) weakly protruding dorsally in lateral view; dorsal endosomal process (
<emphasis id="B965EAFEFFA5FFFC21FAFB875E991335" box="[199,243,1065,1091]" italics="true" pageId="31" pageNumber="32">dep</emphasis>
) in dorsal view subtriangular with lateral margins nearly straight and rather wide median slit in distal portion.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA5FFFC2180FB235A7C126A" pageId="31" pageNumber="32" type="biology_ecology">
<paragraph id="8BAE36ECFFA5FFFC2180FB235F7613D1" blockId="31.[189,284,1165,1191]" box="[189,284,1165,1191]" pageId="31" pageNumber="32">
<heading id="D0E68180FFA5FFFC2180FB235F7613D1" bold="true" box="[189,284,1165,1191]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B965EAFEFFA5FFFC2180FB235F7613D1" bold="true" box="[189,284,1165,1191]" pageId="31" pageNumber="32">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180FB125A7C126A" blockId="31.[189,1399,1212,1308]" pageId="31" pageNumber="32">
The specimens from
<collectingRegion id="49D5F80EFFA5FFFC2089FB125C5913A0" box="[436,563,1212,1238]" country="Cambodia" name="Siem Reap" pageId="31" pageNumber="32">Siem Reap</collectingRegion>
were attracted to a light trap in a strongly anthropized area while those from Phnom Aural were found sitting on the trunk of an unidentified tree (
<figureCitation id="132A2A69FFA5FFFC2578FB715ABE138F" box="[1093,1236,1247,1273]" captionStart="Fig" captionStartId="28.[185,226,1960,1986]" captionTargetBox="[213,1370,261,1921]" captionTargetId="figure-36@28.[213,1370,258,1921]" captionTargetPageId="28" captionText="Fig. 17. Zanna limbourgi Constant sp. nov., holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018. A. Habitus, lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi="http://doi.org/10.5281/zenodo.13837078" httpUri="https://zenodo.org/record/13837078/files/figure.png" pageId="31" pageNumber="32">Fig. 17A–B</figureCitation>
) about
<quantity id="4CE99B09FFA5FFFC240CFB715B1D138C" box="[1329,1399,1247,1274]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="1.5">1.5 m</quantity>
above the ground in rather heavily disturbed secondary forest (
<figureCitation id="132A2A69FFA5FFFC22A0FAAC5A63126A" box="[925,1033,1282,1308]" captionStart="Fig" captionStartId="28.[185,226,1960,1986]" captionTargetBox="[213,1370,261,1921]" captionTargetId="figure-36@28.[213,1370,258,1921]" captionTargetPageId="28" captionText="Fig. 17. Zanna limbourgi Constant sp. nov., holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018. A. Habitus, lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi="http://doi.org/10.5281/zenodo.13837078" httpUri="https://zenodo.org/record/13837078/files/figure.png" pageId="31" pageNumber="32">Fig. 17C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA5FFFC2180FAED5DFE12FB" box="[189,916,1347,1421]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BAE36ECFFA5FFFC2180FAED5F3C122B" blockId="31.[189,342,1347,1373]" box="[189,342,1347,1373]" pageId="31" pageNumber="32">
<heading id="D0E68180FFA5FFFC2180FAED5F3C122B" bold="true" box="[189,342,1347,1373]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B965EAFEFFA5FFFC2180FAED5F3C122B" bold="true" box="[189,342,1347,1373]" pageId="31" pageNumber="32">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180FADC5DFE12FB" blockId="31.[189,916,1394,1421]" box="[189,916,1394,1421]" pageId="31" pageNumber="32">
<collectingCountry id="F306767CFFA5FFFC2180FADC5F5012FA" box="[189,314,1394,1420]" name="Cambodia" pageId="31" pageNumber="32">Cambodia</collectingCountry>
:
<collectingRegion id="49D5F80EFFA5FFFC2078FADD5F9112FA" box="[325,507,1394,1421]" country="Cambodia" name="Kampong Speu" pageId="31" pageNumber="32">Kampong Speu</collectingRegion>
and
<collectingRegion id="49D5F80EFFA5FFFC2309FADC5CD912FB" box="[564,691,1394,1421]" country="Cambodia" name="Siem Reap" pageId="31" pageNumber="32">Siem Reap</collectingRegion>
provinces (
<figureCitation id="132A2A69FFA5FFFC2200FADC5DEF12FB" box="[829,901,1394,1421]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="31" pageNumber="32">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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