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<document id="1ECD3D4C47207D52AB0601DC25506C2C" ID-CLB-Dataset="102843" ID-DOI="10.11646/zootaxa.5168.2.5" ID-GBIF-Dataset="16df2448-1e2c-4410-ac7b-af3623ff401f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6877267" ID-ZooBank="36831FDB-E9BE-4255-82B9-BFFCDCD40098" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658420700313" checkinUser="plazi" docAuthor="Zhao, Ming-Zhi &amp; Zorn, Carsten" docDate="2022" docId="03B887EBDF7EFFA917AC06670A4DFE9C" docLanguage="en" docName="zootaxa.5168.2.5.pdf" docOrigin="Zootaxa 5168 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala inclinata Zhao &amp; Zorn 2022, new species" docType="treatment" docUuid="F3ACD87D-EF5F-4B35-9F1E-463CF72BAE12" docUuidSource="ZooBank" docVersion="6" lastPageNumber="191" masterDocId="FF81FF93DF72FFB9173B045C0836FF81" masterDocTitle="Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan" masterLastPageNumber="195" masterPageNumber="175" pageNumber="187" updateTime="1699361553354" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8534ECB1AE950A0659BE5AF6AFA4C36E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B8D2538069453AEDD4D95B6113F4EFE8">
<mods:title id="EB5CFE6E06485B297C717D74A8A16050">Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan</mods:title>
</mods:titleInfo>
<mods:name id="1D84DA6405296ADA82ABA0103FEF31FF" type="personal">
<mods:role id="8412E08658424B33C553E7BDC6833759">
<mods:roleTerm id="F98189BB60BBF563E9FE1857389ED60C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AEA857192F85FA2A554BF038DC94AFAE">Zhao, Ming-Zhi</mods:namePart>
<mods:nameIdentifier id="5B324DC52ED59B5D4830732D4D230E97" type="ORCID">0000-0002-3158-6177</mods:nameIdentifier>
<mods:affiliation id="C4E0BE2877FB0BA92865E093C2B1BFE5">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, P. R. China. zhaomzhai @ 126. com; https: // orcid. org / 0000 - 0002 - 3158 - 6177</mods:affiliation>
</mods:name>
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<mods:roleTerm id="72538E595F4B7BBE51295361C44BEFE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C208D35C31CAAE3C3C52693461EDBEDF">Zorn, Carsten</mods:namePart>
<mods:affiliation id="1D45263691DB41FC8C4F510134ADBE0B">Rostocker Strasse 1 a, Gnoien, 17179, Germany.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="DE18B4E481E64A12330D560621FE787C">text</mods:typeOfResource>
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<mods:title id="9C56763B77CD5FC1F1B3075728E8E67D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3675F0C76AE2CB176D3541ED926B2D1E">
<mods:date id="2144FCD0D3D4963372966301C10AABDB">2022</mods:date>
<mods:detail id="41C24047DA05865691855DCBD1AD53A7" type="pubDate">
<mods:number id="F875983324C1D73121AA06D6D53FB9C6">2022-07-21</mods:number>
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<mods:detail id="A3D022DF5B14A646CE2E2844B7137C09" type="volume">
<mods:number id="C07ED121A74BC4D9276FB36E20515FE2">5168</mods:number>
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<mods:classification id="195688B9C415F25A8CA84FDD3337481C">journal article</mods:classification>
<mods:identifier id="E8135672483B30BE1EF46CCC4DFCE0C8" type="CLB-Dataset">102843</mods:identifier>
<mods:identifier id="82C817A73B0F5BD27441F618C683385B" type="DOI">10.11646/zootaxa.5168.2.5</mods:identifier>
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<mods:identifier id="2A4DE8F9FA3E4004C4B54A1C1D127B48" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B887EBDF7EFFA917AC06670A4DFE9C" ID-DOI="http://doi.org/10.5281/zenodo.6877217" ID-GBIF-Taxon="197841318" ID-Zenodo-Dep="6877217" LSID="urn:lsid:zoobank.org:act:F3ACD87D-EF5F-4B35-9F1E-463CF72BAE12" httpUri="http://treatment.plazi.org/id/03B887EBDF7EFFA917AC06670A4DFE9C" lastPageId="16" lastPageNumber="191" pageId="12" pageNumber="187">
<subSubSection id="C30B6576DF7EFFB517AC06670C05FDD7" box="[151,1075,571,598]" pageId="12" pageNumber="187" type="nomenclature">
<paragraph id="8BAE36FDDF7EFFB517AC06670C05FDD7" blockId="12.[151,1075,571,635]" box="[151,1075,571,598]" pageId="12" pageNumber="187">
<heading id="D0E68191DF7EFFB517AC06670C05FDD7" bold="true" box="[151,1075,571,598]" fontSize="11" level="1" pageId="12" pageNumber="187" reason="1">
<emphasis id="B965EAEFDF7EFFB517AC06670C05FDD7" bold="true" box="[151,1075,571,598]" pageId="12" pageNumber="187">
<taxonomicName id="4C114D7EDF7EFFB517AC06670A1CFDD7" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[151,554,571,598]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata" status="sp. nov.">
<emphasis id="B965EAEFDF7EFFB517AC0667094FFDD7" bold="true" box="[151,377,571,598]" italics="true" pageId="12" pageNumber="187">Anomala inclinata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<taxonomicNameLabel id="A2565794DF7EFFB5150C06600AF1FDD7" box="[567,711,572,598]" pageId="12" pageNumber="187" rank="species">new species</taxonomicNameLabel>
[DZṜ异fflŵoi/Øae條ŵff]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF7EFFB517AC063C09D8FDFB" box="[151,494,608,635]" pageId="12" pageNumber="187" type="description">
<paragraph id="8BAE36FDDF7EFFB517AC063C09D8FDFB" blockId="12.[151,1075,571,635]" box="[151,494,608,635]" pageId="12" pageNumber="187">
(
<figureCitation id="132A2A78DF7EFFB517A4063C08D9FDFA" box="[159,239,608,635]" captionStart="FIGURE 7" captionStartId="13.[151,250,1974,1998]" captionTargetBox="[181,1411,404,1936]" captionTargetId="figure-106@13.[158,1430,381,1950]" captionTargetPageId="13" captionText="FIGURE 7. Habitus of Anomala inclinata Zhao &amp; Zorn, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877281" httpUri="https://zenodo.org/record/6877281/files/figure.png" pageId="12" pageNumber="187">Figs. 7</figureCitation>
,
<figureCitation id="132A2A78DF7EFFB517C0063C0974FDFB" box="[251,322,608,634]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="12" pageNumber="187">8G–H</figureCitation>
,
<figureCitation id="132A2A78DF7EFFB51676063C09D3FDFB" box="[333,485,608,634]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="12" pageNumber="187">9K–M, 9P–Q</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF7EFFB517AC06F409C8FD67" pageId="12" pageNumber="187" type="materials_examined">
<paragraph id="8BAE36FDDF7EFFB517AC06F409C8FD67" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517AC06F4097EFD43" bold="true" box="[151,328,680,706]" pageId="12" pageNumber="187">Type material.</emphasis>
<materialsCitation id="3B793CA0DF7EFFB5166A06F40CDBFD43" ID-GBIF-Occurrence="3864839325" box="[337,1261,680,707]" collectionCode="NMNS" country="Taiwan Island" location="Taitung" municipality="Taimali" pageId="12" pageNumber="187" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B965EAEFDF7EFFB5166A06F409F1FD43" bold="true" box="[337,455,680,706]" pageId="12" pageNumber="187">
<typeStatus id="54AA885FDF7EFFB5166A06F409F4FD43" box="[337,450,680,706]" pageId="12" pageNumber="187" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D17FD74DF7EFFB516EA06F509D0FD42" box="[465,486,681,707]" pageId="12" pageNumber="187" type="male">♂</specimenCount>
(
<collectionCode id="ED00AE38DF7EFFB516CC06F40A7BFD43" box="[503,589,680,706]" country="Taiwan" name="National Museum of Natural Science" pageId="12" pageNumber="187">NMNS</collectionCode>
),
<collectingCountry id="F306766DDF7EFFB5155806F40B33FD43" box="[611,773,680,706]" name="Taiwan Island" pageId="12" pageNumber="187">Taiwan Island</collectingCountry>
,
<location id="8ECE6026DF7EFFB5142806F40B5DFD43" LSID="urn:lsid:plazi:treatment:03B887EBDF7EFFA917AC06670A4DFE9C:8ECE6026DF7EFFB5142806F40B5DFD43" box="[787,875,680,706]" country="Taiwan Island" municipality="Taimali" name="Taitung" pageId="12" pageNumber="187">Taitung</location>
,
<collectingMunicipality id="6BCAAC87DF7EFFB5144306F40BFBFD43" box="[888,973,680,706]" pageId="12" pageNumber="187">Taimali</collectingMunicipality>
[=
<emphasis id="B965EAEFDF7EFFB514C206F50C7BFD41" bold="true" box="[1017,1101,681,704]" pageId="12" pageNumber="187">kDzae</emphasis>
],
<date id="FFAF103DDF7EFFB5135C06F40CDBFD43" box="[1127,1261,680,706]" pageId="12" pageNumber="187" value="2008-04-12">2008-IV-12</date>
</materialsCitation>
.
<materialsCitation id="3B793CA0DF7EFFB513C206F409CCFD67" ID-GBIF-Occurrence="3864839327" collectionCode="CCPC" country="Taiwan Island" location="Taitung" municipality="Taimali" pageId="12" pageNumber="187" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B965EAEFDF7EFFB513C206F40D46FD43" bold="true" box="[1273,1392,680,706]" pageId="12" pageNumber="187">
<typeStatus id="54AA885FDF7EFFB513C206F40D5CFD43" box="[1273,1386,680,706]" pageId="12" pageNumber="187" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D17FD74DF7EFFB5124206F40DAAFD43" box="[1401,1436,680,706]" pageId="12" pageNumber="187" type="female">1♀</specimenCount>
(
<collectionCode id="ED00AE38DF7EFFB517A4069008DAFD67" box="[159,236,716,742]" pageId="12" pageNumber="187">CCPC</collectionCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF7EFFB617FC06AC0D2BF879" lastPageId="15" lastPageNumber="190" pageId="12" pageNumber="187" type="description">
<paragraph id="8BAE36FDDF7EFFB517FC06AC0DA7FC8A" blockId="12.[151,1437,680,2039]" box="[199,1425,752,779]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC06AC0A38FC8B" bold="true" box="[199,526,752,778]" pageId="12" pageNumber="187">
Description.
<typeStatus id="54AA885FDF7EFFB5165B06AC09FBFC8B" box="[352,461,752,778]" pageId="12" pageNumber="187" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="132A2A78DF7EFFB5152706AC0A97FC8B" box="[540,673,752,778]" captionStart="FIGURE 7" captionStartId="13.[151,250,1974,1998]" captionTargetBox="[181,1411,404,1936]" captionTargetId="figure-106@13.[158,1430,381,1950]" captionTargetPageId="13" captionText="FIGURE 7. Habitus of Anomala inclinata Zhao &amp; Zorn, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877281" httpUri="https://zenodo.org/record/6877281/files/figure.png" pageId="12" pageNumber="187">Figs. 7A–C</figureCitation>
,
<figureCitation id="132A2A78DF7EFFB5159706AC0AE7FC8B" box="[684,721,752,778]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="12" pageNumber="187">8G</figureCitation>
,
<figureCitation id="132A2A78DF7EFFB515E606AC0B51FC8B" box="[733,871,752,778]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="12" pageNumber="187">9H–J, 9P–Q</figureCitation>
). Body length: 13.0 mm, greatest width:
<quantity id="4CE99B18DF7EFFB5120F06AC0DBBFC8A" box="[1332,1421,752,779]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="12" pageNumber="187" unit="mm" value="7.3">7.3 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC07480976FCD3" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC074809F6FCAF" bold="true" box="[199,448,788,814]" pageId="12" pageNumber="187">General appearance.</emphasis>
Body shape elongated ovoid, weakly convex. Entirely blackish brown, with weak green metallic sheen.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC0700097BFB86" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC07000931FCF7" bold="true" box="[199,263,860,886]" pageId="12" pageNumber="187">Head</emphasis>
(
<figureCitation id="132A2A78DF7EFFB5162C07000959FCF7" box="[279,367,860,886]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="12" pageNumber="187">Fig. 8G</figureCitation>
). Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin somewhat rounded and strongly reflexed; surface densely rugopunctate. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons densely rugopunctate, other portions of frons and vertex with irregularly distributed, dense small punctures. Antennal club longer than antennomeres 2–6 combined. Inner margin of eye with several moderately long setae.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC004C0B2DFB16" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC004C0971FBAB" bold="true" box="[199,327,1040,1066]" pageId="12" pageNumber="187">Pronotum.</emphasis>
Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle round. Basal marginal line interrupted before scutellum, other marginal lines complete. With dense small punctures, punctures being coalescent at marginal portions, impunctate before scutellum; with a longitudinal medial furrow not reaching the base. Lateral margin with several long setae.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC00FC0B53FB5E" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC00FC0970FB3B" bold="true" box="[199,326,1184,1210]" pageId="12" pageNumber="187">Scutellum.</emphasis>
Subtriangular, lateral margin arched. Margins with polished appearance, disc with irregularly distributed coarse punctures. Disc with some semierect short setae.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC00B40A83FA5B" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC00B4092CFA83" bold="true" box="[199,282,1256,1282]" pageId="12" pageNumber="187">Elytra.</emphasis>
Intervals strongly convex; strial punctures dense to very dense, annulated and large, punctures of secondary striae unequal in spaces; the whole surface with scattered minute punctures. Interstices I to III each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria shortly doubled in basal third; all secondary striae reaching the level of apical protuberance and usually interrupted, secondary stria of interstice II obsolete from the level of scutellar apex to apical two fifth. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina indistinct. Epipleura with a row of moderately dense long setae. Marginal membrane complete.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC01B80D67FA7F" blockId="12.[151,1437,680,2039]" box="[199,1361,1508,1535]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC01B8095DFA7F" bold="true" box="[199,363,1508,1534]" pageId="12" pageNumber="187">Propygidium.</emphasis>
With dense transverse punctures, usually coalescent into transverse striolation; glabrous.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC025409D1F9EA" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC02540974F9A3" bold="true" box="[199,322,1544,1570]" pageId="12" pageNumber="187">Pygidium.</emphasis>
Distinctly bulging, posterior margin protruding. Disc with dense ovoid large punctures, punctures more transverse near all margins. With several short setae distributed along posterior margin, posterior margin with a row of long setae near apex.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC02280D41F97B" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC022809D8F90F" bold="true" box="[199,494,1652,1678]" pageId="12" pageNumber="187">Ventral thoracic surface.</emphasis>
Hypomeron with dense coalescent striolation and moderately dense long setae. Collar of mesosternite transversely striolate with dense recumbent short setae. Other portions of mesosternite with small punctures and somewhat rugopunctate, with dense short setae. Ventral metathoracic surface with dense annulated small punctures and dense, rather long setae at each side; with minute punctures and glabrous medially.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB517FC03580A50F80B" blockId="12.[151,1437,680,2039]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC035809F1F89F" bold="true" box="[199,455,1796,1822]" pageId="12" pageNumber="187">Abdominal ventrites.</emphasis>
Ventrites 1–2 and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with irregularly distributed small punctures, punctures coalescent into striolation in ventrite 6. Ventrites 2–5 each with a transverse row of sparse and moderately long setae, interrupted medially in ventrites 3–4; ventrite 6 with a complete row of long setae along posterior margin.
</paragraph>
<paragraph id="8BAE36FDDF7EFFB417FC03C80920FEE4" blockId="12.[151,1437,680,2039]" lastBlockId="13.[151,1436,150,357]" lastPageId="13" lastPageNumber="188" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC03C80930F82F" bold="true" box="[199,262,1940,1966]" pageId="12" pageNumber="187">Legs.</emphasis>
Protibia tridentate, apical and second teeth acute at apices, proximal tooth small; apical tooth extending to level of halfway of protarsomere 2. Inner spur inserted at the level indention between second and proximal tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches longer and wider, the lower branch of inner protarsal claw concave internobasally. Each tarsomere 5 with an internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows, one at anterior margin and another situated behind middle, the row at posterior margin usually with shorter setae. Metafemoral surface with three transverse rows of sparse long setae, the first row at anterior margin, a second row situated behind middle with robuster setae, and a third row at posterior margin with short setae.
</paragraph>
<caption id="DF6E6675DF7FFFB417AC03EA0AF9F870" ID-DOI="http://doi.org/10.5281/zenodo.6877281" ID-Zenodo-Dep="6877281" httpUri="https://zenodo.org/record/6877281/files/figure.png" pageId="13" pageNumber="188" startId="13.[151,250,1974,1998]" targetBox="[181,1411,404,1936]" targetPageId="13">
<paragraph id="8BAE36FDDF7FFFB417AC03EA0AF9F870" blockId="13.[151,1436,1974,2034]" pageId="13" pageNumber="188">
<emphasis id="B965EAEFDF7FFFB417AC03EA0924F84F" bold="true" box="[151,274,1974,1998]" pageId="13" pageNumber="188">FIGURE 7.</emphasis>
Habitus of
<taxonomicName id="4C114D7EDF7FFFB416BC03EA0AF9F84C" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[391,719,1974,1998]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="188" phylum="Arthropoda" rank="species" species="inclinata" status="sp. nov.">
<emphasis id="B965EAEFDF7FFFB416BC03EA0A74F84F" box="[391,578,1974,1998]" italics="true" pageId="13" pageNumber="188">Anomala inclinata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7FFFB415E203EA0B63F84F" bold="true" box="[729,853,1974,1998]" pageId="13" pageNumber="188">
<taxonomicNameLabel id="A2565794DF7FFFB415E203EA0B63F84F" box="[729,853,1974,1998]" pageId="13" pageNumber="188" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="DF6E6675DF7CFFB717AC030508EFF85C" ID-DOI="http://doi.org/10.5281/zenodo.6877285" ID-Zenodo-Dep="6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="14" pageNumber="189" startId="14.[151,250,1881,1905]" targetBox="[151,1426,181,1857]" targetPageId="14">
<paragraph id="8BAE36FDDF7CFFB717AC030508EFF85C" blockId="14.[151,1437,1881,2013]" pageId="14" pageNumber="189">
<emphasis id="B965EAEFDF7CFFB717AC03050923F8F0" bold="true" box="[151,277,1881,1905]" pageId="14" pageNumber="189">FIGURE 8.</emphasis>
Head of
<taxonomicName id="4C114D7EDF7CFFB7164203050A17F8F0" box="[377,545,1881,1905]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B965EAEFDF7CFFB71642030509E2F8F0" box="[377,468,1881,1905]" italics="true" pageId="14" pageNumber="189">Anomala</emphasis>
spp. A
</taxonomicName>
,
<taxonomicName id="4C114D7EDF7CFFB7151503060B63F8F0" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[558,853,1881,1905]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="linwenhsini" status="sp. nov.">
<emphasis id="B965EAEFDF7CFFB7151503060AF7F8F0" box="[558,705,1881,1905]" italics="true" pageId="14" pageNumber="189">A. linwenhsini</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7CFFB7145A03050BD6F8F0" bold="true" box="[865,992,1881,1905]" pageId="14" pageNumber="189">
<taxonomicNameLabel id="A2565794DF7CFFB7145A03050BD6F8F0" box="[865,992,1881,1905]" pageId="14" pageNumber="189" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; B, ditto, female paratype; C,
<taxonomicName id="4C114D7EDF7CFFB712BD030609A1F814" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="wutaiensis" status="sp. nov.">
<emphasis id="B965EAEFDF7CFFB712BD03060934F814" italics="true" pageId="14" pageNumber="189">A. wutaiensis</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7CFFB7169803210A14F814" bold="true" box="[419,546,1917,1941]" pageId="14" pageNumber="189">
<taxonomicNameLabel id="A2565794DF7CFFB7169803210A14F814" box="[419,546,1917,1941]" pageId="14" pageNumber="189" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; D, ditto, female paratype; E,
<taxonomicName id="4C114D7EDF7CFFB714F303220D3AF814" authority="Zorn &amp; Zhao" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[968,1292,1917,1941]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="kanshireiensis" status="sp. nov.">
<emphasis id="B965EAEFDF7CFFB714F303220C41F814" box="[968,1143,1917,1941]" italics="true" pageId="14" pageNumber="189">A. kanshireiensis</emphasis>
Zorn &amp; Zhao
</taxonomicName>
,
<emphasis id="B965EAEFDF7CFFB7122C03210DA0F814" bold="true" box="[1303,1430,1917,1941]" pageId="14" pageNumber="189">
<taxonomicNameLabel id="A2565794DF7CFFB7122C03210DA0F814" box="[1303,1430,1917,1941]" pageId="14" pageNumber="189" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; F, ditto, female paratype; G,
<taxonomicName id="4C114D7EDF7CFFB7151A03FE0B10F838" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[545,806,1953,1977]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="inclinata" status="sp. nov.">
<emphasis id="B965EAEFDF7CFFB7151A03FE0AA1F838" box="[545,663,1953,1977]" italics="true" pageId="14" pageNumber="189">A. inclinata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7CFFB7140B03FD0B9BF838" bold="true" box="[816,941,1953,1977]" pageId="14" pageNumber="189">
<taxonomicNameLabel id="A2565794DF7CFFB7140B03FD0B9BF838" box="[816,941,1953,1977]" pageId="14" pageNumber="189" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; H, ditto, female paratype. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="DF6E6675DF7DFFB617AC027308F0F956" ID-DOI="http://doi.org/10.5281/zenodo.6877289" ID-Zenodo-Dep="6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="15" pageNumber="190" startId="15.[151,250,1583,1607]" targetBox="[163,1411,190,1552]" targetPageId="15">
<paragraph id="8BAE36FDDF7DFFB617AC027308F0F956" blockId="15.[151,1437,1583,1751]" pageId="15" pageNumber="190">
<emphasis id="B965EAEFDF7DFFB617AC02730922F9C6" bold="true" box="[151,276,1583,1607]" pageId="15" pageNumber="190">FIGURE 9.</emphasis>
Male genitalia (A–M) and everted endophallus (N–Q) of
<taxonomicName id="4C114D7EDF7DFFB61455026C0C22F9C9" box="[878,1044,1584,1608]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B965EAEFDF7DFFB61455026C0BFFF9C9" box="[878,969,1584,1608]" italics="true" pageId="15" pageNumber="190">Anomala</emphasis>
spp. A
</taxonomicName>
–C,
<taxonomicName id="4C114D7EDF7DFFB61304026C0D53F9C6" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[1087,1381,1584,1608]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="linwenhsini" status="sp. nov.">
<emphasis id="B965EAEFDF7DFFB61304026C0CE4F9C9" box="[1087,1234,1584,1608]" italics="true" pageId="15" pageNumber="190">A. linwenhsini</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7DFFB6124A026C08D4F9EA" bold="true" pageId="15" pageNumber="190">
<taxonomicNameLabel id="A2565794DF7DFFB6124A026C08D4F9EA" pageId="15" pageNumber="190" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; D–G,
<taxonomicName id="4C114D7EDF7DFFB616AD02080A87F9EA" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[406,689,1620,1644]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="wutaiensis" status="sp. nov.">
<emphasis id="B965EAEFDF7DFFB616AD02080A29F9EA" box="[406,543,1620,1643]" italics="true" pageId="15" pageNumber="190">A. wutaiensis</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7DFFB6158702080B0DF9EA" bold="true" box="[700,827,1619,1644]" pageId="15" pageNumber="190">
<taxonomicNameLabel id="A2565794DF7DFFB6158702080B0DF9EA" box="[700,827,1619,1644]" pageId="15" pageNumber="190" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; H–J, N–O,
<taxonomicName id="4C114D7EDF7DFFB6131E02080D50F9ED" authority="Zorn &amp; Zhao" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[1061,1382,1620,1644]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="kanshireiensis" status="sp. nov.">
<emphasis id="B965EAEFDF7DFFB6131E02080CE5F9ED" box="[1061,1235,1620,1644]" italics="true" pageId="15" pageNumber="190">A. kanshireiensis</emphasis>
Zorn &amp; Zhao
</taxonomicName>
,
<emphasis id="B965EAEFDF7DFFB6124A020808D4F90E" bold="true" pageId="15" pageNumber="190">
<taxonomicNameLabel id="A2565794DF7DFFB6124A020808D4F90E" pageId="15" pageNumber="190" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; K–M, P–Q,
<taxonomicName id="4C114D7EDF7DFFB616EA02240AE0F90E" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[465,726,1656,1680]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="inclinata" status="sp. nov.">
<emphasis id="B965EAEFDF7DFFB616EA02240A71F911" box="[465,583,1656,1680]" italics="true" pageId="15" pageNumber="190">A. inclinata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7DFFB615DB02240B6BF90E" bold="true" box="[736,861,1655,1680]" pageId="15" pageNumber="190">
<taxonomicNameLabel id="A2565794DF7DFFB615DB02240B6BF90E" box="[736,861,1655,1680]" pageId="15" pageNumber="190" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm.
</paragraph>
</caption>
<paragraph id="8BAE36FDDF7DFFB617FC035A0DADF8A1" blockId="15.[151,1437,1797,2040]" box="[199,1435,1797,1824]" pageId="15" pageNumber="190">
<emphasis id="B965EAEFDF7DFFB617FC035A094FF89E" bold="true" box="[199,377,1797,1824]" pageId="15" pageNumber="190">Male genitalia.</emphasis>
See
<figureCitation id="132A2A78DF7DFFB61694035A0A19F8A1" box="[431,559,1798,1824]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="15" pageNumber="190">Fig. 9K–M</figureCitation>
. Parameres asymmetric, right paramere longer. Ventral plate flat, apex arched.
</paragraph>
<paragraph id="8BAE36FDDF7DFFB617FC03760D2BF879" blockId="15.[151,1437,1797,2040]" pageId="15" pageNumber="190">
<emphasis id="B965EAEFDF7DFFB617FC037609AEF8C5" bold="true" box="[199,408,1834,1860]" pageId="15" pageNumber="190">
<typeStatus id="54AA885FDF7DFFB617FC03760900F8C5" box="[199,310,1834,1860]" pageId="15" pageNumber="190" type="paratype">Paratype</typeStatus>
. Female
</emphasis>
(
<figureCitation id="132A2A78DF7DFFB6169C03760A13F8C5" box="[423,549,1834,1860]" captionStart="FIGURE 7" captionStartId="13.[151,250,1974,1998]" captionTargetBox="[181,1411,404,1936]" captionTargetId="figure-106@13.[158,1430,381,1950]" captionTargetPageId="13" captionText="FIGURE 7. Habitus of Anomala inclinata Zhao &amp; Zorn, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877281" httpUri="https://zenodo.org/record/6877281/files/figure.png" pageId="15" pageNumber="190">Figs. 7D–F</figureCitation>
,
<figureCitation id="132A2A78DF7DFFB6151403760A64F8C5" box="[559,594,1834,1860]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="15" pageNumber="190">8H</figureCitation>
). Body length:
<quantity id="4CE99B18DF7DFFB615C403760B52F8C5" box="[767,868,1834,1861]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="15" pageNumber="190" unit="mm" value="13.5">13.5 mm</quantity>
, greatest width:
<quantity id="4CE99B18DF7DFFB6132203760C47F8C5" box="[1049,1137,1834,1860]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="15" pageNumber="190" unit="mm" value="6.9">6.9 mm</quantity>
. Generally similar to male. Antennal club almost equally as long as antennomeres 2–6 combined. Pygidium more bulging, shorter than in male. Protibia and protarsomeres thinner than in male; apical protibial tooth tongue-shaped, almost extends to level of apex of protarsomere 2; other two teeth blunt at apex, the third one indistinct; inner spur situated at the level of the third tooth; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length. Ventral metathoracic surface less setose than in male.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF62FFA917FC04CA0D6DFF30" box="[199,1371,150,177]" pageId="16" pageNumber="191" type="diagnosis">
<paragraph id="8BAE36FDDF62FFA917FC04CA0D6DFF30" blockId="16.[151,1436,150,285]" box="[199,1371,150,177]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC04CA09F8FF31" bold="true" box="[199,462,150,176]" pageId="16" pageNumber="191">Differential diagnosis.</emphasis>
See differential diagnosis of
<taxonomicName id="4C114D7EDF62FFA9142D04CB0C88FF30" authority="Zorn &amp; Zhao" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[790,1214,151,177]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="191" phylum="Arthropoda" rank="species" species="kanshireiensis" status="sp. nov.">
<emphasis id="B965EAEFDF62FFA9142D04CB0C17FF30" box="[790,1057,151,177]" italics="true" pageId="16" pageNumber="191">Anomala kanshireiensis</emphasis>
Zorn &amp; Zhao
</taxonomicName>
,
<emphasis id="B965EAEFDF62FFA913F104CB0D65FF31" bold="true" box="[1226,1363,150,177]" pageId="16" pageNumber="191">
<taxonomicNameLabel id="A2565794DF62FFA913F104CB0D65FF31" box="[1226,1363,150,177]" pageId="16" pageNumber="191" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF62FFA917FC04E7098EFF78" pageId="16" pageNumber="191" type="etymology">
<paragraph id="8BAE36FDDF62FFA917FC04E7098EFF78" blockId="16.[151,1436,150,285]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC04E70978FF54" bold="true" box="[199,334,187,213]" pageId="16" pageNumber="191">Etymology.</emphasis>
The specific epithet derives from the Latin adjective “
<emphasis id="B965EAEFDF62FFA9148A04E70C48FF55" box="[945,1150,187,213]" italics="true" pageId="16" pageNumber="191">inclinatus, -a, -um</emphasis>
”, alluding to the inclined parameres of this species.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF62FFA917FC055E0A4DFE9C" box="[199,635,258,285]" pageId="16" pageNumber="191" type="distribution">
<paragraph id="8BAE36FDDF62FFA917FC055E0A4DFE9C" blockId="16.[151,1436,150,285]" box="[199,635,258,285]" pageId="16" pageNumber="191">
<emphasis id="B965EAEFDF62FFA917FC055E0957FE9D" bold="true" box="[199,353,258,284]" pageId="16" pageNumber="191">Distribution.</emphasis>
<collectingCountry id="F306766DDF62FFA9165C055F0A32FE9C" box="[359,516,259,285]" name="Taiwan Island" pageId="16" pageNumber="191">Taiwan Island</collectingCountry>
(Taitung).
</paragraph>
</subSubSection>
</treatment>
</document>
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