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<document id="51A86EAC108DE1CBAB31EE07061D2306" ID-DOI="10.26107/RBZ-2022-0023" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174433" ID-ZooBank="E4F49C16-DB5E-4593-AB8E-03D8859A59D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665290687137" checkinUser="felipe" docAuthor="Páll-Gergely, Barna &amp; Hunyadi, András" docDate="2022" docId="03B887D7230F2671AE8637272715FEF2" docLanguage="en" docName="RafflesBZool.70.417439.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Clostophis thinbowguensis Páll-Gergely &amp; Hunyadi 2022, new species" docType="treatment" docVersion="4" lastPageNumber="427" masterDocId="FF81FFAF2305267BAE6F347A2327FFA7" masterDocTitle="New and little-known species of Clostophis Benson, 1860 from Southeast Asia (Gastropoda: Eupulmonata: Hypselostomatidae)" masterLastPageNumber="439" masterPageNumber="417" pageNumber="427" updateTime="1699379781661" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B8A8C6DBAA4D5D80807D791A1312F1B6">New and little-known species of Clostophis Benson, 1860 from Southeast Asia (Gastropoda: Eupulmonata: Hypselostomatidae)</mods:title>
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<mods:name id="685940833D6D436F23CC41EB302D0AA7" type="personal">
<mods:role id="9D0BFE8A79377E307632387C165B9CFE">
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<mods:namePart id="94563B57AB6599F0F6E4A6375D1AB875">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="1C39A7489E0EF5D5B21B3D3B2DF6DA1C">Hunyadi, András</mods:namePart>
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<mods:typeOfResource id="2DFA2D30E2D3BA0986FE318A21D3A3BC">text</mods:typeOfResource>
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<mods:title id="D92C79EBF4F2AAA5C9B60020CB084F50">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="FAD241F164D7406150C489414508E8C4">
<mods:date id="F6A5BD3616B3B858595B8DCCDCD0BAED">2022</mods:date>
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<mods:number id="EBA6AEA4CE8D79DC82931F6D821F68C1">2022-09-09</mods:number>
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<mods:number id="D4BAA46C8CA7FA1FCCDCD84943834704">70</mods:number>
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<mods:identifier id="63F8E04F88BD13BAF4BB93523E3688A9" type="DOI">10.26107/RBZ-2022-0023</mods:identifier>
<mods:identifier id="1D0EB9CC0FDEF1F416CEA4F573712B8B" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03B887D7230F2671AE8637272715FEF2" ID-DOI="http://doi.org/10.5281/zenodo.7171123" ID-Zenodo-Dep="7171123" LSID="urn:lsid:plazi:treatment:03B887D7230F2671AE8637272715FEF2" httpUri="http://treatment.plazi.org/id/03B887D7230F2671AE8637272715FEF2" lastPageNumber="427" pageId="10" pageNumber="427">
<subSubSection id="C30B654A230F2671AE8637272181FCD2" box="[233,678,861,885]" pageId="10" pageNumber="427" type="nomenclature">
<paragraph id="8BAE36C1230F2671AE8637272181FCD2" blockId="10.[233,678,861,917]" box="[233,678,861,885]" pageId="10" pageNumber="427">
<heading id="D0E681AD230F2671AE8637272181FCD2" bold="true" box="[233,678,861,885]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="427" reason="2">
<emphasis id="B965EAD3230F2671AE8637272181FCD2" bold="true" box="[233,678,861,885]" pageId="10" pageNumber="427">
<taxonomicName id="4C114D42230F2671AE8637272136FCD2" ID-CoL="f8afcf90-4b8f-4c6b-a454-ac24eaaf7a30" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[233,529,861,885]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="species" species="thinbowguensis" status="sp. nov.">
<emphasis id="B965EAD3230F2671AE8637272136FCD2" bold="true" box="[233,529,861,885]" italics="true" pageId="10" pageNumber="427">Clostophis thinbowguensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A8230F2671AC4E37272181FCD2" box="[545,678,861,885]" pageId="10" pageNumber="427" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30B654A230F2671AFF1370722D7FC32" box="[414,496,893,917]" pageId="10" pageNumber="427" type="description">
<paragraph id="8BAE36C1230F2671AFF1370722D7FC32" blockId="10.[233,678,861,917]" box="[414,496,893,917]" pageId="10" pageNumber="427">
(
<figureCitation id="132A2A44230F2671AFC9370722CFFC32" box="[422,488,893,917]" captionStart="Fig" captionStartId="11.[128,163,2093,2115]" captionTargetBox="[129,1436,189,2077]" captionTargetId="figure-19@11.[129,1436,189,2077]" captionTargetPageId="11" captionText="Fig. 9. Clostophis thinbowguensis, new species (holotype, MNHN-IM-2000-38057)." figureDoi="http://doi.org/10.5281/zenodo.7174453" httpUri="https://zenodo.org/record/7174453/files/figure.png" pageId="10" pageNumber="427">Fig. 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B654A230F2671AEE737C721A8FB92" pageId="10" pageNumber="427" type="materials_examined">
<paragraph id="8BAE36C1230F2671AEE737C721A8FB92" blockId="10.[136,775,957,1077]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE737C72211FC72" bold="true" box="[136,310,957,981]" pageId="10" pageNumber="427">Type material.</emphasis>
<materialsCitation id="3B793C9C230F2671AF2F37C721A8FB92" collectingDate="2015-02-20" collectorName="F. Brehier" country="Myanmar" latitude="11.189667" location="Phayahran Camp" longLatPrecision="13" longitude="99.17167" municipality="Thin Bow Gu Cave" pageId="10" pageNumber="427" specimenCount="1" stateProvince="Taninthayi" typeStatus="holotype">
<typeStatus id="54AA8863230F2671AF2F37C72282FC72" box="[320,421,957,981]" pageId="10" pageNumber="427" type="holotype">Holotype</typeStatus>
(SH:
<quantity id="4CE99B24230F2671AF8337C72175FC72" box="[492,594,957,981]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="427" unit="mm" value="1.27">1.27 mm</quantity>
, SW:
<quantity id="4CE99B24230F2671ACF537C721D8FC72" box="[666,767,957,981]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="427" unit="mm" value="1.25">1.25 mm</quantity>
) (MNHN-IM-2000-38057),
<collectingCountry id="F3067651230F2671AFDD37A4213AFC51" box="[434,541,990,1014]" name="Myanmar" pageId="10" pageNumber="427">Myanmar</collectingCountry>
,
<collectingRegion id="49D5F823230F2671AC4437A72024FC52" box="[555,771,989,1013]" country="Myanmar" name="Taninthayi" pageId="10" pageNumber="427">Tanintharyi Region</collectingRegion>
,
<collectingMunicipality id="6BCAACBB230F2671AEE737872259FBB2" box="[136,382,1021,1045]" pageId="10" pageNumber="427">Thin Bow Gu Cave</collectingMunicipality>
,
<location id="8ECE601A230F2671AFFF37872143FBB2" LSID="urn:lsid:plazi:treatment:03B887D7230F2671AE8637272715FEF2:8ECE601A230F2671AFFF37872143FBB2" box="[400,612,1021,1045]" country="Myanmar" latitude="11.189667" longLatPrecision="13" longitude="99.17167" municipality="Thin Bow Gu Cave" name="Phayahran Camp" pageId="10" pageNumber="427" stateProvince="Taninthayi">Phayahran Camp</location>
;
<geoCoordinate id="EE255006230F2671AC1937862026FBB2" box="[630,769,1020,1045]" degrees="11" direction="north" minutes="11.38" orientation="latitude" pageId="10" pageNumber="427" precision="9" value="11.189667">11°11.38N</geoCoordinate>
,
<geoCoordinate id="EE255006230F2671AEE530662223FB92" box="[138,260,1052,1077]" degrees="99" direction="east" minutes="10.30" orientation="longitude" pageId="10" pageNumber="427" precision="9" value="99.17167">99°10.30E</geoCoordinate>
; coll.
<collectorName id="26E45317230F2671AF263064229DFB92" box="[329,442,1053,1077]" pageId="10" pageNumber="427">F. Bréhier</collectorName>
,
<date id="FFAF1001230F2671AFA9306721ACFB92" box="[454,651,1053,1077]" pageId="10" pageNumber="427" value="2015-02-20">
<collectingDate id="EFEBE9E9230F2671AFA9306721ACFB92" box="[454,651,1053,1077]" pageId="10" pageNumber="427" value="2015-02-20">20 February 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B654A230F2671AEE73027217AFB52" pageId="10" pageNumber="427" type="diagnosis">
<paragraph id="8BAE36C1230F2671AEE73027217AFB52" blockId="10.[136,777,1117,1269]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE7302723DAFBD2" bold="true" box="[136,253,1117,1141]" pageId="10" pageNumber="427">Diagnosis.</emphasis>
Among species of
<taxonomicName id="4C114D42230F2671AFBA30272160FBD2" authorityName="Benson" authorityYear="1860" box="[469,583,1117,1141]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="genus">
<emphasis id="B965EAD3230F2671AFBA30272160FBD2" box="[469,583,1117,1141]" italics="true" pageId="10" pageNumber="427">Clostophis</emphasis>
</taxonomicName>
with the last part of the penultimate whorl attached to the penultimate whorl,
<taxonomicName id="4C114D42230F2671AEE630E4226AFB12" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[137,333,1181,1205]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="species" species="thinbowguensis" status="sp. nov.">
<emphasis id="B965EAD3230F2671AEE630E4226AFB12" box="[137,333,1181,1205]" italics="true" pageId="10" pageNumber="427">C. thinbowguensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A8230F2671AF3430E422F9FB12" box="[347,478,1181,1205]" pageId="10" pageNumber="427" rank="species">new species</taxonomicNameLabel>
, differs by the incurved last part of the last whorl, in combination with the presence of a single tooth (a parietalis) in the aperture.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A230F2671AEE731672020F892" pageId="10" pageNumber="427" type="description">
<paragraph id="8BAE36C1230F2671AEE7316722A9F952" blockId="10.[135,776,1309,1781]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE731672234FA92" bold="true" box="[136,275,1309,1333]" pageId="10" pageNumber="427">Description.</emphasis>
Shell conical, width and height equal or almost equal. Whorls rounded. Last whorl not detached from penultimate whorl, although aperture slightly protruding (parietal callus elevated from penultimate whorl). Protoconch consisting of slightly less than 1.5 whorls, its sculpture is not discernible due to corrosion of shell surface. Entire shell with 4.75 whorls. Umbilicus normally wide, ca. one third of shell width. Entire teleoconch corroded, except for the characteristic presence of dense spiral striation on the apertural and ventral side of the body whorl (counting them is not possible). Some weak, irregular radial growth lines are present. Aperture subcircular, sinulus wide and weakly isolated due to low, deeply situated parietal tooth. No additional apertural barriers present. Peristome slightly expanded, not reflected.
</paragraph>
<paragraph id="8BAE36C1230F2671AEE733672020F892" blockId="10.[136,775,1821,1845]" box="[136,775,1821,1845]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE7336722BFF892" bold="true" box="[136,408,1821,1845]" pageId="10" pageNumber="427">Measurements (in mm).</emphasis>
SH = 1.27, SW = 1.25 (
<typeStatus id="54AA8863230F2671ACF7336721DBF892" box="[664,764,1821,1845]" pageId="10" pageNumber="427" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B654A230F2671AEE73327260CFF52" pageId="10" pageNumber="427" type="diagnosis">
<paragraph id="8BAE36C1230F2671AEE73327260CFF52" blockId="10.[136,777,1885,2037]" lastBlockId="10.[835,1474,189,245]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE7332722B7F8D2" bold="true" box="[136,400,1885,1909]" pageId="10" pageNumber="427">Differential diagnosis.</emphasis>
<taxonomicName id="4C114D42230F2671AFF4332721EFF8D2" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[411,712,1885,1909]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="species" species="thinbowguensis" status="sp. nov.">
<emphasis id="B965EAD3230F2671AFF4332721EFF8D2" box="[411,712,1885,1909]" italics="true" pageId="10" pageNumber="427">Clostophis thinbowguensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A8230F2671ACB6332423FFF832" pageId="10" pageNumber="427" rank="species">new species</taxonomicNameLabel>
, is similar to some
<taxonomicName id="4C114D42230F2671AFDD330721ADF832" box="[434,650,1917,1941]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AFDD33072114F832" box="[434,563,1917,1941]" italics="true" pageId="10" pageNumber="427">Angustopila</emphasis>
species
</taxonomicName>
, but differs from them by the generally larger size and the denser spiral striation. The most similar species is probably
<taxonomicName id="4C114D42230F2671ACE733C723C5F852" baseAuthorityName="F. G. Thompson &amp; Upatham" baseAuthorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="species" species="concava">
<emphasis id="B965EAD3230F2671ACE733C723C5F852" italics="true" pageId="10" pageNumber="427">Angustopila concava</emphasis>
</taxonomicName>
due to the similar strongly oblique aperture, but that species is smaller, has a concave conical shell, and a more anteriorly situated lower parietal tooth.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A230F2671AD2C35672715FEF2" pageId="10" pageNumber="427" type="etymology">
<paragraph id="8BAE36C1230F2671AD2C35672715FEF2" blockId="10.[834,1474,285,341]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AD2C356720E2FE92" bold="true" box="[835,965,285,309]" pageId="10" pageNumber="427">Etymology.</emphasis>
This new species is named after its
<typeStatus id="54AA8863230F2671AB513564264BFE91" box="[1342,1388,286,310]" pageId="10" pageNumber="427">type</typeStatus>
locality (Thin Bow Gu Cave).
</paragraph>
</subSubSection>
</treatment>
</document>