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<document id="1E4427A1490AB9CC2828929FCD2D9E02" ID-CLB-Dataset="300725" ID-DOI="10.11646/phytotaxa.650.2.7" ID-GBIF-Dataset="ccaf79df-e5e1-4e3f-a9fb-06f3f306daf2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13216366" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722602792570" checkinUser="felipe" docAuthor="Lee, Gaik Ee &amp; He, Xiaolan" docDate="2024" docId="8F217579FB689D6551B5FDEAFC9ECFC4" docLanguage="en" docName="phytotaxa.650.2.7.pdf" docOrigin="Phytotaxa 650 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.650.2.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Lejeunea timokoponenii G. E. Lee &amp; X. L. He, sp. nov." docType="treatment" docVersion="2" lastPageNumber="197" masterDocId="73180D01FB699D66513DFF9EFFBDCD49" masterDocTitle="A new species of Lejeunea (Lejeuneaceae) from Papua New Guinea" masterLastPageNumber="198" masterPageNumber="194" pageNumber="195" updateTime="1722809251276" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="102D27C6E7D62A7DD8D0259DFF79FDBE">A new species of Lejeunea (Lejeuneaceae) from Papua New Guinea</mods:title>
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<mods:namePart id="42A24866AC52E63BC52C7A1552BF69DB">Lee, Gaik Ee</mods:namePart>
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<mods:affiliation id="F1DB21BEB935A7091D9CF98CD4217D66">Gaik Ee Lee, Institute of Tropical Biodiversity and Sustainable Development, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia.</mods:affiliation>
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<mods:namePart id="2EF0194F02FF5DA8F3357991D9128899">He, Xiaolan</mods:namePart>
<mods:nameIdentifier id="2F7307FE63A34080407F240660BEB9A4" type="ORCID">0000-0002-7754-2233</mods:nameIdentifier>
<mods:affiliation id="859A2CCB21F77D68461B3E52386D87DD">Xiaolan He, Botany Unit, Finnish Museum of Natural History, P. O. Box 7. 00014 University of Helsinki, Finland.</mods:affiliation>
<mods:nameIdentifier id="9760235772158D6B9CB7C69DBB79F0AB" type="email">xiao-lan.he@helsinki.fi</mods:nameIdentifier>
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<mods:title id="2464383DCE54C50DD0DE2FF14766D000">Phytotaxa</mods:title>
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<mods:date id="08E84DE434ED00C746C5C8E1B0A95FE5">2024</mods:date>
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<treatment id="8F217579FB689D6551B5FDEAFC9ECFC4" ID-DOI="http://doi.org/10.5281/zenodo.13214569" ID-GBIF-Taxon="237652090" ID-Zenodo-Dep="13214569" LSID="urn:lsid:plazi:treatment:8F217579FB689D6551B5FDEAFC9ECFC4" httpUri="http://treatment.plazi.org/id/8F217579FB689D6551B5FDEAFC9ECFC4" lastPageId="3" lastPageNumber="197" pageId="1" pageNumber="195">
<subSubSection id="4F9297E4FB689D6751B5FDEAFCFDCFC7" box="[136,832,628,655]" pageId="1" pageNumber="195" type="nomenclature">
<paragraph id="0737C46FFB689D6751B5FDEAFCFDCFC7" blockId="1.[136,832,628,655]" box="[136,832,628,655]" pageId="1" pageNumber="195">
<heading id="5C7F7303FB689D6751B5FDEAFCFDCFC7" box="[136,832,628,655]" fontSize="11" level="2" pageId="1" pageNumber="195" reason="3">
<taxonomicName id="C088BFECFB689D6751B5FDEAFD30CFC6" authority="G.E. Lee &amp; X.L. He" authorityName="G. E. Lee &amp; X. L. He" box="[136,653,628,655]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="1" pageNumber="195" phylum="Marchantiophyta" rank="species" species="timokoponenii" status="sp. nov.">
<emphasis id="35FC187DFB689D6751B5FDEAFE1DCFC6" bold="true" box="[136,416,628,655]" italics="true" pageId="1" pageNumber="195">Lejeunea timokoponenii</emphasis>
G.E. Lee &amp; X.L. He
</taxonomicName>
,
<taxonomicNameLabel id="2ECFA506FB689D6753A4FDEAFD4DCFC6" box="[665,752,628,655]" pageId="1" pageNumber="195" rank="species">
<emphasis id="35FC187DFB689D6753A4FDEAFD57CFC7" bold="true" box="[665,746,628,654]" italics="true" pageId="1" pageNumber="195">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<figureCitation id="9FB3D8EAFB689D6753C2FDEAFCFDCFC7" box="[767,832,628,654]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[205,1383,197,1919]" captionTargetId="figure-17@2.[196,1391,187,1926]" captionTargetPageId="2" captionText="FIGURE 1. Lejeunea timokoponenii G.E. Lee &amp; X.L. He. A. Part of plant in ventral view. B,C. Leaves. D,E. Underleaves. F. Upper portion of perianth. G. Perianth. H. Median cells of leaf. I. Stem portion and leaf lobules. J. Cross-section of stem. K,L. Female bracts. M. Upper part of leaf lobules when flattened (hyaline papillae and disc cell shown in gray and asterisk, “*”, respectively). N. Perianth with bracts and bracteole. All from the holotype, Timo Koponen 34857a (H). Illustrations drawn by G.E. Lee." figureDoi="http://doi.org/10.5281/zenodo.13216368" httpUri="https://zenodo.org/record/13216368/files/figure.png" pageId="1" pageNumber="195">Fig. 1</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4F9297E4FB689D6751B5FD02FCE1CA7B" pageId="1" pageNumber="195" type="materials_examined">
<paragraph id="0737C46FFB689D6751B5FD02FAE0CF9C" blockId="1.[136,1451,668,726]" pageId="1" pageNumber="195">
Type:—
<collectingCountry id="7F9F84FFFB689D6751EEFD02FE78CFF8" box="[211,453,668,689]" name="Papua New Guinea" pageId="1" pageNumber="195">PAPUA NEW GUINEA</collectingCountry>
.
<collectingRegion id="C54C0A8DFB689D6750F2FD02FDC7CFF8" box="[463,634,668,690]" country="Papua New Guinea" name="West Sepik" pageId="1" pageNumber="195">West Sepik Prov.</collectingRegion>
: Frieda River prospecting area of Frieda Copper Co. Frieda
<collectionCode id="61995CAAFB689D6755FBFD02FB64CFF8" box="[1222,1241,668,689]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="195">R</collectionCode>
. Base Camp at Horse Creek, open gardens and lawns,
<quantity id="C070698AFB689D6750C8FD5EFD8FCF9C" box="[501,562,704,725]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="195" unit="m" value="400.0">400 m</quantity>
alt.,
<geoCoordinate id="62BCA2A8FB689D67535CFD5EFD1BCF9C" box="[609,678,704,725]" degrees="4" direction="south" minutes="42" orientation="latitude" pageId="1" pageNumber="195" precision="925" value="-4.7">4°42S</geoCoordinate>
,
<geoCoordinate id="62BCA2A8FB689D67538DFD5EFCB3CF9C" box="[688,782,704,725]" degrees="141" direction="east" minutes="47" orientation="longitude" pageId="1" pageNumber="195" precision="925" value="141.78334">141°47E</geoCoordinate>
,
<date id="7336E2AFFB689D675224FD5EFC1BCF9C" box="[793,934,704,725]" pageId="1" pageNumber="195" value="1981-08-01">1 August 1981</date>
,
<emphasis id="35FC187DFB689D67528DFD5EFB36CF9C" box="[944,1163,704,725]" italics="true" pageId="1" pageNumber="195">Timo Koponen 34857a</emphasis>
(
<typeStatus id="D8337ACDFB689D6755A5FD5EFB53CF9F" box="[1176,1262,704,726]" pageId="1" pageNumber="195" type="holotype">holotype</typeStatus>
<collectionCode id="61995CAAFB689D6755CEFD5EFAB9CF9C" box="[1267,1284,704,725]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="195" type="Herbarium">H</collectionCode>
[c.per.]).
</paragraph>
<paragraph id="0737C46FFB689D6751B5FC9AFC78CBEA" blockId="1.[136,1452,772,1987]" pageId="1" pageNumber="195">
<emphasis id="35FC187DFB689D6751B5FC9AFF6ECE57" bold="true" box="[136,211,772,798]" pageId="1" pageNumber="195">Plants</emphasis>
dioicous (only gynoecium seen),
<quantity id="C070698AFB689D675358FC9AFD4CCE56" box="[613,753,772,799]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="195" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 mm</quantity>
wide, irregularly and loosely branched, seldom bipinnately branched, branches spreading, with three small basal collar lobes.
<emphasis id="35FC187DFB689D675243FCB6FC06CE0B" bold="true" box="[894,955,808,834]" pageId="1" pageNumber="195">Stem</emphasis>
ca.
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in diameter, about 6 cells wide in cross-section; epidermal cells 7, 2840 µm wide, medullary cells 1315, 620 µm wide, ventral merophyte 2 cells wide.
<emphasis id="35FC187DFB689D675034FCEEFE38CEC3" bold="true" box="[265,389,880,906]" pageId="1" pageNumber="195">Leaf lobes</emphasis>
<quantity id="C070698AFB689D6750B0FCEEFDA8CEC2" box="[397,533,880,907]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="195" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
<emphasis id="35FC187DFB689D675320FCEFFD8DCEC3" box="[541,560,881,906]" italics="true" pageId="1" pageNumber="195">×</emphasis>
<quantity id="C070698AFB689D67537CFCEEFD58CEC2" box="[577,741,880,907]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="1" pageNumber="195" unit="mm" value="0.475" valueMax="0.55" valueMin="0.4">0.400.55 mm</quantity>
(when flattened), imbricate, ovate; leaf apex broadly rounded, with wavy margin curled upwards; leaf margin crenulate; the ventral margin forming an angle of
<superScript id="F0FD6927FB689D6755D3FC0AFAAFCEEB" attach="none" box="[1262,1298,916,942]" fontSize="6" pageId="1" pageNumber="195">800</superScript>
<superScript id="F0FD6927FB689D67541CFC0AFAF9CEEB" attach="left" box="[1313,1348,916,942]" fontSize="6" pageId="1" pageNumber="195">900</superScript>
with the keel when flattened; insertion on dorsal side 1012 lobe cells long.
<emphasis id="35FC187DFB689D67529FFC26FBABCE9B" bold="true" box="[930,1046,952,978]" pageId="1" pageNumber="195">Leaf cells</emphasis>
isodiametric; apical cells 1620
<emphasis id="35FC187DFB689D6754A5FC27FA16CE9B" box="[1432,1451,953,978]" italics="true" pageId="1" pageNumber="195">×</emphasis>
1520 µm, median cells 2030
<emphasis id="35FC187DFB689D6750D0FC43FDBDCEBF" box="[493,512,989,1014]" italics="true" pageId="1" pageNumber="195">×</emphasis>
1620 µm, basal cells 3343
<emphasis id="35FC187DFB689D675269FC43FCDACEBF" box="[852,871,989,1014]" italics="true" pageId="1" pageNumber="195">×</emphasis>
2026 µm; cell walls hyaline, with well-developed trigones and frequently with intermediate thickenings, 12 per cell, 01 between 2 adjacent trigones. Cuticle smooth to weakly papillose. Oil bodies not seen.
<emphasis id="35FC187DFB689D67535CFBBAFD49C977" bold="true" box="[609,756,1060,1086]" pageId="1" pageNumber="195">Leaf lobules</emphasis>
seldom reduced (i.e. differing from its typical form, see Fig. 2
<collectionCode id="61995CAAFB689D6751ABFBD6FF1BC92B" box="[150,166,1096,1122]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="1" pageNumber="195" type="Herbarium">F</collectionCode>
,
<collectionCode id="61995CAAFB689D675197FBD6FF02C92B" box="[170,191,1096,1122]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="1" pageNumber="195" type="Herbarium">G</collectionCode>
in
<bibRefCitation id="6319B99EFB689D6751DEFBD7FE30C92B" author="Lee, G. E. &amp; Bechteler, J. &amp; Pocs, T. &amp; Schafer-Verwimp, A. &amp; Tang, H. Y. &amp; Chia, P. W." box="[227,397,1096,1123]" pageId="1" pageNumber="195" refId="ref2962" refString="Lee, G. E., Bechteler, J., Pocs, T., Schafer-Verwimp, A., Tang, H. Y. &amp; Chia, P. W. (2022) Integrative taxonomy reveals a new species of the genus Lejeunea (Marchantiophyta: Lejeuneaceae) from Peninsular Malaysia. Plants 11: 1642. https: // doi. org / 10.3390 / plants 11131642" type="journal volume" year="2022">
Lee
<emphasis id="35FC187DFB689D675028FBD7FEF3C92B" box="[277,334,1096,1122]" italics="true" pageId="1" pageNumber="195">et al.</emphasis>
2022
</bibRefCitation>
under
<taxonomicName id="C088BFECFB689D6750E1FBD7FD76C92B" box="[476,715,1096,1122]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="1" pageNumber="195" phylum="Marchantiophyta" rank="species" species="malaysiana">
<emphasis id="35FC187DFB689D6750E1FBD7FD76C92B" box="[476,715,1096,1122]" italics="true" pageId="1" pageNumber="195">Lejeunea malaysiana</emphasis>
</taxonomicName>
), 0.250.35
<emphasis id="35FC187DFB689D67526EFBD7FCDBC92B" box="[851,870,1097,1122]" italics="true" pageId="1" pageNumber="195">×</emphasis>
<quantity id="C070698AFB689D675250FBD6FBAFC92A" box="[877,1042,1096,1123]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="195" unit="mm" value="0.175" valueMax="0.2" valueMin="0.15">0.150.20 mm</quantity>
, to 1/22/5 the length of the lobe, at an angle of
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to the stem, ovate-triangular, inflated along the keel; apex obliquely truncate; keel curved; free margin incurved fully and abruptly flattened towards the apex; first tooth 2528 µm long, oblong, apex obtuse; margin between tooth and sinus 3 cells long, large rectangular disc cell present (indicated by “*” in
<figureCitation id="9FB3D8EAFB689D67559EFB2AFB43C987" box="[1187,1278,1204,1230]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[205,1383,197,1919]" captionTargetId="figure-17@2.[196,1391,187,1926]" captionTargetPageId="2" captionText="FIGURE 1. Lejeunea timokoponenii G.E. Lee &amp; X.L. He. A. Part of plant in ventral view. B,C. Leaves. D,E. Underleaves. F. Upper portion of perianth. G. Perianth. H. Median cells of leaf. I. Stem portion and leaf lobules. J. Cross-section of stem. K,L. Female bracts. M. Upper part of leaf lobules when flattened (hyaline papillae and disc cell shown in gray and asterisk, “*”, respectively). N. Perianth with bracts and bracteole. All from the holotype, Timo Koponen 34857a (H). Illustrations drawn by G.E. Lee." figureDoi="http://doi.org/10.5281/zenodo.13216368" httpUri="https://zenodo.org/record/13216368/files/figure.png" pageId="1" pageNumber="195">Fig. 1M</figureCitation>
or refer to Lee 2020 for further details), 4045 µm long; hyaline papilla at the proximal side of the first tooth.
<emphasis id="35FC187DFB689D675581FB46FAEDC9BB" bold="true" box="[1212,1360,1240,1266]" pageId="1" pageNumber="195">Underleaves</emphasis>
0.30.4
<emphasis id="35FC187DFB689D6751B5FB63FF26C85F" box="[136,155,1277,1302]" italics="true" pageId="1" pageNumber="195">×</emphasis>
<quantity id="C070698AFB689D67519FFB62FE91C85E" box="[162,300,1276,1303]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="195" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, to 67 times wider than the stem, imbricate to approximate, reniform; fully covering the leaf lobules; bilobed, lobes to 1/31/4 of underleaf length, about 68 cells wide, triangular; sinus broad, obtuse, U-shaped; tips acute to obtuse; underleaf margin slightly crenulate; two large basal underleaf cells differentiated; base margin straight, insertion line curved.
<emphasis id="35FC187DFB689D6750BDFAF6FE41C8CB" bold="true" box="[384,508,1384,1410]" pageId="1" pageNumber="195">Androecia</emphasis>
not seen.
<emphasis id="35FC187DFB689D67534FFAF6FD5FC8CB" bold="true" box="[626,738,1384,1410]" pageId="1" pageNumber="195">Gynoecia</emphasis>
on short lateral branches, female bracts loosely arranged, with one innovation, with 12 gynoecia in a lateral position. Female bracts larger than the leaf, erect-spreading when moist, slightly enveloping the perianth. Lobes ca. 1.0
<emphasis id="35FC187DFB689D6753A7FA2FFD10C883" box="[666,685,1457,1482]" italics="true" pageId="1" pageNumber="195">×</emphasis>
<quantity id="C070698AFB689D675388FA2EFCE6C882" box="[693,859,1456,1483]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="1" pageNumber="195" unit="mm" value="0.475" valueMax="0.55" valueMin="0.4">0.400.55 mm</quantity>
, obovate, apex broadly to narrowly rounded, margin crenulate. Lobules 0.40.5
<emphasis id="35FC187DFB689D675083FA4BFE6CC8A7" box="[446,465,1493,1518]" italics="true" pageId="1" pageNumber="195">×</emphasis>
<quantity id="C070698AFB689D6750E4FA4AFDD9C8A6" box="[473,612,1492,1519]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="195" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.10.2 mm</quantity>
, rarely reduced, 1/3 the width and 1/22/3 the length of the lobe, oblong, apex obtuse, keels straight, smooth,
<quantity id="C070698AFB689D675321FA66FD1ECB5A" box="[540,675,1528,1555]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="195" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.40.5 mm</quantity>
long. Female bracteoles 0.70.8
<emphasis id="35FC187DFB689D675528FA67FB95CB5B" box="[1045,1064,1529,1554]" italics="true" pageId="1" pageNumber="195">×</emphasis>
<quantity id="C070698AFB689D675513FA66FB6ECB5A" box="[1070,1235,1528,1555]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="1" pageNumber="195" unit="mm" value="0.525" valueMax="0.6" valueMin="0.45">0.450.60 mm</quantity>
, 3/4 of the perianth length, ovate with tips acute, lobes to 1/8 of female bracteole length, distant, sinus narrow, acute, margin crenulate. Perianths 0.71.2
<emphasis id="35FC187DFB689D67506AF9DFFED7CB13" box="[343,362,1601,1626]" italics="true" pageId="1" pageNumber="195">×</emphasis>
<quantity id="C070698AFB689D67504EF9DEFE42CB12" box="[371,511,1600,1627]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="195" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.40.5 mm</quantity>
, emergent to 1/7 of the perianth length, oblong, with 5 keels, margin of the keels sometimes lobed, undulate, cells of the perianth at the keels irregularly bulging on surface; beak 56 cells long; stalklike elongation lacking.
<emphasis id="35FC187DFB689D6750AAF916FD9CCBEB" bold="true" box="[407,545,1672,1698]" pageId="1" pageNumber="195">Sporophyte</emphasis>
and
<emphasis id="35FC187DFB689D67536AF916FCE7CBEB" bold="true" box="[599,858,1672,1698]" pageId="1" pageNumber="195">vegetative propagules</emphasis>
not seen.
</paragraph>
<paragraph id="0737C46FFB689D675180F932FCE1CA7B" blockId="1.[136,1452,772,1987]" pageId="1" pageNumber="195">
<emphasis id="35FC187DFB689D675180F932FDFFCB8F" bold="true" box="[189,578,1708,1734]" pageId="1" pageNumber="195">Additional specimens examined:</emphasis>
<collectingRegion id="C54C0A8DFB689D675374F933FCB3CB8E" box="[585,782,1708,1735]" country="Papua New Guinea" name="West Sepik" pageId="1" pageNumber="195">West Sepik Prov.</collectingRegion>
: Frieda River prospecting area of Frieda Copper Co. Frieda
<collectionCode id="61995CAAFB689D6751B5F94FFF23CBA2" box="[136,158,1745,1771]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="195">R</collectionCode>
. Base Camp at Horse Creek, open gardens and lawns,
<quantity id="C070698AFB689D6753C7F94EFCFCCBA2" box="[762,833,1744,1771]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="195" unit="m" value="400.0">400 m</quantity>
alt.,
<geoCoordinate id="62BCA2A8FB689D67524AF94EFC75CBA3" box="[887,968,1744,1770]" degrees="4" direction="south" minutes="42" orientation="latitude" pageId="1" pageNumber="195" precision="925" value="-4.7">4°42S</geoCoordinate>
,
<geoCoordinate id="62BCA2A8FB689D6752EEF94EFBFCCBA3" box="[979,1089,1744,1770]" degrees="141" direction="east" minutes="47" orientation="longitude" pageId="1" pageNumber="195" precision="925" value="141.78334">141°47E</geoCoordinate>
,
<date id="7336E2AFFB689D675571F94EFB4DCBA3" box="[1100,1264,1744,1770]" pageId="1" pageNumber="195" value="1981-08-01">1 August 1981</date>
, on a bush,
<emphasis id="35FC187DFB689D67544EF94FFE86CA47" italics="true" pageId="1" pageNumber="195">Timo Koponen 34826</emphasis>
(
<typeStatus id="D8337ACDFB689D675076F96AFE10CA47" box="[331,429,1780,1806]" pageId="1" pageNumber="195" type="paratype">paratype</typeStatus>
<collectionCode id="61995CAAFB689D675088F96BFE74CA46" box="[437,457,1781,1807]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="195" type="Herbarium">H</collectionCode>
[c.per.]), ibid.,
<emphasis id="35FC187DFB689D675346F96BFCD3CA47" box="[635,878,1780,1806]" italics="true" pageId="1" pageNumber="195">Timo Koponen 34824</emphasis>
(
<typeStatus id="D8337ACDFB689D675243F96AFC5DCA47" box="[894,992,1780,1806]" pageId="1" pageNumber="195" type="paratype">paratype</typeStatus>
<collectionCode id="61995CAAFB689D6752D5F96AFC42CA47" box="[1000,1023,1780,1806]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="195" type="Herbarium">H</collectionCode>
), ibid., on cliff,
<emphasis id="35FC187DFB689D675585F96BFA16CA47" box="[1208,1451,1780,1806]" italics="true" pageId="1" pageNumber="195">Timo Koponen 34841</emphasis>
(
<typeStatus id="D8337ACDFB689D6751B2F886FF4CCA7B" box="[143,241,1816,1842]" pageId="1" pageNumber="195" type="paratype">paratype</typeStatus>
<collectionCode id="61995CAAFB689D6751C4F886FEADCA7B" box="[249,272,1816,1842]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="195" type="Herbarium">H</collectionCode>
), ibid., on soil,
<emphasis id="35FC187DFB689D675083F887FD7DCA7B" box="[446,704,1816,1842]" italics="true" pageId="1" pageNumber="195">
Timo Koponen 34818
<collectionCode id="61995CAAFB689D675392F886FD7DCA7B" box="[687,704,1816,1842]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="195" type="Herbarium">A</collectionCode>
</emphasis>
(
<typeStatus id="D8337ACDFB689D6753F3F886FC8DCA7B" box="[718,816,1816,1842]" pageId="1" pageNumber="195" type="paratype">paratype</typeStatus>
<collectionCode id="61995CAAFB689D675205F886FCEDCA7B" box="[824,848,1816,1842]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="195" type="Herbarium">H</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="4F9297E4FB689D675180F8A2FF48CA32" pageId="1" pageNumber="195" type="distribution">
<paragraph id="0737C46FFB689D675180F8A2FF48CA32" blockId="1.[136,1452,772,1987]" pageId="1" pageNumber="195">
<emphasis id="35FC187DFB689D675180F8A2FE55CA1F" bold="true" box="[189,488,1852,1878]" pageId="1" pageNumber="195">Distribution and habitat:</emphasis>
Only known from
<collectingCountry id="7F9F84FFFB689D675387F8A2FD48CA1F" box="[698,757,1852,1878]" name="Papua New Guinea" pageId="1" pageNumber="195">PNG</collectingCountry>
, growing in an open, dry slope on a bush or soil at an elevation of
<specimenCount id="118E0FE6FB689D67519BF8FEFF48CA32" box="[166,245,1888,1915]" count="400" pageId="1" pageNumber="195" type="generic">
<quantity id="C070698AFB689D67519BF8FEFF4DCA32" box="[166,240,1888,1915]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="195" unit="m" value="400.0">400 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="4F9297E4FB689D675180F81AFD0ACA8B" pageId="1" pageNumber="195" type="etymology">
<paragraph id="0737C46FFB689D675180F81AFD0ACA8B" blockId="1.[136,1452,772,1987]" pageId="1" pageNumber="195">
<emphasis id="35FC187DFB689D675180F81AFEF5CAD7" bold="true" box="[189,328,1924,1950]" pageId="1" pageNumber="195">Etymology:</emphasis>
The species is named in honour of Professor Timo Koponen, a prominent bryologist, who collected the
<typeStatus id="D8337ACDFB689D67518CF837FF5CCA8A" box="[177,225,1961,1987]" pageId="1" pageNumber="195">type</typeStatus>
specimens in
<collectingRegion id="C54C0A8DFB689D6750BDF837FDD6CA8B" box="[384,619,1960,1987]" country="Papua New Guinea" name="West Sepik" pageId="1" pageNumber="195">West Sepik Province</collectingRegion>
,
<collectingCountry id="7F9F84FFFB689D67534AF836FD0ECA8B" box="[631,691,1960,1986]" name="Papua New Guinea" pageId="1" pageNumber="195">PNG</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="53F794E7FB6B9D6451B5F83FFBCEC56B" ID-DOI="http://doi.org/10.5281/zenodo.13216368" ID-Zenodo-Dep="13216368" httpUri="https://zenodo.org/record/13216368/files/figure.png" pageId="2" pageNumber="196" startId="2.[136,229,1953,1975]" targetBox="[205,1383,197,1919]" targetPageId="2" targetType="figure">
<paragraph id="0737C46FFB6B9D6451B5F83FFBCEC56B" blockId="2.[136,1452,1953,2083]" pageId="2" pageNumber="196">
<emphasis id="35FC187DFB6B9D6451B5F83FFEBDCAFF" bold="true" box="[136,256,1953,1975]" pageId="2" pageNumber="196">FIGURE 1.</emphasis>
<taxonomicName id="C088BFECFB6B9D645035F83CFCA5CAFF" authority="G. E. Lee &amp; X. L. He. A. Part" authorityName="G. E. Lee &amp; X. L. He. A. Part" box="[264,792,1953,1975]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="2" pageNumber="196" phylum="Marchantiophyta" rank="species" species="timokoponenii">
<emphasis id="35FC187DFB6B9D645035F83CFE4CCAFF" box="[264,497,1953,1975]" italics="true" pageId="2" pageNumber="196">Lejeunea timokoponenii</emphasis>
G.E. Lee &amp; X.L. He. A. Part
</taxonomicName>
of plant in ventral view. B,C. Leaves. D,E. Underleaves. F. Upper portion of perianth. G. Perianth. H. Median cells of leaf. I. Stem portion and leaf lobules. J. Cross-section of stem. K,L. Female bracts. M. Upper part of leaf lobules when flattened (hyaline papillae and disc cell shown in gray and asterisk, “*”, respectively). N. Perianth with bracts and bracteole. All from the holotype,
<emphasis id="35FC187DFB6B9D645312F790FCB7C56B" box="[559,778,2061,2083]" italics="true" pageId="2" pageNumber="196">Timo Koponen 34857a</emphasis>
(H). Illustrations drawn by G.E. Lee.
</paragraph>
</caption>
<subSubSection id="4F9297E4FB6A9D655180FF00FC9ECFC4" pageId="3" pageNumber="197" type="discussion">
<paragraph id="0737C46FFB6A9D655180FF00FC9ECFC4" blockId="3.[136,1452,158,653]" pageId="3" pageNumber="197">
<emphasis id="35FC187DFB6A9D655180FF00FEF8CDF1" bold="true" box="[189,325,158,184]" pageId="3" pageNumber="197">Discussion:</emphasis>
The distinguishing characteristics of
<taxonomicName id="C088BFECFB6A9D6553DBFF01FC4ECDF1" box="[742,1011,158,184]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="timokoponenii">
<emphasis id="35FC187DFB6A9D6553DBFF01FC4ECDF1" box="[742,1011,158,184]" italics="true" pageId="3" pageNumber="197">Lejeunea timokoponenii</emphasis>
</taxonomicName>
are: 1) the leaf margins curled upwards (particularly at the apex) including the female bracts, 2) the large, reniform underleaves 67 times wider than the stem, 3) the leaf cells with well-developed trigones and frequently with conspicuous intermediate thickenings, 4) the fully incurved free margins of the lobules, 5) the large disc cell below the first tooth of the leaf lobules, 6) the perianths with undulate keels and a 56 cells long beak. In its undulate-keeled perianth, leaf cells and outline of leaf lobules,
<taxonomicName id="C088BFECFB6A9D6551B5FECDFEFBCC25" box="[136,326,338,364]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="timokoponenii">
<emphasis id="35FC187DFB6A9D6551B5FECDFEFBCC25" box="[136,326,338,364]" italics="true" pageId="3" pageNumber="197">L. timokoponenii</emphasis>
</taxonomicName>
resembles the previously documented Malesian species,
<taxonomicName id="C088BFECFB6A9D6552FAFECDFAFCCC24" authority="(Eifrig 1937: 96) Lee" authorityName="Lee" baseAuthorityName="Eifrig" baseAuthorityPageNumber="96" baseAuthorityYear="1937" box="[967,1345,338,365]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="dipterota">
<emphasis id="35FC187DFB6A9D6552FAFECDFBF6CC25" box="[967,1099,338,364]" italics="true" pageId="3" pageNumber="197">L. dipterota</emphasis>
(
<bibRefCitation id="6319B99EFB6A9D655564FECCFABACC25" author="Eifrig, H." box="[1113,1287,338,364]" pageId="3" pageNumber="197" pagination="73 - 114" refId="ref2249" refString="Eifrig, H. (1937) Monografische Studien u ¨ ber die Indomalayischen Arten von Taxilejeunea. Annales Bryologici 9: 73 - 114." type="journal article" year="1937">Eifrig 1937: 96</bibRefCitation>
) Lee
</taxonomicName>
in
<bibRefCitation id="6319B99EFB6A9D655458FECDFECCCCD9" author="Lee, G. E. &amp; Gradstein, S. R." pageId="3" pageNumber="197" pagination="59 - 69" refId="ref2739" refString="Lee, G. E. &amp; Gradstein, S. R. (2013) Distribution and habitat of the Malaysian species of Lejeunea (Marchantiophyta: Lejeuneaceae), with description of Lejeunea tamaspocsii sp. nov. Polish Botanical Journal 58 (1): 59 - 69. https: // doi. org / 10.2478 / pbj- 2013 - 0007" type="journal article" year="2013">Lee &amp; Gradstein (2013: 61)</bibRefCitation>
(recently recorded in
<collectingCountry id="7F9F84FFFB6A9D65535AFEE8FD22CCD9" box="[615,671,374,400]" name="Papua New Guinea" pageId="3" pageNumber="197">PNG</collectingCountry>
fide
<bibRefCitation id="6319B99EFB6A9D6553E5FEE8FC2ECCD9" author="Maul, K. &amp; Wei, Y. - M. &amp; Iskandar, E. A. P. &amp; Chantanaorrapint, S. &amp; Ho, B. - C. &amp; Quandt, D. &amp; Kessler, M." box="[728,915,374,400]" pageId="3" pageNumber="197" refId="ref3035" refString="Maul, K., Wei, Y. - M., Iskandar, E. A. P., Chantanaorrapint, S., Ho, B. - C., Quandt, D. &amp; Kessler, M. (2023) Liverworts show a globally consistent mid-elevation richness peak. Ecology and Evolution 13: e 9862. https: // doi. org / 10.1002 / ece 3.9862" type="journal volume" year="2023">
Maul
<emphasis id="35FC187DFB6A9D655227FEE9FCEFCCD9" box="[794,850,374,400]" italics="true" pageId="3" pageNumber="197">et al.</emphasis>
2023
</bibRefCitation>
,
<bibRefCitation id="6319B99EFB6A9D6552A3FEE8FBE9CCD9" author="Ellis, L. T. &amp; Alegro, A. &amp; Alvarez, D. J. &amp; Aponte Rojas, A. M. &amp; Ashouri, A. &amp; Atwood, J. J. &amp; Bednarek-Ochyra, H. &amp; Burghardt, M. &amp; Dhyani, A. &amp; Erzberger, P. &amp; Espinoza-Prieto, B. &amp; Fedosov, V. E. &amp; Gradstein, S. R. &amp; He, X. - L. &amp; Hodgetts, N. G. &amp; Hugonnot, V., Ku &amp; Lee, G. E. &amp; Mahdigholi, K. &amp; Manju, C. N. &amp; Mufeed, B. &amp; Norhazrina, N. &amp; O'Leary, S. V. &amp; Rajesh, K. P. &amp; Rimac, A. &amp; Skurko, A. V. &amp; Spitale, D. &amp; Sruthi, O. M. &amp; Syazwana, N. &amp; Uniyal, P. L. &amp; Vineesha, P. M. &amp; Winter, G." box="[926,1108,374,400]" pageId="3" pageNumber="197" pagination="321 - 338" refId="ref2275" refString="Ellis, L. T., Alegro, A., Alvarez, D. J., Aponte Rojas, A. M., Ashouri, A., Atwood, J. J., Bednarek-Ochyra, H., Bi ˆ rsan, C. - C., Burghardt, M., Dhyani, A., Erzberger, P., Espinoza-Prieto, B., Fedosov, V. E., Gradstein, S. R., He, X. - L., Hodgetts, N. G., Hugonnot, V., Ku ¨ rschner, H., Lee, G. E., Mahdigholi, K., Manju, C. N., Mir-Rossello´, P. M., Mufeed, B., Norhazrina, N., O'Leary, S. V., Rajesh, K. P., Rimac, A., Scha ¨ fer-Verwimp, A., S ˇ egota, V., Se´rgio, C., Skurko, A. V., Spitale, D., Sruthi, O. M., S ̦ tefa ˘ nut ¸, S., Sua´rez, G. M., Syazwana, N., Szu &quot; cs, P., Uniyal, P. L., Vineesha, P. M., Vonc ˇ ina, G. &amp; Winter, G. (2024) New national and regional bryophytes, 75. Journal of Bryology 45 (4): 321 - 338. https: // doi. org / 10.1080 / 03736687.2024.2313380" type="journal article" year="2024">
Ellis
<emphasis id="35FC187DFB6A9D6552E5FEE9FBADCCD9" box="[984,1040,374,400]" italics="true" pageId="3" pageNumber="197">et al.</emphasis>
2024
</bibRefCitation>
). However,
<taxonomicName id="C088BFECFB6A9D6555E4FEE9FAE1CCD9" authorityName="Lee" baseAuthorityName="Eifrig" baseAuthorityYear="1937" box="[1241,1372,374,400]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="dipterota">
<emphasis id="35FC187DFB6A9D6555E4FEE9FAE1CCD9" box="[1241,1372,374,400]" italics="true" pageId="3" pageNumber="197">L. dipterota</emphasis>
</taxonomicName>
differs in the perianth character by having perianth keels with 2 wings and a beak that is only 23 cells long. On the other hand, the perianth of
<taxonomicName id="C088BFECFB6A9D655011FE21FE5ACC91" box="[300,487,446,472]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="timokopenenii">
<emphasis id="35FC187DFB6A9D655011FE21FE5ACC91" box="[300,487,446,472]" italics="true" pageId="3" pageNumber="197">L. timokopenenii</emphasis>
</taxonomicName>
lacks wings and has a long beak, which is 56 cells long. Furthermore,
<taxonomicName id="C088BFECFB6A9D6555CDFE21FA16CC91" box="[1264,1451,446,472]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="timokoponenii">
<emphasis id="35FC187DFB6A9D6555CDFE21FA16CC91" box="[1264,1451,446,472]" italics="true" pageId="3" pageNumber="197">L. timokoponenii</emphasis>
</taxonomicName>
is also distinguished by its distinct wavy leaf margins and large, reniform underleaves. This characteristic of large, reniform underleaves covering the leaf lobules is also observed in several species of
<emphasis id="35FC187DFB6A9D655505FD99FB1BCF69" box="[1080,1190,519,544]" italics="true" pageId="3" pageNumber="197">
<taxonomicName id="C088BFECFB6A9D655505FD99FB1CCF69" authorityName="Libert" authorityYear="1820" box="[1080,1185,519,544]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="genus">Lejeunea</taxonomicName>
,
</emphasis>
such as
<taxonomicName id="C088BFECFB6A9D655438FD99FF66CF0D" authority="Lee &amp; Pocs" authorityName="Lee &amp; Pocs" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="heinarii">
<emphasis id="35FC187DFB6A9D655438FD99FAC6CF69" box="[1285,1403,518,544]" italics="true" pageId="3" pageNumber="197">L. heinarii</emphasis>
Lee &amp; Pócs
</taxonomicName>
in
<bibRefCitation id="6319B99EFB6A9D65503FFDB5FE41CF0D" author="Lee, G. E. &amp; Pocs, T. &amp; Koid, C. W. &amp; He, X. - L." box="[258,508,554,581]" pageId="3" pageNumber="197" pagination="492 - 501" refId="ref2897" refString="Lee, G. E., Pocs, T., Koid, C. W. &amp; He, X. - L. (2020) New species of Lejeunea (Porellales: Lejeuneaceae) from Papua New Guinea. The Bryologist 123: 492 - 501. https: // doi. org / 10.1639 / 0007 - 2745 - 123.3.492" type="journal article" year="2020">
Lee
<emphasis id="35FC187DFB6A9D655008FDB5FED2CF0D" box="[309,367,554,580]" italics="true" pageId="3" pageNumber="197">et al.</emphasis>
(2020: 493)
</bibRefCitation>
,
<taxonomicName id="C088BFECFB6A9D655331FDB5FC3CCF0D" authority="Hurlimann (1993: 12)" authorityName="Hurlimann" authorityPageNumber="12" authorityYear="1993" box="[524,897,554,580]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="mimula">
<emphasis id="35FC187DFB6A9D655331FDB5FDC2CF0D" box="[524,639,554,580]" italics="true" pageId="3" pageNumber="197">L. mimula</emphasis>
<bibRefCitation id="6319B99EFB6A9D6553B5FDB4FC3CCF0D" author="Hurlimann, H." box="[648,897,554,580]" pageId="3" pageNumber="197" pagination="3 - 17" refId="ref2605" refString="Hurlimann, H. (1993) Hepaticae aus dem Gebiete des sudlichen Pazifik XII. Bauhinia 11: 3 - 17." type="journal article" year="1993">Hürlimann (1993: 12)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C088BFECFB6A9D6552ADFDB5FF06CF21" authority="(Stephani 1914: 562) Mizutani (1970: 243)" authorityName="Mizutani" authorityPageNumber="243" authorityYear="1970" baseAuthorityName="Stephani" baseAuthorityPageNumber="562" baseAuthorityYear="1914" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="leratii">
<emphasis id="35FC187DFB6A9D6552ADFDB5FC4ECF0D" box="[912,1011,554,580]" italics="true" pageId="3" pageNumber="197">L. leratii</emphasis>
(
<bibRefCitation id="6319B99EFB6A9D655539FDB4FB58CF0C" author="Stephani, F." box="[1028,1253,554,581]" pageId="3" pageNumber="197" pagination="449 - 704" refId="ref3622" refString="Stephani, F. (1914) Species Hepaticarum 5. George &amp; Cie, Gene`ve &amp; Bale, pp. 449 - 704." type="book chapter" year="1914">Stephani 1914: 562</bibRefCitation>
)
<bibRefCitation id="6319B99EFB6A9D6555C8FDB4FF06CF21" author="Mizutani, M." pageId="3" pageNumber="197" pagination="225 - 265" refId="ref3110" refString="Mizutani, M. (1970) Lejeuneaceae, subfamilies Lejeuneoideae and Cololejeuneoideae from Sabah (North Borneo). Journal of Hattori Botanical Laboratory 33: 225 - 265." type="journal article" year="1970">Mizutani (1970: 243)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="C088BFECFB6A9D6551C0FDD1FC83CF20" authority="(Nees 1830: 41) Montagne (1840: 335)" authorityName="Montagne" authorityPageNumber="335" authorityYear="1840" baseAuthorityName="Nees" baseAuthorityPageNumber="41" baseAuthorityYear="1830" box="[253,830,590,617]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="sordida">
<emphasis id="35FC187DFB6A9D6551C0FDD1FECFCF21" box="[253,370,590,616]" italics="true" pageId="3" pageNumber="197">L. sordida</emphasis>
(
<bibRefCitation id="6319B99EFB6A9D6550BEFDD0FD96CF21" author="Nees, C. G." box="[387,555,590,616]" pageId="3" pageNumber="197" refId="ref3180" refString="Nees, C. G. (1830) Enumeratio plantarum cryptogamicarum Javae et insularum adiacentium. Fasciculus prior, hepaticas complectens, ab editore illustrates. Grass, Barth &amp; Co., Breslau, 86 pp." type="book" year="1830">Nees 1830: 41</bibRefCitation>
)
<bibRefCitation id="6319B99EFB6A9D655300FDD1FC83CF20" author="Montagne, C." box="[573,830,590,617]" pageId="3" pageNumber="197" pagination="321 - 350" refId="ref3141" refString="Montagne, C. (1840) Seconde centurie de plantes cellulaires exotiques nouvelles, Decades VI, VII et VIII. Annales des Sciences Naturelle; Botanique, ser. 2, 14: 321 - 350." type="journal article" year="1840">Montagne (1840: 335)</bibRefCitation>
</taxonomicName>
. However, none of these species exhibits the wavy or upward curling of leaf margins present in
<taxonomicName id="C088BFECFB6A9D655363FDEDFCA1CFC5" box="[606,796,626,652]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Lejeunea" kingdom="Plantae" order="Porellales" pageId="3" pageNumber="197" phylum="Marchantiophyta" rank="species" species="timokoponenii">
<emphasis id="35FC187DFB6A9D655363FDEDFCA1CFC5" box="[606,796,626,652]" italics="true" pageId="3" pageNumber="197">L. timokoponenii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>