364 lines
45 KiB
XML
364 lines
45 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.885.38980" ID-GBIF-Dataset="5fe8e574-2edb-4095-8b3e-c6d856bb07f2" ID-PMC="PMC6848233" ID-Pensoft-Pub="1313-2970-885-1" ID-Pensoft-UUID="870C0613A13A5C1EA54511B2C64DEBE4" ID-PubMed="31736615" ID-ZooBank="A950D3CB4C914E62AF6631B76D73880C" ModsDocID="1313-2970-885-1" checkinTime="1572912889973" checkinUser="pensoft" docAuthor="Sutcharit, Chirasak, Naggs, Fred, Ablett, Jonathan, Sang, Pham Van, Luong Van Hao, & Panha, Somsak" docDate="2019" docId="45F5B6E687BC5CBB83E299909A7EBF3D" docLanguage="en" docName="ZooKeys 885: 1-14" docOrigin="ZooKeys 885" docSource="http://dx.doi.org/10.3897/zookeys.885.38980" docTitle="Bertia cambojiensis Reeve 1860" docType="treatment" docVersion="4" id="870C0613A13A5C1EA54511B2C64DEBE4" lastPageNumber="1" masterDocId="870C0613A13A5C1EA54511B2C64DEBE4" masterDocTitle="Notes on the sinistral helicoid snail Bertia cambojiensis (Reeve, 1860) from Vietnam (Eupulmonata, Dyakiidae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="1" updateTime="1668168000113" updateUser="ExternalLinkService">
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<mods:title>Notes on the sinistral helicoid snail Bertia cambojiensis (Reeve, 1860) from Vietnam (Eupulmonata, Dyakiidae)</mods:title>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:namePart>Naggs, Fred</mods:namePart>
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<mods:namePart>Ablett, Jonathan</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sang, Pham Van</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Luong Van Hao,</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>885</mods:number>
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<mods:start>1</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.885.38980</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.885.38980</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-885-1</mods:identifier>
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<mods:identifier type="ZooBank">A950D3CB4C914E62AF6631B76D73880C</mods:identifier>
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<treatment ID-GBIF-Taxon="160134102" LSID="urn:lsid:plazi:treatment:45F5B6E687BC5CBB83E299909A7EBF3D" httpUri="http://treatment.plazi.org/id/45F5B6E687BC5CBB83E299909A7EBF3D" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="45F5B6E6-87BC-5CBB-83E2-99909A7EBF3D" authority="Reeve, 1860" authorityName="Reeve" authorityYear="1860" class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia cambojiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambojiensis">Bertia cambojiensis (Reeve, 1860)</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shells of Bertia cambojiensis A lectotype NHMUK 20130220 and B specimen NHMUK 20130833." figureDoi="10.3897/zookeys.885.38980.figure1" httpUri="https://binary.pensoft.net/fig/352321" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure1">Figs 1</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix mouhoti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mouhoti">Helix mouhoti</taxonomicName>
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Reeve, 1860a [Sep.]: 203, 204 [not
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="133 - 141" refId="B29" refString="Pfeiffer, L, 1860a. Descriptions of thirty-six new species of land-shells from Mr. H. Cuming's collection. Proceedings of the Zoological Society of London 28: 133 - 141" title="Descriptions of thirty-six new species of land-shells from Mr. H. Cuming's collection." volume="28" year="1860 a">Pfeiffer 1860a</bibRefCitation>
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[May]: 136, pl. 50 fig. 5]. Type locality: Cambojia [Cambodia].
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B30" refString="Pfeiffer, L, 1860b. Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 2. Verlag von Theodor Fischer, Cassel, 139-303. [pls 37-72. p. 139-160, pls 37-42 (1860] [Published in parts, dates follow Johnson (1969)]" title="Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 2. Verlag von Theodor Fischer, Cassel, 139 - 303. [pls 37 - 72. p. 139 - 160, pls 37 - 42 (1860] [Published in parts, dates follow Johnson (1969)]" year="1860 b">Pfeiffer 1860b</bibRefCitation>
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: 173, 174, pl. 47, figs 1, 2.
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Brockhaus, Lipsiae" pageId="0" pageNumber="1" refId="B31" refString="Pfeiffer, L, 1868. Monographia Heliceorum Viventum, Supplement Tertium. Volume 5. F.A. Brockhaus, Lipsiae" title="Monographia Heliceorum Viventum, Supplement Tertium. Volume 5. F. A." year="1868">Pfeiffer 1868</bibRefCitation>
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: 64.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix cambojiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambojiensis">Helix cambojiensis</taxonomicName>
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Reeve, 1860b [Dec.]: 455 [new replacement name].
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<bibRefCitation author="Martens, E von" journalOrPublisher="Zoologischer Teil, II; Koenigliche Geheime Ober-Hofdruckerei, Berlin" pageId="0" pageNumber="1" refId="B22" refString="Martens, E von, 1867. Die Landschnecken. Die Preussische Expedition nach Ost-Asien. Zoologischer Teil, II; Koenigliche Geheime Ober-Hofdruckerei, Berlin" title="Die Landschnecken. Die Preussische Expedition nach Ost-Asien." year="1867">Martens 1867</bibRefCitation>
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: 76.
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Brockhaus, Lipsiae" pageId="0" pageNumber="1" refId="B31" refString="Pfeiffer, L, 1868. Monographia Heliceorum Viventum, Supplement Tertium. Volume 5. F.A. Brockhaus, Lipsiae" title="Monographia Heliceorum Viventum, Supplement Tertium. Volume 5. F. A." year="1868">Pfeiffer 1868</bibRefCitation>
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: 64.
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<bibRefCitation author="Daniel, F" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="126 - 128" refId="B8" refString="Daniel, F, 1869. Note sur la provenance exacte de ľ Helixcambojiensis, Reeve. Journal de Conchyliologie 17: 126 - 128" title="Note sur la provenance exacte de ľ Helixcambojiensis, Reeve." volume="17" year="1869">Daniel 1869</bibRefCitation>
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: 126-128.
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<bibRefCitation author="Morelet, A" journalOrPublisher="Zoologischer Teil, II; Koenigliche Geheime Ober-Hofdruckerei, Berlin" pageId="0" pageNumber="1" refId="B23" refString="Morelet, A, 1875. Series Conchyliologiques, comprenant Ľenumeration de mollusques terrestres et fluviatites, recueillis pendant le cours de differents voyages, ainsi que la description, de plusieurs especes nouvelles. IV. 4e livraison Indo Chine, 227-377." title="Series Conchyliologiques, comprenant Ľenumeration de mollusques terrestres et fluviatites, recueillis pendant le cours de differents voyages, ainsi que la description, de plusieurs especes nouvelles. IV. 4 e livraison Indo Chine, 227 - 377." year="1875">Morelet 1875</bibRefCitation>
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: 250.
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Brockhaus, Lipsiae" pageId="0" pageNumber="1" refId="B32" refString="Pfeiffer, L, 1876. Monographia Heliceorum Viventum, Supplement Tertium. Volume 7. F.A. Brockhaus, Lipsiae" title="Monographia Heliceorum Viventum, Supplement Tertium. Volume 7. F. A." year="1876">Pfeiffer 1876</bibRefCitation>
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: 78, 79.
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Brockhaus, Lipsiae" pageId="0" pageNumber="1" refId="B33" refString="Pfeiffer, L, Kobelt, W, 1881. Die Schnirkelschnecken nebst den zunaechst verwandten Gattungen. In: Systematisches Conchylien-Cabinet von Martini und Chemnitz, Ersten Bandes, zwoelfte Abtheilung, vierter Theil 1(12) (4): 595-610. [pls. 173-176]" title="Die Schnirkelschnecken nebst den zunaechst verwandten Gattungen. In: Systematisches Conchylien-Cabinet von Martini und Chemnitz, Ersten Bandes, zwoelfte Abtheilung, vierter Theil 1 (12) (4): 595 - 610. [pls. 173 - 176]" year="1881">Pfeiffer and Kobelt 1881</bibRefCitation>
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: 604, pl. 176, figs 1, 2.
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<bibRefCitation author="Tryon, Jr GW" journalOrPublisher="Academy of Natural Science Philadelphia, Philadelphia" pageId="0" pageNumber="1" refId="B46" refString="Tryon, Jr GW, 1886. Manual of Conchology, Structure and Systematic, with Illustrations of the Species, 2nd Series, Volume 2. Academy of Natural Science Philadelphia, Philadelphia" title="Manual of Conchology, Structure and Systematic, with Illustrations of the Species, 2 nd Series, Volume 2." year="1886">Tryon 1886</bibRefCitation>
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: 18, pl. 6, fig. 9.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix cambodjensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambodjensis">Helix cambodjensis</taxonomicName>
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:
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<bibRefCitation author="Mabille, J" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="117 - 138" refId="B21" refString="Mabille, J, le Mesle, G, 1869. Observation sur la fanue malacologique de la Cochinchine et du Cambodje, comprenant la description des especes nouvelles. Journal de Conchyliologie 14: 117 - 138" title="Observation sur la fanue malacologique de la Cochinchine et du Cambodje, comprenant la description des especes nouvelles." volume="14" year="1869">Mabille and le Mesle 1869</bibRefCitation>
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: 128 [incorrect subsequent spelling].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina cambodgiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambodgiensis">Nanina cambodgiensis</taxonomicName>
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:
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<bibRefCitation author="Ancey, C-F" journalOrPublisher=", Melbourne, Florida" pageId="0" pageNumber="1" refId="B3" refString="Ancey, C-F, 1887. Description of new genera or subgenera of Helicidae. The Conchologist Exchange 1: 53, 54." title="Description of new genera or subgenera of Helicidae. The Conchologist Exchange 1: 53, 54." year="1887">Ancey 1887</bibRefCitation>
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: 53 [incorrect subsequent spelling].
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<bibRefCitation author="Fischer, H" journalOrPublisher="In: Mission Pavie, Etudes diverses, Volume" pageId="0" pageNumber="1" pagination="390 - 450" refId="B10" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Mission Pavie, Etudes diverses, Volume 3: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volume="3" year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 393.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Ariophanta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariophanta (Rhyssota) cambojiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambojiensis" subGenus="Rhyssota">Ariophanta (Rhyssota) cambojiensis</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Imprimerie Dejussieu Pere et Fils, Autun" pageId="0" pageNumber="1" refId="B11" refString="Fischer, P, 1891. Catalogue et distribution geographique des Mollusques terrestres, fluviatilies & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Imprimerie Dejussieu Pere et Fils, Autun" title="Catalogue et distribution geographique des Mollusques terrestres, fluviatilies & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" year="1891">Fischer 1891</bibRefCitation>
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: 23.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia cambodjiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambodjiensis">Bertia cambodjiensis</taxonomicName>
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:
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<bibRefCitation author="Abbott, RT" journalOrPublisher=", Melbourne, Florida" pageId="0" pageNumber="1" refId="B1" refString="Abbott, RT, 1989. Compendium of Landshells. American Malacologists Inc., Melbourne, Florida" title="Compendium of Landshells. American Malacologists Inc." year="1989">Abbott 1989</bibRefCitation>
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: 127 with text figure [incorrect subsequent spelling].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia cambojiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambojiensis">Bertia cambojiensis</taxonomicName>
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:
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<bibRefCitation author="Schileyko, AA" journalOrPublisher="Annals and Magazine of Natural History, Series" pageId="0" pageNumber="1" refId="B37" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309-1466." title="Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309 - 1466." year="2003">Schileyko 2003</bibRefCitation>
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: 1362, fig. 1777.
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<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="1 - 68" refId="B38" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011</bibRefCitation>
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: 37.
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<bibRefCitation author="Thach, NN" journalOrPublisher="48 HrBooks Company, Ohio" pageId="0" pageNumber="1" refId="B44" refString="Thach, NN, 2016. Vietnamese new mollusks with 59 new species. 48HrBooks Company, Ohio" title="Vietnamese new mollusks with 59 new species." year="2016">Thach 2016</bibRefCitation>
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: 142, fig. 271.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Two syntypes from the Cuming collection. The specimen figured by
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<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B30" refString="Pfeiffer, L, 1860b. Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 2. Verlag von Theodor Fischer, Cassel, 139-303. [pls 37-72. p. 139-160, pls 37-42 (1860] [Published in parts, dates follow Johnson (1969)]" title="Novitates Conchologicae. Series Prima. Mollusca Extramarina. Band 2. Verlag von Theodor Fischer, Cassel, 139 - 303. [pls 37 - 72. p. 139 - 160, pls 37 - 42 (1860] [Published in parts, dates follow Johnson (1969)]" year="1860 b">Pfeiffer (1860b</bibRefCitation>
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: pl. 47, figs 1, 2) can be recognised by two broken growth lines; one on the border of penultimate and last whorls, and one on the last whorl close to the apertural lip (seen from umbilical view) and is here designated as the lectotype NMUK 20130220 (height 54.6 mm, width 76.2 mm;
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shells of Bertia cambojiensis A lectotype NHMUK 20130220 and B specimen NHMUK 20130833." figureDoi="10.3897/zookeys.885.38980.figure1" httpUri="https://binary.pensoft.net/fig/352321" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure1">Fig. 1A</figureCitation>
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). The remaining paralectotype NHMUK 20130219 measures height 54.4 mm and width 73.6 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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All additional specimens were from the area of Cat Tien National Park, Dong Nai Province, Vietnam (11°26.147"N, 107°25.643"E): NHMUK 20130818 from Lodge Gardens (4 specimens + 2 juveniles); NHMUK 20130833 from Cave Site (7 specimens + 1 juvenile;
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shells of Bertia cambojiensis A lectotype NHMUK 20130220 and B specimen NHMUK 20130833." figureDoi="10.3897/zookeys.885.38980.figure1" httpUri="https://binary.pensoft.net/fig/352321" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure1">Fig. 1B</figureCitation>
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; COI accession no. MN296022, 16sRNA accession no. MN296390); NHMUK 20130874 from Near Lake (6 specimens + 3 juveniles; COI accession no. MN296023, 16sRNA accession no. MN296391 and 28S accession no. MN296349). Measurements: shell height 49.9-55.7 mm, average 52.6 mm; shell width 64.4-74.5 mm, average 69.7 mm.
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</paragraph>
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<caption doi="10.3897/zookeys.885.38980.figure1" httpUri="https://binary.pensoft.net/fig/352321" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
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Shells of
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia cambojiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambojiensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Bertia cambojiensis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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lectotype NHMUK 20130220 and
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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specimen NHMUK 20130833.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shell sinistral, large, dome shape and thickened. Whorls 6 to 7, increasing regularly, slightly convex, and with wide and shallow suture. Periostracum thin to slightly thickened, corneous. Spire convex, apex obtuse, embryonic shell large with smooth surface, following whorls possess a series of thin nodules on growth lines. Last whorl well rounded. Upper shell surface glossy, varying from rich cream and brownish to blackish bands; narrow pale white to yellowish spiral band on periphery; narrow reddish-brown to dark subsutural band. Below periphery always with reddish-brown to dark colour and with broad white spiral band surrounding umbilicus. Umbilical area usually reddish-brown or darker. Aperture ovate; parietal callus translucent whitish; columella thickened and slightly dilated. Lip simple or slightly thickened in old adults. Umbilicus narrowly perforate to rimate and deep.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genital organs.</emphasis>
|
||
Atrium (at) very short (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 2). Penis (p) long cylindrical tube. Penial retractor muscle (pr) short, thickened and attached distally to penis. Epiphallus (e) cylindrical tube, about half of penis length and slightly smaller diameter than distal penis. Vas deferens (vd) relatively smaller diameter and thin tube extending from free oviduct (fo) and entering epiphallus, apically; flagellum absent (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2A</figureCitation>
|
||
). Internal wall of penis with large penial pilasters (pp) for nearly entire length of chamber; proximally with smooth surface and distally with very thin crenellations on surface; penial verge absent (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Gametolytic organ (go; duct and sac undifferentiated) proximal to genital opening about one-fourth of amatorial organ length. Slightly swollen proximally, then tapering to small, long cylindrical tube and attached to ovary with thin connective tissues. Amatorial organ (am) well-developed enlarged cylinder; proximally attached to atrium. Amatorial organ glands (amg) enlarged, composed distally of four major lobes bounded to amatorial organ by thin connective tissue. Each of four major lobes of the amatorial organ gland extend proximally into thick amatorial organ ducts (amd) that are twisted together and bound with thin connective tissue before entering the distal tip of the amatorial organ (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2A</figureCitation>
|
||
). Internal wall sculpture of amatorial organ: proximally smooth surface for about one-third of chamber; distally consists of smooth surface of enlarged longitudinal amatorial organ pilasters (amp). Amatorial organ papilla small, short and conical, tipped by a large and long blackish spike (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Vagina (v) long enlarged cylinder, about the same length as penis. Free oviduct (fo) cylindrical tube; oviduct (ov) long with lobules; prostate gland (pg) bound to oviduct. Most of albumen gland, hermaphroditic duct and gland missing from figured specimen (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2A</figureCitation>
|
||
). Internal wall of vagina sculptured with uniform scale-like or triangular lingulate pilasters (vp), varying in size from small to large (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<taxonomicName class="Bivalvia" family="Limidae" genus="Radula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radula" order="Limida" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Teeth arranged in wide angle V-shaped rows with approximately 245 teeth with formula ((124-118)-1-(120-122)). Central tooth symmetric unicuspid and triangular. Lateral and marginal teeth undifferentiated, slightly curved unicuspid, triangular, and inclined towards central tooth (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2E</figureCitation>
|
||
). Inner teeth similar in shape and size to central teeth and then gradually reducing to slender and elongate sword-shaped with pointed cusp; outermost teeth slightly shorter with pointed tip (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Genitalia and radula of Bertia cambojiensis, NHMUK 20130833. A overview of genital system B-D internal wall sculpture of B penis C amatorial organ and D vagina E, F SEM image of radula E central and inner latero-marginal teeth and F outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by ' C'." figureDoi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.885.38980.figure2">Fig. 2F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.885.38980.figure2" httpUri="https://binary.pensoft.net/fig/352322" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
|
||
Genitalia and radula of
|
||
<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia cambojiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Bertia cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
, NHMUK 20130833.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
overview of genital system
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<normalizedToken originalValue="B–D">B-D</normalizedToken>
|
||
</emphasis>
|
||
internal wall sculpture of
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
penis
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
amatorial organ and
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
vagina
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||
SEM image of radula
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
central and inner latero-marginal teeth and
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
outermost teeth. Numbers indicate tooth order from lateral to marginal end; central tooth indicated by
|
||
<normalizedToken originalValue="‘C’">'C'</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">External features</emphasis>
|
||
.
|
||
</emphasis>
|
||
Living snail with long, blackish-brown tentacles. Skin reticulated brown with black reticulations around head. Foot sole relatively elongated, broad and unipartite. Sole of foot brownish to orange and unspotted; side of body brownish. Tail curved mid-dorsally, tall dome-shaped in cross section. Caudal horn not overhanging; caudal foss long vertical slit in tail above sole margin. Typical aulacopoda with well-defined pedal groove.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Mantle collar large and shell lobes thickened, shell lappet absent. Right dorsal lobe (right side of anus) large and thickened. Left dorsal lobe (left side of anus) composed of thick crescentric anterior left dorsal lobe and thin elongated posterior left dorsal lobe. Pulmonary cavity typically sigmurethran.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The range of
|
||
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
is likely to be confined to localised forest patches within Dong Nai, Binh Thuan, Lam Dong and Binh Phuc provinces. Records far outside of this range such as Ba Vi National Park (
|
||
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="1 - 68" refId="B38" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011</bibRefCitation>
|
||
) are very doubtful and to date, no specific localities have been recorded for Cambodia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The specimens from southern Vietnam that we examined exhibit clear differences in genital anatomy from those described by
|
||
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Annals and Magazine of Natural History, Series" pageId="0" pageNumber="1" refId="B37" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309-1466." title="Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309 - 1466." year="2003">Schileyko (2003</bibRefCitation>
|
||
: 1362, fig. 1777b, c) that were based on a specimen in the
|
||
<normalizedToken originalValue="Muséum">Museum</normalizedToken>
|
||
National
|
||
<normalizedToken originalValue="ďHistoire">ďHistoire</normalizedToken>
|
||
Naturelle, Paris, recorded from Cambodia. The Paris specimen exhibits a long, cylindrical gametolytic duct (spermathecal stalk in Schileyko) and a bulbous gametolytic sac (receptaculum seminalis in Schileyko), distally pointed. The amatorial organ contains numerous amatorial organ ducts. In the NHM specimens the gametolytic duct is short and the amatorial organ includes four amatorial organ ducts. On the basis of numerous studies in the reproductive anatomy of pulmonate snails, these character differences strongly suggest species level distinction. The shell of the Paris specimen was not figured by Schileyko, his figured shell of
|
||
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
(fig. 1777a) being from the Naturalis Museum, Leiden, which was also recorded as being from Cambodia.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
In December 1858, the French explorer and naturalist Henri Mouhot set off from Bangkok by sea to the port of Komput (Kampot), Cambodia, from where he continued on his destination to Brelum in Annam (Vietnam), He reached the Catholic Mission in Brelum in August 1859.
|
||
<bibRefCitation author="Stuart, BL" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="129 - 155" refId="B41" refString="Stuart, BL, Sok, K, Neang, T, 2006. A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology 54: 129 - 155" title="A collection of amphibians and reptiles from hilly eastern Cambodia." volume="54" year="2006">Stuart et al. (2006)</bibRefCitation>
|
||
state that Brelum was considered to be within the confines of Cambodia at the time of
|
||
<normalizedToken originalValue="Mouhot’s">Mouhot's</normalizedToken>
|
||
visit. However,
|
||
<bibRefCitation author="Mouhot, H" journalOrPublisher="Murray, London, Volume 1" pageId="0" pageNumber="1" refId="B25" refString="Mouhot, H, 1864a. Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860. Murray, London, Volume 1" title="Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860." year="1864 a">Mouhot (1864a</bibRefCitation>
|
||
: 237) unequivocally described how 'It took us two long
|
||
<normalizedToken originalValue="days’">days'</normalizedToken>
|
||
journey to reach
|
||
<normalizedToken originalValue="Brelum’">Brelum'</normalizedToken>
|
||
after having passed the Cambodian border town of Pump-Ka-Daye. Brelum was
|
||
<normalizedToken originalValue="Mouhot’s">Mouhot's</normalizedToken>
|
||
collecting base for the following three months before setting off on his return journey in November (
|
||
<bibRefCitation author="Mouhot, H" journalOrPublisher="Murray, London, Volume 1" pageId="0" pageNumber="1" refId="B25" refString="Mouhot, H, 1864a. Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860. Murray, London, Volume 1" title="Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860." year="1864 a">Mouhot 1864a</bibRefCitation>
|
||
). During his stay in Brelum his host, the missionary R.P. Guillon, wanted Mouhot to sample the local escargot, which proved to be
|
||
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
. Staggered by the beauty of this snail Mouhot set about obtaining examples for his collections from the local area, where it was abundant (
|
||
<bibRefCitation author="Daniel, F" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="126 - 128" refId="B8" refString="Daniel, F, 1869. Note sur la provenance exacte de ľ Helixcambojiensis, Reeve. Journal de Conchyliologie 17: 126 - 128" title="Note sur la provenance exacte de ľ Helixcambojiensis, Reeve." volume="17" year="1869">Daniel 1869</bibRefCitation>
|
||
).
|
||
<bibRefCitation author="Daniel, F" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="126 - 128" refId="B8" refString="Daniel, F, 1869. Note sur la provenance exacte de ľ Helixcambojiensis, Reeve. Journal de Conchyliologie 17: 126 - 128" title="Note sur la provenance exacte de ľ Helixcambojiensis, Reeve." volume="17" year="1869">Daniel (1869)</bibRefCitation>
|
||
went on to state that
|
||
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
had not been found in Cambodia. Historical records may all relate to the mistaken assumption that the type locality was Cambodia. Mouhot was dependant on the sale of his collections to finance his fieldwork and it is likely that he shipped a large number of specimens of
|
||
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
to his agent in London, Samuel Stevens, who would have sold them on to museums and collectors throughout Europe, presumably labelled with the locality
|
||
<normalizedToken originalValue="‘Cambodia’">'Cambodia'</normalizedToken>
|
||
. The field and travel conditions impacted severely on
|
||
<normalizedToken originalValue="Mouhot’s">Mouhot's</normalizedToken>
|
||
collections and he clearly had insufficient time to arrange them in good order before they were shipped. Writing to Stevens from Pinhalu, near Phnom Penh, on 20th December, 1859
|
||
<bibRefCitation author="Mouhot, H" journalOrPublisher="Murray, London, Volume 2" pageId="0" pageNumber="1" refId="B26" refString="Mouhot, H, 1864b. Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860. Murray, London, Volume 2" title="Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860." year="1864 b">Mouhot (1864b</bibRefCitation>
|
||
: 248) commented:
|
||
<normalizedToken originalValue="‘…">'...</normalizedToken>
|
||
I have little time to give you any details as to what I despatch from Komput and
|
||
<normalizedToken originalValue="Singapore’">Singapore'</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
According to
|
||
<bibRefCitation author="Ashburton, J" editor="Mouhot, H" journalOrPublisher="Murray, London, Volume" pageId="0" pageNumber="1" pagination="296 - 300" refId="B4" refString="Ashburton, J, 1864. Paper read at the Royal Geographical Society 10th March, 1862. In: Mouhot, H, Ed., Travels in the Central Parts of Indo-China (Siam), Cambodia, and Laos, During the Years 1858, 1859, and 1860. . Murray, London, Volume 2: 296 - 300" title="Paper read at the Royal Geographical Society 10 th March, 1862." volume="2" volumeTitle="Travels in the Central Parts of Indo-China (Siam), Cambodia, and Laos, During the Years 1858, 1859, and 1860." year="1864">Ashburton (1864)</bibRefCitation>
|
||
, the location of Brelum, the type locality, is 11°58"N, 107°12"E, which is at an altitude of 460 m (Google Earth). This location is 30 km north of the north-western boundary of Cat Tien National Park and some 94 km north and west of the entrance to Cat Tien National Park. However, there remains some doubt as to the exact location of Brelum. Under the heading of a letter to Stevens,
|
||
<bibRefCitation author="Mouhot, H" journalOrPublisher="Murray, London, Volume 2" pageId="0" pageNumber="1" refId="B26" refString="Mouhot, H, 1864b. Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860. Murray, London, Volume 2" title="Travels in the central parts of Indo-China (Siam), Cambodia, and Laos, during the years 1858, 1859, and 1860." year="1864 b">Mouhot (1864b</bibRefCitation>
|
||
: 241) stated: 'Brelum, among the savage
|
||
<normalizedToken originalValue="Stiêns">Stiens</normalizedToken>
|
||
, lat,
|
||
<geoCoordinate degrees="11" direction="north" minutes="46" orientation="latitude" precision="15" seconds="30" value="11.775">11°46'30"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="103" direction="west" minutes="3" orientation="longitude" precision="925" value="-103.05">103°3'W</geoCoordinate>
|
||
merid. of Paris, 15th October, 1859'. This is clearly wrong because in addition to W (west) it should be E (east) and in the following text he gives exactly the same geo-reference for Pinhalu that he states to be about nine miles from the capital, Phnom Penh
|
||
<geoCoordinate degrees="11" direction="north" minutes="46" orientation="latitude" precision="15" seconds="58" value="11.782778">11°46'58"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="104" direction="east" minutes="22" orientation="longitude" precision="15" seconds="59" value="104.38306">104°22'59"E</geoCoordinate>
|
||
(Google Earth) whereas
|
||
<normalizedToken originalValue="Mouhot’s">Mouhot's</normalizedToken>
|
||
reading
|
||
<geoCoordinate degrees="11" direction="north" minutes="46" orientation="latitude" precision="15" seconds="30" value="11.775">11°46'30"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="103" direction="west" minutes="3" orientation="longitude" precision="925" value="-103.05">103°3'W</geoCoordinate>
|
||
Paris (= 2°20'14.03" east of Greenwich) = 105°23'14"E, is some 54 km to the east and north of Phnom Penh (Google Earth).
|
||
<bibRefCitation author="Kottelat, M" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="0" pageNumber="1" pagination="1 - 19" refId="B18" refString="Kottelat, M, Tan, HH, 2017. Three new species of archerfishes from the freshwaters of Southeast Asia (Teleostei: Toxotidae) and notes on Henri Mouhot's fish collections. Ichthyological Exploration of Freshwaters 952: 1 - 19" title="Three new species of archerfishes from the freshwaters of Southeast Asia (Teleostei: Toxotidae) and notes on Henri Mouhot's fish collections." volume="952" year="2017">Kottelat and Tan (2017)</bibRefCitation>
|
||
identify Bro Lam Phe,
|
||
<geoCoordinate degrees="11" direction="north" minutes="56" orientation="latitude" precision="925" value="11.933333">11°56'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="106" direction="east" minutes="40" orientation="longitude" precision="925" value="106.666664">106°40'E</geoCoordinate>
|
||
, in Loc Ninh District, Binh Phuoc Province, Vietnam, as the location of Brelum. This places Brelum at 100 m altitude and only 2 km from the closest boundary with Cambodia, which at this point follows the course of a meandering tributary of the Mekong. It is some 70 km from the boundary of Cat Tien National Park and 100 km from the main park entrance. It seems that Mouhot would have been most unlikely to retrace his steps and cross the river into Cambodia before his return journey and we conclude that he only collected
|
||
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
from the vicinity of Brelum in Vietnam. Nevertheless, being in close proximity to the Cambodian border it does seem likely that
|
||
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
|
||
</taxonomicName>
|
||
will occur in this area of Cambodia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |