464 lines
68 KiB
XML
464 lines
68 KiB
XML
<document id="876A7D6EDF57EAECF0394D7859E7A4BF" ID-CLB-Dataset="26847" ID-DOI="10.11646/zootaxa.4604.1.1" ID-GBIF-Dataset="b5902a9d-c38b-462f-ad8a-95693c1822dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="2783674" ID-ZooBank="E6D82300-8DC6-4560-A945-8F29D3B3D501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1557723545002" checkinUser="plazi" docAuthor="Ruta, Rafał" docDate="2019" docId="71140E6CF26B143E73B5E9BD50332E91" docLanguage="en" docName="zootaxa.4604.1.1.pdf" docOrigin="Zootaxa 4604 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvariopsis Ruta 2019, gen. nov." docType="treatment" docVersion="12" lastPageNumber="9" masterDocId="8D2D7614F26914367322EA4454252806" masterDocTitle="Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="3" updateTime="1698728494285" updateUser="ExternalLinkService">
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<mods:title id="32FBF55C62A79C0EB2097BD857701395">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<mods:name id="988B255FB015CB61254329CAE67AAAE9" type="personal">
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<mods:namePart id="1F4DBB09D93F4EC7876CF62638992311">Ruta, Rafał</mods:namePart>
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<mods:typeOfResource id="999DF69F3412E141921DD53F6680E28A">text</mods:typeOfResource>
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<mods:title id="A0F6912B912C63F6B49054EA0421F436">Zootaxa</mods:title>
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<mods:part id="76BC95DDB249C5E6B8EF357F1B7683C3">
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<mods:date id="F6831EDD7CE8B034D80CD6658523351D">2019</mods:date>
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<mods:number id="3C177A2119BC0085250A7FE212669C7A">2019-05-13</mods:number>
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<mods:number id="BBB0182FAB5FB057F82C593F1FA152E7">4604</mods:number>
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<mods:identifier id="6B155F354BB75FCBC5DF2D9351DB033F" type="DOI">10.11646/zootaxa.4604.1.1</mods:identifier>
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<mods:identifier id="38CE81F99B5D27FC31263C54310CDBDA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="71140E6CF26B143E73B5E9BD50332E91" ID-GBIF-Taxon="157135220" LSID="urn:lsid:plazi:treatment:71140E6CF26B143E73B5E9BD50332E91" httpUri="http://treatment.plazi.org/id/71140E6CF26B143E73B5E9BD50332E91" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3">
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<subSubSection id="B1A7ECF1F26B143473B5E9BD55DF2C15" box="[151,506,1017,1044]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="F902BF7AF26B143473B5E9BD55DF2C15" blockId="2.[151,506,1017,1079]" box="[151,506,1017,1044]" pageId="2" pageNumber="3">
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<heading id="A24A0816F26B143473B5E9BD55DF2C15" bold="true" box="[151,506,1017,1044]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="CBC96368F26B143473B5E9BD55DF2C15" bold="true" box="[151,506,1017,1044]" pageId="2" pageNumber="3">
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Genus
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<taxonomicName id="3EBDC4F9F26B143473CCE9BD55A12C12" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[238,388,1017,1044]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="CBC96368F26B143473CCE9BD55A12C12" bold="true" box="[238,388,1017,1044]" italics="true" pageId="2" pageNumber="3">Calvariopsis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D0FADE13F26B143472AEE9BD55DF2C15" box="[396,506,1017,1043]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F26B143473B5EE5955E42C31" box="[151,449,1052,1079]" pageId="2" pageNumber="3" type="description">
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<paragraph id="F902BF7AF26B143473B5EE5955E42C31" blockId="2.[151,506,1017,1079]" box="[151,449,1052,1079]" pageId="2" pageNumber="3">
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(
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<figureCitation id="6186A3FFF26B143473BDEE5955372C31" box="[159,274,1052,1079]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="." captionStart-12="FIGURE 13" captionStart-13="FIGURE 14" captionStart-14="FIGURE 15" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="3.[151,250,1767,1792]" captionStartId-1="4.[151,250,1057,1082]" captionStartId-10="14.[151,250,1377,1402]" captionStartId-11="15.[404,408,1541,1564]" captionStartId-12="17.[151,250,1690,1715]" captionStartId-13="18.[151,250,1731,1756]" captionStartId-14="20.[151,250,1137,1162]" captionStartId-2="5.[151,250,1905,1930]" captionStartId-3="6.[151,250,1424,1449]" captionStartId-4="7.[151,250,1512,1537]" captionStartId-5="8.[151,250,1158,1183]" captionStartId-6="9.[151,250,1631,1656]" captionStartId-7="10.[151,250,1352,1377]" captionStartId-8="12.[151,250,1323,1348]" captionStartId-9="13.[151,250,1849,1874]" captionTargetBox-0="[183,1404,196,1722]" captionTargetBox-1="[164,1423,199,1014]" captionTargetBox-10="[153,1435,181,1322]" captionTargetBox-11="[153,1435,668,1486]" captionTargetBox-12="[153,1435,396,1639]" captionTargetBox-13="[184,1403,897,1694]" captionTargetBox-14="[186,1401,251,1088]" captionTargetBox-2="[165,1422,181,1857]" captionTargetBox-3="[178,1410,181,1383]" captionTargetBox-4="[165,1403,204,1476]" captionTargetBox-5="[153,1435,181,1105]" captionTargetBox-6="[165,1422,181,1592]" captionTargetBox-7="[153,1435,181,1302]" captionTargetBox-8="[158,1430,181,1270]" captionTargetBox-9="[178,1410,181,1810]" captionTargetId-0="figure@3.[183,1404,196,1722]" captionTargetId-1="figure@4.[164,1423,199,1014]" captionTargetId-10="figure@14.[153,1438,181,1322]" captionTargetId-11="figure@15.[153,1438,668,1488]" captionTargetId-12="figure@17.[153,1438,417,1639]" captionTargetId-13="figure@18.[181,1405,897,1694]" captionTargetId-14="figure@20.[186,1403,272,1088]" captionTargetId-2="figure@5.[165,1424,181,1857]" captionTargetId-3="figure@6.[177,1411,179,1383]" captionTargetId-4="figure@7.[165,1424,181,1476]" captionTargetId-5="figure@8.[153,1438,181,1105]" captionTargetId-6="figure@9.[165,1424,181,1592]" captionTargetId-7="figure@10.[153,1438,181,1302]" captionTargetId-8="figure@12.[158,1433,181,1270]" captionTargetId-9="figure@13.[178,1412,181,1810]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="14" captionTargetPageId-11="17" captionTargetPageId-12="17" captionTargetPageId-13="18" captionTargetPageId-14="20" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." captionText-1="FIGURE 2. Calvariopsis gen. nov., females, dorsal view.A) C. sp., Colombia; B) C. sp., Costa Rica; C) C. sp., Ecuador; D) C. sp., Ecuador; E) C. sp., French Guiana; F) C. sp., Panama; G) C. sp., Paraguay; H) C. sp., Peru. Scale bars = 1.0 mm." captionText-10="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." captionText-11="." captionText-12="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." captionText-13="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." captionText-14="FIGURE 15. Calvariopsis pittieri sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." captionText-2="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." captionText-3="FIGURE 4. Calvariopsis gen. nov., mouthparts.A–C) C. yanayacuense sp. nov.; D–F) C. sp., group II, female (French Guiana). A, D) mandible; B, E) maxilla; C, F) labium." captionText-4="FIGURE 5. Calvariopsis gen. nov., wing. A) C. yanayacuense sp. nov.; B) C. bituberculata sp. nov." captionText-5="FIGURE 6. Calvariopsis yanayacuense sp. nov., abdomen, tergites and ventrites." captionText-6="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." captionText-7="FIGURE 8. Calvariopsis gen. nov., group I (A, C, E–F) and group II (B, D), females, apical portion of elytra, excitators. A) C. pittieri; B) C. brasiliense, an arrow indicates patch of modified setae; C) C. sp., Colombia, Gorgona Island; D) C. sp., Ecuador, an arrow indicates patch of modified setae; E) C. sp., Paraguay; F) C. sp., Peru." captionText-8="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." captionText-9="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri-0="https://zenodo.org/record/2783682/files/figure.png" httpUri-1="https://zenodo.org/record/2783686/files/figure.png" httpUri-10="https://zenodo.org/record/2783738/files/figure.png" httpUri-11="https://zenodo.org/record/2783740/files/figure.png" httpUri-12="https://zenodo.org/record/2783740/files/figure.png" httpUri-13="https://zenodo.org/record/2783746/files/figure.png" httpUri-14="https://zenodo.org/record/2783750/files/figure.png" httpUri-2="https://zenodo.org/record/2783692/files/figure.png" httpUri-3="https://zenodo.org/record/2783698/files/figure.png" httpUri-4="https://zenodo.org/record/2783704/files/figure.png" httpUri-5="https://zenodo.org/record/2783710/files/figure.png" httpUri-6="https://zenodo.org/record/2783718/files/figure.png" httpUri-7="https://zenodo.org/record/2783720/files/figure.png" httpUri-8="https://zenodo.org/record/2783724/files/figure.png" httpUri-9="https://zenodo.org/record/2783730/files/figure.png" pageId="2" pageNumber="3">Figs 1–15</figureCitation>
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,
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<figureCitation id="6186A3FFF26B1434723CEE59551F2C31" box="[286,314,1053,1079]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="2" pageNumber="3">17</figureCitation>
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–18, 20–27)
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F26B143473B5EE2157C42C79" box="[151,993,1124,1151]" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="F902BF7AF26B143473B5EE2157C42C79" blockId="2.[151,1437,1124,1799]" box="[151,993,1124,1151]" pageId="2" pageNumber="3">
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<emphasis id="CBC96368F26B143473B5EE2155112C79" bold="true" box="[151,308,1124,1151]" pageId="2" pageNumber="3">
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<typeStatus id="260601D8F26B143473B5EE2154F72C79" box="[151,210,1125,1151]" pageId="2" pageNumber="3">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="3EBDC4F9F26B1434721EEE21564A2C79" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[316,623,1125,1151]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
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<emphasis id="CBC96368F26B1434721EEE21564A2C79" box="[316,623,1125,1151]" italics="true" pageId="2" pageNumber="3">Calvariopsis yanayacuense</emphasis>
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</taxonomicName>
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<emphasis id="CBC96368F26B14347154EE2156EA2C79" bold="true" box="[630,719,1125,1151]" pageId="2" pageNumber="3">
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<taxonomicNameLabel id="D0FADE13F26B14347154EE2156EA2C79" box="[630,719,1125,1151]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F26B143473E5EECC501B2D7D" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="F902BF7AF26B143473E5EECC501B2D7D" blockId="2.[151,1437,1124,1799]" pageId="2" pageNumber="3">
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<emphasis id="CBC96368F26B143473E5EECC55672CA5" bold="true" box="[199,322,1160,1187]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
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Head strongly transverse, eyes not protuberant, subantennal grooves wide and deep, antennae filiform, antennomere 1 wide, with sharp anterior edge, labrum narrow, prosternal process elongate, lanceolate with rounded or subtriangular apical portion, mesoventral process as long as wide to transverse. Mesoventrite and basal portion of metaventrite with sphaerical cuticular structures (
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<figureCitation id="6186A3FFF26B14347065EEB157852D09" box="[839,928,1269,1295]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="2" pageNumber="3">Fig. 3C</figureCitation>
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), apical portions of female elytra often with excitators. Hind legs not saltatorial, femora not swollen. Penis elongated, with distinct parameroids and well developed trigonium. Tegmen elongate, U-shaped, parameres free or fused in apical portion. Tergite VIII with various modifications of apical margin. Bursa with single sclerite of species-specific shape.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F26B143E73E5EFC0505B2DE5" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3" type="description">
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<paragraph id="F902BF7AF26B143473E5EFC055222DE1" blockId="2.[151,1437,1124,1799]" pageId="2" pageNumber="3">
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<emphasis id="CBC96368F26B143473E5EFC0557C2D99" bold="true" box="[199,345,1412,1439]" pageId="2" pageNumber="3">Description.</emphasis>
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Male. Body oval, TL
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<quantity id="3E45129FF26B14347172EFC156FC2D99" box="[592,729,1413,1439]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.6" metricValueMin="1.7" pageId="2" pageNumber="3" unit="mm" value="2.65" valueMax="3.6" valueMin="1.7">1.7–3.6 mm</quantity>
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, strongly convex, usually brown to black, in some species with contrasting orange and brown or black pattern, covered with brownish, suberect setae, punctation of dorsum not granulate.
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</paragraph>
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<paragraph id="F902BF7AF26B143273E5EFB555952E99" blockId="2.[151,1437,1124,1799]" lastBlockId="4.[151,1437,1164,1983]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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Head (
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<figureCitation id="6186A3FFF26B14347230EFB555A92E0D" box="[274,396,1521,1547]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="2" pageNumber="3">Figs 3B, E</figureCitation>
|
||
) small, wider than long; eyes large, oval, not protuberant, finely facetted; tempora relatively short, distance from posterior margin of eye to occipital ridge ca. 1/2 diameter of the eye, strongly converging basally; distance from ventral margin of eye to subgenal ridge small, equal to ca. 1/6 diameter of the eye when observed from ventral side; supraantennal ridges not elevated, joined mesally to clypeal margin; frons slightly convex; subantennal grooves deep, ventrally reaching subgenal ridge; indistinct elevated area runs obliquely between margin of eye and subgenal ridge. Frontoclypeal suture absent; clypeus transverse, ca. 4× wider than long, with straight anterior margin, lateral margins subtly converging posteriorly. Anteclypeus present, shorter than clypeus. Ventral portion of head between subgenal ridges slightly concave, gular area convex, gular ridges well developed. Subgenal ridges well marked, without buttonhole configuration (
|
||
<emphasis id="CBC96368F26D143271DDEEC957182CA0" box="[767,829,1165,1190]" italics="true" pageId="4" pageNumber="5">sensu</emphasis>
|
||
Zwick 2013b). Antennae filiform, reaching basal portion of elytra; antennomere 1 widest, subcylindrical, slightly flattened, with sharp ridge on anterior margin; antennomere 2 subcylindrical, narrower and ca. 2× shorter than antennomere 1; antennomere 3 very narrow, subconical, ca. 2× shorter than antennomere 2; antennomeres 4–11 slightly widening apically, antennomere 4 slightly longer than antennomeres 5–11, which have similar length. Labrum narrower than clypeus, transverse, ca. 2.0× wider than long, anterior margin slightly emarginated, anterolateral angles rounded. Mandibles (
|
||
<figureCitation id="6186A3FFF26D1432777DEF0450C52D5D" box="[1119,1248,1344,1371]" captionStart="FIGURE 4" captionStartId="6.[151,250,1424,1449]" captionTargetBox="[178,1410,181,1383]" captionTargetId="figure@6.[177,1411,179,1383]" captionTargetPageId="6" captionText="FIGURE 4. Calvariopsis gen. nov., mouthparts.A–C) C. yanayacuense sp. nov.; D–F) C. sp., group II, female (French Guiana). A, D) mandible; B, E) maxilla; C, F) labium." httpUri="https://zenodo.org/record/2783698/files/figure.png" pageId="4" pageNumber="5">Figs 4A, D</figureCitation>
|
||
) symmetrical or subtly asymmetrical, ca. 2× longer than wide at bases, abruptly curved in apical 1/4, apices unidentate, outer margin covered with setae; inner margin of mandibles with sharp ridge and subtriangular denticle; mola smoothly rounded, without spines. Maxillary palpi (
|
||
<figureCitation id="6186A3FFF26D14327128EFE856AF2DC1" box="[522,650,1452,1479]" captionStart="FIGURE 4" captionStartId="6.[151,250,1424,1449]" captionTargetBox="[178,1410,181,1383]" captionTargetId="figure@6.[177,1411,179,1383]" captionTargetPageId="6" captionText="FIGURE 4. Calvariopsis gen. nov., mouthparts.A–C) C. yanayacuense sp. nov.; D–F) C. sp., group II, female (French Guiana). A, D) mandible; B, E) maxilla; C, F) labium." httpUri="https://zenodo.org/record/2783698/files/figure.png" pageId="4" pageNumber="5">Figs 4B, E</figureCitation>
|
||
): palpomere 1 narrow, subconical, 2 longer, subcylindrical, 3 longest and widest, subconical, 4 narrower and shorter than 3, subconical. Galea with subparallel sides, slightly shorter than lacinia, with two fan-shaped rows of setae at apex; inner margin of lacinia with straight setae, setae on apical portion curved. Mentum (
|
||
<figureCitation id="6186A3FFF26D143272B6EC5C56282E35" box="[404,525,1560,1587]" captionStart="FIGURE 4" captionStartId="6.[151,250,1424,1449]" captionTargetBox="[178,1410,181,1383]" captionTargetId="figure@6.[177,1411,179,1383]" captionTargetPageId="6" captionText="FIGURE 4. Calvariopsis gen. nov., mouthparts.A–C) C. yanayacuense sp. nov.; D–F) C. sp., group II, female (French Guiana). A, D) mandible; B, E) maxilla; C, F) labium." httpUri="https://zenodo.org/record/2783698/files/figure.png" pageId="4" pageNumber="5">Figs 4C, F</figureCitation>
|
||
) slightly transverse (W/L = 1.4), subtrapezoidal. Ligula narrow at base, widened apically, transversely oval, apical margin membranous, rounded, covered with setae. Labial palpomeres: palpomere 1 long, narrow, palpomere 2 wider, subconical, slightly curved; apical palpomere narrow, subconical, arising subapically from preapical one.
|
||
</paragraph>
|
||
<caption id="ADC2EFF2F26A143573B5ECA350D42FD1" httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,250,1767,1792]" targetBox="[183,1404,196,1722]" targetPageId="3">
|
||
<paragraph id="F902BF7AF26A143573B5ECA350D42FD1" blockId="3.[151,1436,1767,2008]" pageId="3" pageNumber="4">
|
||
<emphasis id="CBC96368F26A143573B5ECA355362F06" bold="true" box="[151,275,1767,1792]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
|
||
<taxonomicName id="3EBDC4F9F26A14357238ECAC55B92F06" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[282,412,1768,1792]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CBC96368F26A14357238ECAC55B92F06" box="[282,412,1768,1792]" italics="true" pageId="3" pageNumber="4">Calvariopsis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357280ECAC56252F06" bold="true" box="[418,512,1768,1792]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357280ECAC56252F06" box="[418,512,1768,1792]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal view. A)
|
||
<taxonomicName id="3EBDC4F9F26A1435718CECAC57152F06" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[686,816,1768,1792]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A1435718CECAC57152F06" box="[686,816,1768,1792]" italics="true" pageId="3" pageNumber="4">C. borowieci</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357015ECAC57AD2F06" bold="true" box="[823,904,1768,1792]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357015ECAC57AD2F06" box="[823,904,1768,1792]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; B)
|
||
<taxonomicName id="3EBDC4F9F26A143570D6ECAC50502F06" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1012,1141,1768,1792]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143570D6ECAC50502F06" box="[1012,1141,1768,1792]" italics="true" pageId="3" pageNumber="4">C. borowieci</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A1435775EECAC50E82F06" bold="true" box="[1148,1229,1768,1792]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A1435775EECAC50E82F06" box="[1148,1229,1768,1792]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female; C)
|
||
<emphasis id="CBC96368F26A1435766EECAC54DD2F22" italics="true" pageId="3" pageNumber="4">C. four- gassiense</emphasis>
|
||
<emphasis id="CBC96368F26A14357222ED4855772F22" bold="true" box="[256,338,1804,1828]" pageId="3" pageNumber="4">sp. nov.</emphasis>
|
||
, male; D)
|
||
<taxonomicName id="3EBDC4F9F26A143572E0ED4856762F22" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[450,595,1804,1828]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="panamense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143572E0ED4856762F22" box="[450,595,1804,1828]" italics="true" pageId="3" pageNumber="4">C. panamense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357178ED4856892F22" bold="true" box="[602,684,1804,1828]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357178ED4856892F22" box="[602,684,1804,1828]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; E)
|
||
<taxonomicName id="3EBDC4F9F26A14357038ED4857BA2F22" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[794,927,1804,1828]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A14357038ED4857BA2F22" box="[794,927,1804,1828]" italics="true" pageId="3" pageNumber="4">C. peruviana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357085ED4857DD2F22" bold="true" box="[935,1016,1804,1828]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357085ED4857DD2F22" box="[935,1016,1804,1828]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Dept. Cusco, male; F)
|
||
<taxonomicName id="3EBDC4F9F26A143577D3ED4851522F22" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1265,1399,1804,1828]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143577D3ED4851522F22" box="[1265,1399,1804,1828]" italics="true" pageId="3" pageNumber="4">C. peruviana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A1435765CED4854E62F4E" bold="true" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A1435765CED4854E62F4E" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Callanga, male; G)
|
||
<taxonomicName id="3EBDC4F9F26A143572BBED74563B2F4E" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[409,542,1840,1864]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143572BBED74563B2F4E" box="[409,542,1840,1864]" italics="true" pageId="3" pageNumber="4">C. peruviana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357106ED7456502F4E" bold="true" box="[548,629,1840,1864]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357106ED7456502F4E" box="[548,629,1840,1864]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Dept. Junín, male; H)
|
||
<taxonomicName id="3EBDC4F9F26A14357044ED7457922F4E" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[870,951,1840,1864]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="picta" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A14357044ED7457922F4E" box="[870,951,1840,1864]" italics="true" pageId="3" pageNumber="4">C. picta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A1435709CED74502A2F4E" bold="true" box="[958,1039,1840,1864]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A1435709CED74502A2F4E" box="[958,1039,1840,1864]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Callanga, male; I)
|
||
<taxonomicName id="3EBDC4F9F26A143577F9ED74511B2F4E" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1243,1342,1840,1864]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143577F9ED74511B2F4E" box="[1243,1342,1840,1864]" italics="true" pageId="3" pageNumber="4">C. pittieri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357666ED7451B02F4E" bold="true" box="[1348,1429,1840,1864]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357666ED7451B02F4E" box="[1348,1429,1840,1864]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; J)
|
||
<taxonomicName id="3EBDC4F9F26A143573CCED1055752F6A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[238,336,1876,1900]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143573CCED1055752F6A" box="[238,336,1876,1900]" italics="true" pageId="3" pageNumber="4">C. pittieri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357274ED1055832F6A" bold="true" box="[342,422,1876,1900]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357274ED1055832F6A" box="[342,422,1876,1900]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female; K)
|
||
<taxonomicName id="3EBDC4F9F26A14357107ED10571E2F6A" authority="(Pic, 1931)" authorityName="Pic" authorityYear="1931" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[549,827,1875,1900]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sculptipenne">
|
||
<emphasis id="CBC96368F26A14357107ED1056E72F6A" box="[549,706,1876,1900]" italics="true" pageId="3" pageNumber="4">C. sculptipenne</emphasis>
|
||
(Pic, 1931)
|
||
</taxonomicName>
|
||
, female; L)
|
||
<taxonomicName id="3EBDC4F9F26A14357096ED1050702F6A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[948,1109,1876,1900]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A14357096ED1050702F6A" box="[948,1109,1876,1900]" italics="true" pageId="3" pageNumber="4">C. venezuelense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357779ED10508E2F6A" bold="true" box="[1115,1195,1876,1900]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357779ED10508E2F6A" box="[1115,1195,1876,1900]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; M–O)
|
||
<emphasis id="CBC96368F26A14357618ED1054CC2F96" italics="true" pageId="3" pageNumber="4">C. yanay- acuense</emphasis>
|
||
<emphasis id="CBC96368F26A143573CDED3C551A2F96" bold="true" box="[239,319,1912,1936]" pageId="3" pageNumber="4">sp. nov.</emphasis>
|
||
, males; P)
|
||
<taxonomicName id="3EBDC4F9F26A14357292ED3C567E2F96" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[432,603,1912,1936]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A14357292ED3C567E2F96" box="[432,603,1912,1936]" italics="true" pageId="3" pageNumber="4">C. yanayacuense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A14357142ED3C56952F96" bold="true" box="[608,688,1912,1936]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A14357142ED3C56952F96" box="[608,688,1912,1936]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female; Q)
|
||
<taxonomicName id="3EBDC4F9F26A1435700DED3C57F12F96" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[815,980,1912,1936]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bituberculata" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A1435700DED3C57F12F96" box="[815,980,1912,1936]" italics="true" pageId="3" pageNumber="4">C. bituberculata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A143570F8ED3C500F2F96" bold="true" box="[986,1066,1912,1936]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A143570F8ED3C500F2F96" box="[986,1066,1912,1936]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; R)
|
||
<taxonomicName id="3EBDC4F9F26A143577B6ED3C51BD2F96" authority="(Pic, 1916)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[1172,1432,1911,1936]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="brasiliense">
|
||
<emphasis id="CBC96368F26A143577B6ED3C51052F96" box="[1172,1312,1912,1936]" italics="true" pageId="3" pageNumber="4">C. brasiliense</emphasis>
|
||
(Pic, 1916)
|
||
</taxonomicName>
|
||
, male; S)
|
||
<taxonomicName id="3EBDC4F9F26A143573D6EDD855A42FB2" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[244,385,1948,1972]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="brasiliense">
|
||
<emphasis id="CBC96368F26A143573D6EDD855A42FB2" box="[244,385,1948,1972]" italics="true" pageId="3" pageNumber="4">C. brasiliense</emphasis>
|
||
</taxonomicName>
|
||
, female; T)
|
||
<taxonomicName id="3EBDC4F9F26A143572DDEDD856AD2FB2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[511,648,1948,1972]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guyanense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143572DDEDD856AD2FB2" box="[511,648,1948,1972]" italics="true" pageId="3" pageNumber="4">C. guyanense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A143571ADEDD856C52FB2" bold="true" box="[655,736,1948,1972]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A143571ADEDD856C52FB2" box="[655,736,1948,1972]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; U)
|
||
<taxonomicName id="3EBDC4F9F26A1435706CEDD857E72FB2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[846,962,1948,1972]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kawense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A1435706CEDD857E72FB2" box="[846,962,1948,1972]" italics="true" pageId="3" pageNumber="4">C. kawense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A143570EAEDD8503C2FB2" bold="true" box="[968,1049,1948,1972]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A143570EAEDD8503C2FB2" box="[968,1049,1948,1972]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; V)
|
||
<taxonomicName id="3EBDC4F9F26A143577A5EDD850FD2FB2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1159,1240,1948,1972]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143577A5EDD850FD2FB2" box="[1159,1240,1948,1972]" italics="true" pageId="3" pageNumber="4">C. nana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A143577FDEDD851152FB2" bold="true" box="[1247,1328,1948,1972]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A143577FDEDD851152FB2" box="[1247,1328,1948,1972]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; W)
|
||
<taxonomicName id="3EBDC4F9F26A143573B5ED8455172FDE" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,306,1984,2008]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="saopaulense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143573B5ED8455172FDE" box="[151,306,1984,2008]" italics="true" pageId="3" pageNumber="4">C. saopaulense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A1435721AED8455AC2FDE" bold="true" box="[312,393,1984,2008]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A1435721AED8455AC2FDE" box="[312,393,1984,2008]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; X)
|
||
<taxonomicName id="3EBDC4F9F26A143572D4ED8456422FDE" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[502,615,1984,2008]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="wittmeri" status="sp. nov.">
|
||
<emphasis id="CBC96368F26A143572D4ED8456422FDE" box="[502,615,1984,2008]" italics="true" pageId="3" pageNumber="4">C. wittmeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26A1435714CED84569B2FDE" bold="true" box="[622,702,1984,2008]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="D0FADE13F26A1435714CED84569B2FDE" box="[622,702,1984,2008]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male; Y)
|
||
<emphasis id="CBC96368F26A14357008ED8457672FD1" box="[810,834,1984,2007]" italics="true" pageId="3" pageNumber="4">C.</emphasis>
|
||
sp., Bolivia, female. Scale bars = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="ADC2EFF2F26D143273B5EE65511E2C5B" httpUri="https://zenodo.org/record/2783686/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,250,1057,1082]" targetBox="[164,1423,199,1014]" targetPageId="4">
|
||
<paragraph id="F902BF7AF26D143273B5EE65511E2C5B" blockId="4.[151,1437,1057,1118]" pageId="4" pageNumber="5">
|
||
<emphasis id="CBC96368F26D143273B5EE6555362C3C" bold="true" box="[151,275,1057,1082]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
|
||
<taxonomicName id="3EBDC4F9F26D1432723BEE6655BE2C3C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[281,411,1058,1082]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CBC96368F26D1432723BEE6655BE2C3C" box="[281,411,1058,1082]" italics="true" pageId="4" pageNumber="5">Calvariopsis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26D14327282EE6655DB2C3C" bold="true" box="[416,510,1058,1082]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="D0FADE13F26D14327282EE6655DB2C3C" box="[416,510,1058,1082]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, females, dorsal view. A)
|
||
<emphasis id="CBC96368F26D14327027EE6657382C3F" box="[773,797,1058,1081]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., Colombia; B)
|
||
<emphasis id="CBC96368F26D143270FEEE6657D62C3F" box="[988,1011,1058,1081]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., Costa Rica; C)
|
||
<emphasis id="CBC96368F26D1432779FEE6650F12C3F" box="[1213,1236,1058,1081]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., Ecuador; D)
|
||
<emphasis id="CBC96368F26D143276A7EE6651B82C3F" box="[1413,1437,1058,1081]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., Ecuador; E)
|
||
<emphasis id="CBC96368F26D14327263EE02557D2C5B" box="[321,344,1094,1117]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., French Guiana; F)
|
||
<emphasis id="CBC96368F26D1432716AEE0256452C5B" box="[584,608,1094,1117]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., Panama; G)
|
||
<emphasis id="CBC96368F26D1432702CEE0257002C5B" box="[782,805,1094,1117]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., Paraguay; H)
|
||
<emphasis id="CBC96368F26D143270C3EE0257DC2C5B" box="[993,1017,1094,1117]" italics="true" pageId="4" pageNumber="5">C.</emphasis>
|
||
sp., Peru. Scale bars = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F902BF7AF26D143273E5ECED50A22F29" blockId="4.[151,1437,1164,1983]" pageId="4" pageNumber="5">Pronotum small, transverse, narrower than base of elytra, ca. 2× wider than long; lateral carinae almost straight, converging anteriorly; disc convex; anterior margin rounded; anterolateral angles slightly projecting; posterior angles obtuse, well marked; base bisinuate. Prosternum reduced anteriorly; prosternal process narrow, elongate, lanceolate with subtriangular apex, ca. 2.7× longer than wide, covered with delicate setae.</paragraph>
|
||
<paragraph id="F902BF7AF26D143273E5ED7C56002FB9" blockId="4.[151,1437,1164,1983]" pageId="4" pageNumber="5">Scutellar shield small, equilaterally triangular, flat, with pointed apex; anterior margin straight. Elytra oval to elongated, distinctly wider at base than base of pronotum; humeri well marked, apices regularly rounded. Epipleura wide at base, as wide as metanepisternum, regularly narrowing to apex. Punctation of elytra irregular, not granulate, punctures of the same sizes, sparse.</paragraph>
|
||
<caption id="ADC2EFF2F26C143373B5ED3557662FD4" httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,250,1905,1930]" targetBox="[165,1422,181,1857]" targetPageId="5">
|
||
<paragraph id="F902BF7AF26C143373B5ED3557662FD4" blockId="5.[151,1437,1905,2002]" pageId="5" pageNumber="6">
|
||
<emphasis id="CBC96368F26C143373B5ED3555312F8C" bold="true" box="[151,276,1905,1930]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
|
||
<taxonomicName id="3EBDC4F9F26C1433723EED3655BB2F8C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[284,414,1906,1930]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CBC96368F26C1433723EED3655BB2F8C" box="[284,414,1906,1930]" italics="true" pageId="5" pageNumber="6">Calvariopsis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26C14337287ED3656202F8C" bold="true" box="[421,517,1906,1930]" pageId="5" pageNumber="6">
|
||
<taxonomicNameLabel id="D0FADE13F26C14337287ED3656202F8C" box="[421,517,1906,1930]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, SEM images of morphological details, A–C)
|
||
<taxonomicName id="3EBDC4F9F26C143370C4ED3650DB2F8F" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[998,1278,1906,1930]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26C143370C4ED3650DB2F8F" box="[998,1278,1906,1930]" italics="true" pageId="5" pageNumber="6">Calvariopsis yanayacuense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26C14337624ED3651722F8C" bold="true" box="[1286,1367,1906,1930]" pageId="5" pageNumber="6">
|
||
<taxonomicNameLabel id="D0FADE13F26C14337624ED3651722F8C" box="[1286,1367,1906,1930]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; D–F)
|
||
<taxonomicName id="3EBDC4F9F26C143373B5EDD2558C2FA8" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,425,1942,1966]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bituberculata" status="sp. nov.">
|
||
<emphasis id="CBC96368F26C143373B5EDD2558C2FA8" box="[151,425,1942,1966]" italics="true" pageId="5" pageNumber="6">Calvariopsis bituberculata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26C14337293EDD256212FA8" bold="true" box="[433,516,1942,1966]" pageId="5" pageNumber="6">
|
||
<taxonomicNameLabel id="D0FADE13F26C14337293EDD256212FA8" box="[433,516,1942,1966]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="ADC2EFF2F26F143073B5EFD456452DCB" httpUri="https://zenodo.org/record/2783698/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1424,1449]" targetBox="[178,1410,181,1383]" targetPageId="6">
|
||
<paragraph id="F902BF7AF26F143073B5EFD456452DCB" blockId="6.[151,1437,1424,1485]" pageId="6" pageNumber="7">
|
||
<emphasis id="CBC96368F26F143073B5EFD455372DAF" bold="true" box="[151,274,1424,1449]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
|
||
<taxonomicName id="3EBDC4F9F26F14307235EFD455BC2DAE" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[279,409,1424,1448]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CBC96368F26F14307235EFD455BC2DAE" box="[279,409,1424,1448]" italics="true" pageId="6" pageNumber="7">Calvariopsis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26F143072BFEFD555DE2DAF" bold="true" box="[413,507,1425,1449]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="D0FADE13F26F143072BFEFD555DE2DAF" box="[413,507,1425,1449]" pageId="6" pageNumber="7" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, mouthparts.A–C)
|
||
<taxonomicName id="3EBDC4F9F26F143071E2EFD5574F2DAE" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[704,874,1425,1448]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26F143071E2EFD5574F2DAE" box="[704,874,1425,1448]" italics="true" pageId="6" pageNumber="7">C. yanayacuense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26F1430704DEFD5579B2DAF" bold="true" box="[879,958,1425,1449]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel id="D0FADE13F26F1430704DEFD5579B2DAF" box="[879,958,1425,1449]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; D–F)
|
||
<emphasis id="CBC96368F26F14307724EFD550382DAE" box="[1030,1053,1425,1448]" italics="true" pageId="6" pageNumber="7">C.</emphasis>
|
||
sp., group II, female (French Guiana). A, D) mandible; B, E) maxilla; C, F) labium.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F902BF7AF26F143073E5EFBF54F82E87" blockId="6.[151,1437,1530,2025]" pageId="6" pageNumber="7">
|
||
Hind wings (
|
||
<figureCitation id="6186A3FFF26F14307279EFBF55FC2E13" box="[347,473,1530,1557]" captionStart="FIGURE 5" captionStartId="7.[151,250,1512,1537]" captionTargetBox="[165,1403,204,1476]" captionTargetId="figure@7.[165,1424,181,1476]" captionTargetPageId="7" captionText="FIGURE 5. Calvariopsis gen. nov., wing. A) C. yanayacuense sp. nov.; B) C. bituberculata sp. nov." httpUri="https://zenodo.org/record/2783704/files/figure.png" pageId="6" pageNumber="7">Figs 5A, B</figureCitation>
|
||
) ca. 2× longer than width; radial cell subtriangular; apical portion of r3 well visible, straight; rp-mp2 arcuate, forming oblique angle with MP
|
||
<subScript id="6539BD3FF26F1430702EEC7457002E38" attach="right" box="[780,805,1584,1598]" fontSize="6" pageId="6" pageNumber="7">1+2</subScript>
|
||
; MP
|
||
<subScript id="6539BD3FF26F14307079EC7457462E38" attach="left" box="[859,867,1584,1598]" fontSize="6" pageId="6" pageNumber="7">4</subScript>
|
||
short, joining MP
|
||
<subScript id="6539BD3FF26F1430770EEC7450112E38" attach="left" box="[1068,1076,1584,1598]" fontSize="6" pageId="6" pageNumber="7">3</subScript>
|
||
and CuA+AA
|
||
<subScript id="6539BD3FF26F143077F7EC7450CA2E38" attach="left" box="[1237,1263,1584,1598]" fontSize="6" pageId="6" pageNumber="7">1+2</subScript>
|
||
; anal field with well marked AP
|
||
<subScript id="6539BD3FF26F1430726DEC1055722E64" attach="left" box="[335,343,1620,1634]" fontSize="6" pageId="6" pageNumber="7">3</subScript>
|
||
. Medial field with 3 free veins; pigmented areas in radial cell, medial, radial, central and apical fields.
|
||
</paragraph>
|
||
<paragraph id="F902BF7AF26F143073E5ECCF51702F33" blockId="6.[151,1437,1530,2025]" pageId="6" pageNumber="7">
|
||
Mesoventrite (
|
||
<figureCitation id="6186A3FFF26F1430724FECCF55E12EA3" box="[365,452,1675,1701]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="6" pageNumber="7">Fig. 3C</figureCitation>
|
||
) small; covered with sphaerical cuticular structures; anterior margin with small coxal rests; mesoventral process with well marked longitudinal suture; anterior portion forming oval, deep fossa for recep- tion of prosternal process; mesoventral process completely separating mesocoxae, very short, ca. 1.2–2.0× shorter than wide, sides subparallel, apex bilobed, touching metaventrite; mesanepisternum subrectangular, impunctate, forming coxal rest for procoxa; mesepimeron smaller, impunctate, subtriangular, strongly narrowed mesally.
|
||
</paragraph>
|
||
<paragraph id="F902BF7AF26F143073E5ED7B51732FEF" blockId="6.[151,1437,1530,2025]" pageId="6" pageNumber="7">
|
||
Metaventrite (
|
||
<figureCitation id="6186A3FFF26F14307245ED7B55C22F5F" box="[359,487,1855,1881]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="6" pageNumber="7">Figs 3A, D</figureCitation>
|
||
) moderately long, transverse, convex; basal portion with sphaerical cuticular struc- tures (
|
||
<figureCitation id="6186A3FFF26F143073C2ED2755412F7B" box="[224,356,1891,1917]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="6" pageNumber="7">Figs 3C, E</figureCitation>
|
||
); discrimen present in posterior 1/2–2/3; metanepisternum subtrapezoidal, slightly narrowing posteriorly. Metacoxae moderately large, strongly oblique, ca. 3.0× wider than its length, extending laterally to epipleura; metacoxal plate obliquely narrowing, absent in outermost portion. Metendosternite short, lateral furcal arms relatively wide; ventral processes well marked, subtriangular; anterior furcal arms very wide and short.
|
||
</paragraph>
|
||
<caption id="ADC2EFF2F26E143173B5EFAC50BD2E07" box="[151,1176,1512,1537]" httpUri="https://zenodo.org/record/2783704/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1512,1537]" targetBox="[165,1403,204,1476]" targetPageId="7">
|
||
<paragraph id="F902BF7AF26E143173B5EFAC50BD2E07" blockId="7.[151,1176,1512,1537]" box="[151,1176,1512,1537]" pageId="7" pageNumber="8">
|
||
<emphasis id="CBC96368F26E143173B5EFAC55362E07" bold="true" box="[151,275,1512,1537]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
|
||
<taxonomicName id="3EBDC4F9F26E1431723BEFAC55BE2E06" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[281,411,1512,1536]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CBC96368F26E1431723BEFAC55BE2E06" box="[281,411,1512,1536]" italics="true" pageId="7" pageNumber="8">Calvariopsis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26E14317283EFAD56252E07" bold="true" box="[417,512,1513,1537]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="D0FADE13F26E14317283EFAD56252E07" box="[417,512,1513,1537]" pageId="7" pageNumber="8" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, wing. A)
|
||
<taxonomicName id="3EBDC4F9F26E1431714EEFAD57322E06" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[620,791,1513,1536]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
|
||
<emphasis id="CBC96368F26E1431714EEFAD57322E06" box="[620,791,1513,1536]" italics="true" pageId="7" pageNumber="8">C. yanayacuense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F26E1431703CEFAD574B2E07" bold="true" box="[798,878,1513,1537]" pageId="7" pageNumber="8">
|
||
<taxonomicNameLabel id="D0FADE13F26E1431703CEFAD574B2E07" box="[798,878,1513,1537]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; B)
|
||
<taxonomicName id="3EBDC4F9F26E143170BEEFAD50642E06" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[924,1089,1512,1536]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bituberculata" status="sp. nov.">
|
||
<emphasis id="CBC96368F26E143170BEEFAD50642E06" box="[924,1089,1512,1536]" italics="true" pageId="7" pageNumber="8">C. bituberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0FADE13F26E14317765EFAD50BD2E07" box="[1095,1176,1513,1537]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F902BF7AF26E143173E5EC6B57D72E4F" blockId="7.[151,1437,1583,2041]" box="[199,1010,1583,1609]" pageId="7" pageNumber="8">Legs moderately long, tibiae with paired carinae, without modifications.</paragraph>
|
||
<paragraph id="F902BF7AF26E143173E5EC17563B2EDF" blockId="7.[151,1437,1583,2041]" pageId="7" pageNumber="8">
|
||
Abdomen (
|
||
<figureCitation id="6186A3FFF26E14317267EC1755AD2E68" box="[325,392,1619,1646]" captionStart="FIGURE 6" captionStartId="8.[151,250,1158,1183]" captionTargetBox="[153,1435,181,1105]" captionTargetId="figure@8.[153,1438,181,1105]" captionTargetPageId="8" captionText="FIGURE 6. Calvariopsis yanayacuense sp. nov., abdomen, tergites and ventrites." httpUri="https://zenodo.org/record/2783710/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
|
||
) ca. 1.2–1.3× wider than its length, convex; ventrites 1–4 of similar length; ventrite 5 slightly longer, regularly rounded or subtriangular at apex, occasionally deeply emarginated. Ventrites covered with setae with exception of ventrite 1, which lacks setae on lateral portions. Tergite VII with 2 (or more) transverse rows of dense, fringe-like setae (
|
||
<figureCitation id="6186A3FFF26E1431728BECFB56342EDF" box="[425,529,1727,1753]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="7" pageNumber="8">Fig. 13G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F902BF7AF26E143173E5ECA755D12FFF" blockId="7.[151,1437,1583,2041]" pageId="7" pageNumber="8">
|
||
Sternite VIII very small, widely U-shaped; sternite IX consisting of paired subtriangular hemisternites or oval plate with single apodeme; tergite VIII with moderately long apodemes as long as apical plate or shorter than apical plate, apical margin with various modifications of diagnostic value (processes, emarginations, denticles, indentations etc.); tergite IX narrower than tergite VIII, with membranous apical portion, apodemes as long as apical portion, curved or diverging. Tegmen reduced, smaller than penis, of two
|
||
<typeStatus id="260601D8F26E143170BDED3757FB2F8B" box="[927,990,1907,1933]" pageId="7" pageNumber="8">types</typeStatus>
|
||
: either elongated, with setose parameres fused at apex or shorter, U-shaped with free parameres. Penis symmetrical, dorsoventrally flattened, in some species curved in lateral view; trigonium elongated, pointed at apex, parameroids narrow, apices rounded, slightly longer or slightly shorter than trigonium.
|
||
</paragraph>
|
||
<caption id="ADC2EFF2F261143E73B5EEC257F22C99" box="[151,983,1158,1183]" httpUri="https://zenodo.org/record/2783710/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1158,1183]" targetBox="[153,1435,181,1105]" targetPageId="8">
|
||
<paragraph id="F902BF7AF261143E73B5EEC257F22C99" blockId="8.[151,983,1158,1183]" box="[151,983,1158,1183]" pageId="8" pageNumber="9">
|
||
<emphasis id="CBC96368F261143E73B5EEC255362C99" bold="true" box="[151,275,1158,1183]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
|
||
<taxonomicName id="3EBDC4F9F261143E723BEEC2560A2C98" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[281,559,1158,1182]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
|
||
<emphasis id="CBC96368F261143E723BEEC2560A2C98" box="[281,559,1158,1182]" italics="true" pageId="8" pageNumber="9">Calvariopsis yanayacuense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F261143E7114EEC256A32C98" bold="true" box="[566,646,1158,1182]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="D0FADE13F261143E7114EEC256A32C98" box="[566,646,1158,1182]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, abdomen, tergites and ventrites.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F902BF7AF261143E73E5EE89505B2DE5" blockId="8.[151,1437,1229,1687]" pageId="8" pageNumber="9">
|
||
Female. TL 1.9–3.7. Apical portion of elytra often with excitators (
|
||
<emphasis id="CBC96368F261143E70E7EE8950262CE0" box="[965,1027,1229,1254]" italics="true" pageId="8" pageNumber="9">sensu</emphasis>
|
||
<bibRefCitation id="9D2CC28BF261143E772EEE8950A42CEE" author="Ruta, R." box="[1036,1153,1229,1256]" pageId="8" pageNumber="9" pagination="49 - 68" refId="ref23212" refString="Ruta, R. (2008) Contribution to the knowledge of Seychellois Scirtidae (Coleoptera: Scirtoidea). Zootaxa, 1913, 49 - 68." type="journal article" year="2008">Ruta 2008</bibRefCitation>
|
||
)—presumably excretory structures connected with exocrine glands. Hind wings functional, fully developed. Sternite VIII (
|
||
<figureCitation id="6186A3FFF261143E77EDEEB5510D2D0D" box="[1231,1320,1265,1291]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="8" pageNumber="9">
|
||
Fig.
|
||
<date id="8D0399BAF261143E7620EEB5510D2D0D" box="[1282,1320,1265,1291]" pageId="8" pageNumber="9" value="2010-01">10I</date>
|
||
</figureCitation>
|
||
) elongate, Y-shaped, with single apodeme, apex with sparse setae. Tergite VIII very long; apical portion subtrapezoidal, with sparse setation on apical margin; apodemes long, several times longer than apical portion. Ovipositor long; branchlets absent; gonocoxites either strongly sclerotized, widely triangular, not covered with setae, gonostyli arising subapically, with two tufts of setae at apex (
|
||
<figureCitation id="6186A3FFF261143E715CEFC556FA2D9D" box="[638,735,1409,1435]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="8" pageNumber="9">Fig. 10F</figureCitation>
|
||
) or gonocoxites narrower, membranous, with gonostyli arising from apex (
|
||
<figureCitation id="6186A3FFF261143E7235EFE1555A2DB9" box="[279,383,1445,1471]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="8" pageNumber="9">Fig. 10K</figureCitation>
|
||
). Apex of gonostylus with tuft of setae. Bursa membranous, with single sclerite present. Bursal sclerite oval to rod-like, with small subconical or digitiform outgrowth directed posterad.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1A7ECF1F261143E73E5EFA856E42E49" pageId="8" pageNumber="9" type="biology_ecology">
|
||
<paragraph id="F902BF7AF261143E73E5EFA856E42E49" blockId="8.[151,1437,1229,1687]" pageId="8" pageNumber="9">
|
||
<emphasis id="CBC96368F261143E73E5EFA855022E01" bold="true" box="[199,295,1516,1543]" pageId="8" pageNumber="9">Biology.</emphasis>
|
||
In most cases only general data on collecting circumstances is available. Individuals of
|
||
<taxonomicName id="3EBDC4F9F261143E762FEFA955112E2C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
|
||
<emphasis id="CBC96368F261143E762FEFA955112E2C" italics="true" pageId="8" pageNumber="9">Calvariopsis yanayacuense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBC96368F261143E7219EC5555B12E2D" bold="true" box="[315,404,1553,1579]" pageId="8" pageNumber="9">
|
||
<taxonomicNameLabel id="D0FADE13F261143E7219EC5555B12E2D" box="[315,404,1553,1579]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were observed in great numbers on trees densely covered by lichens on the edges of Andean cloud forest in
|
||
<collectingRegion id="3B797198F261143E7262EC7155CC2E49" box="[320,489,1589,1615]" country="Ecuador" name="Napo" pageId="8" pageNumber="9">Napo province</collectingRegion>
|
||
,
|
||
<collectingCountry id="81AAFFEAF261143E72D6EC7156742E49" box="[500,593,1589,1615]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="6186A3FFF261143E7142EC7156912E49" box="[608,692,1589,1615]" captionStart="FIGURE 19" captionStartId="25.[151,250,842,867]" captionTargetBox="[153,1435,254,799]" captionTargetId="figure@25.[151,1436,254,799]" captionTargetPageId="25" captionText="FIGURE 19. Calvariopsis yanayacuense sp. nov., habitat. A) Ecuador, prov. Napo, road from Yanayacu Biological Station to Rio Aliso, sideroad; B) tree trunk covered with lichens and mosses, where C. yanayacuense was collected in great numbers." httpUri="https://zenodo.org/record/2783766/files/figure.png" pageId="8" pageNumber="9">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1A7ECF1F261143E73E5EC1D50332E91" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="F902BF7AF261143E73E5EC1D50332E91" blockId="8.[151,1437,1229,1687]" pageId="8" pageNumber="9">
|
||
<emphasis id="CBC96368F261143E73E5EC1D556B2E75" bold="true" box="[199,334,1625,1651]" pageId="8" pageNumber="9">Etymology.</emphasis>
|
||
Combination of the genus name
|
||
<taxonomicName id="3EBDC4F9F261143E71E3EC1D571C2E75" authorityName="Pic. Gender" authorityYear="1918" box="[705,825,1625,1651]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBC96368F261143E71E3EC1D571C2E75" box="[705,825,1625,1651]" italics="true" pageId="8" pageNumber="9">Calvarium</emphasis>
|
||
</taxonomicName>
|
||
and the Greek word ópsis (όψις) meaning appearance, view. Reference to the similarity to the genus
|
||
<taxonomicName id="3EBDC4F9F261143E71BBEC3957852E91" authority="Pic. Gender" authorityName="Pic. Gender" authorityYear="1918" box="[665,928,1661,1687]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBC96368F261143E71BBEC3957342E91" box="[665,785,1661,1687]" italics="true" pageId="8" pageNumber="9">Calvarium</emphasis>
|
||
Pic. Gender
|
||
</taxonomicName>
|
||
: feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |