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<document id="94D3C80B9D40EC0154CBEA7A0EFD5DDB" ID-CLB-Dataset="26847" ID-DOI="10.11646/zootaxa.4604.1.1" ID-GBIF-Dataset="b5902a9d-c38b-462f-ad8a-95693c1822dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="2783674" ID-ZooBank="E6D82300-8DC6-4560-A945-8F29D3B3D501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1557723545002" checkinUser="plazi" docAuthor="Ruta, Rafał" docDate="2019" docId="71140E6CF265143973B5EFFE573A2918" docLanguage="en" docName="zootaxa.4604.1.1.pdf" docOrigin="Zootaxa 4604 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvariopsis fourgassiense Ruta 2019, sp. nov." docType="treatment" docVersion="12" lastPageNumber="16" masterDocId="8D2D7614F26914367322EA4454252806" masterDocTitle="Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="13" updateTime="1698728494285" updateUser="ExternalLinkService">
<mods:mods id="7EF0350EB40C0F9465FB3889D9F27E95" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4D0F7D95CACAFADE28397ACC4B40AFE4">
<mods:title id="3C0705F6ED669A3FF6FDBCB975F1750D">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
</mods:titleInfo>
<mods:name id="66BC2F9038A7E78647B2F96F015F4E65" type="personal">
<mods:role id="CC720957D5B5B9A43866471329F075F8">
<mods:roleTerm id="5C1D1C2B6EE056A1EEA7DBD09FAE0FDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E30242297B3B30B9E1BB6BBABDED9456">Ruta, Rafał</mods:namePart>
</mods:name>
<mods:typeOfResource id="60647A5E33B1A3D5C25E450A203429D2">text</mods:typeOfResource>
<mods:relatedItem id="0B20B467E1993CCFF1A1FE880A275DC0" type="host">
<mods:titleInfo id="4962A1FC7C04103B5FB1821971362AB4">
<mods:title id="F5DBB6753634EAA13A8EBBB0B831831A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="912F9EACB704B0550BFA3B33561CA7B7">
<mods:date id="696BA5ED2E32176FC533A606B2DD699B">2019</mods:date>
<mods:detail id="54CDAEC261C40283BE35EB457D9C1744" type="pubDate">
<mods:number id="582AA0CA1B0C2718DD98CBF2A1F53D0F">2019-05-13</mods:number>
</mods:detail>
<mods:detail id="74B56EFD0FF75708F5556E92CA479147" type="volume">
<mods:number id="E3FE23F682B0B83CDFED64000ED15C5F">4604</mods:number>
</mods:detail>
<mods:detail id="3C5FB42F56C0728410D2B9760A606E35" type="issue">
<mods:number id="5339B0464838DF5D1A0C72130EB7395F">1</mods:number>
</mods:detail>
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<mods:start id="3BEBD6CAB239B20556B46644901804DD">1</mods:start>
<mods:end id="0D8F5C13612147705FFB198FBAD1CF07">41</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6AAC4D92DB9E2AF7E775665DA799BFF8">journal article</mods:classification>
<mods:identifier id="ADFE7DE9387BADCCD4D942DD737A40A8" type="CLB-Dataset">26847</mods:identifier>
<mods:identifier id="49D5EDCD5C20B05A424F32FE79F58BB4" type="DOI">10.11646/zootaxa.4604.1.1</mods:identifier>
<mods:identifier id="F3B8A77D24142395C0FA72787B305622" type="GBIF-Dataset">b5902a9d-c38b-462f-ad8a-95693c1822dc</mods:identifier>
<mods:identifier id="CE2EBC4A354CC4ED8807D95D9FB54FDC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A35112A8B9036EF3E19C6DAA6705070F" type="Zenodo-Dep">2783674</mods:identifier>
<mods:identifier id="2E175D65511EAD42882EA2B71FB3121F" type="ZooBank">E6D82300-8DC6-4560-A945-8F29D3B3D501</mods:identifier>
</mods:mods>
<treatment id="71140E6CF265143973B5EFFE573A2918" ID-GBIF-Taxon="157135246" LSID="urn:lsid:plazi:treatment:71140E6CF265143973B5EFFE573A2918" httpUri="http://treatment.plazi.org/id/71140E6CF265143973B5EFFE573A2918" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
<subSubSection id="B1A7ECF1F265143A73B5EFFE56652DD2" box="[151,576,1466,1493]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="F902BF7AF265143A73B5EFFE56652DD2" blockId="12.[151,576,1466,1528]" box="[151,576,1466,1493]" pageId="12" pageNumber="13">
<heading id="A24A0816F265143A73B5EFFE56652DD2" bold="true" box="[151,576,1466,1493]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="CBC96368F265143A73B5EFFE56652DD2" bold="true" box="[151,576,1466,1493]" pageId="12" pageNumber="13">
<taxonomicName id="3EBDC4F9F265143A73B5EFFE55FF2DD2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,474,1466,1493]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F265143A73B5EFFE55FF2DD2" bold="true" box="[151,474,1466,1493]" italics="true" pageId="12" pageNumber="13">Calvariopsis fourgassiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F265143A72C0EFFE56652DD2" box="[482,576,1466,1492]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143A73B5EF9A550E2DFE" box="[151,299,1502,1528]" pageId="12" pageNumber="13" type="description">
<paragraph id="F902BF7AF265143A73B5EF9A550E2DFE" blockId="12.[151,576,1466,1528]" box="[151,299,1502,1528]" pageId="12" pageNumber="13">
(
<figureCitation id="6186A3FFF265143A73BDEF9A54DF2DFE" box="[159,250,1502,1528]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="12" pageNumber="13">Figs 1C</figureCitation>
,
<figureCitation id="6186A3FFF265143A7224EF9A55042DFE" box="[262,289,1502,1528]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="12" pageNumber="13">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143A73B5EC6257742E62" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="F902BF7AF265143A73B5EC6257742E62" blockId="12.[151,1437,1573,2032]" pageId="12" pageNumber="13">
<emphasis id="CBC96368F265143A73B5EC6255602E46" bold="true" box="[151,325,1573,1600]" pageId="12" pageNumber="13">Type material.</emphasis>
<materialsCitation id="49D5B527F265143A7269EC6255062E62" ID-GBIF-Occurrence="2239706639" collectingDate="1995-01-10" collectionCode="MNHG" collectorName="M. Kocian" country="French Guiana" county="Fourgassie" location="Francaise" municipality="Montagne Kaw" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F265143A7269EC6255932E46" box="[331,438,1574,1600]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF265143A7124EC6256462E46" box="[518,611,1574,1600]" pageId="12" pageNumber="13">MNHG</collectionCode>
): “
<collectingCountry id="81AAFFEAF265143A715DEC6256FC2E46" box="[639,729,1574,1600]" name="French Guiana" pageId="12" pageNumber="13">Guyane</collectingCountry>
<location id="FC62E9A1F265143A71C2EC62576E2E46" LSID="urn:lsid:plazi:treatment:71140E6CF265143973B5EFFE573A2918:FC62E9A1F265143A71C2EC62576E2E46" box="[736,843,1574,1600]" country="French Guiana" county="Fourgassie" municipality="Montagne Kaw" name="Francaise" pageId="12" pageNumber="13">Francaise</location>
\
<collectingMunicipality id="19662500F265143A7042EC62502B2E46" box="[864,1038,1574,1600]" pageId="12" pageNumber="13">Montagne Kaw</collectingMunicipality>
\
<collectingCounty id="1063C7F6F265143A7701EC6250B92E46" box="[1059,1180,1574,1600]" pageId="12" pageNumber="13">Fourgassié</collectingCounty>
\ 15.9.–
<date id="8D0399BAF265143A77D6EC6151422E46" box="[1268,1383,1573,1600]" pageId="12" pageNumber="13" value="1995-01-10">
<collectingDate id="9D476052F265143A77D6EC6151422E46" box="[1268,1383,1573,1600]" pageId="12" pageNumber="13" value="1995-01-10">1.10.1995</collectingDate>
</date>
\
<collectorName id="5448DAACF265143A765EEC6254CD2E62" pageId="12" pageNumber="13">M. Kocian</collectorName>
lgt.”
</materialsCitation>
.
<materialsCitation id="49D5B527F265143A720CEC0E57682E62" ID-GBIF-Occurrence="2239706629" box="[302,845,1610,1636]" collectingDate="1995-01-10" collectionCode="MNHG" collectorName="M. Kocian" country="French Guiana" county="Fourgassie" location="Francaise" municipality="Montagne Kaw" pageId="12" pageNumber="13" specimenCount="1" typeStatus="paratype">
<typeStatus id="260601D8F265143A720CEC0E55B62E62" box="[302,403,1610,1636]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
, male (
<collectionCode id="9FAC27BFF265143A72C1EC0E56652E62" box="[483,576,1610,1636]" pageId="12" pageNumber="13">MNHG</collectionCode>
): same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143A73E5EC2955A22F1E" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="F902BF7AF265143A73E5EC2955A22F1E" blockId="12.[151,1437,1573,2032]" pageId="12" pageNumber="13">
<emphasis id="CBC96368F265143A73E5EC2955672E8E" bold="true" box="[199,322,1645,1672]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
Small (TL
<quantity id="3E45129FF265143A72E0EC2A56392E8E" box="[450,540,1646,1672]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="12" pageNumber="13" unit="mm" value="2.3">2.3 mm</quantity>
), dark brown, with rounded sides of elytra, externally resembling some group II species of
<taxonomicName id="3EBDC4F9F265143A722EECD655BE2EAA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[268,411,1682,1708]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F265143A722EECD655BE2EAA" box="[268,411,1682,1708]" italics="true" pageId="12" pageNumber="13">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F265143A7283ECD656222EAA" box="[417,519,1682,1708]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
Penis very long and narrow, strongly narrowed in its middle, apical plate of tergite VIII with M-shaped, shallowly notched median process and distinct denticles in lateral portions. Genitalia similar to
<taxonomicName id="3EBDC4F9F265143A7396EC9E55662EF2" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[180,323,1754,1780]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F265143A7396EC9E55662EF2" box="[180,323,1754,1780]" italics="true" pageId="12" pageNumber="13">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F265143A7268EC9E55862EF2" bold="true" box="[330,419,1754,1780]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="D0FADE13F265143A7268EC9E55862EF2" box="[330,419,1754,1780]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but median process of tergite VIII is distinctly shorter and more weakly developed than in
<taxonomicName id="3EBDC4F9F265143A73B5ECBA55022F1E" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,295,1790,1816]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F265143A73B5ECBA55022F1E" box="[151,295,1790,1816]" italics="true" pageId="12" pageNumber="13">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F265143A720CECBA55A22F1E" bold="true" box="[302,391,1790,1816]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="D0FADE13F265143A720CECBA55A22F1E" box="[302,391,1790,1816]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F265143973E5ED6556EA28D1" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="F902BF7AF265143873E5ED6555312E54" blockId="12.[151,1437,1573,2032]" lastBlockId="14.[151,1437,1484,2014]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<emphasis id="CBC96368F265143A73E5ED65557C2F3A" bold="true" box="[199,345,1825,1852]" pageId="12" pageNumber="13">Description.</emphasis>
Male. Body oval, convex, covered with brown, suberect setae. Head brown with yellowish frons and clypeus, pronotum and elytra dark brown, venter brown, legs yellowish-brown, antennae yellowish. Head very broad, 1.8× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by 3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 2.0–3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</paragraph>
<caption id="ADC2EFF2F264143B73B5ED7D55832FE4" httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1849,1874]" targetBox="[178,1410,181,1810]" targetPageId="13">
<paragraph id="F902BF7AF264143B73B5ED7D55832FE4" blockId="13.[151,1437,1849,2018]" pageId="13" pageNumber="14">
<emphasis id="CBC96368F264143B73B5ED7D55062F54" bold="true" box="[151,291,1849,1874]" pageId="13" pageNumber="14">FIGURE 10.</emphasis>
<taxonomicName id="3EBDC4F9F264143B720FED7E558A2F54" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[301,431,1850,1874]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F264143B720FED7E558A2F54" box="[301,431,1850,1874]" italics="true" pageId="13" pageNumber="14">Calvariopsis</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B729AED7E563F2F54" bold="true" box="[440,538,1850,1874]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B729AED7E563F2F54" box="[440,538,1850,1874]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C)
<taxonomicName id="3EBDC4F9F264143B7614ED1B54992F9C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F264143B7614ED1B54992F9C" italics="true" pageId="13" pageNumber="14">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B73E0EDC655362F9C" bold="true" box="[194,275,1922,1946]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B73E0EDC655362F9C" box="[194,275,1922,1946]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; D)
<taxonomicName id="3EBDC4F9F264143B7263EDC755812F9C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[321,420,1923,1946]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
<emphasis id="CBC96368F264143B7263EDC755812F9C" box="[321,420,1923,1946]" italics="true" pageId="13" pageNumber="14">C. pittieri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B7288EDC655DE2F9C" bold="true" box="[426,507,1922,1946]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B7288EDC655DE2F9C" box="[426,507,1922,1946]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E–H)
<taxonomicName id="3EBDC4F9F264143B7164EDC756C12F9C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[582,740,1923,1946]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="yanyacuense" status="sp. nov.">
<emphasis id="CBC96368F264143B7164EDC756C12F9C" box="[582,740,1923,1946]" italics="true" pageId="13" pageNumber="14">C. yanyacuense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F264143B71C9EDC6571E2F9C" bold="true" box="[747,827,1922,1946]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="D0FADE13F264143B71C9EDC6571E2F9C" box="[747,827,1922,1946]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; I–K)
<taxonomicName id="3EBDC4F9F264143B705DEDC750602F9C" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[895,1093,1921,1946]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F264143B705DEDC750292F9C" box="[895,1036,1922,1946]" italics="true" pageId="13" pageNumber="14">C. brasiliense</emphasis>
(Pic)
</taxonomicName>
, specimen from Sao Paulo; L–M)
<taxonomicName id="3EBDC4F9F264143B73B5EDE355782FB8" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[151,349,1957,1982]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F264143B73B5EDE355012FB8" box="[151,292,1958,1982]" italics="true" pageId="13" pageNumber="14">C. brasiliense</emphasis>
(Pic)
</taxonomicName>
, specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor.
</paragraph>
</caption>
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<emphasis id="CBC96368F267143873B5EF25553A2D7C" bold="true" box="[151,287,1377,1402]" pageId="14" pageNumber="15">FIGURE 11.</emphasis>
<taxonomicName id="3EBDC4F9F26714387207EF2556122D7C" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[293,567,1377,1402]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F26714387207EF2556122D7C" box="[293,567,1377,1402]" italics="true" pageId="14" pageNumber="15">Calvariopsis fourgassiense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F2671438711CEF2656AA2D7C" bold="true" box="[574,655,1378,1402]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="D0FADE13F2671438711CEF2656AA2D7C" box="[574,655,1378,1402]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="F902BF7AF267143873E5EC1857CC2F74" blockId="14.[151,1437,1484,2014]" pageId="14" pageNumber="15">
Male genitalia. Penis (L
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,
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W
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mm,
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A–B) very long and narrow, widest in basal half, wid- ened also in apical portion, parameroids short, narrow, curved, trigonium subtriangular, narrow, pointed at apex. Tegmen (L
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,
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W
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mm,
<figureCitation id="6186A3FFF2671438713FECE0565A2EB8" box="[541,639,1700,1726]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15">Fig. 11C</figureCitation>
) very long, sides diverging, apex with short lateral lobes, covered with setae. Sternite IX (L
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,
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W
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mm,
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) subrectangular, with paired apodemes, apical margin rounded, apical portion covered with short, sparse setae. Tergite VIII (L
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,
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W
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mm,
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) with short apical plate, central portion of apical plate produced, with small notch at apex, lateral portions of apical margin with a few denticles in mesal part, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
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,
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mm,
<figureCitation id="6186A3FFF267143873B5ED1C54D32F74" box="[151,246,1880,1906]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="14" pageNumber="15">Fig. 11F</figureCitation>
) with membranous apical plate, apodemes longer than apical plate.
</paragraph>
<paragraph id="F902BF7AF267143873E5ED3855BB2F90" blockId="14.[151,1437,1484,2014]" box="[199,414,1916,1942]" pageId="14" pageNumber="15">Female. Unknown.</paragraph>
<paragraph id="F902BF7AF267143873E5EDE450FC2FBC" blockId="14.[151,1437,1484,2014]" box="[199,1241,1952,1978]" pageId="14" pageNumber="15">
Variability.
<typeStatus id="260601D8F26714387268EDE455892FBC" box="[330,428,1952,1978]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
has a small, orange spot along adsutural portion in central part of elytra.
</paragraph>
<paragraph id="F902BF7AF267143973E5ED8056EA28D1" blockId="14.[151,1437,1484,2014]" lastBlockId="15.[151,1436,152,287]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="CBC96368F267143873E5ED8055DA2FD8" bold="true" box="[199,511,1987,2014]" pageId="14" pageNumber="15">Measurements and ratios.</emphasis>
Males (n = 2) TL 2.27–2.30 (2.28) mm, PL 0.47–0.50 (0.48) mm, PW 1.03–1.07 (1.05) mm, EL 1.83–1.93 (1.88) mm, EW 1.47–1.50 (1.48) mm, TL/EW 1.53–1.55 (1.54), PW/PL 2.07–2.29 (2.18), EL/EW 1.22–1.32 (1.27), EL/PL 3.67–4.14 (3.90).
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F266143973E5EAA4579028FC" box="[199,949,224,251]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="F902BF7AF266143973E5EAA4579028FC" blockId="15.[151,1436,152,287]" box="[199,949,224,251]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973E5EAA4554428FD" bold="true" box="[199,353,224,251]" pageId="15" pageNumber="16">Distribution.</emphasis>
Known only from its
<typeStatus id="260601D8F2661439717BEAA556AC28FD" box="[601,649,225,251]" pageId="15" pageNumber="16">type</typeStatus>
locality in
<collectingCountry id="81AAFFEAF2661439702AEAA4579428FC" box="[776,945,224,250]" name="French Guiana" pageId="15" pageNumber="16">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F266143973E5EB41573A2918" box="[199,799,260,287]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="F902BF7AF266143973E5EB41573A2918" blockId="15.[151,1436,152,287]" box="[199,799,260,287]" pageId="15" pageNumber="16">
<emphasis id="CBC96368F266143973E5EB41556B2919" bold="true" box="[199,334,261,287]" pageId="15" pageNumber="16">Etymology.</emphasis>
Named after
<typeStatus id="260601D8F266143972C5EB4156322919" box="[487,535,261,287]" pageId="15" pageNumber="16">type</typeStatus>
locality of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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