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treatments-xml/data/71/14/0E/71140E6CF262143D73B5EAA551AF2F75.xml
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<document id="91E88849003780C0E485E9768E16D1BC" ID-CLB-Dataset="26847" ID-DOI="10.11646/zootaxa.4604.1.1" ID-GBIF-Dataset="b5902a9d-c38b-462f-ad8a-95693c1822dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="2783674" ID-ZooBank="E6D82300-8DC6-4560-A945-8F29D3B3D501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1557723545002" checkinUser="plazi" docAuthor="Ruta, Rafał" docDate="2019" docId="71140E6CF262143D73B5EAA551AF2F75" docLanguage="en" docName="zootaxa.4604.1.1.pdf" docOrigin="Zootaxa 4604 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvariopsis borowieci Ruta 2019, sp. nov." docType="treatment" docVersion="12" lastPageNumber="12" masterDocId="8D2D7614F26914367322EA4454252806" masterDocTitle="Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="12" updateTime="1698728494285" updateUser="ExternalLinkService">
<mods:mods id="865E4A9499345F8A94FED25F06C97251" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A84665BE27D1FBC4AE9F72B68577FE59">
<mods:title id="438A3293C2882A23BB9B8E19D0F34492">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
</mods:titleInfo>
<mods:name id="BFD2A10EC618C42634126668A27C033A" type="personal">
<mods:role id="049B96C8B675F21583C0D596CF43A852">
<mods:roleTerm id="1F753063BB594D013C5DD672C366B394">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD4E158E8C70B6322A43C43C27FBBCA1">Ruta, Rafał</mods:namePart>
</mods:name>
<mods:typeOfResource id="7AF8A18666603DAB68120F04670E2045">text</mods:typeOfResource>
<mods:relatedItem id="AC523C3ACF1535D09D652104819A1FDE" type="host">
<mods:titleInfo id="FCC6E270852A9E454BEAB79654E8CE32">
<mods:title id="B6EE858AF34264826D540065FDE67D58">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="94AF9DD1E5AF94DAAC0561FE824F8169">
<mods:date id="3413046D0DC77ACE2F69F4E148C60430">2019</mods:date>
<mods:detail id="B09AEC6213B5DBFA4C5E58B4322F5605" type="pubDate">
<mods:number id="2BF09C3894AAECF2691BBF3A57F5A084">2019-05-13</mods:number>
</mods:detail>
<mods:detail id="C4265C3370C25BDB0B7ABD338C011FDD" type="volume">
<mods:number id="50E7E73838FDD0A2579FAF45B88A0FDC">4604</mods:number>
</mods:detail>
<mods:detail id="44448ECA631D8F013C434785271749F1" type="issue">
<mods:number id="37092CAE264D7AD629422664874829D7">1</mods:number>
</mods:detail>
<mods:extent id="FBC8EF3388E49B66741A37EC4742AE9B" unit="page">
<mods:start id="8EF9072834A15E927D7AECA970D6F624">1</mods:start>
<mods:end id="DDE640ED1C36492542204BB2C7C62186">41</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="29479C46DA0975B4409C499256B568B6">journal article</mods:classification>
<mods:identifier id="BBAD08BFF56E3FB284AEC61EF1A6A46B" type="CLB-Dataset">26847</mods:identifier>
<mods:identifier id="C74286DEE8CEBCCB385C1171761CDD45" type="DOI">10.11646/zootaxa.4604.1.1</mods:identifier>
<mods:identifier id="8168106CD87B328A5204DE8EBA437F8C" type="GBIF-Dataset">b5902a9d-c38b-462f-ad8a-95693c1822dc</mods:identifier>
<mods:identifier id="FE38D48567288AECD5EC18772DA1AB0D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E102864FFA3FE66DA92B24AE274EB6FA" type="Zenodo-Dep">2783674</mods:identifier>
<mods:identifier id="B232B7790D8FCE9122DEBC63BEAE69CF" type="ZooBank">E6D82300-8DC6-4560-A945-8F29D3B3D501</mods:identifier>
</mods:mods>
<treatment id="71140E6CF262143D73B5EAA551AF2F75" ID-GBIF-Taxon="157135245" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75" httpUri="http://treatment.plazi.org/id/71140E6CF262143D73B5EAA551AF2F75" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="B1A7ECF1F262143D73B5EAA5562A28FA" box="[151,527,225,252]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="F902BF7AF262143D73B5EAA5562A28FA" blockId="11.[151,527,225,287]" box="[151,527,225,252]" pageId="11" pageNumber="12">
<heading id="A24A0816F262143D73B5EAA5562A28FA" bold="true" box="[151,527,225,252]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="CBC96368F262143D73B5EAA5562A28FA" bold="true" box="[151,527,225,252]" pageId="11" pageNumber="12">
<taxonomicName id="3EBDC4F9F262143D73B5EAA5558C28FA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[151,425,225,252]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F262143D73B5EAA5558C28FA" bold="true" box="[151,425,225,252]" italics="true" pageId="11" pageNumber="12">Calvariopsis borowieci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F262143D7293EAA6562A28FA" box="[433,527,226,252]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73B5EB4155052919" box="[151,288,261,287]" pageId="11" pageNumber="12" type="description">
<paragraph id="F902BF7AF262143D73B5EB4155052919" blockId="11.[151,527,225,287]" box="[151,288,261,287]" pageId="11" pageNumber="12">
(
<figureCitation id="6186A3FFF262143D73BDEB4154D92919" box="[159,252,261,287]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="11" pageNumber="12">Figs 1A</figureCitation>
,
<figureCitation id="6186A3FFF262143D722AEB4155322919" box="[264,279,261,287]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73B5EB0A55BF2AC9" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="F902BF7AF262143D73B5EB0A55BF2AC9" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73B5EB0A5560296E" bold="true" box="[151,325,333,360]" pageId="11" pageNumber="12">Type material.</emphasis>
<materialsCitation id="49D5B527F262143D7269EB0954ED298D" ID-GBIF-Occurrence="2239706624" collectingDate="2009-11-30" collectionCode="DBET" collectorName="Lech" country="Ecuador" location="Yanayacu-Rio" municipality="Aliso" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<typeStatus id="260601D8F262143D7269EB0955932961" box="[331,438,333,359]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="9FAC27BFF262143D7124EB0956732961" box="[518,598,333,359]" pageId="11" pageNumber="12">DBET</collectionCode>
): “
<collectingCountry id="81AAFFEAF262143D7153EB0956DE2961" box="[625,763,333,359]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
\ road
<location id="FC62E9A1F262143D706AEB0957C22961" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D706AEB0957C22961" box="[840,999,333,359]" country="Ecuador" municipality="Aliso" name="Yanayacu-Rio" pageId="11" pageNumber="12">Yanayacu-Rio</location>
\
<collectingMunicipality id="19662500F262143D70D8EB09501C2961" box="[1018,1081,333,359]" pageId="11" pageNumber="12">Aliso</collectingMunicipality>
,
<date id="8D0399BAF262143D7767EB0950E62961" box="[1093,1219,333,360]" pageId="11" pageNumber="12" value="2009-11-30">
<collectingDate id="9D476052F262143D7767EB0950E62961" box="[1093,1219,333,360]" pageId="11" pageNumber="12" value="2009-11-30">30 XI 2009</collectingDate>
</date>
\ leg.
<collectorName id="5448DAACF262143D762AEB0951652961" box="[1288,1344,333,359]" pageId="11" pageNumber="12">Lech</collectorName>
Borow- iec”
</materialsCitation>
.
<materialsCitation id="49D5B527F262143D73F3EB36571129A9" ID-GBIF-Occurrence="2239706610" collectingDate="2009-12-17" collectionCode="DBET" collectorName="R. Ruta" country="Ecuador" elevation="2100" location="Male" pageId="11" pageNumber="12" specimenCount="8" specimenCount-female="8" stateProvince="Napo" typeStatus="paratype">
<typeStatus id="260601D8F262143D73F3EB365567298A" box="[209,322,370,396]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
.
<location id="FC62E9A1F262143D7268EB3555A1298D" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D7268EB3555A1298D" box="[330,388,369,395]" country="Ecuador" name="Male" pageId="11" pageNumber="12" stateProvince="Napo">Male</location>
(
<collectionCode id="9FAC27BFF262143D72B3EB3555C4298D" box="[401,481,369,395]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 77,
<collectingRegion id="3B797198F262143D71ABEB3656E3298A" box="[649,710,370,396]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Yanayacu \ Station,
<quantity id="3E45129FF262143D77B1EB35510C298D" box="[1171,1321,369,395]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F262143D77B1EB35510C298D" box="[1171,1321,369,395]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, \
<date id="8D0399BAF262143D720AEBD1559129A9" box="[296,436,405,432]" pageId="11" pageNumber="12" value="2009-12-17">
<collectingDate id="9D476052F262143D720AEBD1559129A9" box="[296,436,405,432]" pageId="11" pageNumber="12" value="2009-12-17">17 XII 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF262143D72D2EBD2566929A9" box="[496,588,405,432]" pageId="11" pageNumber="12">R. Ruta</collectorName>
”.
<specimenCount id="EFBB74F3F262143D7142EBD156E929A9" box="[608,716,405,431]" pageId="11" pageNumber="12" type="female">8 females</specimenCount>
(
<collectionCode id="9FAC27BFF262143D71FEEBD1570929A9" box="[732,812,405,431]" pageId="11" pageNumber="12">DBET</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="49D5B527F262143D701CEBD256EF29F1" ID-GBIF-Occurrence="2239706640" collectingDate="9 XII" collectionCode="DBET" collectorName="Rafal Ruta" country="Ecuador" elevation="2125" latitude="-0.6051111" location="Female" longLatPrecision="1" longitude="-77.88583" municipality="Aliso" pageId="11" pageNumber="12" specimenCount="2" specimenCount-female="2" stateProvince="Napo" typeStatus="paratype">
same data as holotype.
<location id="FC62E9A1F262143D7766EBD150B329A9" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D7766EBD150B329A9" box="[1092,1174,405,431]" latitude="-0.6051111" longLatPrecision="1" longitude="-77.88583" name="Female" pageId="11" pageNumber="12" stateProvince="Napo">Female</location>
(
<collectionCode id="9FAC27BFF262143D7784EBD150D329A9" box="[1190,1270,405,431]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 64,
<collectingRegion id="3B797198F262143D73B5EBFE54F129D2" box="[151,212,442,468]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Yanayacu \ Station, Bamboo hill trail, \
<quantity id="3E45129FF262143D708BEBFD502629D2" box="[937,1027,441,468]" metricMagnitude="3" metricUnit="m" metricValue="2.125" pageId="11" pageNumber="12" unit="m" value="2125.0">
<elevation id="72905849F262143D708BEBFD502629D2" box="[937,1027,441,468]" metricMagnitude="3" metricUnit="m" metricValue="2.125" pageId="11" pageNumber="12" unit="m" value="2125.0">2125 m</elevation>
</quantity>
,
<geoCoordinate id="9C89D9BDF262143D7733EBFD509129D5" box="[1041,1204,441,467]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="-0.6051111">S 00º36’18.4’’</geoCoordinate>
\
<geoCoordinate id="9C89D9BDF262143D77EDEBFE51A529D2" box="[1231,1408,441,468]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="-77.88583">W 77º53’09.0’’</geoCoordinate>
,
<date id="8D0399BAF262143D76ACEBFD549B29FE" pageId="11" pageNumber="12">
<collectingDate id="9D476052F262143D76ACEBFD549B29FE" pageId="11" pageNumber="12">9 XII</collectingDate>
</date>
\ 2009, leg.
<collectorName id="5448DAACF262143D7275EB9955FC29F1" box="[343,473,477,503]" pageId="11" pageNumber="12">Rafał Ruta</collectorName>
”.
<specimenCount id="EFBB74F3F262143D72D2EB99564529F1" box="[496,608,477,503]" pageId="11" pageNumber="12" type="female">2 females</specimenCount>
(
<collectionCode id="9FAC27BFF262143D7150EB9956E729F1" box="[626,706,477,503]" pageId="11" pageNumber="12">DBET</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="49D5B527F262143D71FAEB9955B32AC9" ID-GBIF-Occurrence="2239706633" collectingDate="1999-11-05" collectingDateMax="2009-12-17" collectingDateMin="1999-11-05" collectionCode="DBET, SEMC" collectorName="L. Borowiec &amp; Rafal Ruta &amp; Z. H. Fallin" country="Ecuador" elevation="2100" location="Female" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
“
<collectingCountry id="81AAFFEAF262143D71C3EB99574B29F1" box="[737,878,477,503]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
,
<collectingRegion id="3B797198F262143D705FEB9A579F29FE" box="[893,954,478,504]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov. \ Cosanga vic., Yanayacu \ Station,
<quantity id="3E45129FF262143D73B5E845550B2A1D" box="[151,302,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F262143D73B5E845550B2A1D" box="[151,302,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’ S, 77°53’W, \
<date id="8D0399BAF262143D7115E84557342A1D" box="[567,785,513,540]" pageId="11" pageNumber="12" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">
<collectingDate id="9D476052F262143D7115E84557342A1D" box="[567,785,513,540]" pageId="11" pageNumber="12" value="2009-11-23" valueMax="2009-12-17" valueMin="2009-11-23">23 XI–17 XII 2009</collectingDate>
</date>
, \ leg.
<collectorName id="5448DAACF262143D7079E84657C82A1D" box="[859,1005,513,540]" pageId="11" pageNumber="12">L. Borowiec</collectorName>
”.
<location id="FC62E9A1F262143D7723E84550762A1D" LSID="urn:lsid:plazi:treatment:71140E6CF262143D73B5EAA551AF2F75:FC62E9A1F262143D7723E84550762A1D" box="[1025,1107,513,539]" country="Ecuador" name="Female" pageId="11" pageNumber="12" stateProvince="Napo">Female</location>
(
<collectionCode id="9FAC27BFF262143D7741E84550962A1D" box="[1123,1203,513,539]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 19,
<collectingRegion id="3B797198F262143D7642E84651B82A1A" box="[1376,1437,514,540]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Yanayacu \ Station, moist forest near \ the road,
<quantity id="3E45129FF262143D7093E861506D2A39" box="[945,1096,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
<elevation id="72905849F262143D7093E861506D2A39" box="[945,1096,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="11" pageNumber="12" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200m</elevation>
</quantity>
, \ 00°35’S / 77°53’W, 27 \
<date id="8D0399BAF262143D765DE86254F72A65" pageId="11" pageNumber="12" value="2009-11">
<collectingDate id="9D476052F262143D765DE86254F72A65" pageId="11" pageNumber="12" value="2009-11">XI 2009</collectingDate>
</date>
, leg.
<collectorName id="5448DAACF262143D722CE80D55A82A65" box="[270,397,585,611]" pageId="11" pageNumber="12">Rafał Ruta</collectorName>
”. Female (
<collectionCode id="9FAC27BFF262143D7126E80D56722A65" box="[516,599,585,611]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="12" type="Museum">SEMC</collectionCode>
): “
<collectingCountry id="81AAFFEAF262143D7156E80D57262A65" box="[628,771,585,611]" name="Ecuador" pageId="11" pageNumber="12">ECUADOR</collectingCountry>
:
<collectingRegion id="3B797198F262143D702FE80E57692A62" box="[781,844,586,612]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
, Cosanga \ 1.0 km N on Baeza-Tena \ Road then
<quantity id="3E45129FF262143D765BE80D549E2A81" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="km" value="1.5">1.5 km</quantity>
SW on road to \ Sierra Azul,
<quantity id="3E45129FF262143D712CE82956792A8E" box="[526,604,621,648]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="11" pageNumber="12" unit="m" value="2150.0">
<elevation id="72905849F262143D712CE82956792A8E" box="[526,604,621,648]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="11" pageNumber="12" unit="m" value="2150.0">2150m</elevation>
</quantity>
\ 0°35’24”S, 77°53’19”W \
<date id="8D0399BAF262143D7082E82950742A81" box="[928,1105,621,648]" pageId="11" pageNumber="12" value="1999-11-05" valueMax="1999-11-07" valueMin="1999-11-05">
<collectingDate id="9D476052F262143D7082E82950742A81" box="[928,1105,621,648]" pageId="11" pageNumber="12" value="1999-11-05" valueMax="1999-11-07" valueMin="1999-11-05">5–7 NOV 1999</collectingDate>
</date>
;
<collectorName id="5448DAACF262143D777FE82A50C72A81" box="[1117,1250,621,648]" pageId="11" pageNumber="12">Z. H. Fallin</collectorName>
\ ECU1F99 124
<collectingMethod id="20FCC76DF262143D73B5E8D555502AAD" box="[151,373,657,683]" pageId="11" pageNumber="12">flight intercept trap</collectingMethod>
”. Female (
<collectionCode id="9FAC27BFF262143D72CEE8D556192AAD" box="[492,572,657,683]" pageId="11" pageNumber="12">DBET</collectionCode>
): “Ecuador 41,
<collectingRegion id="3B797198F262143D71C8E8D657022AAA" box="[746,807,658,684]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo</collectingRegion>
prov., \ Cosanga vic., Antisana \ trail, Yanayacu,
<date id="8D0399BAF262143D7673E8D551A82AAA" box="[1361,1421,657,684]" pageId="11" pageNumber="12">
<collectingDate id="9D476052F262143D7673E8D551A82AAA" box="[1361,1421,657,684]" pageId="11" pageNumber="12">3 XII</collectingDate>
</date>
\ 2009, leg.
<collectorName id="5448DAACF262143D722FE8F155A92AC9" box="[269,396,693,719]" pageId="11" pageNumber="12">Rafał Ruta</collectorName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5E89D563D2B85" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="F902BF7AF262143D73E5E89D563D2B85" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5E89D55672AF2" bold="true" box="[199,322,729,756]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Moderately large (TL
<quantity id="3E45129FF262143D711FE89D56E22AF2" box="[573,711,729,756]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="11" pageNumber="12" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
), resembling black individuals of
<taxonomicName id="3EBDC4F9F262143D771CE89E50DE2AF5" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1086,1275,730,756]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
<emphasis id="CBC96368F262143D771CE89E50DE2AF5" box="[1086,1275,730,756]" italics="true" pageId="11" pageNumber="12">C. yanayacuense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D7623E89E517F2AF2" bold="true" box="[1281,1370,730,756]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D7623E89E517F2AF2" box="[1281,1370,730,756]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Penis very long and narrow, apical plate of tergite VIII with M-shaped median process flanked with shallow emarginations covered with distinct denticles. Genitalia similar to that of
<taxonomicName id="3EBDC4F9F262143D700EE96657C22B3A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[812,999,802,828]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F262143D700EE96657C22B3A" box="[812,999,802,828]" italics="true" pageId="11" pageNumber="12">C. fourgassiense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D70CCE966506D2B3A" bold="true" box="[1006,1096,802,828]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D70CCE966506D2B3A" box="[1006,1096,802,828]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but median process of tergite VIII is longer and better developed in
<taxonomicName id="3EBDC4F9F262143D7172E90256C42B59" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[592,737,837,864]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F262143D7172E90256C42B59" box="[592,737,837,864]" italics="true" pageId="11" pageNumber="12">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D71C8E90257632B66" bold="true" box="[746,838,838,864]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D71C8E90257632B66" box="[746,838,838,864]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Similar to
<taxonomicName id="3EBDC4F9F262143D70EAE90250592B59" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[968,1148,837,864]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
<emphasis id="CBC96368F262143D70EAE90250592B59" box="[968,1148,837,864]" italics="true" pageId="11" pageNumber="12">C. venezuelense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F262143D77A7E90250C42B66" bold="true" box="[1157,1249,838,864]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="D0FADE13F262143D77A7E90250C42B66" box="[1157,1249,838,864]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but body is less elongated and its sides are curved.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5E9C955732F01" pageId="11" pageNumber="12" type="description">
<paragraph id="F902BF7AF262143D73E5E9C9511A2CCE" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5E9C9557C2BAE" bold="true" box="[199,345,909,936]" pageId="11" pageNumber="12">Description.</emphasis>
Male. Body oblong oval, convex, covered with brown, suberect setae. Body uniformly dark brown, venter brown, legs yellowish-brown, antennae yellowish, darkening towards apex. Head very broad, 1.6× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by a 3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 2.0–3.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, with very subtle punctation. Elytra oblong, without traces of longitudinal ridges, sides subparallel in basal 1/2, curved and regularly converging to apex in posterior 1/2. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.
</paragraph>
<paragraph id="F902BF7AF262143D73E5EE9550302DE1" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
Male genitalia. Penis (L
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,
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W
<quantity id="3E45129FF262143D71A2EE9556942CED" box="[640,689,1233,1259]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="mm" value="0.1">0.10</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF262143D71D5EE95571B2CED" box="[759,830,1233,1259]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="11" pageNumber="12">Figs 9</figureCitation>
A–B) very long and narrow, widest in basal portion, parameroids short, very narrow, curved, rounded at apices, trigonium subtriangular, narrow, pointed at apex. Teg- men (L
<quantity id="3E45129FF262143D73CCEF5D55762D32" box="[238,339,1305,1332]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="11" pageNumber="12" unit="mm" value="0.78">0.78 mm</quantity>
,
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W
<quantity id="3E45129FF262143D725CEF5D558A2D35" box="[382,431,1305,1331]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="12" unit="mm" value="0.21">0.21</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF262143D72CCEF5D56662D35" box="[494,579,1305,1331]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="11" pageNumber="12">Fig. 9C</figureCitation>
) very long, sides diverging, apex with short lateral lobes, covered with setae. Sternite IX (L
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,
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W
<quantity id="3E45129FF262143D72F3EF7956272D5E" box="[465,514,1341,1368]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="12" unit="mm" value="0.25">0.25</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF262143D7166EF7956B92D51" box="[580,668,1341,1367]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="11" pageNumber="12">Fig. 9D</figureCitation>
) subrectangular, with paired apodemes, apical margin rounded, api- cal portion covered with short, sparse setae. Tergite VIII (L
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,
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W
<quantity id="3E45129FF262143D70EAEF2557DC2D7D" box="[968,1017,1377,1403]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="11" pageNumber="12" unit="mm" value="0.37">0.37</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF262143D7718EF2550AB2D7D" box="[1082,1166,1377,1403]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="11" pageNumber="12">Fig. 9E</figureCitation>
) with short apical plate, central portion of apical plate produced, M-shaped, with V-shaped emargination at apex, lateral portions of apical margin with a few denticles, apodemes distinctly longer than apical plate, diverging. Tergite IX (L
<quantity id="3E45129FF262143D77FEEFED51642DC2" box="[1244,1345,1449,1476]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="12" unit="mm" value="0.45">0.45 mm</quantity>
,
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W
<quantity id="3E45129FF262143D7649EFED51B92DC5" box="[1387,1436,1449,1475]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="12" unit="mm" value="0.31">0.31</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF262143D73F3EF8955062DE1" box="[209,291,1485,1511]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="11" pageNumber="12">Fig. 9F</figureCitation>
) with membranous apical plate, apodemes longer than apical plate.
</paragraph>
<paragraph id="F902BF7AF262143D73E5EFB557202E9D" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
Female. Body (
<figureCitation id="6186A3FFF262143D7254EFB555E92E0D" box="[374,460,1521,1547]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="11" pageNumber="12">Fig. 1B</figureCitation>
) dark brown to black, with no orange pattern on dorsal portion, TL (n = 5) 2.60–3.10 (2.88) mm. Apical portion of each elytron with two concavities, shallow one and deeper one, situated closer to suture and covered with longer setae, forming a peculiar hooked tuft of setae (
<figureCitation id="6186A3FFF262143D773EEC7D505D2E52" box="[1052,1144,1593,1620]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="11" pageNumber="12">Fig. 7A</figureCitation>
). Gonocoxites (
<figureCitation id="6186A3FFF262143D760EEC7D51B12E55" box="[1324,1428,1593,1619]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="11" pageNumber="12">Fig. 10A</figureCitation>
) broadly triangular, gonostyli subapical, bursal sclerite (L
<quantity id="3E45129FF262143D7034EC19575C2E7E" box="[790,889,1629,1656]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="mm" value="0.35">0.35 mm</quantity>
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W
<quantity id="3E45129FF262143D70BDEC1957F52E71" box="[927,976,1629,1655]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="mm" value="0.13">0.13</quantity>
</geoCoordinate>
mm,
<figureCitation id="6186A3FFF262143D7732EC1950472E71" box="[1040,1122,1629,1655]" captionStart="FIGURE 10" captionStartId="13.[151,250,1849,1874]" captionTargetBox="[178,1410,181,1810]" captionTargetId="figure@13.[178,1412,181,1810]" captionTargetPageId="13" captionText="FIGURE 10. Calvariopsis gen. nov., female genitalia, ascl—anterior portion of bursal sclerite, bsa—bursa, bsl—bursella, gla—gland, oscl—outgrowth of bursal sclerite, pscl—posterior portion of bursal sclerite, scl—bursal sclerite. A–C) C. borowieci sp. nov.; D) C. pittieri sp. nov.; E–H) C. yanyacuense sp. nov.; I–K) C. brasiliense (Pic), specimen from Sao Paulo; L–M) C. brasiliense (Pic), specimen from Santo Amaro. A, D, E, L) genital tract; B, C, G, H, M) bursal sclerite; F, K) gonocoxites; I) sternite VIII; J) ovipositor." httpUri="https://zenodo.org/record/2783730/files/figure.png" pageId="11" pageNumber="12">Figs 10</figureCitation>
B–C) elongated, widened in posterior portion, with subconical outgrowth in middle.
</paragraph>
<paragraph id="F902BF7AF262143D73E5ECE255732F01" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5ECE255DA2EC6" bold="true" box="[199,511,1701,1728]" pageId="11" pageNumber="12">Measurements and ratios.</emphasis>
Males (n = 2) TL 2.60–3.13 (2.87) mm, PL 0.50–0.57 (0.53) mm, PW 1.07–1.23 (1.15) mm, EL 2.33–2.67 (2.50) mm, EW 1.60–1.93 (1.77) mm, TL/EW 1.6, PW/PL 2.1–2.2 (2.2), EL/EW 1.4–1.5 (1.4), EL/PL 4.7.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5ED5557BB2F2D" box="[199,926,1809,1836]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="F902BF7AF262143D73E5ED5557BB2F2D" blockId="11.[151,1437,333,1908]" box="[199,926,1809,1836]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5ED5555442F2A" bold="true" box="[199,353,1809,1836]" pageId="11" pageNumber="12">Distribution.</emphasis>
Known from localities in
<collectingRegion id="3B797198F262143D71A5ED5657152F2D" box="[647,816,1809,1836]" country="Ecuador" name="Napo" pageId="11" pageNumber="12">Napo province</collectingRegion>
,
<collectingCountry id="81AAFFEAF262143D7019ED5557BF2F2D" box="[827,922,1809,1835]" name="Ecuador" pageId="11" pageNumber="12">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F262143D73E5ED7251AF2F75" pageId="11" pageNumber="12" type="etymology">
<paragraph id="F902BF7AF262143D73E5ED7251AF2F75" blockId="11.[151,1437,333,1908]" pageId="11" pageNumber="12">
<emphasis id="CBC96368F262143D73E5ED72556B2F56" bold="true" box="[199,334,1846,1872]" pageId="11" pageNumber="12">Etymology.</emphasis>
Named after Prof. Lech Borowiec, an expert in
<taxonomicName id="3EBDC4F9F262143D707DED7150BF2F56" authority="" box="[863,1178,1845,1872]" class="Insecta" family="Chrysomelidae" genus="Cassidinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Chrysomelidae">Cassidinae (Chrysomelidae)</taxonomicName>
systematics, my teach- er, member of the expedition to
<collectingCountry id="81AAFFEAF262143D72DCED1D567E2F75" box="[510,603,1881,1907]" name="Ecuador" pageId="11" pageNumber="12">Ecuador</collectingCountry>
in 2009 and collector of many specimens examined in the present study.
</paragraph>
</subSubSection>
</treatment>
</document>
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