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<document id="7F4651876B3EF300894C71232D777B01" ID-CLB-Dataset="26847" ID-DOI="10.11646/zootaxa.4604.1.1" ID-GBIF-Dataset="b5902a9d-c38b-462f-ad8a-95693c1822dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="2783674" ID-ZooBank="E6D82300-8DC6-4560-A945-8F29D3B3D501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1557723545002" checkinUser="plazi" docAuthor="Ruta, Rafał" docDate="2019" docId="71140E6CF24F141173B5EFBD51B92AFF" docLanguage="en" docName="zootaxa.4604.1.1.pdf" docOrigin="Zootaxa 4604 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvariopsis Ruta 2019, gen. nov." docType="key" docVersion="12" lastPageNumber="40" masterDocId="8D2D7614F26914367322EA4454252806" masterDocTitle="Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="39" updateTime="1698728494285" updateUser="ExternalLinkService">
<mods:mods id="E6C3E12F93A54B6C2E71D25905A9CD5B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6BCB1EB3B58AEB48458BDC909826C6B6">
<mods:title id="A403E9C9A22DD8FA01F40853D19E379F">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
</mods:titleInfo>
<mods:name id="6A3806DB64B4377521817C695C4276A2" type="personal">
<mods:role id="813096B43EAB9B00EDEC06A04A977E2D">
<mods:roleTerm id="A56DFF3B830288BE5A5CAFBA6A4C1A1A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE0B72F5E4BBC6C1B5D5D481ADC8339E">Ruta, Rafał</mods:namePart>
</mods:name>
<mods:typeOfResource id="78215F9C7BD46689FA932C495CB092C8">text</mods:typeOfResource>
<mods:relatedItem id="9B522E08D80D002AF9A32B53E49A7CC8" type="host">
<mods:titleInfo id="516DE821F6A068858148BDE534AA6EA9">
<mods:title id="0E5E8BBAD50C494D23E06F0D87D8D43E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="00BC0B6EA88265C3746DAC5363060356">
<mods:date id="96B540EA769903606500FDE1553EF399">2019</mods:date>
<mods:detail id="2E3A5E40E95EE1DAF28BB8DFB9AF2387" type="pubDate">
<mods:number id="0A8FBE2BBDD76A3D70F1E800544BDB82">2019-05-13</mods:number>
</mods:detail>
<mods:detail id="04B44D5E19994105994EE491231387E4" type="volume">
<mods:number id="EF4C294A690B3369F3D4607E2778337E">4604</mods:number>
</mods:detail>
<mods:detail id="51E87AF225668C4F55B6700D3C5F4411" type="issue">
<mods:number id="F2E806BA0699399AF2F3786446B27A86">1</mods:number>
</mods:detail>
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<mods:start id="DB5553E2F1065DC98EE7F4DE203AC155">1</mods:start>
<mods:end id="1D5C30ECC65254797D7941736A29F5B2">41</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="50D61354A11806C3DBEE518C5CC9E78E">journal article</mods:classification>
<mods:identifier id="271FCC33EE985512CD62003B0E4856B8" type="CLB-Dataset">26847</mods:identifier>
<mods:identifier id="28458BDE030BE83AC57F3FFFC412948E" type="DOI">10.11646/zootaxa.4604.1.1</mods:identifier>
<mods:identifier id="0C75AA233E62D1A5D06E33B439C936B3" type="GBIF-Dataset">b5902a9d-c38b-462f-ad8a-95693c1822dc</mods:identifier>
<mods:identifier id="1253BB8ABBD3FA65320935507E95651A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3E9AD8BDA790B023653D9FAB81DACF86" type="Zenodo-Dep">2783674</mods:identifier>
<mods:identifier id="D4B6BF7B8B79CB18DBECD6E4B1C206BD" type="ZooBank">E6D82300-8DC6-4560-A945-8F29D3B3D501</mods:identifier>
</mods:mods>
<treatment id="71140E6CF24F141173B5EFBD51B92AFF" ID-GBIF-Taxon="157135221" LSID="urn:lsid:plazi:treatment:71140E6CF24F141173B5EFBD51B92AFF" httpUri="http://treatment.plazi.org/id/71140E6CF24F141173B5EFBD51B92AFF" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<subSubSection id="B1A7ECF1F24F141073B5EFBD51392E30" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="F902BF7AF24F141073B5EFBD57032E15" blockId="38.[151,1308,1528,1591]" box="[151,806,1528,1555]" pageId="38" pageNumber="39">
<heading id="A24A0816F24F141073B5EFBD57032E15" bold="true" box="[151,806,1528,1555]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="CBC96368F24F141073B5EFBD57032E15" bold="true" box="[151,806,1528,1555]" pageId="38" pageNumber="39">
Key to identification for males of
<taxonomicName id="3EBDC4F9F24F14107104EFBC56902E15" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[550,693,1528,1555]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CBC96368F24F14107104EFBC56902E15" bold="true" box="[550,693,1528,1555]" italics="true" pageId="38" pageNumber="39">Calvariopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0FADE13F24F1410719FEFBD57032E15" box="[701,806,1529,1555]" pageId="38" pageNumber="39" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F902BF7AF24F141073B5EC5851392E30" blockId="38.[151,1308,1528,1591]" box="[151,1308,1564,1591]" pageId="38" pageNumber="39">
Please note that
<emphasis id="CBC96368F24F1410726CEC5855F12E30" box="[334,468,1564,1590]" italics="true" pageId="38" pageNumber="39">Calvaropsis</emphasis>
<taxonomicName id="3EBDC4F9F24F141072F9EC58568D2E31" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[475,680,1564,1591]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sculptipenne">
<emphasis id="CBC96368F24F141072F9EC58564C2E30" box="[475,617,1564,1590]" italics="true" pageId="38" pageNumber="39">sculptipenne</emphasis>
(Pic)
</taxonomicName>
, with only the female known, is not included in the key.
</paragraph>
</subSubSection>
<subSubSection id="B1A7ECF1F24F141173B5EC2551B92AFF" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="key">
<keyStep id="4249A7DFF24F141073B5EC2551B92E92" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5EC2551B92E71" blockId="38.[151,1437,1633,2037]" box="[151,1436,1633,1655]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EC2551B92E71" box="[151,1436,1633,1655]" pageId="38" pageNumber="39">
1. Sternite IX with paired apodemes (
<figureCitation id="6186A3FFF24F14107110EC2556AD2E71" box="[562,648,1633,1655]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="38" pageNumber="39">Fig. 11D</figureCitation>
)........................................................ 2 (group I)
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5EC3B51B92E92" blockId="38.[151,1437,1633,2037]" box="[151,1436,1662,1685]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EC3B51B92E92" box="[151,1436,1662,1685]" pageId="38" pageNumber="39">
-. Sternite IX with a single apodeme (
<figureCitation id="6186A3FFF24F14107114EC3A56A92E93" box="[566,652,1662,1685]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="38" pageNumber="39">Fig. 21C</figureCitation>
)....................................................... 9 (group II)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24F141073B5ECD851B92EEA" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5ECD851B92EB7" blockId="38.[151,1437,1633,2037]" box="[151,1436,1692,1714]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ECD851B92EB7" box="[151,1436,1692,1714]" pageId="38" pageNumber="39">
2. Basal half of penis distinctly wider than apical half (
<figureCitation id="6186A3FFF24F141071FCECD857132EB4" box="[734,822,1692,1714]" captionStart="." captionStartId="15.[404,408,1541,1564]" captionTargetBox="[153,1435,668,1486]" captionTargetId="figure@15.[153,1438,668,1488]" captionText="." pageId="38" pageNumber="39">Fig. 12A</figureCitation>
)................................
<taxonomicName id="3EBDC4F9F24F141077E0ECD8516F2EB7" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1218,1354,1692,1713]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="panamense" status="sp. nov.">
<emphasis id="CBC96368F24F141077E0ECD8516F2EB7" box="[1218,1354,1692,1713]" italics="true" pageId="38" pageNumber="39">C. panamense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672ECD851B92EB7" bold="true" box="[1360,1436,1692,1713]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672ECD851B92EB7" box="[1360,1436,1692,1713]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5ECFE51B92EEA" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ECFE51B92EEA" pageId="38" pageNumber="39">
-. Basal portion of penis as wide as apical half (
<figureCitation id="6186A3FFF24F141071BFECFD56D02EC9" box="[669,757,1721,1743]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="38" pageNumber="39">Fig. 14A</figureCitation>
) or less than basal 1/3 of penis is slightly wider than apical half (Figs 13A, 15A)...........................................................................................3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24F141073B5ECB051B92F42" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5ECB051B92F0F" blockId="38.[151,1437,1633,2037]" box="[151,1436,1780,1802]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ECB051B92F0F" box="[151,1436,1780,1802]" pageId="38" pageNumber="39">
3. Penis moderately long, 6× longer than wide (
<figureCitation id="6186A3FFF24F141071B0ECB056CB2F0F" box="[658,750,1780,1802]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="38" pageNumber="39">Figs 14A</figureCitation>
,
<figureCitation id="6186A3FFF24F141071DAECB057012F0C" box="[760,804,1780,1802]" captionStart="FIGURE 15" captionStartId="20.[151,250,1137,1162]" captionTargetBox="[186,1401,251,1088]" captionTargetId="figure@20.[186,1403,272,1088]" captionTargetPageId="20" captionText="FIGURE 15. Calvariopsis pittieri sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783750/files/figure.png" pageId="38" pageNumber="39">15A</figureCitation>
), with subparallel or subtly sinuate lateral margins............4
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5ED5651B92F42" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ED5651B92F42" pageId="38" pageNumber="39">
-. Penis very long and narrow, more than 8–10× longer than wide (
<figureCitation id="6186A3FFF24F14107068ED5557802F20" box="[842,933,1809,1831]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="38" pageNumber="39">Figs 11A</figureCitation>
,
<figureCitation id="6186A3FFF24F1410708DED5557FE2F21" box="[943,987,1809,1831]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="38" pageNumber="39">13A</figureCitation>
), widened in basal portion and narrowed in the middle...............................................................................................5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24F141073B5ED0851B92F9A" pageId="38" pageNumber="39">
<paragraph id="F902BF7AF24F141073B5ED0851B92F78" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ED0851B92F78" pageId="38" pageNumber="39">
4. Penis with sinuate lateral margins, trigonium narrow, with subparallel sides (
<figureCitation id="6186A3FFF24F141070E0ED08503F2F64" box="[962,1050,1868,1890]" captionStart="FIGURE 15" captionStartId="20.[151,250,1137,1162]" captionTargetBox="[186,1401,251,1088]" captionTargetId="figure@20.[186,1403,272,1088]" captionTargetPageId="20" captionText="FIGURE 15. Calvariopsis pittieri sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783750/files/figure.png" pageId="38" pageNumber="39">Fig. 15A</figureCitation>
), known only from
<collectingCountry id="81AAFFEAF24F141077FAED08511E2F64" box="[1240,1339,1868,1890]" name="Venezuela" pageId="38" pageNumber="39">Venezuela</collectingCountry>
.............................................................................................
<taxonomicName id="3EBDC4F9F24F141077CEED2D516C2F79" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1260,1353,1897,1919]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
<emphasis id="CBC96368F24F141077CEED2D516C2F79" box="[1260,1353,1897,1919]" italics="true" pageId="38" pageNumber="39">C. pittieri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672ED2D51B92F78" bold="true" box="[1360,1436,1897,1918]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672ED2D51B92F78" box="[1360,1436,1897,1918]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5EDC351B92F9A" blockId="38.[151,1437,1633,2037]" box="[151,1436,1926,1949]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EDC351B92F9A" box="[151,1436,1926,1949]" pageId="38" pageNumber="39">
-. Penis with subparallel sides, trigonium wide, subtriangular (
<figureCitation id="6186A3FFF24F14107001EDC257592F9B" box="[803,892,1926,1949]" captionStart="FIGURE 14" captionStartId="18.[151,250,1731,1756]" captionTargetBox="[184,1403,897,1694]" captionTargetId="figure@18.[181,1405,897,1694]" captionTargetPageId="18" captionText="FIGURE 14. Calvariopsis picta sp. nov., male.A) penis; B) tegmen; C) sternite IX; D) tergite VIII and sternite VIII; E) tergite IX; F) terminal segments and genitalia, not separated. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783746/files/figure.png" pageId="38" pageNumber="39">Fig. 14A</figureCitation>
), known only from
<collectingCountry id="81AAFFEAF24F14107718EDC250A42F9A" box="[1082,1153,1926,1948]" name="Bolivia" pageId="38" pageNumber="39">Bolivia</collectingCountry>
.........
<taxonomicName id="3EBDC4F9F24F141077DCEDC3516F2F9A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1278,1354,1927,1948]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="picta" status="sp. nov.">
<emphasis id="CBC96368F24F141077DCEDC3516F2F9A" box="[1278,1354,1927,1948]" italics="true" pageId="38" pageNumber="39">C. picta</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672EDC351B92F9A" bold="true" box="[1360,1436,1927,1948]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672EDC351B92F9A" box="[1360,1436,1927,1948]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="F902BF7AF24F141073B5EDE051B92FD0" blockId="38.[151,1437,1633,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5EDE051B92FD0" pageId="38" pageNumber="39">
<keyStep id="4249A7DFF24F141073B5EDE051B92FD0" pageId="38" pageNumber="39">
5. TL
<quantity id="3E45129FF24F14107225EDE055702FBF" box="[263,341,1956,1977]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="38" pageNumber="39" unit="mm" value="2.3">2.3 mm</quantity>
, median process of tergite VIII not flanked with deep emarginations (Fig.
<figureCitation id="6186A3FFF24F14107719EDE050B42FBC" box="[1083,1169,1956,1978]" captionStart="FIGURE 11" captionStartId="14.[151,250,1377,1402]" captionTargetBox="[153,1435,181,1322]" captionTargetId="figure@14.[153,1438,181,1322]" captionTargetPageId="14" captionText="FIGURE 11. Calvariopsis fourgassiense sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783738/files/figure.png" pageId="38" pageNumber="39">Fig. 11E</figureCitation>
), known only from
<collectingCountry id="81AAFFEAF24F14107678EDE055022FD1" name="French Guiana" pageId="38" pageNumber="39">French Guiana</collectingCountry>
...........................................................................
<taxonomicName id="3EBDC4F9F24F14107788ED85516C2FD0" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1194,1353,1985,2006]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fourgassiense" status="sp. nov.">
<emphasis id="CBC96368F24F14107788ED85516C2FD0" box="[1194,1353,1985,2006]" italics="true" pageId="38" pageNumber="39">C. fourgassiense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24F14107672ED8551B92FD0" bold="true" box="[1360,1436,1985,2006]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="D0FADE13F24F14107672ED8551B92FD0" box="[1360,1436,1985,2006]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24F141073B5ED9B51B92FF2" blockId="38.[151,1437,1633,2037]" box="[151,1436,2014,2037]" pageId="38" pageNumber="39">
<keyLead id="424C1C4FF24F141073B5ED9B51B92FF2" box="[151,1436,2014,2037]" pageId="38" pageNumber="39">
-. TL
<quantity id="3E45129FF24F14107226ED9A555F2FF2" box="[260,378,2014,2036]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.6" metricValueMin="2.5" pageId="38" pageNumber="39" unit="mm" value="3.05" valueMax="3.6" valueMin="2.5">2.5–3.6 mm</quantity>
, median process of tergite VIII flanked with deep emarginations (
<figureCitation id="6186A3FFF24F141070CEED9A501C2FF2" box="[1004,1081,2014,2036]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="38" pageNumber="39">Figs 9E</figureCitation>
,
<figureCitation id="6186A3FFF24F14107760ED9B504E2FF2" box="[1090,1131,2015,2036]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="38" pageNumber="39">13E</figureCitation>
,
<figureCitation id="6186A3FFF24F14107756ED9B50BA2FF2" box="[1140,1183,2015,2036]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="38" pageNumber="39">17C</figureCitation>
, 18D), known from Andes
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173C0EADE51B928CA" blockId="39.[151,1437,153,762]" pageId="39" pageNumber="40">
(
<collectingCountry id="81AAFFEAF24E141173CBEADE553C28A9" box="[233,281,154,175]" name="Peru" pageId="39" pageNumber="40">Peru</collectingCountry>
,
<collectingCountry id="81AAFFEAF24E14117200EADD555128A9" box="[290,372,153,175]" name="Ecuador" pageId="39" pageNumber="40">Ecuador</collectingCountry>
,
<collectingCountry id="81AAFFEAF24E1411725EEADD55C528A9" box="[380,480,153,175]" name="Venezuela" pageId="39" pageNumber="40">Venezuela</collectingCountry>
).............................................................................. 6
<keyLead id="424C1C4FF24E141173B5EAF351B928CA" box="[151,1436,183,205]" pageId="39" pageNumber="40">
<keyStep id="4249A7DFF24E141173B5EAF351B928CA" box="[151,1436,183,205]" pageId="39" pageNumber="40">
6. Median process of tergite VIII short, flanked by deep, semicircular emarginations (
<figureCitation id="6186A3FFF24E141170DFEAF3507428CB" box="[1021,1105,183,205]" captionStart="FIGURE 13" captionStartId="17.[151,250,1690,1715]" captionTargetBox="[153,1435,396,1639]" captionTargetId="figure@17.[153,1438,417,1639]" captionTargetPageId="17" captionText="FIGURE 13. Calvariopsis peruviana sp. nov., male. A) penis; B) apex of penis, lateral view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX; G) tergite VII. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783740/files/figure.png" pageId="39" pageNumber="40">Fig. 13E</figureCitation>
)........
<taxonomicName id="3EBDC4F9F24E141177EEEAF3516C28CA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1228,1353,183,204]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="CBC96368F24E141177EEEAF3516C28CA" box="[1228,1353,183,204]" italics="true" pageId="39" pageNumber="40">C. peruviana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EAF351B928CA" bold="true" box="[1360,1436,183,204]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EAF351B928CA" box="[1360,1436,183,204]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EA9151B928EC" blockId="39.[151,1437,153,762]" box="[151,1436,212,234]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EA9151B928EC" box="[151,1436,212,234]" pageId="39" pageNumber="40">
-. Median process of tergite VIII longer, flanked by shallow and wide emarginations (
<figureCitation id="6186A3FFF24E14117723EA90506A28EF" box="[1025,1103,212,234]" captionStart="FIGURE 9" captionStartId="12.[151,250,1323,1348]" captionTargetBox="[158,1430,181,1270]" captionTargetId="figure@12.[158,1433,181,1270]" captionTargetPageId="12" captionText="FIGURE 9. Calvariopsis borowieci sp. nov., male. A) penis; B) apex of penis; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783724/files/figure.png" pageId="39" pageNumber="40">Figs 9E</figureCitation>
,
<figureCitation id="6186A3FFF24E1411777BEA9050A628EF" box="[1113,1155,212,233]" captionStart="FIGURE 17" captionStartId="22.[151,250,1621,1646]" captionTargetBox="[152,1422,215,1572]" captionTargetId="figure@22.[165,1424,237,1572]" captionTargetPageId="22" captionText="FIGURE 17. Calvariopsis venezuelense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783760/files/figure.png" pageId="39" pageNumber="40">17C</figureCitation>
, 18D)................. 7
</keyLead>
</paragraph>
<keyStep id="4249A7DFF24E141173B5EAB651B92922" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EAB651B92901" blockId="39.[151,1437,153,762]" box="[151,1436,241,264]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EAB651B92901" box="[151,1436,241,264]" pageId="39" pageNumber="40">
7. Median process of tergite VIII wide, short, M-shaped, emarginations without denticles (Fig. 18D).
<taxonomicName id="3EBDC4F9F24E14117785EAB6516F2901" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1191,1354,242,263]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="yanayacuense" status="sp. nov.">
<emphasis id="CBC96368F24E14117785EAB6516F2901" box="[1191,1354,242,263]" italics="true" pageId="39" pageNumber="40">C. yanayacuense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EAB651B92901" bold="true" box="[1360,1436,242,263]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EAB651B92901" box="[1360,1436,242,263]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EB4B51B92922" blockId="39.[151,1437,153,762]" box="[151,1436,271,293]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB4B51B92922" box="[151,1436,271,293]" pageId="39" pageNumber="40">-. Median process of tergite VIII longer, emarginations covered with denticles........................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EB6851B92959" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EB6851B92947" blockId="39.[151,1437,153,762]" box="[151,1436,300,322]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB6851B92947" box="[151,1436,300,322]" pageId="39" pageNumber="40">
8. Body oblong oval, with rounded sides (
<figureCitation id="6186A3FFF24E1411717EEB68568D2944" box="[604,680,300,322]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="39" pageNumber="40">Fig. 1A</figureCitation>
), TL/EW 1.6, known only from
<collectingCountry id="81AAFFEAF24E141170C2EB6850152944" box="[992,1072,300,322]" name="Ecuador" pageId="39" pageNumber="40">Ecuador</collectingCountry>
............
<taxonomicName id="3EBDC4F9F24E141177ECEB68516C2947" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1230,1353,300,321]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="borowieci" status="sp. nov.">
<emphasis id="CBC96368F24E141177ECEB68516C2947" box="[1230,1353,300,321]" italics="true" pageId="39" pageNumber="40">C. borowieci</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB6851B92947" bold="true" box="[1360,1436,300,321]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB6851B92947" box="[1360,1436,300,321]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EB0E51B92959" blockId="39.[151,1437,153,762]" box="[151,1436,329,352]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB0E51B92959" box="[151,1436,329,352]" pageId="39" pageNumber="40">
-. Body elongate, with subparallel sides (
<figureCitation id="6186A3FFF24E14117174EB0D56BA2966" box="[598,671,329,352]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="39" pageNumber="40">Fig. 1L</figureCitation>
), TL/EW 1.8, known only from
<collectingCountry id="81AAFFEAF24E141170F4EB0D501C2959" box="[982,1081,329,351]" name="Venezuela" pageId="39" pageNumber="40">Venezuela</collectingCountry>
.........
<taxonomicName id="3EBDC4F9F24E14117792EB0E516C2958" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1200,1353,329,351]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="venezuelense" status="sp. nov.">
<emphasis id="CBC96368F24E14117792EB0E516C2958" box="[1200,1353,329,351]" italics="true" pageId="39" pageNumber="40">C. venezuelense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB0E51B92959" bold="true" box="[1360,1436,330,351]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB0E51B92959" box="[1360,1436,330,351]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EB2351B9299F" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EB2351B9297A" blockId="39.[151,1437,153,762]" box="[151,1436,359,381]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB2351B9297A" box="[151,1436,359,381]" pageId="39" pageNumber="40">
9. Last abdominal ventrite with paired tubercles (
<figureCitation id="6186A3FFF24E14117180EB2356C9297A" box="[674,748,359,381]" captionStart="FIGURE 3" captionStartId="5.[151,250,1905,1930]" captionTargetBox="[165,1422,181,1857]" captionTargetId="figure@5.[165,1424,181,1857]" captionTargetPageId="5" captionText="FIGURE 3. Calvariopsis gen. nov., SEM images of morphological details, A–C) Calvariopsis yanayacuense sp. nov.; D–F) Calvariopsis bituberculata sp. nov. A, D) ventral portion; B, E) head and prosternum; C) mesoventrite, sphaerical cuticular structures are marked with an arrow; F) apical portion of ventrite 5." httpUri="https://zenodo.org/record/2783692/files/figure.png" pageId="39" pageNumber="40">Figs 3F</figureCitation>
,
<figureCitation id="6186A3FFF24E141171D4EB23573B297B" box="[758,798,359,381]" captionStart="FIGURE 20" captionStartId="26.[151,250,1241,1266]" captionTargetBox="[153,1435,361,1198]" captionTargetId="figure@26.[151,1436,361,1198]" captionTargetPageId="26" captionText="FIGURE 20. Calvariopsis bituberculata sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX; E) apex of abdomen. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783774/files/figure.png" pageId="39" pageNumber="40">20E</figureCitation>
)................................
<taxonomicName id="3EBDC4F9F24E1411778FEB23516C297A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1197,1353,359,380]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bituberculata" status="sp. nov.">
<emphasis id="CBC96368F24E1411778FEB23516C297A" box="[1197,1353,359,380]" italics="true" pageId="39" pageNumber="40">C. bituberculata</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB2351B9297A" bold="true" box="[1360,1436,359,380]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB2351B9297A" box="[1360,1436,359,380]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EBC151B9299F" blockId="39.[151,1437,153,762]" box="[151,1436,388,410]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBC151B9299F" box="[151,1436,388,410]" pageId="39" pageNumber="40">-. Last abdominal ventrite without tubercles................................................................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EBE651B829D2" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EBE651B929B1" blockId="39.[151,1437,153,762]" box="[151,1436,417,439]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBE651B929B1" box="[151,1436,417,439]" pageId="39" pageNumber="40">
10. Tergite VIII without lateral spines, tegmen without basal apodemes (
<figureCitation id="6186A3FFF24E14117051EBE557EB29B1" box="[883,974,417,439]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="39" pageNumber="40">Figs 25B</figureCitation>
,
<figureCitation id="6186A3FFF24E141170FAEBE5502429B1" box="[984,1025,417,439]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="39" pageNumber="40">26C</figureCitation>
)................................11
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EBFB51B829D2" blockId="39.[151,1437,153,762]" box="[151,1437,447,469]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBFB51B829D2" box="[151,1437,447,469]" pageId="39" pageNumber="40">
-. Tergite VIII with lateral spines, tegmen with paired basal apodemes (
<figureCitation id="6186A3FFF24E1411705AEBFB57F629D2" box="[888,979,447,469]" captionStart="FIGURE 20" captionStartId="26.[151,250,1241,1266]" captionTargetBox="[153,1435,361,1198]" captionTargetId="figure@26.[151,1436,361,1198]" captionTargetPageId="26" captionText="FIGURE 20. Calvariopsis bituberculata sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX; E) apex of abdomen. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783774/files/figure.png" pageId="39" pageNumber="40">Figs 20B</figureCitation>
,
<figureCitation id="6186A3FFF24E141170FFEBFB502229D2" box="[989,1031,447,468]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">21B</figureCitation>
,
<figureCitation id="6186A3FFF24E14117733EBFB501E29D2" box="[1041,1083,447,468]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">22B</figureCitation>
,
<figureCitation id="6186A3FFF24E14117767EBFB504B29D3" box="[1093,1134,447,469]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="39" pageNumber="40">23B</figureCitation>
).......................12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5EB9851B92A09" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5EB9851B929F7" blockId="39.[151,1437,153,762]" box="[151,1436,476,498]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EB9851B929F7" box="[151,1436,476,498]" pageId="39" pageNumber="40">
11. Tegmen V-shaped (
<figureCitation id="6186A3FFF24E141172B9EB9855D629F4" box="[411,499,476,498]" captionStart="FIGURE 25" captionStartId="32.[151,250,946,971]" captionTargetBox="[153,1435,195,907]" captionTargetId="figure@32.[151,1436,195,907]" captionTargetPageId="32" captionText="FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783792/files/figure.png" pageId="39" pageNumber="40">Fig. 25B</figureCitation>
), penis not curved in lateral view...............................
<taxonomicName id="3EBDC4F9F24E14117795EB98516F29F7" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1207,1354,476,497]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="saopaulense" status="sp. nov.">
<emphasis id="CBC96368F24E14117795EB98516F29F7" box="[1207,1354,476,497]" italics="true" pageId="39" pageNumber="40">C. saopaulense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EB9851B929F7" bold="true" box="[1360,1436,476,497]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EB9851B929F7" box="[1360,1436,476,497]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5EBBE51B92A09" blockId="39.[151,1437,153,762]" box="[151,1436,505,527]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5EBBE51B92A09" box="[151,1436,505,527]" pageId="39" pageNumber="40">
-. Tegmen subrectangular (
<figureCitation id="6186A3FFF24E141172EDEBBD56022A09" box="[463,551,505,527]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="39" pageNumber="40">Fig. 26C</figureCitation>
), penis curved in lateral view (
<figureCitation id="6186A3FFF24E1411706DEBBD57832A09" box="[847,934,505,527]" captionStart="FIGURE 26" captionStartId="33.[151,250,1439,1464]" captionTargetBox="[153,1435,505,1397]" captionTargetId="figure@33.[151,1436,505,1397]" captionTargetPageId="33" captionText="FIGURE 26. Calvariopsis wittmeri sp. nov., male. A) penis, lateral view; B) penis, dorsal view; C) tegmen; D) sternite IX; E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783798/files/figure.png" pageId="39" pageNumber="40">Fig. 26A</figureCitation>
).........................
<taxonomicName id="3EBDC4F9F24E141177FFEBBE516F2A09" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1245,1354,506,527]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="wittmeri" status="sp. nov.">
<emphasis id="CBC96368F24E141177FFEBBE516F2A09" box="[1245,1354,506,527]" italics="true" pageId="39" pageNumber="40">C. wittmeri</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672EBBE51B92A09" bold="true" box="[1360,1436,506,527]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672EBBE51B92A09" box="[1360,1436,506,527]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5E85351B92A4F" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5E85351B92A2A" blockId="39.[151,1437,153,762]" box="[151,1436,535,557]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E85351B92A2A" box="[151,1436,535,557]" pageId="39" pageNumber="40">
12. Tergite VIII with T-shaped emargination in central portion (
<figureCitation id="6186A3FFF24E1411703CE85357502A2B" box="[798,885,535,557]" captionStart="FIGURE 23" captionStartId="29.[151,250,1583,1608]" captionTargetBox="[153,1435,885,1572]" captionTargetId="figure@29.[153,1438,885,1572]" captionTargetPageId="29" captionText="FIGURE 23. Calvariopsis kawense sp. nov., male. A) penis; B) tegmen; C) sternite IX; D) tergite VIII; E) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/3403757/files/figure.png" pageId="39" pageNumber="40">Fig. 23D</figureCitation>
).............................
<taxonomicName id="3EBDC4F9F24E141177FEE853516C2A2A" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1244,1353,535,556]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kawense" status="sp. nov.">
<emphasis id="CBC96368F24E141177FEE853516C2A2A" box="[1244,1353,535,556]" italics="true" pageId="39" pageNumber="40">C. kawense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672E85351B92A2A" bold="true" box="[1360,1436,535,556]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672E85351B92A2A" box="[1360,1436,535,556]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5E87151B92A4F" blockId="39.[151,1437,153,762]" box="[151,1436,564,586]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E87151B92A4F" box="[151,1436,564,586]" pageId="39" pageNumber="40">
-. Tergite VIII with V-shaped emargination in central portion (
<figureCitation id="6186A3FFF24E14117002E87057592A4F" box="[800,892,564,586]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">Figs 21D</figureCitation>
,
<figureCitation id="6186A3FFF24E141170A4E87057952A4F" box="[902,944,564,585]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">22C</figureCitation>
,
<figureCitation id="6186A3FFF24E14117098E87057C12A4C" box="[954,996,564,586]" captionStart="FIGURE 24" captionStartId="30.[151,250,1766,1791]" captionTargetBox="[153,1435,971,1727]" captionTargetId="figure@30.[151,1436,971,1727]" captionTargetPageId="30" captionText="FIGURE 24. Calvariopsis nana sp. nov., male. A) penis; B) tergite VIII; C) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783786/files/figure.png" pageId="39" pageNumber="40">24B</figureCitation>
)...................................13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5E81651B92AB9" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5E81651B92A83" blockId="39.[151,1437,153,762]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E81651B92A83" pageId="39" pageNumber="40">
13. Trigonium of penis shorter, reaching 1/4 of length of penis, penis with subparallel sides (
<figureCitation id="6186A3FFF24E1411776EE81550822A6E" box="[1100,1191,593,616]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">Fig. 21A</figureCitation>
), emargination of central portion of tergite VIII deep (
<figureCitation id="6186A3FFF24E141172D1E82B566F2A83" box="[499,586,623,645]" captionStart="FIGURE 21" captionStartId="27.[151,250,1432,1457]" captionTargetBox="[153,1435,616,1388]" captionTargetId="figure@27.[151,1436,616,1388]" captionTargetPageId="27" captionText="FIGURE 21. Calvariopsis brasiliense (Pic), male, A–D), F)—specimen from NMPC; E)—holotype. A) penis; B) tegmen; C) sternite IX; D–E) tergite VIII; F) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783778/files/figure.png" pageId="39" pageNumber="40">Fig. 21D</figureCitation>
)......................................................
<taxonomicName id="3EBDC4F9F24E141177C0E82B51B92A83" authority="(Pic)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1916" box="[1250,1436,623,645]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="brasiliense">
<emphasis id="CBC96368F24E141177C0E82B51422A82" box="[1250,1383,623,644]" italics="true" pageId="39" pageNumber="40">C. brasiliense</emphasis>
(Pic)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5E8C951B92AB9" blockId="39.[151,1437,153,762]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E8C951B92AB9" pageId="39" pageNumber="40">
-. Trigonium of penis relatively long, reaching 1/3 of length of penis, penis with curved sides (
<figureCitation id="6186A3FFF24E1411777DE8C8509E2AA7" box="[1119,1211,652,674]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">Figs 22A</figureCitation>
,
<figureCitation id="6186A3FFF24E141177E4E8C850D72AA4" box="[1222,1266,652,674]" captionStart="FIGURE 24" captionStartId="30.[151,250,1766,1791]" captionTargetBox="[153,1435,971,1727]" captionTargetId="figure@30.[151,1436,971,1727]" captionTargetPageId="30" captionText="FIGURE 24. Calvariopsis nana sp. nov., male. A) penis; B) tergite VIII; C) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783786/files/figure.png" pageId="39" pageNumber="40">24A</figureCitation>
), emargination of central portion of tergite VIII shallow (
<figureCitation id="6186A3FFF24E1411717AE8ED56972AB9" box="[600,690,681,703]" captionStart="FIGURE 22" captionStartId="28.[151,250,1786,1811]" captionTargetBox="[153,1435,899,1737]" captionTargetId="figure@28.[153,1438,921,1737]" captionTargetPageId="28" captionText="FIGURE 22. Calvariopsis guyanense sp. nov., male. A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783782/files/figure.png" pageId="39" pageNumber="40">Figs 22C</figureCitation>
,
<figureCitation id="6186A3FFF24E1411719EE8EE56C32AC6" box="[700,742,682,704]" captionStart="FIGURE 24" captionStartId="30.[151,250,1766,1791]" captionTargetBox="[153,1435,971,1727]" captionTargetId="figure@30.[151,1436,971,1727]" captionTargetPageId="30" captionText="FIGURE 24. Calvariopsis nana sp. nov., male. A) penis; B) tergite VIII; C) tergite IX. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/2783786/files/figure.png" pageId="39" pageNumber="40">24B</figureCitation>
)....................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4249A7DFF24E141173B5E88351B92AFF" pageId="39" pageNumber="40">
<paragraph id="F902BF7AF24E141173B5E88351B92ADA" blockId="39.[151,1437,153,762]" box="[151,1436,711,733]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E88351B92ADA" box="[151,1436,711,733]" pageId="39" pageNumber="40">
14. TL
<quantity id="3E45129FF24E14117226E88355752ADA" box="[260,336,711,732]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="39" pageNumber="40" unit="mm" value="1.7">1.7 mm</quantity>
, known from
<collectingCountry id="81AAFFEAF24E141172F4E88356342ADB" box="[470,529,711,733]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
.............................................................
<taxonomicName id="3EBDC4F9F24E141177DCE883516F2ADA" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1278,1354,711,732]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="nana" status="sp. nov.">
<emphasis id="CBC96368F24E141177DCE883516F2ADA" box="[1278,1354,711,732]" italics="true" pageId="39" pageNumber="40">C. nana</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672E88351B92ADA" bold="true" box="[1360,1436,711,732]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672E88351B92ADA" box="[1360,1436,711,732]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F902BF7AF24E141173B5E8A151B92AFF" blockId="39.[151,1437,153,762]" box="[151,1436,740,762]" pageId="39" pageNumber="40">
<keyLead id="424C1C4FF24E141173B5E8A151B92AFF" box="[151,1436,740,762]" pageId="39" pageNumber="40">
-. TL 2.0 mm, known from
<collectingCountry id="81AAFFEAF24E141172F4E8A0563E2AFC" box="[470,539,740,762]" name="Guyana" pageId="39" pageNumber="40">Guiana</collectingCountry>
........................................................
<taxonomicName id="3EBDC4F9F24E141177EAE8A0516C2AFC" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[1224,1353,740,762]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="guyanense" status="sp. nov.">
<emphasis id="CBC96368F24E141177EAE8A0516C2AFC" box="[1224,1353,740,762]" italics="true" pageId="39" pageNumber="40">C. guyanense</emphasis>
</taxonomicName>
<emphasis id="CBC96368F24E14117672E8A051B92AFF" bold="true" box="[1360,1436,740,761]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="D0FADE13F24E14117672E8A051B92AFF" box="[1360,1436,740,761]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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