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<mods:title id="CFBB539CEA7ACA3D5C4615A038BF8655">Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora)</mods:title>
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<mods:namePart id="561B0D56370550C24F17A6CEDDD55E8A">Pall-Gergely, Barna</mods:namePart>
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<mods:namePart id="10A1DF52F3EAD5A2B3680B593F729683">Hunyadi, Andras</mods:namePart>
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<mods:namePart id="28A89800F9B581A007FB61C2E3A0C816">Ablett, Jonathan</mods:namePart>
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<mods:namePart id="D1632713EE763A4CB8609700E83D546A">Lương, Hao Văn</mods:namePart>
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<mods:namePart id="3108D3C8DF42CA582744776972A077EF">Fred Naggs,</mods:namePart>
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<treatment id="9C47E4E439A1BB906BE5DD87518399A9" ID-GBIF-Taxon="152057815" LSID="urn:lsid:plazi:treatment:9C47E4E439A1BB906BE5DD87518399A9" httpUri="http://treatment.plazi.org/id/9C47E4E439A1BB906BE5DD87518399A9" lastPageId="67" lastPageNumber="68" pageId="61" pageNumber="62">
<subSubSection id="FEEFC32408D173DB82A51E1F78056714" pageId="61" pageNumber="62" type="multiple">
<paragraph id="0751DCC787D778D282D6EAEFE9381C15" pageId="61" pageNumber="62">Taxon classification Animalia Stylommatophora Plectopylidae</paragraph>
</subSubSection>
<subSubSection id="76C80B31F256B63937CBED9930FDC324" pageId="61" pageNumber="62" type="nomenclature">
<paragraph id="035060E16621ED85E7F54497A9578D20" pageId="61" pageNumber="62">
<taxonomicName id="3F223F6BFD76FA918058B46D1196B869" ID-CoL="4X762" authority="Gude, 1908" authorityName="Gude" authorityYear="1908" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus mansuyi" order="Stylommatophora" pageId="61" pageNumber="62" phylum="Mollusca" rank="species" species="mansuyi">Sicradiscus mansuyi (Gude, 1908)</taxonomicName>
Figures 2A, 9H, 11
<normalizedToken id="416602ACB5C14B5144CD9D050A44C958" originalValue="AB">A-B</normalizedToken>
, 27, 31A, 36
<normalizedToken id="6584892369EB34C49A3DF07E1F57FE63" originalValue="JL">J-L</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="E86DCFF2DD8C7B9D32D507CFD873781F" pageId="62" pageNumber="63" type="reference_group">
<paragraph id="B70EE94D780028D8F0B8BF6E3DD4258E" pageId="62" pageNumber="63">
<taxonomicName id="A3D311063487C9BDC861766A0D1FE317" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis Mansuyi" order="Stylommatophora" pageId="62" pageNumber="63" phylum="Mollusca" rank="species" species="Mansuyi">
<pageBreakToken id="EBDE102F82303DC90DDFFBB8A7DB5580" pageId="62" pageNumber="63" start="start">Plectopylis</pageBreakToken>
Mansuyi
</taxonomicName>
Gude 1908, Journal de Conchyliologie, 55: 347, 348-351., Figs 2
<normalizedToken id="58DCF81986181AF5C6AAD44AAAF08E5F" originalValue="ae">a-e</normalizedToken>
, Plate 7., Figs 1-3. [&quot;Ha-Lang, Tonkin&quot;]
</paragraph>
<paragraph id="29D497A230AA8ACECA65E5E0919DA237" pageId="62" pageNumber="63">
<taxonomicName id="F0095EFC3E5BE8C79522950725892238" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus mansuyi" order="Stylommatophora" pageId="62" pageNumber="63" phylum="Mollusca" rank="species" species="mansuyi">Sicradiscus mansuyi</taxonomicName>
, -
<normalizedToken id="705C6932844C951661AF086AF499A363" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi 2013, Archiv
<normalizedToken id="F64064AB6AB6387EA9041118C3F6886F" originalValue="für">fuer</normalizedToken>
Molluskenkunde, 142 (1): 50.
</paragraph>
</subSubSection>
<subSubSection id="B4B9F5AAACECABB39029A5CD4B45E3CA" pageId="62" pageNumber="63" type="types examined">
<paragraph id="7BDDC8E4249DE6A9D46740BEC4A41A21" pageId="62" pageNumber="63">Types examined.</paragraph>
<paragraph id="E7B1C3B0C687F5DC4BF855A14F508C52" pageId="62" pageNumber="63">Tonkin, Ha-Lang, leg. Mansuy, NHMUK 1907.2.20.19 (syntype, Figure 2A).</paragraph>
</subSubSection>
<subSubSection id="670613C789B09A8A7F00D67487483D81" pageId="62" pageNumber="63" type="museum material examined">
<paragraph id="CD3C7D94BE6E9184D302F5A256FE0CF7" pageId="62" pageNumber="63">Museum material examined.</paragraph>
<paragraph id="9F2F24459BC85D36CB09F4DF3191E998" pageId="62" pageNumber="63">Ha-Lang, coll. Mansuy, MNHN-IM-2012-2365/6; Ha-Lang, leg. Mansuy, MNHN-IM-2012-2384/7; HaLang, Tonkin, coll. Steenberg, ZMUC-GAS-1808/2.</paragraph>
</subSubSection>
<subSubSection id="B3D8AEAD27CCE5C6DDFD9753927C09E6" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63" type="new material examined">
<paragraph id="7DA1A5F11FD99C47D838B41407BC759F" pageId="62" pageNumber="63">New material examined.</paragraph>
<paragraph id="916E2B1CC5DB0DFBFD42A5FF4B646DEE" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63">
20081116C Cao
<normalizedToken id="A8AE0D698DD1E89091835F3F112C658E" originalValue="Bằng">Bằng</normalizedToken>
Province,
<normalizedToken id="3B49B8382C3AC54E1B14766F6D550642" originalValue="Trùng">Trung</normalizedToken>
<normalizedToken id="27F76EAE048262DD48EFF8876C7F3D28" originalValue="Khánh">Khanh</normalizedToken>
District,
<normalizedToken id="1184A60B1F8D1BA4F9F4E9E1475F0E08" originalValue="Cảnh">Cảnh</normalizedToken>
<normalizedToken id="4EF3AFED1AD5657ECC3B95CF8771A96F" originalValue="Tiên">Tien</normalizedToken>
Commune,
<normalizedToken id="348B01DDD7AA170140D1D0714290C6DB" originalValue="Pắc">Pắc</normalizedToken>
<normalizedToken id="EF8DB0A7B462C9D394B592D9AB042A86" originalValue="Rảo">Rảo</normalizedToken>
Village, ca 545 m,
<geoCoordinate id="FC27421717285EF75E082DCCE084BC1D" direction="north" orientation="latitude" precision="1" value="22.815683">22°48.941'N</geoCoordinate>
,
<geoCoordinate id="0033800AF315D846B4F01E3028C00648" direction="east" orientation="longitude" precision="1" value="106.50915">106°30.549'E</geoCoordinate>
, leg. Ohara, K., 16.11.2008., OK/66, PGB/5; 2011/81 Cao
<normalizedToken id="09ABBC8CB480C71E04002E86F8FD63B4" originalValue="Bằng">Bằng</normalizedToken>
Province,
<normalizedToken id="E9270B493CCD2B0D6C419534F28A10A9" originalValue="Đèo">Đeo</normalizedToken>
<normalizedToken id="E2AF2CEF48622C66831371FF96FE3522" originalValue="Mã">Ma</normalizedToken>
<normalizedToken id="B1E7488F3BAFB52E15DE10D8B2CD706C" originalValue="Phục">Phục</normalizedToken>
(pass) 500 m towards
<normalizedToken id="6DA9CDB8BA6CB96F07DA515260D92009" originalValue="Quảng">Quảng</normalizedToken>
<normalizedToken id="4CFEC7D42583F901033321BFBE38BBEF" originalValue="Uyên">Uyen</normalizedToken>
, left side of the road, rock cavern, 610 m,
<geoCoordinate id="61479B47BEE6EB6AB12FDE868F0E4C00" direction="north" orientation="latitude" precision="1" value="22.733017">22°43.981'N</geoCoordinate>
,
<geoCoordinate id="B03C8ADD014B7107E81B32BFE44B4DB3" direction="east" orientation="longitude" precision="1" value="106.33888">106°20.333'E</geoCoordinate>
, leg. Hunyadi, A., 14.11.2011., HA/10; 2012/43 Cao
<normalizedToken id="D1BEF047A5EE197712733FA4556AF957" originalValue="Bằng">Bằng</normalizedToken>
Province,
<normalizedToken id="2DF4DC73E989451397153A7F0604811E" originalValue="Pắc">Pắc</normalizedToken>
<normalizedToken id="F8EBE486F6D8BBD95252F5E26DCF46C2" originalValue="Rảo">Rảo</normalizedToken>
,
<normalizedToken id="88F3A01EBCBA03AD751927E3185DCCE7" originalValue="Cảnh">Cảnh</normalizedToken>
<normalizedToken id="86D715C54321A48BEC4A90DDFA10E47F" originalValue="Tiên">Tien</normalizedToken>
Commune cross, 300 m towards
<normalizedToken id="900CFA39A1AA9DD15E15E988FB93B376" originalValue="Trùng">Trung</normalizedToken>
<pageBreakToken id="6643F5DD77D2DCBE8B17F54A850F8378" pageId="63" pageNumber="64" start="start">
<normalizedToken id="FC939C92213601998D98C4639ACC0FFE" originalValue="Khánh">Khanh</normalizedToken>
</pageBreakToken>
, right side of the road, 530 m,
<geoCoordinate id="C55DB4DD2A36B12BCA6957A684D25DEE" direction="north" orientation="latitude" precision="1" value="22.823084">22°49.385'N</geoCoordinate>
,
<geoCoordinate id="878F5A9EFE7CD59A6695BDF6F900050F" direction="east" orientation="longitude" precision="1" value="106.51237">106°30.742'E</geoCoordinate>
, leg. Hunyadi, A., 28.05.2012., HA/9+5 jb; 2012/44 Cao
<normalizedToken id="1D895C332D74633019AFDF73E63E7D6D" originalValue="Bằng">Bằng</normalizedToken>
Province, southern edge of
<normalizedToken id="8970C594FB5D0599CBEFA35857E157A5" originalValue="Pắc">Pắc</normalizedToken>
<normalizedToken id="4B2EB2E009A7F727ECE6CCB0546BFA20" originalValue="Rảo">Rảo</normalizedToken>
,
<normalizedToken id="08BC0D64E026872B527E7CD45414EF85" originalValue="Trùng">Trung</normalizedToken>
<normalizedToken id="768A0CD5B32BC2B71D9D2EE6F9C576D0" originalValue="Khánh">Khanh</normalizedToken>
3 km towards
<normalizedToken id="AC971168CFCEE5715BFB4E369DC0EB19" originalValue="Quảng">Quảng</normalizedToken>
<normalizedToken id="323D75BFF630EB8003226AF510BFDAE2" originalValue="Uyên">Uyen</normalizedToken>
, left side of the road, 570 m,
<geoCoordinate id="1738F1E76E67C22AD17DF8C7EB20398E" direction="north" orientation="latitude" precision="1" value="22.816017">22°48.961'N</geoCoordinate>
,
<geoCoordinate id="B1CD47DB7C4500372A2E5C49B7CC79D6" direction="east" orientation="longitude" precision="1" value="106.50888">106°30.533'E</geoCoordinate>
, leg. Hunyadi, A., 28.05.2012., HA/226; 2012/47
<normalizedToken id="D5B6F5AC06B0B188F9B04A459C7CDC00" originalValue="Hà">Ha</normalizedToken>
Giang Province,
<normalizedToken id="A44786C1A7154FBC23BC4E9186BD6919" originalValue="Hà">Ha</normalizedToken>
Giang 105.5 km towards
<normalizedToken id="9830BFBEC4AA9D03A782F5469BC555B2" originalValue="Ðồng">Ðồng</normalizedToken>
<normalizedToken id="D1492BC918510735DF3266D78543A5D3" originalValue="Văn">Văn</normalizedToken>
,
<normalizedToken id="6D4A268411892D65ACFD5618F2ACD63F" originalValue="Vân">Van</normalizedToken>
<normalizedToken id="54D81DEECD63E9462654ABECD816E241" originalValue="Chải">Chải</normalizedToken>
Commune, left side of the road 4C,
<geoCoordinate id="E31F8AC917E1AA266DE709DD1DC854BA" direction="north" orientation="latitude" precision="1" value="23.1514">23°09.084'N</geoCoordinate>
,
<geoCoordinate id="D95697C513F68401914F6673D24C2D73" direction="east" orientation="longitude" precision="1" value="105.179565">105°10.774'E</geoCoordinate>
, leg. Hunyadi, A., 31.05.2012., HA/4; Vn11-141
<normalizedToken id="9239571CFE73B49CCB5BBC4B487A9409" originalValue="Hà">Ha</normalizedToken>
<pageBreakToken id="4118E71B3D5CD26C4AAEC8A02BF95D75" pageId="64" pageNumber="65" start="start">Giang</pageBreakToken>
Province, km 105.5 on road 4c, between
<normalizedToken id="06C392C8EAB297CCBF8B0195B070ECA1" originalValue="Yên">Yen</normalizedToken>
Minh and
<normalizedToken id="93F72EB0E4510E77B8929B3F7660B079" originalValue="Đồng">Đồng</normalizedToken>
<normalizedToken id="0D5DA0CAE2828F29985C0CD04523657A" originalValue="Văn">Văn</normalizedToken>
(NE of
<normalizedToken id="880569521F38A325E75BFA38861C4FBE" originalValue="Hà">Ha</normalizedToken>
Giang town),
<geoCoordinate id="F09E291B9A95AE2593E1384CFEC3483C" direction="north" orientation="latitude" precision="1" value="23.149933">23°08.996'N</geoCoordinate>
,
<geoCoordinate id="CB93C291C57D9F008286E908ECFF6C7C" direction="east" orientation="longitude" precision="1" value="105.1722">105°10.332'E</geoCoordinate>
, leg. Hemmen, Ch., 21.03.2011., HE/6; Vn11-143
<normalizedToken id="DCC8D89FDED5BB38A3F6DE71CFC439D1" originalValue="Hà">Ha</normalizedToken>
Giang Province, km 120 on road 4c, between
<normalizedToken id="723645DAC6D42EAC1C3CF97E4DA71586" originalValue="Yên">Yen</normalizedToken>
Minh and
<normalizedToken id="A33D937F685B1148C6BBF2B866D8C269" originalValue="Đồng">Đồng</normalizedToken>
<normalizedToken id="577E1EB9E3D1CD654F95EC7227DC86B7" originalValue="Văn">Văn</normalizedToken>
(NE of
<normalizedToken id="A25A768DCA58AF194EC805EE34FD07FA" originalValue="Hà">Ha</normalizedToken>
Giang town), no GPS-data, leg. Hemmen, Ch. &amp; J., 22.03.2011., HE/3; Vn10-60 Cao
<normalizedToken id="7DB577D9A247CD735CA788AA4A23D029" originalValue="Bằng">Bằng</normalizedToken>
Province, ca. 6.5 km from
<normalizedToken id="F567AB2EB390FED179853DFE8D11AC6A" originalValue="Quảng">Quảng</normalizedToken>
<normalizedToken id="586601730122C541FD9EAA0D371D725F" originalValue="Uyênto">Uyento</normalizedToken>
<normalizedToken id="44A34C6694335FFC8627290F5B5FF909" originalValue="Mã">Ma</normalizedToken>
<normalizedToken id="A2980FEC7E8C6EF7949D2BEACE4A5244" originalValue="Phục">Phục</normalizedToken>
(left off road),
<geoCoordinate id="8D4F78823F6650EE9E6DA90566DEA985" direction="north" orientation="latitude" precision="1" value="22.688217">22°41.293'N</geoCoordinate>
,
<geoCoordinate id="0A3FEECB134AD11B1D0C71198E690AEB" direction="east" orientation="longitude" precision="1" value="106.390366">106°23.422'E</geoCoordinate>
, leg. Hemmen, Ch. &amp; J., 24.03.2010., HE/2; 20050327A China, Guangxi (广西), Daxin Xian (大新), Xialei Zhen (下雷鎮), Detianpubu (德天瀑布) (Detian waterfalls), leg. Ohara, K. &amp; Moriya Shigeki, 27.03.2005., PGB/1 (with glossy dorsal surface and without denticles posterior to the palatal plicae).
</paragraph>
</subSubSection>
<subSubSection id="C1231406AEEF3A313BD010A187607FA0" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="9F1267728A33E05C54817CC0A35472F6" pageId="65" pageNumber="66">
<pageBreakToken id="9330026079C149E038E116B8CE753EE1" pageId="65" pageNumber="66" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph id="E967C2A7692CEBE0A273B6971C001D11" pageId="65" pageNumber="66">
A very small species with reticulated dorsal and glossy ventral surface, elevated spire, elevated, sharp callus and well-developed apertural fold connected to the callus (Figure 9H). Parietal wall with two lamellae, the anterior one separated from both the lower and upper plicae; middle palatal plicae short, connected with a ridge and sometimes ornamented with small denticles posteriorly (Figures 11
<normalizedToken id="E58E893238C0D581D540111F73C891CC" originalValue="AB">A-B</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection id="B1651870D9FFCDB65FE00D287822EDF2" pageId="65" pageNumber="66" type="measurements">
<paragraph id="BB2E6A26734FC56691C2A3B6015DC38D" pageId="65" pageNumber="66">Measurements</paragraph>
<paragraph id="21619F44C62FF421F7AD891DC1621AEF" pageId="65" pageNumber="66">(in mm). D = 6.7-7, H = 3.4-3.9 (n=4, 20081116C).</paragraph>
</subSubSection>
<subSubSection id="363B2EEB6C29AFE2C25DA92F8A579136" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="differential diagnosis">
<paragraph id="E8424F5CE49F4719A0260B73FD9A2122" pageId="65" pageNumber="66">Differential diagnosis.</paragraph>
<paragraph id="F3AB2332D6234B6DF893763000F63643" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
All other similar congeners inhabit China.
<taxonomicName id="7575018598CB50958C3332D3274522BB" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus feheri" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="feheri">
<pageBreakToken id="3A81D4B30DCD040A88CB02C90D976791" pageId="66" pageNumber="67" start="start">Sicradiscus</pageBreakToken>
feheri
</taxonomicName>
<normalizedToken id="ACBD43F93F71FB60C64864F5F2A57C23" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, 2013 is larger, flatter with a wider umbilicus and a shinier dorsal surface, has a longer, horizontal palatal plicae without additional posterior denticles, and has a more elevated and longer apertural fold.
<taxonomicName id="EFCA48D9BDC36CE25102BFE39F2EA0D2" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus transitus" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="transitus">Sicradiscus transitus</taxonomicName>
<normalizedToken id="9171B65F19C570E17C1FF3FA385DB317" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, 2013 has a lower spire and a wider umbilicus with slightly shouldered whorls, sometimes strong radial lines on the ventral surface, and a more elevated callus. Moreover, the anterior lamella of
<taxonomicName id="7ABBEF5FF8A874CA6393550A070B6EFC" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus transitus" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="transitus">Sicradiscus transitus</taxonomicName>
is in contact with both the upper and the lower plicae, which are free from the lamella in
<taxonomicName id="9CE319915D789D4E02A35C90CCB1B9B2" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus mansuyi" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="mansuyi">Sicradiscus mansuyi</taxonomicName>
.
<taxonomicName id="25528753AC34173023F890456B57C325" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus invius" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="invius">Sicradiscus invius</taxonomicName>
is flatter (has shallower umbilicus) with only the protoconch elevated from the dorsal surface; it has weaker dorsal sculpture resulting in a glossy surface (
<taxonomicName id="0ACC86613097536784682E2B91E1946D" class="Insecta" family="Hybotidae" genus="Euthyneura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euthyneura mansuyi" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mansuyi">mansuyi</taxonomicName>
is densely reticulated), and lacks the additional small denticles posterior to the palatal plicae, which are usually present in
<taxonomicName id="62A7610770EC744F5B91AD96CEDC44D7" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus mansuyi" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="mansuyi">Sicradiscus mansuyi</taxonomicName>
.
<taxonomicName id="E682EB3572AB57F7447E005FF18B208A" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus anceyi" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="anceyi">Gudeodiscus anceyi</taxonomicName>
is larger and has a ribbed shell with spiral lines on the whole shell. Species possessing a glossy ventral surface (
<taxonomicName id="4E4F37DBBF0CAF8B4F13DAA6BD9FB0ED" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus cyrtochilus" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="cyrtochilus">Gudeodiscus cyrtochilus</taxonomicName>
,
<taxonomicName id="49763828626726D134A16081230E3746" class="Gastropoda" family="Plectopylidae" genus="Gudeodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gudeodiscus fischeri" order="Stylommatophora" pageId="66" pageNumber="67" phylum="Mollusca" rank="species" species="fischeri">Gudeodiscus fischeri</taxonomicName>
) are also larger and have weaker or no apertural fold.
</paragraph>
</subSubSection>
<subSubSection id="29F9D35E1BBEDC7542E1A8BEF9948D86" pageId="66" pageNumber="67" type="intraspecific diversity">
<paragraph id="7C32AAA9456D2FA7F141938747C92FAD" pageId="66" pageNumber="67">Intraspecific diversity.</paragraph>
<paragraph id="4FC33C2B7AD8B73C2B3AC2F7359F48DA" pageId="66" pageNumber="67">Low; shell characters stable. The species is easily recognisable and can be separated from other plectopylid species without difficulty.</paragraph>
</subSubSection>
<subSubSection id="4B9C9BF4CC27958EA99D8E4671953BED" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="description of the genitalia">
<paragraph id="0A0914921C6365E9110CEF2CC815E7CC" pageId="66" pageNumber="67">Description of the genitalia.</paragraph>
<paragraph id="3569F82BB320981B32E972C0A2EE1560" pageId="66" pageNumber="67">
Two specimens were anatomically examined (Cao
<normalizedToken id="A640E3D6E134826FB46297B83235A27C" originalValue="Bằng">Bằng</normalizedToken>
Province, southern edge of
<normalizedToken id="C24583B7851D31250DA9C458E7B69BC0" originalValue="Pắc">Pắc</normalizedToken>
<normalizedToken id="F0F81FD4A1EC1E146C736754EB10A3B6" originalValue="Rảo">Rảo</normalizedToken>
,
<normalizedToken id="CC13DE42F2F4B988EF46370D8ED57A44" originalValue="Trùng">Trung</normalizedToken>
<normalizedToken id="572A85F22D35065C670243AA2F1424A1" originalValue="Khánh">Khanh</normalizedToken>
3 km towards
<normalizedToken id="31F51B1868261E67C864C64D5301C72C" originalValue="Quảng">Quảng</normalizedToken>
<normalizedToken id="3654B0757963E4F44BC66BA1ABFBDBD0" originalValue="Uyên">Uyen</normalizedToken>
, left side of the road, 570 m,
<geoCoordinate id="04A02C890E909C86365E1F9C33614969" direction="north" orientation="latitude" precision="1" value="22.816017">22°48.961'N</geoCoordinate>
,
<geoCoordinate id="C9B321B25B3E781076512DC6EB43C55D" direction="east" orientation="longitude" precision="1" value="106.50888">106°30.533'E</geoCoordinate>
, leg. Hunyadi, A., 28.05.2012. (Figures 27, 31A).
</paragraph>
<paragraph id="26027A0E42BD52AE2F3C0EAB76490A41" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
Penis with a shorter, slimmer proximal section and a thinner, somewhat longer distal portion; internally with parallel folds which are more elevated in the thinner distal portion, forming pocket-like structures (similar to that of
<taxonomicName id="808539DDD3BF5867A56FB277D773DD58" class="Gastropoda" family="Plectopylidae" genus="Sicradiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicradiscus transitus" order="Stylommatophora" pageId="67" pageNumber="68" phylum="Mollusca" rank="species" species="transitus">
<pageBreakToken id="A70426A6D6973D0F0DA3C76EE4DB465A" pageId="67" pageNumber="68" start="start">Sicradiscus</pageBreakToken>
transitus
</taxonomicName>
, see
<bibRefCitation id="4121E18595C30D432FAAAB54AFFED840" author="Pall-Gergely, B" journalOrPublisher="Genus" pageId="83" pageNumber="84" pagination="527 - 564" title="Additional information on the distribution, anatomy and systematics of living and fossil Chinese Plectopylidae (Gastropoda: Pulmonata)." volume="25" year="2014">
<normalizedToken id="D6FF4A64E85C9459300896B937131203" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
and Asami 2014
</bibRefCitation>
); these
<normalizedToken id="D3F4F0C55EF0139B55B5594106056553" originalValue="“pockets”">&quot;pockets&quot;</normalizedToken>
did not contain granules; epiphallus approximately as long as the penis but much slimmer; internally penis and epiphallus wall with longitudinal, parallel folds; retractor muscle short, inserts on the penis-epiphallus transition; penial caecum absent. Vagina long, with distal vaginal bulb; vaginal bulb and other parts of the vagina with approximately 8, more or less parallel, serrulate folds (Figure 31A); vas deferens long, thicker distally and more slender proximally; gametolytic sac and diverticulum are of equal length, in parallel.
</paragraph>
</subSubSection>
<subSubSection id="4374488874D7174A70EDD710D2DE9153" pageId="67" pageNumber="68" type="radula">
<paragraph id="3294D94D40B367B404FB17F8CEFE2204" pageId="67" pageNumber="68">Radula.</paragraph>
<paragraph id="06052E72665C5BDD779D83E46C356294" pageId="67" pageNumber="68">
See Table 6 and Figures 36
<normalizedToken id="198C4CBDBBFB412727581E0374A0378B" originalValue="JL">J-L</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="E586A4B6BC8D160F54FC0948C255CD41" pageId="67" pageNumber="68" type="distribution">
<paragraph id="343357FF7AD3D400B70919FF5C16552E" pageId="67" pageNumber="68">Distribution.</paragraph>
<paragraph id="ED61AB326998296E2709E76F3D810FE7" pageId="67" pageNumber="68">
This species was described from
<normalizedToken id="DF546F422CE39FA1840942C8CFA67B6D" originalValue="Hạ">Hạ</normalizedToken>
Lang (eastern part of Cao
<normalizedToken id="5B889D83F30A11C6A1261E80B2223CFB" originalValue="Bằng">Bằng</normalizedToken>
Province, see Figure 39). We have seen newly collected material from northern
<normalizedToken id="A0534020BA3F9261184C30CA9E2C1456" originalValue="Hà">Ha</normalizedToken>
Giang and Cao
<normalizedToken id="77B85238A51E1554967877C696265035" originalValue="Bằng">Bằng</normalizedToken>
provinces. The first occurrence of the species from China is reported. This locality is situated very close to the Vietnamese border.
</paragraph>
</subSubSection>
</treatment>
</document>