treatments-xml/data/9C/42/87/9C4287CFFFB9FF92CD3FFDD10CC1F84D.xml
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<mods:title id="17A4595D643135A19B697C12C4E37C97">Nine new species of Scorpiops Peters, 1861 (Scorpiones: Scorpiopidae) from China, India, Nepal, and Pakistan</mods:title>
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<taxonomicName id="D3EB4D5AFFB9FF92CD3FFDD10B49FD47" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[294,517,636,662]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="grosseri" status="sp. nov.">Scorpiops grosseri</taxonomicName>
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(
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,
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,
<tableCitation id="59690362FFB9FF92CE18FD330B02FD67" box="[513,590,670,694]" captionStart="Table 1" captionStartId="3.[113,169,985,1006]" captionText="Table 1. Comparative measurements of adults of Scorpiops furai sp. n., S. grosseri sp. n., S. harmsi sp. n. and S. hofereki sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="7" pageNumber="6">Table 1</tableCitation>
)
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), leg.
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ETYMOLOGY. The species epithet is a patronym honoring a Czech entomologist Walter Grosser, the collector of
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of the new species.
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DIAGNOSIS (
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). Total length
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long. Base color uniformly reddish brown to black, telson and legs yellow to yellowish brown. Pectinal teeth number
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female, fulcra absent; a pecten forms one compact unit with an incomplete furrow between places for marginal and middle lamellae. Patella of pedipalp with 17 (5
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, 2
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, 2
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, 4
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) external and 7 ventral trichobothria. Chela of pedipalp with 4 ventral trichobothria located on ventral surface. Chelal trichobothrium
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Eb
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is located in proximal half of external surface between trichobothria
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and
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. Fingers of pedipalps flexed in female. Chela length to width ratio
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female. Pedipalp movable finger with 3436 IAD, which form a second row, parallel to MD (
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number); there are also 4 ID and 8 OD present. Tarsomere II of legs with 46 stout median ventral spines in a row and two other parallel spines. Telson bulbous and finely granulate, length to depth ratio
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female; annular ring present.
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DESCRIPTION (
<specimenCount id="02EDFD50FFB9FF92CD13F97C0857F938" box="[266,283,1745,1769]" pageId="7" pageNumber="6" type="female"></specimenCount>
). Total length of female
<typeStatus id="CB50887BFFB9FF92CE03F97C0B3BF938" box="[538,631,1745,1769]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
<quantity id="D3139B3CFFB9FF92CE65F97C0B8BF939" box="[636,711,1745,1769]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="7" pageNumber="6" unit="mm" value="59.0">59 mm</quantity>
, male unknown. The habitus is shown in
<figureCitation id="8CD02A5CFFB9FF92CDE8F95C0B38F8D8" box="[497,628,1777,1801]" captionStart="Figures 2328" captionStartId="8.[115,192,1986,2008]" captionTargetBox="[120,1467,234,1954]" captionTargetId="figure-51@8.[115,1476,234,1973]" captionTargetPageId="8" captionText="Figures 2328. Scorpiops grosseri sp. n., female holotype. Dorsal (23) and ventral (24) views. Telson lateral (25). Metasoma and telson lateral (26), ventral (27), and dorsal (28) views. Scale bars: 10 mm (2324, 2628)." figureDoi="http://doi.org/10.5281/zenodo.5741549" httpUri="https://zenodo.org/record/5741549/files/figure.png" pageId="7" pageNumber="6">Figs. 2324</figureCitation>
. For position and distribution of trichobothria of pedipalps see
<figureCitation id="8CD02A5CFFB9FF92CE9DF8BC09F0F898" captionStart="Figures 2939" captionStartId="9.[115,192,1957,1979]" captionTargetBox="[125,1474,247,1900]" captionTargetId="figure-71@9.[113,1474,238,1918]" captionTargetPageId="9" captionText="Figures 2939. Scorpiops grosseri sp. n., female holotype, pedipalp segments. Chela dorsal (29), external (30) and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter dorsal (35), internal (36), and ventral (37) views. Movable finger dentition under white light (38) and UV fluorescence (39). Trichobothrial pattern is indicated by white circles." figureDoi="http://doi.org/10.5281/zenodo.5741551" httpUri="https://zenodo.org/record/5741551/files/figure.png" pageId="7" pageNumber="6">Figs. 2934 and 36</figureCitation>
. Fingers of pedipalps are flexed in the female (
<figureCitation id="8CD02A5CFFB9FF92CEB1F89C0BBBF898" box="[680,759,1841,1865]" captionStart="Figures 2939" captionStartId="9.[115,192,1957,1979]" captionTargetBox="[125,1474,247,1900]" captionTargetId="figure-71@9.[113,1474,238,1918]" captionTargetPageId="9" captionText="Figures 2939. Scorpiops grosseri sp. n., female holotype, pedipalp segments. Chela dorsal (29), external (30) and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter dorsal (35), internal (36), and ventral (37) views. Movable finger dentition under white light (38) and UV fluorescence (39). Trichobothrial pattern is indicated by white circles." figureDoi="http://doi.org/10.5281/zenodo.5741551" httpUri="https://zenodo.org/record/5741551/files/figure.png" pageId="7" pageNumber="6">Fig. 30</figureCitation>
).
<emphasis id="269FEACBFFB9FF92CC68F8FC09AEF8B8" bold="true" box="[113,226,1873,1897]" pageId="7" pageNumber="6">Coloration</emphasis>
(
<figureCitation id="8CD02A5CFFB9FF92CCF4F8FC082EF8B8" box="[237,354,1873,1897]" captionStart="Figures 2328" captionStartId="8.[115,192,1986,2008]" captionTargetBox="[120,1467,234,1954]" captionTargetId="figure-51@8.[115,1476,234,1973]" captionTargetPageId="8" captionText="Figures 2328. Scorpiops grosseri sp. n., female holotype. Dorsal (23) and ventral (24) views. Telson lateral (25). Metasoma and telson lateral (26), ventral (27), and dorsal (28) views. Scale bars: 10 mm (2324, 2628)." figureDoi="http://doi.org/10.5281/zenodo.5741549" httpUri="https://zenodo.org/record/5741549/files/figure.png" pageId="7" pageNumber="6">Figs.2324</figureCitation>
).The base color is uniformly reddish brown to black. The telson and legs yellow to yellowish brown. Chelicerae are reddish black and reticulate.
</paragraph>
<paragraph id="145436D9FFB9FF92CF29FF500AABFD87" blockId="7.[816,1475,253,1590]" pageId="7" pageNumber="6">
<emphasis id="269FEACBFFB9FF92CF29FF500D23FEC7" bold="true" box="[816,1135,253,278]" pageId="7" pageNumber="6">Carapace and mesosoma</emphasis>
(
<figureCitation id="8CD02A5CFFB9FF92C894FF530C6FFEC7" box="[1165,1315,254,278]" captionStart="Figures 4048" captionStartId="10.[115,190,1981,2003]" captionTargetBox="[113,1474,238,1943]" captionTargetId="figure-59@10.[113,1474,238,1958]" captionTargetPageId="10" captionText="Figures 4048. Scorpiops grosseri sp. n., female holotype. Carapace and tergites IIII (40), anterior margin of carapace and chelicerae (41), pectine (42), posterior coxosternal area and sternites IIIV (43). Left legs IIV, retrolateral aspect (4447 respectively) and leg III dorsal view (48)." figureDoi="http://doi.org/10.5281/zenodo.5741555" httpUri="https://zenodo.org/record/5741555/files/figure.png" pageId="7" pageNumber="6">Figs. 4043</figureCitation>
). The entire carapace is covered with large and minute granules; carinae are absent. The anterior margin of the carapace is markedly depressed, convex in the middle. The carapace bears three lateral eyes. The mesosoma is granulated, with one median carina. Tergite VII is pentacarinate. The sternites are smooth with two parallel furrows except sternite VII, which bears four sparsely granulate carinae. Pectinal teeth number
<quantity id="D3139B3CFFB9FF92C967FE700C8EFE27" box="[1406,1474,477,502]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="7" pageNumber="6" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">45 in</quantity>
female, fulcra are absent. A pecten forms one compact unit with an incomplete furrow between places for marginal and middle lamellae.
</paragraph>
<paragraph id="145436D9FFB9FF92CF29FDF30DBBFCA4" blockId="7.[816,1475,253,1590]" pageId="7" pageNumber="6">
<emphasis id="269FEACBFFB9FF92CF29FDF30D6BFDA7" bold="true" box="[816,1063,606,630]" pageId="7" pageNumber="6">Metasoma and telson</emphasis>
(
<figureCitation id="8CD02A5CFFB9FF92C82EFDF30DF1FDA4" box="[1079,1213,605,630]" captionStart="Figures 2328" captionStartId="8.[115,192,1986,2008]" captionTargetBox="[120,1467,234,1954]" captionTargetId="figure-51@8.[115,1476,234,1973]" captionTargetPageId="8" captionText="Figures 2328. Scorpiops grosseri sp. n., female holotype. Dorsal (23) and ventral (24) views. Telson lateral (25). Metasoma and telson lateral (26), ventral (27), and dorsal (28) views. Scale bars: 10 mm (2324, 2628)." figureDoi="http://doi.org/10.5281/zenodo.5741549" httpUri="https://zenodo.org/record/5741549/files/figure.png" pageId="7" pageNumber="6">Figs. 2528</figureCitation>
). The metasoma is very sparsely hirsute and granulated, with relatively large granules. Metasomal segment I with 10 carinae, IIIV with 8 carinae, and V with 5 carinae. The median lateral carinae of metasoma V are indicated by isolated granules that may coalesce into carinae. The dorsolateral carinae of segments III and IV granulate with rounded granules, which are not posteriorly terminated with a pronounced tooth. The telson is bulbous and finely granulate, with annular ring present.
</paragraph>
<paragraph id="145436D9FFB9FF92CF29FCD00D66FAE4" blockId="7.[816,1475,253,1590]" pageId="7" pageNumber="6">
<emphasis id="269FEACBFFB9FF92CF29FCD00AD3FC44" bold="true" box="[816,927,893,917]" pageId="7" pageNumber="6">Pedipalps</emphasis>
(
<figureCitation id="8CD02A5CFFB9FF92CFACFCD30D73FC47" box="[949,1087,894,918]" captionStart="Figures 2939" captionStartId="9.[115,192,1957,1979]" captionTargetBox="[125,1474,247,1900]" captionTargetId="figure-71@9.[113,1474,238,1918]" captionTargetPageId="9" captionText="Figures 2939. Scorpiops grosseri sp. n., female holotype, pedipalp segments. Chela dorsal (29), external (30) and ventral (31) views. Patella dorsal (32), external (33) and ventral (34) views. Femur and trochanter dorsal (35), internal (36), and ventral (37) views. Movable finger dentition under white light (38) and UV fluorescence (39). Trichobothrial pattern is indicated by white circles." figureDoi="http://doi.org/10.5281/zenodo.5741551" httpUri="https://zenodo.org/record/5741551/files/figure.png" pageId="7" pageNumber="6">Figs. 2939</figureCitation>
). The pedipalps are very sparsely hirsute. The patella bears 17 (5
<emphasis id="269FEACBFFB9FF92C88BFC330DE7FC67" box="[1170,1195,926,950]" italics="true" pageId="7" pageNumber="6">eb</emphasis>
, 2
<emphasis id="269FEACBFFB9FF92C8C8FC330DB9FC67" box="[1233,1269,926,950]" italics="true" pageId="7" pageNumber="6">esb</emphasis>
, 2
<emphasis id="269FEACBFFB9FF92C905FC330C77FC64" box="[1308,1339,926,949]" italics="true" pageId="7" pageNumber="6">em</emphasis>
, 4
<emphasis id="269FEACBFFB9FF92C97BFC330CCCFC64" box="[1378,1408,926,949]" italics="true" pageId="7" pageNumber="6">est</emphasis>
, 4
<emphasis id="269FEACBFFB9FF92C9BFFC330CF5FC64" box="[1446,1465,926,949]" italics="true" pageId="7" pageNumber="6">et</emphasis>
) external trichobothria and 7 ventral trichobothria. The chela bears 4 ventral trichobothria located on the ventral surface. The femur and patella are granulated. The femur has 5 granulose carinae and the patella has 5 complete carinae with dorsal and ventral patellar spurs present. The manus dorsally bears granules, which are in the central area replaced by large granules forming a longitudinal irregular incomplete carina. The external surface of the chela is covered by minute and larger sparse granules, which form another complete median carina. The movable fingers with 3436 IAD, which form a second row, parallel to MD (
<quantity id="D3139B3CFFB9FF92C87DFB500D88FAC7" box="[1124,1220,1277,1302]" metricMagnitude="0" metricUnit="m" metricValue="1.5875" metricValueMax="1.6509999999999998" metricValueMin="1.524" pageId="7" pageNumber="6" unit="in" value="62.5" valueMax="65.0" valueMin="60.0">6065 in</quantity>
number); there are also 4 ID and 8 OD present.
</paragraph>
<paragraph id="145436D9FFB9FF92CF29FA930D98F9C7" blockId="7.[816,1475,253,1590]" pageId="7" pageNumber="6">
<emphasis id="269FEACBFFB9FF92CF29FA930A29FA87" bold="true" box="[816,869,1342,1366]" pageId="7" pageNumber="6">Legs</emphasis>
(
<figureCitation id="8CD02A5CFFB9FF92CF60FA930D4EFA87" box="[889,1026,1342,1366]" captionStart="Figures 4048" captionStartId="10.[115,190,1981,2003]" captionTargetBox="[113,1474,238,1943]" captionTargetId="figure-59@10.[113,1474,238,1958]" captionTargetPageId="10" captionText="Figures 4048. Scorpiops grosseri sp. n., female holotype. Carapace and tergites IIII (40), anterior margin of carapace and chelicerae (41), pectine (42), posterior coxosternal area and sternites IIIV (43). Left legs IIV, retrolateral aspect (4447 respectively) and leg III dorsal view (48)." figureDoi="http://doi.org/10.5281/zenodo.5741555" httpUri="https://zenodo.org/record/5741555/files/figure.png" pageId="7" pageNumber="6">Figs. 4448</figureCitation>
). The tibia and tarsomeres of legs with several setae not arranged into bristlecombs on dorsal surfaces but with rows of spines on dorsolateral and ventrolateral surface. Tarsomere II of legs IIV with 46 stout median ventral spines in a row and two other parallel spines. The femur bears 34 and patella, 45 incomplete carinae; both femur and patella are finely granulated.
</paragraph>
<paragraph id="145436D9FFB9FF92CF29F9B30D26F9E7" blockId="7.[816,1475,253,1590]" box="[816,1130,1566,1590]" pageId="7" pageNumber="6">
<emphasis id="269FEACBFFB9FF92CF29F9B30A9BF9E7" bold="true" box="[816,983,1566,1590]" pageId="7" pageNumber="6">Measurements</emphasis>
. See
<tableCitation id="59690362FFB9FF92C80BF9B30D28F9E7" box="[1042,1124,1566,1590]" captionStart="Table 1" captionStartId="3.[113,169,985,1006]" captionText="Table 1. Comparative measurements of adults of Scorpiops furai sp. n., S. grosseri sp. n., S. harmsi sp. n. and S. hofereki sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="7" pageNumber="6">Table 1</tableCitation>
.
</paragraph>
<paragraph id="145436D9FFB9FF92CF29F9CC0D7CF889" blockId="7.[816,1474,1633,1881]" pageId="7" pageNumber="6">
AFFINITIES. The described features distinguish
<taxonomicName id="D3EB4D5AFFB9FF92C951F9CC0C8EF9A9" authority="Kovařík, 2020" authorityName="Kovařík" authorityYear="2020" box="[1352,1474,1633,1656]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="grosseri" status="sp. nov.">
<emphasis id="269FEACBFFB9FF92C951F9CC0C8EF9A9" box="[1352,1474,1633,1656]" italics="true" pageId="7" pageNumber="6">S. grosseri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DAC57B0FFB9FF92CF29F92C0A23F949" box="[816,879,1665,1689]" pageId="7" pageNumber="6" rank="species">
<emphasis id="269FEACBFFB9FF92CF29F92C0A05F948" bold="true" box="[816,841,1665,1689]" pageId="7" pageNumber="6">sp</emphasis>
.
<emphasis id="269FEACBFFB9FF92CF40F92C0A24F948" bold="true" box="[857,872,1665,1689]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. The combination of five characters (chelal trichobothrium
<emphasis id="269FEACBFFB9FF92C900F90C0C0CF96E" box="[1305,1344,1697,1727]" italics="true" pageId="7" pageNumber="6">
Eb
<subScript id="886F349CFFB9FF92C921F91C0C0CF96E" attach="left" box="[1336,1344,1713,1727]" fontSize="6" pageId="7" pageNumber="6">3</subScript>
</emphasis>
located in proximal half of external surface between trichobothria
<emphasis id="269FEACBFFB9FF92C9BEF96F0C8EF908" box="[1447,1474,1730,1753]" italics="true" pageId="7" pageNumber="6">Dt</emphasis>
and
<emphasis id="269FEACBFFB9FF92CF47F94C0A33F928" box="[862,895,1761,1785]" italics="true" pageId="7" pageNumber="6">Db</emphasis>
; total length over
<quantity id="D3139B3CFFB9FF92C851F94C0DDBF929" box="[1096,1175,1761,1785]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="6" unit="mm" value="50.0">50 mm</quantity>
; fingers of pedipalps flexed in female; pedipalp movable finger with 3436 IAD; and chela length to width ratio
<quantity id="D3139B3CFFB9FF92C87EF88C0DE4F8E8" box="[1127,1192,1825,1849]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="7" pageNumber="6" unit="in" value="2.6">2.6 in</quantity>
female) is unique in the entire genus
<taxonomicName id="D3EB4D5AFFB9FF92CFA5F8EC0D65F889" authorityName="Peters" authorityYear="1861" box="[956,1065,1857,1880]" class="Arachnida" family="Euscorpiidae" genus="Scorpiops" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="269FEACBFFB9FF92CFA5F8EC0D65F889" box="[956,1065,1857,1880]" italics="true" pageId="7" pageNumber="6">Scorpiops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="145436D9FFB9FF92CF29F8290CC1F84D" blockId="7.[816,1421,1924,1948]" box="[816,1421,1924,1948]" pageId="7" pageNumber="6">
DISTRIBUTION.
<collectingCountry id="6CFC7649FFB9FF92CFC8F8290D46F84D" box="[977,1034,1924,1948]" name="India" pageId="7" pageNumber="6">India</collectingCountry>
,
<collectingRegion id="D62FF83BFFB9FF92C80CF8290C58F84D" box="[1045,1300,1924,1948]" country="India" name="Himachal Pradesh" pageId="7" pageNumber="6">Himachal Pradesh State</collectingRegion>
(
<figureCitation id="8CD02A5CFFB9FF92C93AF8290CCCF84D" box="[1315,1408,1924,1948]" captionStart="Figure 240" captionStartId="41.[113,181,1361,1383]" captionTargetBox="[113,1474,234,1350]" captionTargetId="figure-353@41.[113,1474,234,1350]" captionTargetPageId="41" captionText="Figure 240. Map of distribution of nine new species described here and Scorpiops vonwicki Birula, 1913." figureDoi="http://doi.org/10.5281/zenodo.5741606" httpUri="https://zenodo.org/record/5741606/files/figure.png" pageId="7" pageNumber="6">Fig. 240</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>