226 lines
25 KiB
XML
226 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZBK="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="3FA3C8D0DC101E53AE1D774B9AE63A93" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus inobservatus Bistroem, Nilsson & Bergsten, 2015, sp. n." docType="treatment" docUuid="A01A25B7-A2C0-4B3D-B810-315E70BB68CA" docUuidSource="ZooBank" docVersion="5" lastPageNumber="58" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="55" updateTime="1668162284477" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Bistroem, Olof</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Nilsson, Anders N.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Bergsten, Johannes</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2015</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>542</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>379</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-542-1</mods:identifier>
|
||
<mods:identifier type="ZBK">026407877355425BAB10BF1674510F12</mods:identifier>
|
||
<mods:identifier type="ZooBank">026407877355425BAB10BF1674510F12</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="127862780" LSID="urn:lsid:zoobank.org:act:A01A25B7-A2C0-4B3D-B810-315E70BB68CA" httpUri="http://treatment.plazi.org/id/3FA3C8D0DC101E53AE1D774B9AE63A93" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55">
|
||
<subSubSection pageId="54" pageNumber="55" type="multiple">
|
||
<paragraph pageId="54" pageNumber="55">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="54" pageNumber="55" type="nomenclature">
|
||
<paragraph pageId="54" pageNumber="55">
|
||
<taxonomicName LSID="http://zoobank.org/A01A25B7-A2C0-4B3D-B810-315E70BB68CA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inobservatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="inobservatus">Laccophilus inobservatus</taxonomicName>
|
||
<taxonomicNameLabel pageId="54" pageNumber="55">sp. n.</taxonomicNameLabel>
|
||
Figs 51-52, 251, 406-408, 537
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="54" pageNumber="55" type="type locality">
|
||
<paragraph pageId="54" pageNumber="55">Type locality.</paragraph>
|
||
<paragraph pageId="54" pageNumber="55">Chad: Near Bongor.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55" type="type material">
|
||
<paragraph pageId="54" pageNumber="55">Type material</paragraph>
|
||
<paragraph lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55">
|
||
(234 exs.). Holotype: male: "Chad nr Bongor 27.5. 1973 R. Linnavuori" (MZH; habitus in Fig. 406). - Paratypes: Gambia: "Abuko Nature Reserve, at light at the bamboo pool 18.30-20.30, 18.11. 1977 UTM 28 PCK2181 Loc. 24 / Cederholm-Danielsson-Hammarstedt-Hedquist-Samuelsson" (1 ex. MZLU, 1 ex. NHMB); "Outside Abuko Nature Reserve at waterworks. At light 19.00-22.00, 26.2. 1977 Loc. No. 6 UTM 28 PCK 214812 / Cederholm-Danielsson-Hammarstedt-Hedquist-Samuelsson" (1 ex. MZLU); "Tendeba Camp at light in semiarid veg. near River Gambia 18.30-20.30, 14.11. 1977, UTM 28 POK1285 Loc. 12a / Cederholm-Danielsson-Hammarstedt-Hedquist-Samuelsson" (1 ex. MZLU); "3.5 km S Georgetown, Hilltop at Sankuli Kunda, alt. about 30 m, at light 18.30-20.15, 15.11. 1977 UTM 28PEK2593 Loc. 37 /-Danielsson-Hammarstedt-Hedquist-Samuelsson" (1 ex. MZLU); "2 km S Kitty, 7 km SSW Brikama Road junction. In and at Fresh Water Stream 13.11. 1977 UTM 28PCK 1761 Loc 7/-Danielsson-Hammarstedt-Hedquist-Samuelsson" (1 ex. MZLU, 1 ex. MZH, 1 ex. NHMB); "Gam Bathurst Jan. 68 Palm /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
Reg. det. Sven Persson" (4 exs. MZLU); "Bathurst Jan. 1968 T-E Leiler" (5 exs. NHRS, 1 ex. MZH); "Kuntaur NW Georgetown 21.11. 2003 B. Vondel /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. Rocchi 2004" (1 ex. CSR); Gambia: Bakau 6-26.11. 1984 leg. Palm /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Rég">Reg</normalizedToken>
|
||
. det. Sven Persson" (1 ex. MZLU); "Gambia Oil Palm and mangrove veg. close to the beach, about 5 km SSW Gunjur, at light 18.45-20.30 13.11. 1977 UTM 28PCK0554 Loc. 8 / Cederholm & al." (1 ex. NHMB); "Gambia-southern Senegal
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="13.166667">13°10'N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-16.6">16°36'W</geoCoordinate>
|
||
stream N of Selety 19,2, 1976 M. Holmen" (1 ex. ZMUC). - Senegal: "70 km W Tambacounda
|
||
<geoCoordinate direction="north" orientation="latitude" precision="92" value="13.956667">13°57.4'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="92" value="14.265">14°15.9'E</geoCoordinate>
|
||
, 29.6. 2004 Halada leg." (2 exs. NMPC); "Senegal, Parc National de Niokolo Koba 16.2. 1989" (1 ex.
|
||
<pageBreakToken pageId="55" pageNumber="56" start="start">NHMB</pageBreakToken>
|
||
); "Senegal Cayare II. 46 A. Villiers" (1 ex. NHMB). - Mali: "NW Afr., K. Macina 10.11. 1973 D.R. Reynolds (C. O. R.R.) BM 1974-222" (1 ex. BMNH, 1 ex. MZH); "Kogoni X. 1966 G. Schmitz" (8 exs. MRAC, 2 exs. MZH); "Korioume, Niger Riv. 18.2. 2000
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="18.666666">18°40'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-3.0">3°00'W</geoCoordinate>
|
||
, leg. Komarek & Mayer /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. Wewalka 2001" (1 ex. NMW); "Markala Niger River,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="13.666667">13°40'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-6.0833335">6°05'W</geoCoordinate>
|
||
, leg. Komarek & Mayer 9-4 /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. Wewalka 2001" (1 ex. NMW); "S Tombouctou
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="16.666666">16°40'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-3.0">3°00'W</geoCoordinate>
|
||
, 18.2. 2000 leg. Komarek & Mayer 18-1 /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. Wewalka 2001" (1 ex. NMW, 1 ex. MZH); "Mopti Niger Riv.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="14.5">14°30'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-4.2">4°12'W</geoCoordinate>
|
||
, 21.2. 2000 leg. Komarek & Mayer 21-2 /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. Wewalka 2001" (1 ex. NMW, 1 ex. MZH); "Goundaka, Bandiagara Riv.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="14.483334">14°29'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-3.9333334">3°56'W</geoCoordinate>
|
||
, 12.2. 2000 leg. Komarek & Mayer 12-1 /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. Wewalka 2001" (2 exs. NMW) "SE Douna, Bani Riv.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="13.216666">13°13'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-5.9">5°54'W</geoCoordinate>
|
||
, 10.2. 2000 leg. Komarek & Mayer 10-1 /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. Wewalka 2001" (1 ex. NMW). - Niger: "Nr. Boureimi 9.11. 1973 R. Linnavuori" (1 ex. MZH);
|
||
<normalizedToken originalValue="“Rég">"Reg</normalizedToken>
|
||
. de Zinder, Sultanat de Damagherim Dungass, Mission Tilho Dr. R. Gaillard 1910" (6 exs. NHMB). - Sudan: "Blue Nile Ingessana Mts. 17-22.11. 1962 Linnavuori" (6 exs. MZH); "Nile Blue Nile Singa-Roseiras 15-17.11. 1962 Linnavuori" (1 ex. MZH); "Upper Nile Malakal 5-20.1. 1963 Linnavuori" (6 exs. MZH); "Kordofan Lake Keilak 8-11.2.1963 Linnavuori" (4 exs. MZH); "Bahr el Gazal, Wau 19.2. 1963 Linnavuori" (1 ex. MZH); "Bahr el Abiad
|
||
<normalizedToken originalValue="Trägårdh">Traegardh</normalizedToken>
|
||
/
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régt">Regt</normalizedToken>
|
||
var." (1 ex. NHRS); "Torit 2.7. 1980 Armstrong" (1 ex. USNM); "Nairege River 27.2. 1980 Armstrong" (1 ex. USNM); "Gilo water tank (pumped up from stream) 20.3. 1980 Armstrong (8 exs. USNM, 3 exs. MZH); "Kinyetti River at Imeila 19.3. 1980 Armstrong" (5 exs. USNM, 1 ex. MZH); "Sudan Wad Medani am Bl. Nil, 18.10, 1979 lux, leg. Hieke" (1 ex. NHMB); same but "21.10. 1979" (1 ex. NHMB); same but "29.10. 1979" (1 ex. MZH); same but "12.10. 1979" (1 ex. NHMB); same but "30+31.10. 1979" (1 ex. NHMB); "Tombe 17.1. 1954 Omer-Cooper" (1 ex. AMGS). - Chad: Same data as holotype (5 exs. MZH); "Distr. Kanem
|
||
<normalizedToken originalValue="N’Gouri">N'Gouri</normalizedToken>
|
||
X-XI. 1958 P. Renaud ex. coll. Breuning" (21 exs. MRAC, 2 exs. MZH). - Ethiopia: "Shoa, Awash NP, Filwoha Hot Springs 25.12. 1988, 1500 m leg. S. Persson /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. A. Nilsson" (2 exs. MZLU); "Shoa, Metahara 2.10. 1988 950 m, water hole in lava field, leg. S. Persson /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Régb">Regb</normalizedToken>
|
||
. det. A. Nilsson" (6 exs. MZLU, 2 exs. MZH; habitus in Fig. 407); "Hora Harsadi, Addas 7000 ft 2.12. 1926 J. Omer-Cooper" (1 ex. AMGS). - Burkina Faso:"Pundu Mte Volta1927-1928 Dez.-Juni Olsufiew" (2 exs. NHRS); "Pundu Olsufiew" (38 exs. NHRS, 3 exs. MZH); "Yatenga Gourcy Barrage 300 m 14.1. 1995 Trockengefallen lehmiges Ufer leg. B. Maier /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
<normalizedToken originalValue="Rég">Reg</normalizedToken>
|
||
. det. Rocchi 2002" (1 ex. CSR). - Ivory Coast:
|
||
<normalizedToken originalValue="“Comoé">"Comoe</normalizedToken>
|
||
NP,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5555" value="8.5">N8,5°</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5555" value="-3.5">W3,5°</geoCoordinate>
|
||
leg. N. Reintjes, det. F. Pederzani / 1.3. 1999 rock pool in
|
||
<normalizedToken originalValue="Comoé">Comoe</normalizedToken>
|
||
river bed" (1 ex. NMW).
|
||
<normalizedToken originalValue="“Comoé">"Comoe</normalizedToken>
|
||
NP,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5555" value="8.5">N8,5°</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5555" value="-3.5">W3,5°</geoCoordinate>
|
||
leg. N. Reintjes, det. F. Pederzani / 3.4. 1999 temporary Pond" (1 ex. NMW). - Ghana: "Upper E Prov. Navrongo env. 11-13.6. 2006 Pokorny leg." (5 exs. NMPC, 1 ex. MZH). - Nigeria: "Kano St. Wudil-Kari 17.5. 1973 R. Linnavuori" (1 ex. MZH); "NW St. Gummi-Anka 24.7. 1973 R. Linnavuori" (1 ex. MZH); "NE St. Gombe-Bauchi 27.8. 1973 R. Linnavuori" (4 exs. MZH; habitus in Fig. 408); "Detritus pond, Jos-Bauchi rd 9.4. 1963 JOC"(4 exs. AMGS); "Samaru 17.5. 1959 W.
|
||
<pageBreakToken pageId="56" pageNumber="57" start="start">Sands</pageBreakToken>
|
||
B.M. 1961-525 / Light trap" (1 ex. BMNH); "Marsk, road Katsina-Daura 6.4. 1963 JOC" (14 exs. AMGS). - Cameroon: "Maroua (
|
||
<normalizedToken originalValue="lumière">lumiere</normalizedToken>
|
||
) X-XI. 1965 G. Schmitz" (1 ex. MRAC); "Maroua, Miss. Cath. 26.8. 73" (1 ex. NHMB); "Maroua 5 Aout 71" (1 ex. NHMB); "Tokombere, dint. Maroua, 12.7. 1979 Onore /
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus congener" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="congener">Laccophilus congener</taxonomicName>
|
||
O-C. det. Rocchi 1980" (1 ex. CSR). - Zaire: "P.N.G. Ndelele. K.117/14S, 19.3. 1952 H, De Saeger, 3199" (2 exs. MRAC). P.N.G. II/fd/13, 5.5. 1952 H, De Saeger, 3421" (1 ex. MRAC). - Non-African records: Yemen: "Wadi am Rija W Lahj Al Hulah by road, 13.01.57N, 44.33.30E (GPS) 25-26.10. 2007, 297 m a.s.l., Reitter leg." (7 exs. NMPC, 2 exs. MZH).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="56" pageNumber="57" type="specimens with unclear labelling">
|
||
<paragraph pageId="56" pageNumber="57">Specimens with unclear labelling.</paragraph>
|
||
<paragraph pageId="56" pageNumber="57">
|
||
Two specimens in NHRS labelled
|
||
<normalizedToken originalValue="“Egypten”">"Egypten"</normalizedToken>
|
||
also belong to this species. The material is fairly old and the exact location of these records is somewhat unclear. The specimens are therefore not included in type material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="56" pageNumber="57" type="diagnosis">
|
||
<paragraph pageId="56" pageNumber="57">Diagnosis.</paragraph>
|
||
<paragraph pageId="56" pageNumber="57">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inobservatus" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inobservatus">Laccophilus inobservatus</taxonomicName>
|
||
is closely related especially to
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus simplicistriatus" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="simplicistriatus">Laccophilus simplicistriatus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
. The species can be distinguished by study of penis apex-shape, which is peculiar and stable in all four species. Penis apex of
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inobservatus" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inobservatus">Laccophilus inobservatus</taxonomicName>
|
||
is cut off straight and lacks any signs of anterior processes - resembling species have all at least minor kinds of modifications/processes on penis apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="description">
|
||
<paragraph pageId="56" pageNumber="57">Description.</paragraph>
|
||
<paragraph pageId="56" pageNumber="57">Body length 3.6-4.0 mm, width 1.9-2.2 mm. Colour pattern dorsally, reasonably uniform; rarely reduced so that elytral irrorations are in part fragmentary (Figs 406-408).</paragraph>
|
||
<paragraph pageId="56" pageNumber="57">Head: Pale ferrugineous, lacks darker areas. Slightly mat, finely microsculptured. Reticulation double. Large meshes only slightly more strongly developed in comparison with small meshes. Large meshes may contain 2-6 small meshes. Impunctate, except at eyes, with fine, irregular punctures.</paragraph>
|
||
<paragraph pageId="56" pageNumber="57">Pronotum: Pale ferrugineous. At foremargin between eyes with a dark ferrugineous to blackish, slightly vague marking. At base in middle with two narrow, blackish spots. Sometimes dark areas on pronotum may be reduced. Slightly mat, finely microsculptured. Reticulation double. Large meshes only slightly more strongly developed than small meshes. Large meshes may contain 2-6 small meshes. Impunctate, but at margins except basally in middle with very fine scattered punctures.</paragraph>
|
||
<paragraph pageId="56" pageNumber="57">Elytra: Pale ferrugineous, with dense, blackish to dark ferrugineous irrorations. Irrorations generally quite evenly distributed; sometimes irrorations reduced and at least in part separate irrorations rudimentary. Posterior to middle irrorations can be strongly reduced forming an irregular pale spot on each elytron (Figs 406-408). Slightly mat, finely microsculptured; reticulation double. Large meshes only slightly more strongly developed than small meshes. Large meshes contain 2-6 small meshes. Punctures very fine, sparse and irregularly distributed; on disc, irregular punctures form a vague row of punctures. Pre-apical, lateral row of punctures form a shallow furrow provided with hairs.</paragraph>
|
||
<paragraph lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
|
||
Ventral aspect: Pale ferrugineous, abdomen distinctly darker; dark to blackish ferrugineous. Almost impunctate, except apical ventrite; with some, fine, irregular punctures. Apical ventrite lacks lateral knob (Fig. 51). Rather shiny although finely microsculptured; microsculpture in part indistinct and reduced. Ventrites with fine, slightly curved striae. Metacoxal plates with some 10 almost transversely located, shallow and in
|
||
<pageBreakToken pageId="57" pageNumber="58" start="start">part</pageBreakToken>
|
||
reduced, furrows. Prosternal process slender, posteriorly distinctly extended, apex pointed.
|
||
</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">Legs: Pro- and mesotarsus somewhat extended and enlarged, provided with distinct suckers.</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">Male genitalia: Extreme penis apex, blunt, abruptly broken and lacks any kinds of processes (Fig. 251).</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">Female: Pro- and mesotarsi slender. Apical ventrite as in Fig. 52.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="57" pageNumber="58" type="etymology">
|
||
<paragraph pageId="57" pageNumber="58">Etymology.</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">
|
||
The species name inobservatus is a Latin adjective meaning
|
||
<normalizedToken originalValue="“unobserved”">"unobserved"</normalizedToken>
|
||
. It here refers to the peculiar situation that the species remained overlooked for a long time due to misinterpretation, although it is widespread and common.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="57" pageNumber="58" type="distribution">
|
||
<paragraph pageId="57" pageNumber="58">Distribution.</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">Africa: Gambia, Senegal, Mali, Niger, Sudan, Chad, Ethiopia, Burkina Faso, Ivory Coast, Ghana, Nigeria, Cameroon, Zaire and Asia: Yemen (Fig. 537). Only personally verified records are included in the map. Egypt is also among examined material but exact location of this record is unknown and therefore record is not mapped.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="57" pageNumber="58" type="collecting circumstances">
|
||
<paragraph pageId="57" pageNumber="58">Collecting circumstances.</paragraph>
|
||
<paragraph pageId="57" pageNumber="58">
|
||
Label data provide some information on the living habits of
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inobservatus" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="inobservatus">Laccophilus inobservatus</taxonomicName>
|
||
. Accordingly, in Gambia collected at light at a bamboo pool and in semiarid vegetation near a river. Moreover the species has been collected in and at a fresh water stream and in a rock pool in river bed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |