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911 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="38446D96E8785C33B44B278839E24F7B" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina artata Benson 1856" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<treatment ID-GBIF-Taxon="199839023" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B" httpUri="http://treatment.plazi.org/id/38446D96E8785C33B44B278839E24F7B" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="38446D96-E878-5C33-B44B-278839E24F7B" authority="Benson, 1856" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">Pupina artata Benson, 1856</taxonomicName>
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">Figs 8E-H</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">, 10B</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. External shell morphology of three mainland Southeast Asian Pupina species groups A Pupina artata from the Pupina artata species group, CUMZ 12003 from Ban Ping Khong, Chiang Mai B Pupina peguensis from the Pupina arula species group, CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai C Pupina siamensis from the Pupina arula species group, CUMZ 12052 from Sri Thong Cave, Sra Keo, and D Pupina tchehelensis from the Pupina aureola species group, CUMZ 12136 from limestone mountain, Phang Nga. Red frames focus on the parietal tooth and posterior canal; blue frames focus on the curvature of the apertural lip when observed from lateral view." figureDoi="10.3897/zookeys.1119.85400.figure18" httpUri="https://binary.pensoft.net/fig/735379" pageId="0" pageNumber="1">, 18A</figureCitation>
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">, 21A-M</figureCitation>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">, 22</figureCitation>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">, 23</figureCitation>
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">, 24A</figureCitation>
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">, 25A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">Pupina artata</taxonomicName>
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Benson, 1856: 230. Type locality: Moulmein [Mawlamyine, Mawlamyine Township, Mawlamyine District, Mon State, Myanmar].
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<bibRefCitation author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="245 - 254" refId="B167" refString="Theobald, W, 1858. Notes on the distribution of some of the land and freshwater shells of India. Journal of the Asiatic Society of Bengal 26 (1857): 245 - 254" title="Notes on the distribution of some of the land and freshwater shells of India." volume="26" year="1858">Theobald 1858</bibRefCitation>
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[1857]: 247, 248. Pfeiffer 1860: 142, pl. 37, figs 10-12. Sowerby I 1866:
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<taxonomicName authorityName="L. Pfeiffer" authorityYear="1853" family="Pupinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" rank="family">Pupinidae</taxonomicName>
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, pl. 3 (pl. 265),
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<taxonomicName authorityName="Vignard" authorityYear="1829" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pupina</taxonomicName>
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, figs 1, 2. Hanley and Theobald 1870: 4, pl. 7, fig. 5.
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<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="143 - 259" refId="B150" refString="Stoliczka, F, 1871. Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 259" title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species." volume="40" year="1871">Stoliczka 1871</bibRefCitation>
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: 151, 152.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="1" refId="B104" refString="Nevill, G, 1878. Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
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: 299, 300, Ava [Mandalay Region, Myanmar]; Moulmein; Buket Pondong [Gunung Pondok, Perak State, Malaysia].
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<bibRefCitation author="Reeve, L" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" refId="B139" refString="Reeve, L, 1878. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1-10. Lovell Reeve & Co, London, [without pagination]." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1 - 10. Lovell Reeve & Co, London, [without pagination]." year="1878">Reeve 1878</bibRefCitation>
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:
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<taxonomicName authorityName="L. Pfeiffer" authorityYear="1853" family="Pupinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" rank="family">Pupinidae</taxonomicName>
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, pl. 1, sp. 3.
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<bibRefCitation author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="336 - 340" refId="B26" refString="Crosse, H, 1879. Faune malacologique de Perak (Indo-Chine). Journal de Conchyliologie 27: 336 - 340" title="Faune malacologique de Perak (Indo-Chine)." volume="27" year="1879">Crosse 1879</bibRefCitation>
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: 340. de Morgan 1885: 413, Boukit Pondong,
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<normalizedToken originalValue="Pérak">Perak</normalizedToken>
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; Java [doubtful]; Moulmein; Lahat, Ipoh,
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<normalizedToken originalValue="Gôping">Goping</normalizedToken>
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, Kinta [Perak State, Malaysia].
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<bibRefCitation author="von Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="146 - 156" refId="B192" refString="von Moellendorff, OF, 1894. On a collection of land-shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London 62: 146 - 156" title="On a collection of land-shells from the Samui Islands, Gulf of Siam." volume="62" year="1894">
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von
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1894
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</bibRefCitation>
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: 155, the Samui Islands, Gulf of Siam [Samui Island, Surat Thani Province, Thailand].
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<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen 1897</bibRefCitation>
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: 38, 39, pl. 69, fig. 6, 6a, b.
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<bibRefCitation author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 431, Ile Samui, golfe de Siam [Samui Island, Surat Thani Province, Thailand].
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<bibRefCitation author="van Benthem Jutting, WSS" journalOrPublisher="Basteria" pageId="0" pageNumber="1" pagination="10 - 20" refId="B181" refString="van Benthem Jutting, WSS, 1960. Some notes on land and freshwater Mollusca of Malaya. Basteria 24: 10 - 20" title="Some notes on land and freshwater Mollusca of Malaya." volume="24" year="1960">van Benthem Jutting 1960</bibRefCitation>
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: 12, a hill near the hot springs, near Tandjong Rambutan, N.E. of Ipoh, Perak.
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<bibRefCitation author="Solem, A" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B146" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
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: 12, Chieng Dao, Doi Sutep [Chiang Dao District and Doi Suthep Mountain, Chiang Mai Province, Thailand].
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<bibRefCitation author="Davison, GWH" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="233 - 248" refId="B31" refString="Davison, GWH, 1995. The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993-1994. Malayan Nature Journal 48: 233 - 248" title="The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993 - 1994." volume="48" year="1995">Davison 1995</bibRefCitation>
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: 236, 237, limestone island C, Temengor dam, Perak, Malaysia.
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<bibRefCitation author="Chan, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="1 - 2" refId="B18" refString="Chan, S-Y, 1998b. A brief collecting trip to Perak, West Malaysia, part two. The Papustyla 12: 1 - 2" title="A brief collecting trip to Perak, West Malaysia, part two." volume="12" year="1998 b">Chan 1998b</bibRefCitation>
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: 2, Ipoh, Perak.
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<bibRefCitation author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
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: 39, 40.
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<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
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: 87.
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<bibRefCitation author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
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: fig. 5-13d.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina artata subsp. var. var. blanfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="Artata" subSpecies="var." variety="blanfordiana">Pupina artata var. blanfordiana</taxonomicName>
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Nevill, 1878: 300. Type locality: Thyet Myo [Thayetmyo, Magway Region, Myanmar]; Akoutong [Akauk Taung, Pyay District, Bago Region, Myanmar]; Kamah Hill, Tongoop, & c., Arakan [Toungup, Thandwe District, Rakhine State, Myanmar]; Prome [Pyay, Bago Region, Myanmar].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benson" authorityYear="1860" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peguensis">Pupina peguensis</taxonomicName>
|
|
[non Benson]-
|
|
<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen 1897</bibRefCitation>
|
|
: 40, pl. 69, fig. 3, 3a-d, Kama on the right bank of the Irrawaddy, Pegu [Kamma Township, Thayet District, Magway Region, Myanmar].
|
|
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 93.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata" subGenus="Tylotoechus">Pupina (Tylotoechus) artata</taxonomicName>
|
|
-
|
|
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
|
|
: 306, 307.
|
|
<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
|
|
: 193.
|
|
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
|
: 33.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Pupina) artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata" subGenus="Pupina">Pupina (Pupina) artata</taxonomicName>
|
|
-
|
|
<bibRefCitation author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
|
|
: 39.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Theobald" authorityYear="1864" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina blanfordi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="blanfordi">Pupina blanfordi</taxonomicName>
|
|
[non Theobald]-
|
|
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 89.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Godwin-Austen" authorityYear="1897" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitanea">Pupina limitanea</taxonomicName>
|
|
[non Godwin-Austen]-
|
|
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 90.
|
|
<bibRefCitation author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
|
|
: fig. 5-13g.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Vignard" authorityYear="1829" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pupina</taxonomicName>
|
|
sp.-
|
|
<bibRefCitation author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
|
|
: fig. 5-11a.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="type material">
|
|
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
|
|
UMZC I.102960.A (1 shell; Figs
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21A</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22A</figureCitation>
|
|
) from the R. McAndrew collection, labelled "Bens. col., Moulmein".
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<materialsCitation collectingDate="2008-01-09" collectorName="Figs, Moulmein" country="Myanmar" county="Chiang Rai Province" location="Khao Tham Phra Temple" municipality="Mueang Chiang Rai District" specimenCount="1">
|
|
NHMUK 1906.4.4.28 (6 shells;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21J</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22B</figureCitation>
|
|
) from
|
|
<collectorName>Moulmein</collectorName>
|
|
,
|
|
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
|
|
. CUMZ 12001 (7 shells; Figs
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21H</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22C</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F3458A76246BD6F4E33E000B48596955" country="Myanmar" county="Chiang Rai Province" municipality="Mueang Chiang Rai District" name="Khao Tham Phra Temple">Khao Tham Phra Temple</location>
|
|
,
|
|
<collectingMunicipality>Mueang Chiang Rai District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chiang Rai Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2008-01-09">9 Jan. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2015-10-23" country="Myanmar" county="Chiang Rai Province" location="Luang Cave" municipality="Mae Sai District" specimenCount="1">
|
|
CUMZ 12002 (1 shell) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:75DF4B534B3BCA50C1E226E6EEC83D4A" country="Myanmar" county="Chiang Rai Province" municipality="Mae Sai District" name="Luang Cave">Luang Cave</location>
|
|
,
|
|
<collectingMunicipality>Mae Sai District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chiang Rai Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2015-10-23">23 Oct. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2008-10-08" collectorName="Figs" country="Myanmar" county="Chiang Mai Province" location="Ban Ping Khong" municipality="Chiang Dao District" specimenCount="1">
|
|
CUMZ 12003 (21 shells;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">10B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. External shell morphology of three mainland Southeast Asian Pupina species groups A Pupina artata from the Pupina artata species group, CUMZ 12003 from Ban Ping Khong, Chiang Mai B Pupina peguensis from the Pupina arula species group, CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai C Pupina siamensis from the Pupina arula species group, CUMZ 12052 from Sri Thong Cave, Sra Keo, and D Pupina tchehelensis from the Pupina aureola species group, CUMZ 12136 from limestone mountain, Phang Nga. Red frames focus on the parietal tooth and posterior canal; blue frames focus on the curvature of the apertural lip when observed from lateral view." figureDoi="10.3897/zookeys.1119.85400.figure18" httpUri="https://binary.pensoft.net/fig/735379" pageId="0" pageNumber="1">18A</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21I</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22D</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F926A1F50087751CCDF235571B4F42B5" country="Myanmar" county="Chiang Mai Province" municipality="Chiang Dao District" name="Ban Ping Khong">Ban Ping Khong</location>
|
|
,
|
|
<collectingMunicipality>Chiang Dao District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chiang Mai Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2008-10-08">8 Oct. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2012-11-21" collectorName="Ban Ping Khong" country="Myanmar" location="Chiang Dao District" municipality="Chiang Mai Province" specimenCount="1">
|
|
CUMZ 12193 (4 shells) from
|
|
<collectorName>Ban Ping Khong</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:2E305503310D178ADC931B14CC875208" country="Myanmar" municipality="Chiang Mai Province" name="Chiang Dao District">Chiang Dao District</location>
|
|
,
|
|
<collectingMunicipality>Chiang Mai Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2012-11-21">21 Nov. 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2015-10-25" country="Myanmar" county="Chiang Mai Province" location="Chiang Dao Cave" municipality="Chiang Dao District" specimenCount="1">
|
|
CUMZ 12190 (3 shells) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:BA4500FD7862A675F0A4E6735A73BF34" country="Myanmar" county="Chiang Mai Province" municipality="Chiang Dao District" name="Chiang Dao Cave">Chiang Dao Cave</location>
|
|
,
|
|
<collectingMunicipality>Chiang Dao District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chiang Mai Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2015-10-25">25 Oct. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2017-10-08" country="Myanmar" county="Chiang Mai Province" location="Bua Tong Cave" municipality="Mae Tang District" specimenCount="4">
|
|
CUMZ 12004 (
|
|
<specimenCount type="generic">4 specimens</specimenCount>
|
|
in ethanol) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:8382F5FE09BCAD476EB39D11BFA179EB" country="Myanmar" county="Chiang Mai Province" municipality="Mae Tang District" name="Bua Tong Cave">Bua Tong Cave</location>
|
|
,
|
|
<collectingMunicipality>Mae Tang District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chiang Mai Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2017-10-08">8 Oct. 2017</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2015-10-24" collectorName="Doi Ang Khang" country="Myanmar" location="Fang District" municipality="Chiang Mai Province" specimenCount="1">
|
|
CUMZ 12168 (4 shells) from
|
|
<collectorName>Doi Ang Khang</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F3BF56D495AEF4540AC419134DDB906B" country="Myanmar" municipality="Chiang Mai Province" name="Fang District">Fang District</location>
|
|
,
|
|
<collectingMunicipality>Chiang Mai Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2015-10-24">24 Oct. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2015-01-18" country="Myanmar" county="Mae Hong Son Province" location="Pha Daeng Cave" municipality="Mueang Mae Hong Son District" specimenCount="1">
|
|
CUMZ 12005 (43 shells) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:5BDE773E37E6BE65EEFD1ED90F38E9D4" country="Myanmar" county="Mae Hong Son Province" municipality="Mueang Mae Hong Son District" name="Pha Daeng Cave">Pha Daeng Cave</location>
|
|
,
|
|
<collectingMunicipality>Mueang Mae Hong Son District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Mae Hong Son Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2015-01-18">18 Jan. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-12-03" collectorName="Fig." country="Myanmar" county="Mae Hong Son Province" location="Pha Daeng Cave" municipality="Mueang Mae Hong Son District" specimenCount="1">
|
|
CUMZ 12006 (17 shells and
|
|
<specimenCount type="generic">1 specimen</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Fig.</collectorName>
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8E</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:ADF92299F57A4FCD2A10A8FA06EE05F5" country="Myanmar" county="Mae Hong Son Province" municipality="Mueang Mae Hong Son District" name="Pha Daeng Cave">Pha Daeng Cave</location>
|
|
,
|
|
<collectingMunicipality>Mueang Mae Hong Son District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Mae Hong Son Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2020-12-03">3 Dec. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2008-01-07" collectorName="Tham Nam Pha Pha Ngam" country="Myanmar" location="Mae Phrik District" municipality="Lampang Province" specimenCount="1">
|
|
CUMZ 12007 (7 shells) from
|
|
<collectorName>Tham Nam Pha Pha Ngam</collectorName>
|
|
Temple,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:BE11E04038071036A8ED9000EC34D3A4" country="Myanmar" municipality="Lampang Province" name="Mae Phrik District">Mae Phrik District</location>
|
|
,
|
|
<collectingMunicipality>Lampang Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2008-01-07">7 Jan. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-10-08" collectorName="Fig." country="Myanmar" county="Lampang Province" location="Tham Nam Pha Pha Ngam" municipality="Mae Phrik District" specimenCount="4">
|
|
CUMZ 12008 (3 shells and
|
|
<specimenCount type="generic">4 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Fig.</collectorName>
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8F</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:01D2E8299BF09483B4CD25152962D53F" country="Myanmar" county="Lampang Province" municipality="Mae Phrik District" name="Tham Nam Pha Pha Ngam">Tham Nam Pha Pha Ngam</location>
|
|
Temple,
|
|
<collectingMunicipality>Mae Phrik District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Lampang Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2020-10-08">8 Oct. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2008-10-24" collectorName="Figs" country="Myanmar" location="Phu Sang District" municipality="Phayao Province" specimenCount="1">
|
|
CUMZ 12009 (12 shells;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21G</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22E</figureCitation>
|
|
) from Phu Sang Waterfall,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:4A671163378ACAF960EC3DC94C790046" country="Myanmar" municipality="Phayao Province" name="Phu Sang District">Phu Sang District</location>
|
|
,
|
|
<collectingMunicipality>Phayao Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2008-10-24">24 Oct. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2012-11-19" collectorName="Phu Sang Waterfall" country="Myanmar" location="Phu Sang District" municipality="Phayao Province" specimenCount="1">
|
|
CUMZ 12010 (5 shells) from
|
|
<collectorName>Phu Sang Waterfall</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:AE7A4C53D5517D19EBC953BE71C43A13" country="Myanmar" municipality="Phayao Province" name="Phu Sang District">Phu Sang District</location>
|
|
,
|
|
<collectingMunicipality>Phayao Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2012-11-19">19 Nov. 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2008-07-17" collectorName="Figs" country="Myanmar" county="Nakhon Sawan Province" location="Thep Sathaporn Temple" municipality="Banphot Phisai District" specimenCount="7">
|
|
CUMZ 12011 (12 shells and
|
|
<specimenCount type="generic">7 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22F</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:B492B2CF91C5E75245D5CB450C8E1799" country="Myanmar" county="Nakhon Sawan Province" municipality="Banphot Phisai District" name="Thep Sathaporn Temple">Thep Sathaporn Temple</location>
|
|
,
|
|
<collectingMunicipality>Banphot Phisai District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Nakhon Sawan Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2008-07-17">17 July 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2009-07-08" collectorName="Khao Chuak Charoentham Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12012 (17 shells) from
|
|
<collectorName>Khao Chuak Charoentham Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:85400E052CA5D5D88CAC4AF6F3D456D0" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality>Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2009-07-08">8 July 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2016-08-27" collectorName="Khao Chuak Charoentham Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12013 (3 shells) from
|
|
<collectorName>Khao Chuak Charoentham Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:EB6774EBC5F9D94926C93F51C6460E7D" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality>Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2016-08-27">27 Aug. 2016</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-12-05" collectorName="Khao Chuak Charoentham Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12014 (1 shell) from
|
|
<collectorName>Khao Chuak Charoentham Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:E885D15EB10AF982DE2D45561BFB2C27" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality>Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2020-12-05">5 Dec. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2009-07-08" collectorName="Figs" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12015 (14 shells;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21F</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23A</figureCitation>
|
|
) from Khao Wong Phrommachan Temple,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:2F1B4E38622CC3B8C0F856B2B4B10889" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality>Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2009-07-08">8 July 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-12-05" collectorName="Tham Prathat Mueang Thep Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12016 (5 shells) from
|
|
<collectorName>Tham Prathat Mueang Thep Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:92E29E54A9FE629C337BB66A119EC333" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality>Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2020-12-05">5 Dec. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2006-12-10" country="Myanmar" county="Kanchanaburi Province" location="Krasae Cave" municipality="Sai Yok District" specimenCount="1">
|
|
CUMZ 12017 (13 shells) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:2279B5C2C902810A8233C90779CF9586" country="Myanmar" county="Kanchanaburi Province" municipality="Sai Yok District" name="Krasae Cave">Krasae Cave</location>
|
|
,
|
|
<collectingMunicipality>Sai Yok District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Kanchanaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2006-12-10">10 Dec. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-08-19" collectorName="Tham Charoentham Temple" country="Myanmar" location="Mueang Kanchanaburi District" municipality="Kanchanaburi Province" specimenCount="1">
|
|
CUMZ 12173 (14 shells) from
|
|
<collectorName>Tham Charoentham Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:538EED740A16EE79E004C506B562D599" country="Myanmar" municipality="Kanchanaburi Province" name="Mueang Kanchanaburi District">Mueang Kanchanaburi District</location>
|
|
,
|
|
<collectingMunicipality>Kanchanaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2020-08-19">19 Aug. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2015-06-30" collectorName="Ban Tapoepu-Wakruko" country="Myanmar" location="Umphang District" municipality="Tak Province" specimenCount="1">
|
|
CUMZ 12192 (1 shell) from
|
|
<collectorName>Ban Tapoepu-Wakruko</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:3687A1ADD22DB26ECEC82DDA4146B532" country="Myanmar" municipality="Tak Province" name="Umphang District">Umphang District</location>
|
|
,
|
|
<collectingMunicipality>Tak Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2015-06-30">30 June 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2006-12-09" collectorName="Figs" country="Myanmar" county="Ratchaburi Province" location="Tham Khao Thalu Temple" municipality="Chom Bueang District" specimenCount="10">
|
|
CUMZ 12018 (68 shells and
|
|
<specimenCount type="generic">10 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21M</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23B</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:C7389ABB35BFA1589C509FDF77769369" country="Myanmar" county="Ratchaburi Province" municipality="Chom Bueang District" name="Tham Khao Thalu Temple">Tham Khao Thalu Temple</location>
|
|
,
|
|
<collectingMunicipality>Chom Bueang District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2006-12-09">9 Dec. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2009-12-09" collectorName="Tham Khao Thalu Temple" country="Myanmar" location="Chom Bueang District" municipality="Ratchaburi Province" specimenCount="1">
|
|
CUMZ 12019 (21 shells) from
|
|
<collectorName>Tham Khao Thalu Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:6F38B9E2E6D9A1C4D861B8CAEA87A70C" country="Myanmar" municipality="Ratchaburi Province" name="Chom Bueang District">Chom Bueang District</location>
|
|
,
|
|
<collectingMunicipality>Ratchaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2009-12-09">9 Dec. 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2017-05-08" collectorName="Fig." country="Myanmar" county="Ratchaburi Province" location="Buri Ratchawanaram Temple" municipality="Pak Tho District" specimenCount="6">
|
|
CUMZ 12020 (4 shells and
|
|
<specimenCount type="generic">6 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Fig.</collectorName>
|
|
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">25A</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:05F52406279442C30BF7702955CCBC50" country="Myanmar" county="Ratchaburi Province" municipality="Pak Tho District" name="Buri Ratchawanaram Temple">Buri Ratchawanaram Temple</location>
|
|
,
|
|
<collectingMunicipality>Pak Tho District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2017-05-08">8 May 2017</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-08-18" country="Myanmar" county="Ratchaburi Province" location="Buri Ratchawanaram Temple" municipality="Pak Tho District" specimenCount="1">
|
|
CUMZ 12021 (20 shells and
|
|
<specimenCount type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:923BA5CE72B828BA62D735B8A3D3AFD5" country="Myanmar" county="Ratchaburi Province" municipality="Pak Tho District" name="Buri Ratchawanaram Temple">Buri Ratchawanaram Temple</location>
|
|
,
|
|
<collectingMunicipality>Pak Tho District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2020-08-18">18 Aug. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2019-08-18" collectorName="Fig." country="Myanmar" county="Ratchaburi Province" location="Golden Dragon Cave" municipality="Pak Tho District" specimenCount="5">
|
|
CUMZ 12022 (
|
|
<specimenCount type="generic">5 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Fig.</collectorName>
|
|
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">25B</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:18E4645F1C44CE967CAC46ADC96CE929" country="Myanmar" county="Ratchaburi Province" municipality="Pak Tho District" name="Golden Dragon Cave">Golden Dragon Cave</location>
|
|
,
|
|
<collectingMunicipality>Pak Tho District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2019-08-18">18 Aug. 2019</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2007-04-21" collectorName="Figs" country="Myanmar" location="Bang Saphan District" municipality="Prachub Kirikhan Province" specimenCount="1">
|
|
CUMZ 12023 (17 shells;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21C</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23C</figureCitation>
|
|
) from Tham Khiriwong Temple,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:47CF34F5E00BC3137795A7547A77F428" country="Myanmar" municipality="Prachub Kirikhan Province" name="Bang Saphan District">Bang Saphan District</location>
|
|
,
|
|
<collectingMunicipality>Prachub Kirikhan Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2007-04-21">21 Apr. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2019-07-29" collectorName="Tham Khiriwong Temple" country="Myanmar" location="Bang Saphan District" municipality="Prachub Kirikhan Province" specimenCount="1">
|
|
CUMZ 12024 (147 shells) from
|
|
<collectorName>Tham Khiriwong Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:5D0A27A4465E4E6234BAC370B9F1CB4B" country="Myanmar" municipality="Prachub Kirikhan Province" name="Bang Saphan District">Bang Saphan District</location>
|
|
,
|
|
<collectingMunicipality>Prachub Kirikhan Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2019-07-29">29 July 2019</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-08-24" collectorName="Tham Thep Nimit Temple" country="Myanmar" location="Pak Chong District" municipality="Nakhon Ratchasima Province" specimenCount="1">
|
|
CUMZ 12169 (10 shells) from
|
|
<collectorName>Tham Thep Nimit Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:3482F7EF37CFCEF70560A24194D8CE45" country="Myanmar" municipality="Nakhon Ratchasima Province" name="Pak Chong District">Pak Chong District</location>
|
|
,
|
|
<collectingMunicipality>Nakhon Ratchasima Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2020-08-24">24 Aug. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2006-03-13" country="Myanmar" county="Chonburi Province" location="Tham Khao Cha Ang On Temple" municipality="Bo Thong District" specimenCount="1">
|
|
CUMZ 12025 (28 shells) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:D83DDE694FDE763CD586AC0D9E694C10" country="Myanmar" county="Chonburi Province" municipality="Bo Thong District" name="Tham Khao Cha Ang On Temple">Tham Khao Cha Ang On Temple</location>
|
|
,
|
|
<collectingMunicipality>Bo Thong District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2006-03-13">13 Mar. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2006-08-17" collectorName="Figs" country="Myanmar" county="Chonburi Province" location="Tham Khao Cha Ang On Temple" municipality="Bo Thong District" specimenCount="23">
|
|
CUMZ 12026 (28 shells and
|
|
<specimenCount type="generic">23 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21E</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23D</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:AED7EE91FDB4C53A8EFB9CAFABA20BBD" country="Myanmar" county="Chonburi Province" municipality="Bo Thong District" name="Tham Khao Cha Ang On Temple">Tham Khao Cha Ang On Temple</location>
|
|
,
|
|
<collectingMunicipality>Bo Thong District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2006-08-17">17 Aug. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2008-05-09" country="Myanmar" location="Bo Thong District" municipality="Chonburi Province" specimenCount="1">
|
|
CUMZ 12028 (34 shells) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:046BF8165D0DD2ADEE3B2D3727BCC743" country="Myanmar" municipality="Chonburi Province" name="Bo Thong District">Bo Thong District</location>
|
|
,
|
|
<collectingMunicipality>Chonburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2008-05-09">9 May 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2020-09-19" collectorName="Phromawat Temple" country="Myanmar" location="Si Racha District" municipality="Chonburi Province" specimenCount="2">
|
|
CUMZ 12027 (
|
|
<specimenCount type="generic">2 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName>Phromawat Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:89C438925D5D5ED3E71334F50D8C8A74" country="Myanmar" municipality="Chonburi Province" name="Si Racha District">Si Racha District</location>
|
|
,
|
|
<collectingMunicipality>Chonburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2020-09-19">19 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2010-10-23" collectorName="Tham Khao Loi Temple" country="Myanmar" location="Khao Chamao District" municipality="Rayong Province" specimenCount="1">
|
|
CUMZ 12174 (1 shell) from
|
|
<collectorName>Tham Khao Loi Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:8C5408AEC44EEC57FC6B882C97BDFBDE" country="Myanmar" municipality="Rayong Province" name="Khao Chamao District">Khao Chamao District</location>
|
|
,
|
|
<collectingMunicipality>Rayong Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2010-10-23">23 Oct. 2010</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2012-05-21" collectorName="Fig." country="Myanmar" county="Chachoengsao Province" location="Khao Tham Raet Temple" municipality="Tha Takiap District" specimenCount="10">
|
|
CUMZ 12029 (85 shells and
|
|
<specimenCount type="generic">10 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Fig.</collectorName>
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8G, H</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:AF158BCED7B44D87E9D1E965D9A98C52" country="Myanmar" county="Chachoengsao Province" municipality="Tha Takiap District" name="Khao Tham Raet Temple">Khao Tham Raet Temple</location>
|
|
,
|
|
<collectingMunicipality>Tha Takiap District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Chachoengsao Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2012-05-21">21 May 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2018-03-01" collectorName="Khao Tham Raet Temple" country="Myanmar" location="Tha Takiap District" municipality="Chachoengsao Province" specimenCount="1">
|
|
CUMZ 12030 (3 shells) from
|
|
<collectorName>Khao Tham Raet Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:424F18465441B8527D56966F908844B9" country="Myanmar" municipality="Chachoengsao Province" name="Tha Takiap District">Tha Takiap District</location>
|
|
,
|
|
<collectingMunicipality>Chachoengsao Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2018-03-01">1 Mar. 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2000-04-07" collectorName="Figs" country="Myanmar" location="Khao Chakan District" municipality="Sa Kaeo Province" specimenCount="1">
|
|
CUMZ 12031 (43 shells;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21L</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23E</figureCitation>
|
|
) from Tham Khao Chakan Temple,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:D02B1FBCF5464D182E0BB6676F37BD42" country="Myanmar" municipality="Sa Kaeo Province" name="Khao Chakan District">Khao Chakan District</location>
|
|
,
|
|
<collectingMunicipality>Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2000-04-07">7 Apr. 2000</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2018-02-25" collectorName="Tham Khao Chakan Temple" country="Myanmar" location="Khao Chakan District" municipality="Sa Kaeo Province" specimenCount="7">
|
|
CUMZ 12032 (
|
|
<specimenCount type="generic">7 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName>Tham Khao Chakan Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:EC32765B241B884683E80B23103FDB18" country="Myanmar" municipality="Sa Kaeo Province" name="Khao Chakan District">Khao Chakan District</location>
|
|
,
|
|
<collectingMunicipality>Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2018-02-25">25 Feb. 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2008-11-02" collectorName="Tham Khao Maka Temple" country="Myanmar" location="Mueang Sa Kaeo District" municipality="Sa Kaeo Province" specimenCount="1">
|
|
CUMZ 12033 (2 shells) from
|
|
<collectorName>Tham Khao Maka Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:810B7821D50D150C529009752F02AAAD" country="Myanmar" municipality="Sa Kaeo Province" name="Mueang Sa Kaeo District">Mueang Sa Kaeo District</location>
|
|
,
|
|
<collectingMunicipality>Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2008-11-02">2 Nov. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2018-05-21" collectorName="Khao Pha Pheung Temple" country="Myanmar" location="Klong Had District" municipality="Sra Keo Province" specimenCount="1">
|
|
CUMZ 12034 (2 shells) from
|
|
<collectorName>Khao Pha Pheung Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:C29BB84C9CDABA8AC50EDD26B9CD5A44" country="Myanmar" municipality="Sra Keo Province" name="Klong Had District">Klong Had District</location>
|
|
,
|
|
<collectingMunicipality>Sra Keo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2018-05-21">21 May 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2007-03-04" collectorName="Na Mueang Waterfall" country="Myanmar" location="Ko Samui District" municipality="Surat Thani Province" specimenCount="1">
|
|
CUMZ 12035 (
|
|
<specimenCount type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<collectorName>Na Mueang Waterfall</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:456FA03FBD025C6BA868454C22FCFFE9" country="Myanmar" municipality="Surat Thani Province" name="Ko Samui District">Ko Samui District</location>
|
|
,
|
|
<collectingMunicipality>Surat Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2007-03-04">4 Mar. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2007-03-05" country="Myanmar" county="Surat Thani Province" location="Wua Ta Lap Island" municipality="Ko Samui District" specimenCount="1">
|
|
CUMZ 12036 (10 shells) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:22F7E8E43133B334F33DA398FB233FF5" country="Myanmar" county="Surat Thani Province" municipality="Ko Samui District" name="Wua Ta Lap Island">Wua Ta Lap Island</location>
|
|
,
|
|
<collectingMunicipality>Ko Samui District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Surat Thani Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2007-03-05">5 Mar. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2009-06-06" collectorName="Figs" country="Myanmar" county="Surat Thani Province" location="Wua Ta Lap Island" municipality="Ko Samui District" specimenCount="2">
|
|
CUMZ 12037 (1 shell and
|
|
<specimenCount type="generic">2 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21K</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23F</figureCitation>
|
|
,
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|
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">25C</figureCitation>
|
|
) from
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:7F331EE4AF9DC1C59C0918F27452E889" country="Myanmar" county="Surat Thani Province" municipality="Ko Samui District" name="Wua Ta Lap Island">Wua Ta Lap Island</location>
|
|
,
|
|
<collectingMunicipality>Ko Samui District</collectingMunicipality>
|
|
,
|
|
<collectingCounty>Surat Thani Province</collectingCounty>
|
|
,
|
|
<collectingDate value="2009-06-06">6 June 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2006-10-06" collectorName="Figs" country="Myanmar" location="Mueang Krabi District" municipality="Krabi Province" specimenCount="1">
|
|
CUMZ 12038 (4 shells;
|
|
<collectorName>Figs</collectorName>
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21D</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">24A</figureCitation>
|
|
) from Tham Suea Temple,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F1E617E913559D5635B6CC7F38A6E8CE" country="Myanmar" municipality="Krabi Province" name="Mueang Krabi District">Mueang Krabi District</location>
|
|
,
|
|
<collectingMunicipality>Krabi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2006-10-06">6 Oct. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2015-08-31" collectorName="Khao Noi Phothiyan Temple" country="Myanmar" location="Mueang Satul District" municipality="Satul Province" specimenCount="1">
|
|
CUMZ 12039 (4 shells) from
|
|
<collectorName>Khao Noi Phothiyan Temple</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:C8F64BF6B66466976826ECBB7AC02F35" country="Myanmar" municipality="Satul Province" name="Mueang Satul District">Mueang Satul District</location>
|
|
,
|
|
<collectingMunicipality>Satul Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2015-08-31">31 Aug. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation collectingDate="2007-01-23" collectorName="Khao Rup Chang" country="Myanmar" location="Mueang Songkhla District" municipality="Songkhla Province" specimenCount="1">
|
|
CUMZ 12040 (1 shell) from
|
|
<collectorName>Khao Rup Chang</collectorName>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F603591020FB75FEFD9BBF0B77EBA8AD" country="Myanmar" municipality="Songkhla Province" name="Mueang Songkhla District">Mueang Songkhla District</location>
|
|
,
|
|
<collectingMunicipality>Songkhla Province</collectingMunicipality>
|
|
,
|
|
<collectingDate value="2007-01-23">23 Jan. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">Shell ovate; last whorl ca. three quarters of shell height. Apertural lip slightly thickened, not expanded. Both parietal and columellar teeth fin-shaped and slightly thickened; parietal tooth covering posterior canal; columellar tooth next to slit-like anterior canal.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pupina artata</emphasis>
|
|
</taxonomicName>
|
|
is most similar to
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. pallens" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="pallens">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. pallens</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="limitanea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. limitanea</emphasis>
|
|
</taxonomicName>
|
|
in shell shape, but different from
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. pallens" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="pallens">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. pallens</emphasis>
|
|
</taxonomicName>
|
|
in that the basal position of the apertural lip is not widened, and different from
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="limitanea">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. limitanea</emphasis>
|
|
</taxonomicName>
|
|
by a longer last whorl, and parietal and columellar teeth and apertural lip less thickened.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Peninsular Malaysia, Myanmar (
|
|
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Solem, A" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B146" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
|
), and throughout Thailand except in the northeastern region.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
The type specimen of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata subsp. blanfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" rank="subSpecies" species="Artata" subSpecies="blanfordiana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. artata blanfordiana</emphasis>
|
|
</taxonomicName>
|
|
could not be located, so the validity of this subspecies is still unknown. The specimen identified as
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="peguensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. peguensis</emphasis>
|
|
</taxonomicName>
|
|
and figured in
|
|
<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen (1897</bibRefCitation>
|
|
: pl. 69, fig. 3, 3a-d) from Kama on the right bank of the Irrawaddy River, Pegu is different from the holotype of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="peguensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. peguensis</emphasis>
|
|
</taxonomicName>
|
|
(see
|
|
<bibRefCitation author="Tripathy, B" journalOrPublisher="Records of the Zoological Survey of India" pageId="0" pageNumber="1" pagination="508 - 510" refId="B177" refString="Tripathy, B, Sajan, S, 2019. Pupina peguensis Benson, 1860 (Mollusca: Pupinidae): Tracing a long lost molluscan type. Records of the Zoological Survey of India 119: 508 - 510" title="Pupina peguensis Benson, 1860 (Mollusca: Pupinidae): Tracing a long lost molluscan type." volume="119" year="2019">Tripathy and Sajan 2019</bibRefCitation>
|
|
), but similar to the type specimen of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
. Thus, this specimen is herein identified as
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
The specimen of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
figured in
|
|
<bibRefCitation author="Maassen, WJM" editor="Falkner, M" journalOrPublisher="ConchBooks, Hackenheim" pageId="0" pageNumber="1" pagination="277 - 290" refId="B88" refString="Maassen, WJM, 2002. Remarks on the Pupinidae of Sumatra (Indonesia) with description of two new species (Gastropoda, Prosobranchia: Pupinidae). In: Falkner, M, Groh, K, Speight, MCD, Eds., Collectanea Malacologica Festschrift fuer Gerhard Falkner. ConchBooks, Hackenheim: 277 - 290" title="Remarks on the Pupinidae of Sumatra (Indonesia) with description of two new species (Gastropoda, Prosobranchia: Pupinidae)." volumeTitle="Collectanea Malacologica Festschrift fuer Gerhard Falkner." year="2002">Maassen (2002</bibRefCitation>
|
|
: text-fig. 3) from Sumatra should constitute a different species as it is different from the syntype figured here in having a smaller, sharper parietal tooth revealing the posterior canal and an ear-lobe-shaped columellar tooth covering the anterior canal. Thus, those specimens should belong to the
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. arula" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="arula">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. arula</emphasis>
|
|
</taxonomicName>
|
|
species group instead (see below).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
All specimens from Thailand with a slightly thickened, fin-shaped parietal tooth covering the posterior canal are herein identified as
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
. However, these specimens exhibit a variable shell size (smaller with shell height 5.4 mm, shell width 3.5 mm, to larger with shell height 8.4 mm; shell width 5.9 mm; Fig.
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21A-M</figureCitation>
|
|
). The shell shape is also variable from ovate which is similar to the syntype (Fig.
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21A</figureCitation>
|
|
), to more globose (Fig.
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21F</figureCitation>
|
|
) or more elongate (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21L</figureCitation>
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). In addition, these specimens exhibit a variation in length, outer curvature and thickness of the parietal tooth, and body colour. There is also a case of different shell colour morphs (brown and grey) within the same population (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8G, H</figureCitation>
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). Therefore, DNA data is needed to reveal the extent of genetic differentiation or cryptic diversity within the
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
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<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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morphotype.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |