354 lines
50 KiB
XML
354 lines
50 KiB
XML
<document id="F00F52888190962EC01C20F72AC14845" ID-DOI="10.5281/zenodo.181452" ID-GBIF-Dataset="0e6e9d41-cb9a-4ec3-a951-bf5f2eb06577" ID-ISSN="1175-5326" ID-Zenodo-Dep="181452" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459833984607" checkinUser="plazi" docAuthor="Naruse, Tohru, Ng, Peter K. L. & Guinot, Danièle" docDate="2008" docId="9C7287B09944FFAAC195D071FE7EFEC1" docLanguage="en" docName="zt01739p040.pdf" docOrigin="Zootaxa 1739" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Sulaplax Naruse, Ng & Guinot, 2008, gen. nov." docType="treatment" docVersion="10" lastPageNumber="26" masterDocId="604BFFC89947FFACC102D56EFFB9FFEB" masterDocTitle="Two new genera and two new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on Cancrocaeca Ng, 1991" masterLastPageNumber="40" masterPageNumber="21" pageNumber="24" updateTime="1698223242837" updateUser="plazi">
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<mods:titleInfo id="BB78C08595A2DE4D64DD8709AE98661A">
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<mods:title id="1BEB1FE272200F78494021F185964F38">Two new genera and two new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on Cancrocaeca Ng, 1991</mods:title>
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<mods:namePart id="AA2AC6072BBC42075788983C56C20B05">Naruse, Tohru</mods:namePart>
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<mods:namePart id="A986270E0F0A4683D3630AF1638A6BD6">Ng, Peter K. L.</mods:namePart>
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<mods:roleTerm id="8F4C3E0D1406B14902A83160DE1235E4">Author</mods:roleTerm>
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<mods:namePart id="B8C236E48D5836475D63D75251CAED50">Guinot, Danièle</mods:namePart>
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<mods:typeOfResource id="7CAF240309E94D70BFF56E65F95FADA0">text</mods:typeOfResource>
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<mods:title id="159DDE61B5C08148AEDA600F0B8EFF28">Zootaxa</mods:title>
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<mods:part id="12F75752A43239CE7541F126D297240A">
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<mods:date id="336AB1FB573A0FC5D6C7F8E32B0B6155">2008</mods:date>
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<mods:detail id="AFF4251811DC7C4921BB6FFFB2D92B24" type="volume">
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<mods:number id="EF30E7A1800E0FB677D82CB78EA05EE7">1739</mods:number>
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<mods:classification id="ED457B56D4ACDCA2C225AA26490D7AA8">journal article</mods:classification>
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<mods:identifier id="186D9F64E70F85294450BF59FB5FE182" type="DOI">10.5281/zenodo.181452</mods:identifier>
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<mods:identifier id="C950045B8E0C8F64BE7F98EF8474EF41" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6CB1AF13D18C899759370BFE82B15CDB" type="Zenodo-Dep">181452</mods:identifier>
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<treatment id="9C7287B09944FFAAC195D071FE7EFEC1" ID-DOI="http://doi.org/10.5281/zenodo.6233822" ID-GBIF-Taxon="119358435" ID-Zenodo-Dep="6233822" LSID="urn:lsid:zoobank.org:act:407D63BB-80DA-43DD-868B-2D8AFA16E0AF" httpUri="http://treatment.plazi.org/id/9C7287B09944FFAAC195D071FE7EFEC1" lastPageId="6" lastPageNumber="26" pageId="3" pageNumber="24">
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<subSubSection id="5CC1652D9944FFAFC195D071FECEFAD2" box="[151,375,1311,1337]" pageId="3" pageNumber="24" type="nomenclature">
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<paragraph id="146436A69944FFAFC195D071FECEFAD2" blockId="3.[151,375,1311,1337]" box="[151,375,1311,1337]" pageId="3" pageNumber="24">
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<heading id="4F2C81CA9944FFAFC195D071FECEFAD2" bold="true" box="[151,375,1311,1337]" fontSize="11" level="1" pageId="3" pageNumber="24" reason="1">
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<emphasis id="26AFEAB49944FFAFC195D071FECEFAD2" bold="true" box="[151,375,1311,1337]" pageId="3" pageNumber="24">
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<taxonomicName id="D3DB4D259944FFAFC195D071FEBAFAD2" ID-CoL="7Q3BP" box="[151,259,1311,1337]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="26AFEAB49944FFAFC195D071FEBAFAD2" bold="true" box="[151,259,1311,1337]" italics="true" pageId="3" pageNumber="24">Sulaplax</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3D9C57CF9944FFAFC008D071FECEFAD2" box="[266,375,1311,1337]" pageId="3" pageNumber="24" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5CC1652D9944FFAFC195D004FC2DFA6F" box="[151,916,1386,1412]" pageId="3" pageNumber="24" type="materials_examined">
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<paragraph id="146436A69944FFAFC195D004FC2DFA6F" blockId="3.[151,1437,1386,2012]" box="[151,916,1386,1412]" pageId="3" pageNumber="24">
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<emphasis id="26AFEAB49944FFAFC195D004FE80FA6F" bold="true" box="[151,313,1386,1412]" pageId="3" pageNumber="24">
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<typeStatus id="CB6088049944FFAFC195D004FF6DFA6F" box="[151,212,1386,1412]" pageId="3" pageNumber="24">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="D3DB4D259944FFAFC042D005FE47FA6F" box="[320,510,1387,1412]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="ensifer" status="sp. nov.">
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<emphasis id="26AFEAB49944FFAFC042D005FE47FA6F" box="[320,510,1387,1412]" italics="true" pageId="3" pageNumber="24">Sulaplax ensifer</emphasis>
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</taxonomicName>
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<emphasis id="26AFEAB49944FFAFC307D004FDC4FA6F" bold="true" box="[517,637,1386,1412]" pageId="3" pageNumber="24">
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<taxonomicNameLabel id="3D9C57CF9944FFAFC307D004FDC4FA6F" box="[517,637,1386,1412]" pageId="3" pageNumber="24" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="5CC1652D9944FFA9C1C7D0FCFDB8FE91" lastPageId="5" lastPageNumber="26" pageId="3" pageNumber="24" type="diagnosis">
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<paragraph id="146436A69944FFAFC1C7D0FCFC16F9A7" blockId="3.[151,1437,1386,2012]" pageId="3" pageNumber="24">
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<emphasis id="26AFEAB49944FFAFC1C7D0FCFEFCFA47" bold="true" box="[197,325,1426,1452]" pageId="3" pageNumber="24">Diagnosis.</emphasis>
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Carapace subcircular, wider than long, dorsal surface flat, surrounded by continuous rim, regions clearly separated by grooves; rostrum vestigial, unilobed, deflexed, lateral surface below posterolateral margin without tooth. Anterior surface of cephalothorax (proepistome to epistome, orbit) nearly vertical in lateral view; antennular fossa very shallow, medially separated by thin septum (proepistome); epistome relatively long, with posterior margin produced and concave medially.
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</paragraph>
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<paragraph id="146436A69944FFAFC1C7D334FF05F8D7" blockId="3.[151,1437,1386,2012]" pageId="3" pageNumber="24">Eye slightly visible from dorsal view, base of ocular peduncle fused to orbit, immovable, gradually narrowed distally; cornea without trace of black pigment. Antennule with swollen basal article, largely protruding from fossa. Antenna with thin basal article, base of basal article located at outer lower region at a distance with respect to base of ocular peduncle, flagellum long. Third maxillipeds narrow, forming wide gape when closed; ischium with narrow, long distal inner extension, merus elongated, with inner, outer margins subparallel.</paragraph>
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<paragraph id="146436A69944FFA8C1C7D224FC16FF31" blockId="3.[151,1437,1386,2012]" lastBlockId="4.[151,1436,152,218]" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24">Male abdominal cavity narrow; sternal button for abdominal locking mechanism undiscernible. Female with wide cephalothorax cavity, margin of cavity connected to margin of abdomen-pleotelson by membrane, forming internal brood cavity; vulva of thoracic sternite 6 without basal mount along an imaginary line joining anterior margin of P2 coxae on medial fused plate of thoracic sternum. Chelipeds slender; chela with flat, blade-like fingers. Ambulatory legs extremely slender; dactyli slightly incurving, ending in sharp spine, without subterminal tooth. Penis located anterior to middle of sternite 8.</paragraph>
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<caption id="40A4662E9943FFA8C195D24CFAEFF895" httpUri="https://zenodo.org/record/181454/files/figure.png" pageId="4" pageNumber="25" targetBox="[229,1353,262,1790]" targetPageId="4">
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<paragraph id="146436A69943FFA8C195D24CFAEFF895" blockId="4.[151,1436,1826,1918]" pageId="4" pageNumber="25">
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<emphasis id="26AFEAB49943FFA8C195D24CFEA0F8D1" bold="true" box="[151,281,1826,1850]" pageId="4" pageNumber="25">FIGURE 2.</emphasis>
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<taxonomicName id="D3DB4D259943FFA8C022D24CFD11F8D2" authority="Ng, 1991" authorityName="Ng" authorityYear="1991" box="[288,680,1826,1849]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="xenomorpha">
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<emphasis id="26AFEAB49943FFA8C022D24CFD80F8D2" box="[288,569,1826,1849]" italics="true" pageId="4" pageNumber="25">Cancrocaeca xenomorpha</emphasis>
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Ng, 1991
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</taxonomicName>
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: a, thoracic sternum of female; b–d, female abdomen-pleotelson, outer surface (b and d) and inner surface (c); e–h, second to fifth right pleopods, anterior view. Ovigerous female, ZRC 2007.0118, 4.1 × 5.2 mm. Numbers and “V” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm.
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</paragraph>
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</caption>
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<paragraph id="146436A69943FFA9C1C7D2DCFDB8FE91" blockId="4.[151,1436,1970,2036]" lastBlockId="5.[151,1437,152,2018]" lastPageId="5" lastPageNumber="26" pageId="4" pageNumber="25">Male abdomen-pleotelson 4-segmented, third to fifth segments fused; first segment with protuberances on disto-lateral angles; fused segments narrowed in distal half, proximal half with subparallel lateral margins; pleotelson distinctly narrower than half width of abdomen. G1 strongly bent medially, tapering simply, dorsal side of inner margin lined with long, stiff setae; suture running from dorsal side of proximal part to outer margin of distal part; G2 with dilated base, proximal outer angle slightly visible posteriorly when abdomen closed. Female abdomen with an undivided plate formed by fusion of third segment to pleotelson; pleotelson only demarcated by shallow groove on outer side; second to fifth pleopods short, biramous, entirely enclosed inside abdominal brood cavity.</paragraph>
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</subSubSection>
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<subSubSection id="5CC1652D9942FFA9C1C7D4E9FB2DFE21" pageId="5" pageNumber="26" type="etymology">
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<paragraph id="146436A69942FFA9C1C7D4E9FB2DFE21" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="26">
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<emphasis id="26AFEAB49942FFA9C1C7D4E9FEEBFE4A" bold="true" box="[197,338,391,417]" pageId="5" pageNumber="26">Etymology.</emphasis>
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The genus name,
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<taxonomicName id="D3DB4D259942FFA9C32FD4E6FD2AFE4A" box="[557,659,392,417]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="26AFEAB49942FFA9C32FD4E6FD2AFE4A" box="[557,659,392,417]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
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</taxonomicName>
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, is derived from the Sulawesi, and
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<emphasis id="26AFEAB49942FFA9C538D4E6FBD4FE4A" box="[1082,1133,392,417]" italics="true" pageId="5" pageNumber="26">plax</emphasis>
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, “plate,” the suffix of the closely related genus
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<taxonomicName id="D3DB4D259942FFA9C096D4DEFDECFE22" box="[404,597,432,457]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="26AFEAB49942FFA9C096D4DEFDECFE22" box="[404,597,432,457]" italics="true" pageId="5" pageNumber="26">Neorhynchoplax</emphasis>
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</taxonomicName>
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, alluding to their resemblance. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="5CC1652D9942FFAAC1C7D4B9FE7EFEC1" lastPageId="6" lastPageNumber="27" pageId="5" pageNumber="26" type="discussion">
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<paragraph id="146436A69942FFA9C1C7D4B9FE8CFC11" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="26">
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<emphasis id="26AFEAB49942FFA9C1C7D4B9FE87FE1A" bold="true" box="[197,318,471,497]" pageId="5" pageNumber="26">Remarks.</emphasis>
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Cavernicolous crabs often show varying degrees of characteristic morphological adaptations for a hypogeal lifestyle: long and slender ambulatory legs, depigmentation of the legs and carapace, and reduction of the cornea and eyes (
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<bibRefCitation id="704A4B579942FFA9C322D746FD58FDA9" author="Rodriguez" box="[544,737,552,578]" pageId="5" pageNumber="40" refString="Rodriguez, G. (1985) A new cavernicolous crab (Crustacea, Decapoda, Pseudothelphusidae) from Colombia. Bioloski Vestnik, 2, 73 - 80." type="journal article" year="1985">Rodríguez 1985</bibRefCitation>
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;
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<bibRefCitation id="704A4B579942FFA9C3EFD746FC3AFDA9" author="Guinot" box="[749,899,552,578]" pageId="5" pageNumber="39" refString="Guinot, D. (1988) Les crabes cavernicoles du monde. Memoires de Biospeologie, 15, 3 - 40." type="journal article" year="1988">Guinot 1988</bibRefCitation>
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;
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<bibRefCitation id="704A4B579942FFA9C28DD746FC40FDA9" author="Ng" box="[911,1017,552,578]" pageId="5" pageNumber="39" refString="Chuang, C. T. N. & Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Ng 1991</bibRefCitation>
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).
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<taxonomicName id="D3DB4D259942FFA9C50ED746FBCAFDAA" box="[1036,1139,552,577]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="26AFEAB49942FFA9C50ED746FBCAFDAA" box="[1036,1139,552,577]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
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</taxonomicName>
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<emphasis id="26AFEAB49942FFA9C578D749FB51FDAA" bold="true" box="[1146,1256,551,577]" pageId="5" pageNumber="26">
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<taxonomicNameLabel id="3D9C57CF9942FFA9C578D749FB51FDAA" box="[1146,1256,551,577]" pageId="5" pageNumber="26" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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is superficially similar to
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<taxonomicName id="D3DB4D259942FFA9C010D73EFD91FD81" authority="Ng, 1991" authorityName="Ng" authorityYear="1991" box="[274,552,592,618]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="26AFEAB49942FFA9C010D73EFE14FD82" box="[274,429,592,617]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
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<bibRefCitation id="704A4B579942FFA9C0B4D73EFD91FD81" author="Ng" box="[438,552,592,618]" pageId="5" pageNumber="39" refString="Chuang, C. T. N. & Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Ng, 1991</bibRefCitation>
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</taxonomicName>
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, in these cave-adapted characters.
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<taxonomicName id="D3DB4D259942FFA9C2CAD73EFB96FD82" box="[968,1071,592,617]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="26AFEAB49942FFA9C2CAD73EFB96FD82" box="[968,1071,592,617]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
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</taxonomicName>
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, however, differs significantly from
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<taxonomicName id="D3DB4D259942FFA9C1DBD716FECDFD7A" box="[217,372,632,657]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="26AFEAB49942FFA9C1DBD716FECDFD7A" box="[217,372,632,657]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
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</taxonomicName>
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in its relatively slender and gradually tapering G1 (
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<figureCitation id="8CE02A239942FFA9C2E7D716FB94FD79" box="[997,1069,632,658]" captionStart="FIGURE 5" captionStartId="8.[151,255,1927,1951]" captionTargetBox="[179,1354,202,1883]" captionTargetId="figure@8.[151,1436,194,2013]" captionTargetPageId="8" captionText="FIGURE 5. Sulaplax ensifer spec. nov.: a, left P 3; b, male left chela; c, female left chela; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view. a, c, female paratype, ZRC 2007.0119, 5.0 × 6.5 mm; b, d – f, male holotype," httpUri="https://zenodo.org/record/181457/files/figure.png" pageId="5" pageNumber="26">Fig. 5</figureCitation>
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e, f), abruptly narrowed 4-segmented male abdomen with the third to fifth segments fused (
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<figureCitation id="8CE02A239942FFA9C264D7CEFC15FD51" box="[870,940,672,698]" captionStart="FIGURE 5" captionStartId="8.[151,255,1927,1951]" captionTargetBox="[179,1354,202,1883]" captionTargetId="figure@8.[151,1436,194,2013]" captionTargetPageId="8" captionText="FIGURE 5. Sulaplax ensifer spec. nov.: a, left P 3; b, male left chela; c, female left chela; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view. a, c, female paratype, ZRC 2007.0119, 5.0 × 6.5 mm; b, d – f, male holotype," httpUri="https://zenodo.org/record/181457/files/figure.png" pageId="5" pageNumber="26">Fig. 5</figureCitation>
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d), the female vulva lacking a basal mount (
|
||
<figureCitation id="8CE02A239942FFA9C19DD7A6FF51FD09" box="[159,232,712,738]" captionStart="FIGURE 6" captionStartId="9.[151,255,1593,1617]" captionTargetBox="[156,1436,194,1553]" captionTargetId="figure@9.[151,1436,194,1569]" captionTargetPageId="9" captionText="FIGURE 6. Sulaplax ensifer spec. nov.: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer surface (b and d) and inner surface (c); e – h, second to fifth right pleopods, anterior view. a – d, female paratype, ZRC 2007.0119, 5.0 × 6.5 mm; e – h, female, MNHN-B 30396, 4.0 × 5.0 mm. Scale bars, 1 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively." httpUri="https://zenodo.org/record/181458/files/figure.png" pageId="5" pageNumber="26">Fig. 6</figureCitation>
|
||
a), and the short biramous female pleopods on the second to fifth abdominal segments (
|
||
<figureCitation id="8CE02A239942FFA9C5FBD7A6FAF9FD09" box="[1273,1344,712,738]" captionStart="FIGURE 6" captionStartId="9.[151,255,1593,1617]" captionTargetBox="[156,1436,194,1553]" captionTargetId="figure@9.[151,1436,194,1569]" captionTargetPageId="9" captionText="FIGURE 6. Sulaplax ensifer spec. nov.: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer surface (b and d) and inner surface (c); e – h, second to fifth right pleopods, anterior view. a – d, female paratype, ZRC 2007.0119, 5.0 × 6.5 mm; e – h, female, MNHN-B 30396, 4.0 × 5.0 mm. Scale bars, 1 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively." httpUri="https://zenodo.org/record/181458/files/figure.png" pageId="5" pageNumber="26">Fig. 6</figureCitation>
|
||
e–h). In
|
||
<taxonomicName id="D3DB4D259942FFA9C195D79EFE8BFCE2" box="[151,306,752,777]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C195D79EFE8BFCE2" box="[151,306,752,777]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
, the G1 is stout and with a complex apex (
|
||
<figureCitation id="8CE02A239942FFA9C23AD79EFCC4FCE1" box="[824,893,752,778]" captionStart="FIGURE 1" captionStartId="2.[151,255,1767,1791]" captionTargetBox="[168,1373,457,1723]" captionTargetId="figure@2.[151,1436,431,1746]" captionTargetPageId="2" captionText="FIGURE 1. Cancrocaeca xenomorpha Ng, 1991: a, carapace, dorsal view; b, carapace, frontal view; c, male abdomenpleotelson; d, right G 1, dorsal view; e, distal part of right G 1, ventral view. Male, ZRC 2007.0118, 3.7 × 5.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181453/files/figure.png" pageId="5" pageNumber="26">Fig. 1</figureCitation>
|
||
d, e), the male abdomen is fully 6-segmented without fusion (
|
||
<figureCitation id="8CE02A239942FFA9C057D676FE23FCD9" box="[341,410,792,818]" captionStart="FIGURE 1" captionStartId="2.[151,255,1767,1791]" captionTargetBox="[168,1373,457,1723]" captionTargetId="figure@2.[151,1436,431,1746]" captionTargetPageId="2" captionText="FIGURE 1. Cancrocaeca xenomorpha Ng, 1991: a, carapace, dorsal view; b, carapace, frontal view; c, male abdomenpleotelson; d, right G 1, dorsal view; e, distal part of right G 1, ventral view. Male, ZRC 2007.0118, 3.7 × 5.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181453/files/figure.png" pageId="5" pageNumber="26">Fig. 1</figureCitation>
|
||
c), the female vulva bears a basal mount (
|
||
<figureCitation id="8CE02A239942FFA9C294D676FC66FCD9" box="[918,991,792,818]" captionStart="FIGURE 2" captionStartId="4.[151,255,1826,1850]" captionTargetBox="[229,1353,262,1790]" captionTargetId="figure@4.[229,1354,262,1818]" captionTargetPageId="4" captionText="FIGURE 2. Cancrocaeca xenomorpha Ng, 1991: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer surface (b and d) and inner surface (c); e – h, second to fifth right pleopods, anterior view. Ovigerous female, ZRC 2007.0118, 4.1 × 5.2 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181454/files/figure.png" pageId="5" pageNumber="26">Fig. 2</figureCitation>
|
||
a), and the female pleopods are elongated and prominently setose on the second to fifth abdominal segments (
|
||
<figureCitation id="8CE02A239942FFA9C2F3D62EFB8FFCB1" box="[1009,1078,832,858]" captionStart="FIGURE 2" captionStartId="4.[151,255,1826,1850]" captionTargetBox="[229,1353,262,1790]" captionTargetId="figure@4.[229,1354,262,1818]" captionTargetPageId="4" captionText="FIGURE 2. Cancrocaeca xenomorpha Ng, 1991: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer surface (b and d) and inner surface (c); e – h, second to fifth right pleopods, anterior view. Ovigerous female, ZRC 2007.0118, 4.1 × 5.2 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181454/files/figure.png" pageId="5" pageNumber="26">Fig. 2</figureCitation>
|
||
e–h). In addition,
|
||
<taxonomicName id="D3DB4D259942FFA9C404D62EFAD4FCB2" box="[1286,1389,832,857]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C404D62EFAD4FCB2" box="[1286,1389,832,857]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
can be also distinguished from
|
||
<taxonomicName id="D3DB4D259942FFA9C0D7D606FDC9FC6A" box="[469,624,872,897]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C0D7D606FDC9FC6A" box="[469,624,872,897]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a rostrum (
|
||
<figureCitation id="8CE02A239942FFA9C2D5D606FB90FC69" box="[983,1065,872,898]" captionStart="FIGURE 4" captionStartId="7.[151,255,1962,1986]" captionTargetBox="[154,1420,212,1912]" captionTargetId="figure@7.[151,1436,182,1952]" captionTargetPageId="7" captionText="FIGURE 4. Sulaplax ensifer spec. nov.: a, carapace, dorsal view; b, carapace, frontal view; c, right third maxilliped. Female paratype, ZRC 2007.0119, 5.0 × 6.5 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181456/files/figure.png" pageId="5" pageNumber="26">Figs. 4</figureCitation>
|
||
b) (absent in
|
||
<taxonomicName id="D3DB4D259942FFA9C5C1D606FAE7FC6A" box="[1219,1374,872,897]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C5C1D606FAE7FC6A" box="[1219,1374,872,897]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="8CE02A239942FFA9C46FD606FF1AFC41" captionStart="FIGURE 1" captionStartId="2.[151,255,1767,1791]" captionTargetBox="[168,1373,457,1723]" captionTargetId="figure@2.[151,1436,431,1746]" captionTargetPageId="2" captionText="FIGURE 1. Cancrocaeca xenomorpha Ng, 1991: a, carapace, dorsal view; b, carapace, frontal view; c, male abdomenpleotelson; d, right G 1, dorsal view; e, distal part of right G 1, ventral view. Male, ZRC 2007.0118, 3.7 × 5.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181453/files/figure.png" pageId="5" pageNumber="26">Fig. 1</figureCitation>
|
||
a, b), well-defined dorsal carapace regions (
|
||
<figureCitation id="8CE02A239942FFA9C3CBD6FEFC98FC41" box="[713,801,912,938]" captionStart="FIGURE 3" captionStartId="6.[151,255,1533,1557]" captionTargetBox="[186,1403,965,1525]" captionTargetId="figure@6.[186,1403,965,1526]" captionTargetPageId="6" captionText="FIGURE 3. Sulaplax ensifer spec. nov.: Female paratype, ZRC 2007.0119, 5.0 × 6.5 mm." httpUri="https://zenodo.org/record/181455/files/figure.png" pageId="5" pageNumber="26">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="8CE02A239942FFA9C237D6FEFCFDFC41" box="[821,836,912,938]" captionStart="FIGURE 4" captionStartId="7.[151,255,1962,1986]" captionTargetBox="[154,1420,212,1912]" captionTargetId="figure@7.[151,1436,182,1952]" captionTargetPageId="7" captionText="FIGURE 4. Sulaplax ensifer spec. nov.: a, carapace, dorsal view; b, carapace, frontal view; c, right third maxilliped. Female paratype, ZRC 2007.0119, 5.0 × 6.5 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181456/files/figure.png" pageId="5" pageNumber="26">4</figureCitation>
|
||
a) (ill-defined in
|
||
<taxonomicName id="D3DB4D259942FFA9C51ED6FEFB0EFC42" box="[1052,1207,912,937]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C51ED6FEFB0EFC42" box="[1052,1207,912,937]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="8CE02A239942FFA9C5C9D6FEFAAAFC41" box="[1227,1299,912,938]" captionStart="FIGURE 1" captionStartId="2.[151,255,1767,1791]" captionTargetBox="[168,1373,457,1723]" captionTargetId="figure@2.[151,1436,431,1746]" captionTargetPageId="2" captionText="FIGURE 1. Cancrocaeca xenomorpha Ng, 1991: a, carapace, dorsal view; b, carapace, frontal view; c, male abdomenpleotelson; d, right G 1, dorsal view; e, distal part of right G 1, ventral view. Male, ZRC 2007.0118, 3.7 × 5.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181453/files/figure.png" pageId="5" pageNumber="26">Fig. 1</figureCitation>
|
||
a), and the absence of a longitudinal groove on the side walls of the carapace (
|
||
<figureCitation id="8CE02A239942FFA9C2A4D6D6FC4EFC39" box="[934,1015,952,978]" captionStart="FIGURE 4" captionStartId="7.[151,255,1962,1986]" captionTargetBox="[154,1420,212,1912]" captionTargetId="figure@7.[151,1436,182,1952]" captionTargetPageId="7" captionText="FIGURE 4. Sulaplax ensifer spec. nov.: a, carapace, dorsal view; b, carapace, frontal view; c, right third maxilliped. Female paratype, ZRC 2007.0119, 5.0 × 6.5 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181456/files/figure.png" pageId="5" pageNumber="26">Figs. 4</figureCitation>
|
||
a) (present in
|
||
<taxonomicName id="D3DB4D259942FFA9C595D6D6FA8BFC3A" box="[1175,1330,952,977]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C595D6D6FA8BFC3A" box="[1175,1330,952,977]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="8CE02A239942FFA9C443D6D6FA38FC39" box="[1345,1409,952,978]" captionStart="FIGURE 1" captionStartId="2.[151,255,1767,1791]" captionTargetBox="[168,1373,457,1723]" captionTargetId="figure@2.[151,1436,431,1746]" captionTargetPageId="2" captionText="FIGURE 1. Cancrocaeca xenomorpha Ng, 1991: a, carapace, dorsal view; b, carapace, frontal view; c, male abdomenpleotelson; d, right G 1, dorsal view; e, distal part of right G 1, ventral view. Male, ZRC 2007.0118, 3.7 × 5.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181453/files/figure.png" pageId="5" pageNumber="26">Fig. 1</figureCitation>
|
||
a) (see Table 1).
|
||
</paragraph>
|
||
<paragraph id="146436A69942FFA9C1C7D166FE8CF921" blockId="5.[151,1437,152,2018]" pageId="5" pageNumber="26">
|
||
The brooding habits of
|
||
<taxonomicName id="D3DB4D259942FFA9C0D4D166FD84FBCA" box="[470,573,1032,1057]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C0D4D166FD84FBCA" box="[470,573,1032,1057]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
species also differ remarkably from those of
|
||
<taxonomicName id="D3DB4D259942FFA9C549D166FB5FFBCA" box="[1099,1254,1032,1057]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C549D166FB5FFBCA" box="[1099,1254,1032,1057]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
. The ovigerous females of
|
||
<taxonomicName id="D3DB4D259942FFA9C01BD15EFE77FBA2" box="[281,462,1072,1097]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="xenomorpha">
|
||
<emphasis id="26AFEAB49942FFA9C01BD15EFE77FBA2" box="[281,462,1072,1097]" italics="true" pageId="5" pageNumber="26">C. xenomorpha</emphasis>
|
||
</taxonomicName>
|
||
hold the eggs only in the dome-shaped abdomen, which is marginally sheltered by the exopods of the third to fifth pleopods (
|
||
<figureCitation id="8CE02A239942FFA9C394D136FD66FB99" box="[662,735,1112,1138]" captionStart="FIGURE 2" captionStartId="4.[151,255,1826,1850]" captionTargetBox="[229,1353,262,1790]" captionTargetId="figure@4.[229,1354,262,1818]" captionTargetPageId="4" captionText="FIGURE 2. Cancrocaeca xenomorpha Ng, 1991: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer surface (b and d) and inner surface (c); e – h, second to fifth right pleopods, anterior view. Ovigerous female, ZRC 2007.0118, 4.1 × 5.2 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181454/files/figure.png" pageId="5" pageNumber="26">Fig. 2</figureCitation>
|
||
c, e–h). In contrast, the inner surface of the abdomen of
|
||
<taxonomicName id="D3DB4D259942FFA9C484D136FF5EFB72" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="ensifer" status="sp. nov.">
|
||
<emphasis id="26AFEAB49942FFA9C484D136FF5EFB72" italics="true" pageId="5" pageNumber="26">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="26AFEAB49942FFA9C1EDD111FEDEFB72" bold="true" box="[239,359,1151,1177]" pageId="5" pageNumber="26">
|
||
<taxonomicNameLabel id="3D9C57CF9942FFA9C1EDD111FEDEFB72" box="[239,359,1151,1177]" pageId="5" pageNumber="26" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is covered by a thin membrane and the reduced pleopods are present inside the membrane (
|
||
<figureCitation id="8CE02A239942FFA9C19DD1C6FF50FB29" box="[159,233,1192,1218]" captionStart="FIGURE 6" captionStartId="9.[151,255,1593,1617]" captionTargetBox="[156,1436,194,1553]" captionTargetId="figure@9.[151,1436,194,1569]" captionTargetPageId="9" captionText="FIGURE 6. Sulaplax ensifer spec. nov.: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer surface (b and d) and inner surface (c); e – h, second to fifth right pleopods, anterior view. a – d, female paratype, ZRC 2007.0119, 5.0 × 6.5 mm; e – h, female, MNHN-B 30396, 4.0 × 5.0 mm. Scale bars, 1 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively." httpUri="https://zenodo.org/record/181458/files/figure.png" pageId="5" pageNumber="26">Fig. 6</figureCitation>
|
||
c, e–h). The membrane is connected to, and covers the margins of, the cephalothorax cavity and the eggs are thus kept inside the membrane of the abdomen and the cephalothorax cavities (equivalent to an internal brood cavity). Apart from the cave adaptation, the shape of the G1 and the fusion and shape of the male abdominal segments of
|
||
<taxonomicName id="D3DB4D259942FFA9C0B4D04EFDA4FAD2" box="[438,541,1312,1337]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C0B4D04EFDA4FAD2" box="[438,541,1312,1337]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
suggest a close relationship to
|
||
<taxonomicName id="D3DB4D259942FFA9C29ED04EFBE4FAD2" box="[924,1117,1312,1337]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C29ED04EFBE4FAD2" box="[924,1117,1312,1337]" italics="true" pageId="5" pageNumber="26">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
, whose species dwell in a variety of intertidal to freshwater habitats in the Indo-West Pacific (see
|
||
<bibRefCitation id="704A4B579942FFA9C2F6D026FB31FA89" author="Lucas" box="[1012,1160,1352,1378]" pageId="5" pageNumber="40" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas, 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="704A4B579942FFA9C594D026FA36FA89" author="Ng" box="[1174,1423,1352,1378]" pageId="5" pageNumber="40" refString="Ng, P. K. L. & Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement 3, 1 - 82." type="book chapter" year="1996">Ng & Chuang, 1996</bibRefCitation>
|
||
). Furthermore,
|
||
<taxonomicName id="D3DB4D259942FFA9C03BD01EFE19FA62" box="[313,416,1392,1417]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C03BD01EFE19FA62" box="[313,416,1392,1417]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
and some species of
|
||
<taxonomicName id="D3DB4D259942FFA9C39DD01EFCD9FA62" box="[671,864,1392,1417]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C39DD01EFCD9FA62" box="[671,864,1392,1417]" italics="true" pageId="5" pageNumber="26">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
possess an unusual internal brood cavity, which is otherwise known only in
|
||
<taxonomicName id="D3DB4D259942FFA9C0EFD0F6FC6DFA59" authority="Serene, 1971" authorityName="Serene" authorityYear="1971" box="[493,980,1432,1458]" class="Malacostraca" family="Hymenosomatidae" genus="Crustaenia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="palawanensis">
|
||
<emphasis id="26AFEAB49942FFA9C0EFD0F6FCA3FA5A" box="[493,794,1432,1457]" italics="true" pageId="5" pageNumber="26">Crustaenia palawanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="704A4B579942FFA9C22ED0F6FC75FA59" author="Serene" box="[812,972,1432,1458]" pageId="5" pageNumber="40" refString="Serene, R. (1971) Observations preliminaires sur des Brachyoures nouveaux ou mal connus du Sud-Est Asiatique (Crustacea Decapoda). Bulletin du Museum national d'Histoire naturelle, Paris, (2) 42 (5), 903 - 918." type="journal article" year="1971">Serène, 1971</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and some species of
|
||
<taxonomicName id="D3DB4D259942FFA9C5DDD0F6FE2FFA31" authority="A. Milne Edwards, 1873" authorityName="A. Milne Edwards" authorityYear="1873" class="Malacostraca" family="Hymenosomatidae" genus="Elamenopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C5DDD0F6FACDFA5A" box="[1247,1396,1432,1457]" italics="true" pageId="5" pageNumber="26">Elamenopsis</emphasis>
|
||
A. Milne
|
||
<bibRefCitation id="704A4B579942FFA9C1E7D0AEFE2FFA31" author="Edwards" box="[229,406,1472,1498]" pageId="5" pageNumber="40" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Nouvelles Archives du Museum d'Histoire naturelle, 9, 155 - 322, pls. 4 - 18." type="journal article" year="1873">Edwards, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="704A4B579942FFA9C0A7D0AEFD8DFA31" author="Lucas" box="[421,564,1472,1498]" pageId="5" pageNumber="40" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas 1980</bibRefCitation>
|
||
: 191, fig. 8H;
|
||
<bibRefCitation id="704A4B579942FFA9C3DDD0AEFC60FA31" author="Lucas" box="[735,985,1472,1498]" pageId="5" pageNumber="40" refString="Lucas, J. S. & Davie, P. J. F. (1982) Hymenosomatid crabs of Queensland estuaries and tidal mud flats, including descriptions of four new species of Elamenopsis A. Milne-Edwards and a new species of Amarinus. Memoirs of the Queensland Museum, 20 (3), 401 - 419." type="journal article" year="1982">Lucas & Davie 1982</bibRefCitation>
|
||
: 406, 409;
|
||
<bibRefCitation id="704A4B579942FFA9C55FD0AEFAB5FA31" author="Ali" box="[1117,1292,1472,1498]" pageId="5" pageNumber="39" refString="Ali, M. H., Salman, S. D. & Al-Adhub, A. Y. (1995) Population dynamics of the hymenosomatid crab Elamenopsis kempi in a brackish subtidal region of Basrah, Iraq. Scientia Marina, 59 (1), 1 - 13." type="journal article" year="1995">
|
||
Ali
|
||
<emphasis id="26AFEAB49942FFA9C588D0AEFB7FFA32" box="[1162,1222,1472,1497]" italics="true" pageId="5" pageNumber="26">et al.</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
: 3; Davie & Richer de
|
||
<bibRefCitation id="704A4B579942FFA9C013D086FE1EF9E9" author="Forges" box="[273,423,1512,1538]" pageId="5" pageNumber="39" refString="Davie, P. J. F. & Richer de Forges, B. (1996) Two new species of false spider crabs (Crustacea: Brachyura: Hymenosomatidae) from New Caledonia. Memoirs of the Queensland Museum, 39 (2), 257 - 262." type="journal article" year="1996">Forges 1996</bibRefCitation>
|
||
: 261;
|
||
<bibRefCitation id="704A4B579942FFA9C0EDD086FD65F9E9" author="Ng" box="[495,732,1512,1538]" pageId="5" pageNumber="40" refString="Ng, P. K. L. & Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement 3, 1 - 82." type="book chapter" year="1996">Ng & Chuang 1996</bibRefCitation>
|
||
: 60, figs. 25, 26;
|
||
<bibRefCitation id="704A4B579942FFA9C2A9D086FB31F9E9" author="Naruse" box="[939,1160,1512,1538]" pageId="5" pageNumber="40" refString="Naruse, T., Shokita, S. & Kawahara, T. (2005) Neorhynchoplax yaeyamaensis, a new false spider crab (Decapoda: Brachyura: Hymenosomatidae) from the Yaeyama Group, the Ryukyu Islands, Japan. Zootaxa, 877, 1 - 7." type="journal article" year="2005">
|
||
Naruse
|
||
<emphasis id="26AFEAB49942FFA9C504D086FBFBF9EA" box="[1030,1090,1512,1537]" italics="true" pageId="5" pageNumber="26">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
: 5, fig. 1c–e;
|
||
<bibRefCitation id="704A4B579942FFA9C428D086FEB6F9C1" author="Naruse" pageId="5" pageNumber="40" refString="Naruse, T. & Ng, P. K. L. (2007 a) On a new species of Elamenopsis from Singapore, with notes on Crustaenia palawanensis (Serene, 1971) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 55 (1), 121 - 125." type="journal article" year="2007" yearSuffix="a">Naruse & Ng 2007a</bibRefCitation>
|
||
: 124). In contrast, the G1 and the male and female abdomens of
|
||
<taxonomicName id="D3DB4D259942FFA9C514D37EFB08F9C2" box="[1046,1201,1552,1577]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C514D37EFB08F9C2" box="[1046,1201,1552,1577]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
are more similar to those of
|
||
<taxonomicName id="D3DB4D259942FFA9C1FFD356FE38F9BA" box="[253,385,1592,1617]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C1FFD356FE38F9BA" box="[253,385,1592,1617]" italics="true" pageId="5" pageNumber="26">Limnopilos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3DB4D259942FFA9C0BED356FDDBF9BA" box="[444,610,1592,1617]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C0BED356FDDBF9BA" box="[444,610,1592,1617]" italics="true" pageId="5" pageNumber="26">Hymenicoides</emphasis>
|
||
</taxonomicName>
|
||
(see Remarks under
|
||
<taxonomicName id="D3DB4D259942FFA9C25FD356FC41F9BA" box="[861,1016,1592,1617]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C25FD356FC41F9BA" box="[861,1016,1592,1617]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
and Table 1) than they are to those of
|
||
<taxonomicName id="D3DB4D259942FFA9C1B5D30EFEA4F992" box="[183,285,1632,1657]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C1B5D30EFEA4F992" box="[183,285,1632,1657]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
. These relationships imply that troglobitic
|
||
<taxonomicName id="D3DB4D259942FFA9C21AD30EFCC6F992" box="[792,895,1632,1657]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C21AD30EFCC6F992" box="[792,895,1632,1657]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3DB4D259942FFA9C2B5D30EFBEBF992" box="[951,1106,1632,1657]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C2B5D30EFBEBF992" box="[951,1106,1632,1657]" italics="true" pageId="5" pageNumber="26">Cancrocaeca</emphasis>
|
||
</taxonomicName>
|
||
are derived separately from two different ancestors; their superficial similarity is merely a remarkable example of convergence to a hypogeal lifestyle.
|
||
</paragraph>
|
||
<paragraph id="146436A69942FFAAC1C7D3B6FE7EFEC1" blockId="5.[151,1437,152,2018]" lastBlockId="6.[151,1437,152,298]" lastPageId="6" lastPageNumber="27" pageId="5" pageNumber="26">
|
||
The characters which differentiate
|
||
<taxonomicName id="D3DB4D259942FFA9C360D3B6FD70F91A" box="[610,713,1752,1777]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C360D3B6FD70F91A" box="[610,713,1752,1777]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3DB4D259942FFA9C206D3B6FC7CF91A" box="[772,965,1752,1777]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C206D3B6FC7CF91A" box="[772,965,1752,1777]" italics="true" pageId="5" pageNumber="26">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
are mainly in the cephalothorax and its appendages. The anterior surface of the cephalothorax of
|
||
<taxonomicName id="D3DB4D259942FFA9C24AD26EFC16F8F2" box="[840,943,1792,1817]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C24AD26EFC16F8F2" box="[840,943,1792,1817]" italics="true" pageId="5" pageNumber="26">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
is normal in that it appears flat in lateral view (dorso-anterior margin projecting beyond rest of cephalothorax in
|
||
<taxonomicName id="D3DB4D259942FFA9C2EBD246FB13F8AA" box="[1001,1194,1832,1857]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C2EBD246FB13F8AA" box="[1001,1194,1832,1857]" italics="true" pageId="5" pageNumber="26">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
), the proepistome is relatively long and the epistome is relatively long and wide. The base of the antenna is located at outer lower region at a distance with respect to the base of the ocular peduncle (
|
||
<figureCitation id="8CE02A239942FFA9C2A9D216FC48F879" box="[939,1009,1912,1938]" captionStart="FIGURE 4" captionStartId="7.[151,255,1962,1986]" captionTargetBox="[154,1420,212,1912]" captionTargetId="figure@7.[151,1436,182,1952]" captionTargetPageId="7" captionText="FIGURE 4. Sulaplax ensifer spec. nov.: a, carapace, dorsal view; b, carapace, frontal view; c, right third maxilliped. Female paratype, ZRC 2007.0119, 5.0 × 6.5 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181456/files/figure.png" pageId="5" pageNumber="26">Fig. 4</figureCitation>
|
||
b) (placed on just inner lower part of the base of the ocular peduncle in
|
||
<taxonomicName id="D3DB4D259942FFA9C337D2CEFD4FF852" box="[565,758,1952,1977]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C337D2CEFD4FF852" box="[565,758,1952,1977]" italics="true" pageId="5" pageNumber="26">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
). The merus of the third maxilliped is elongated, and its inner and outer margins are subparallel (inner margin convex in
|
||
<taxonomicName id="D3DB4D259942FFA9C298D2A6FBE2F80A" box="[922,1115,1992,2017]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49942FFA9C298D2A6FBE2F80A" box="[922,1115,1992,2017]" italics="true" pageId="5" pageNumber="26">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
). The dactylus of the third maxilliped is also elongated; its length is about twice the length of its propodus (dactylus clearly shorter than twice the length of the propodus in
|
||
<taxonomicName id="D3DB4D259941FFAAC330D5AEFD4AFF32" box="[562,755,192,217]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49941FFAAC330D5AEFD4AFF32" box="[562,755,192,217]" italics="true" pageId="6" pageNumber="27">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="D3DB4D259941FFAAC209D5AEFCCBFF32" box="[779,882,192,217]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49941FFAAC209D5AEFCCBFF32" box="[779,882,192,217]" italics="true" pageId="6" pageNumber="27">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
can be also easily distinguished from
|
||
<taxonomicName id="D3DB4D259941FFAAC42CD5AEFF4DFEEA" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49941FFAAC42CD5AEFF4DFEEA" italics="true" pageId="6" pageNumber="27">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
by its vestigial rostrum (always well developed in
|
||
<taxonomicName id="D3DB4D259941FFAAC24BD586FBB3FEEA" box="[841,1034,232,257]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49941FFAAC24BD586FBB3FEEA" box="[841,1034,232,257]" italics="true" pageId="6" pageNumber="27">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
) and its reduced eyes (well developed in
|
||
<taxonomicName id="D3DB4D259941FFAAC1F7D47EFE0FFEC2" box="[245,438,272,297]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49941FFAAC1F7D47EFE0FFEC2" box="[245,438,272,297]" italics="true" pageId="6" pageNumber="27">Neorhynchoplax</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |