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<document id="49617C9D8522AAB5DA93073B7350412D" ID-CLB-Dataset="22374" ID-DOI="10.5852/ejt.2015.149" ID-GBIF-Dataset="5f7efcfd-9137-4909-8125-f4cc7d9bae9f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3787554" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588704016992" checkinUser="carolina" docAuthor="Taylor, Paul D. &amp; Tan, Shau-Hwai Aileen" docDate="2015" docId="9C3C87B4BB2DE422FD25FEFCFC9FFA2A" docLanguage="en" docName="ejt-2015-149a.pdf.imf" docOrigin="European Journal of Taxonomy 149" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Parasmittina Osburn 1952" docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="6005FFCCBB3BE434FF8EFF92FFFEFFB2" masterDocTitle="Cheilostome Bryozoa from Penang and Langkawi, Malaysia" masterLastPageNumber="34" masterPageNumber="1" pageNumber="23" updateTime="1698838607080" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FCA314E6D04055DC0B1F3821ED03943E">Cheilostome Bryozoa from Penang and Langkawi, Malaysia</mods:title>
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<mods:namePart id="F4BB5F75E8E73BE89F169B298964130C">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="AB165C2936088529800AD22C178AAA1F">Tan, Shau-Hwai Aileen</mods:namePart>
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<mods:title id="A855123CA5C47A4119ED6851D662BFBA">European Journal of Taxonomy</mods:title>
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<mods:date id="F58A238E375091D03ED2F0DEA775DA4B">2015</mods:date>
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<treatment id="9C3C87B4BB2DE422FD25FEFCFC9FFA2A" ID-DOI="http://doi.org/10.5281/zenodo.3793807" ID-GBIF-Taxon="163963475" ID-Zenodo-Dep="3793807" LSID="urn:lsid:plazi:treatment:9C3C87B4BB2DE422FD25FEFCFC9FFA2A" httpUri="http://treatment.plazi.org/id/9C3C87B4BB2DE422FD25FEFCFC9FFA2A" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="5C8F6529BB2DE422FD25FEFCFC76FE35" box="[683,904,365,392]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="142A36A2BB2DE422FD25FEFCFC76FE35" blockId="22.[683,904,365,427]" box="[683,904,365,392]" pageId="22" pageNumber="23">
<heading id="4F6281CEBB2DE422FD25FEFCFC76FE35" box="[683,904,365,392]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
<emphasis id="26E1EAB0BB2DE422FD25FEFCFC76FE35" bold="true" box="[683,904,365,392]" pageId="22" pageNumber="23">
<taxonomicName id="D3954D21BB2DE422FD25FEFCFCB9FE3A" ID-CoL="84RR9" authorityName="Osburn" authorityYear="1952" box="[683,839,366,392]" class="Gymnolaemata" family="Smittinidae" genus="Parasmittina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="23" phylum="Bryozoa" rank="genus">
<emphasis id="26E1EAB0BB2DE422FD25FEFCFCB9FE3A" bold="true" box="[683,839,366,392]" italics="true" pageId="22" pageNumber="23">Parasmittina</emphasis>
</taxonomicName>
sp. 1
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5C8F6529BB2DE422FD58FE03FCA0FE19" box="[726,862,401,427]" pageId="22" pageNumber="23" type="description">
<paragraph id="142A36A2BB2DE422FD58FE03FCA0FE19" blockId="22.[683,904,365,427]" box="[726,862,401,427]" pageId="22" pageNumber="23">
<figureCitation id="8CAE2A27BB2DE422FD58FE03FCD9FE19" box="[726,807,401,427]" captionStart="Fig" captionStartId="21.[189,232,1677,1703]" captionTargetBox="[188,1399,702,1642]" captionTargetId="figure@21.[173,1411,696,1655]" captionTargetPageId="21" captionText="Fig. 12. Parasmittina spp. — AC. P. raigioidea Liu et al., 2001. Pulau Betong, Penang, MSL BRY020b. A. Part of colony. B. Autozooids and avicularia. C. Orifice with lyrula and proximally directed adventitious avicularium. — DF. Parasmittina sp. 1. Kuah jetty, Penang, MSL BRY021b. D. Colony (bottom) becoming overgrown by a colony of P. winstonae (top). E. Ovicellate zooids and proximally directed vicarious avicularia. F. Detail showing a proximally directed vicarious avicularium (upper left of centre) with a proximally directed spatulate adventitious avicularium to its lower right and a distally directed acuminate adventitious avicularium to the lower right of that. — GI. Parasmittina sp. 2. Kampung Kuala Temoyong, Langkawi, MSL BRY012b; encrusting a large gastropod shell. G. Part of colony. H. Group of zooids. I. Autozooids and an adventitious avicularium. Scale bars: A, H = 500 μm; B, EF = 200 μm; C, I = 100 μm; D, G = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787578" httpUri="https://zenodo.org/record/3787578/files/figure.png" pageId="22" pageNumber="23">Fig. 12</figureCitation>
DF
</paragraph>
</subSubSection>
<subSubSection id="5C8F6529BB2DE422FF33FE40FE9CFD8D" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="142A36A2BB2DE422FF33FE40FED4FE5E" blockId="22.[189,298,466,492]" box="[189,298,466,492]" pageId="22" pageNumber="23">
<heading id="4F6281CEBB2DE422FF33FE40FED4FE5E" bold="true" box="[189,298,466,492]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="26E1EAB0BB2DE422FF33FE40FED4FE5E" bold="true" box="[189,298,466,492]" pageId="22" pageNumber="23">Material</emphasis>
</heading>
</paragraph>
<paragraph id="142A36A2BB2DE422FF33FD93FE9CFD8D" blockId="22.[189,1399,513,575]" pageId="22" pageNumber="23">
<collectingCountry id="6C827632BB2DE422FF33FD93FEA4FDA9" box="[189,346,513,539]" name="Malaysia" pageId="22" pageNumber="23">MALAYSIA</collectingCountry>
: MSL BRY021b, Kuah jetty, Langkawi, on fouled rope hanging from jetty (with
<taxonomicName id="D3954D21BB2DE422FF33FDB4FEAFFD8D" box="[189,337,550,575]" class="Gymnolaemata" family="Smittinidae" genus="Parasmittina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="23" phylum="Bryozoa" rank="species" species="winstonae">
<emphasis id="26E1EAB0BB2DE422FF33FDB4FEAFFD8D" box="[189,337,550,575]" italics="true" pageId="22" pageNumber="23">P. winstonae</emphasis>
</taxonomicName>
).
</paragraph>
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<subSubSection id="5C8F6529BB2DE422FF33FDF7FDB4FB0D" pageId="22" pageNumber="23" type="description">
<paragraph id="142A36A2BB2DE422FF33FDF7FEB0FDCD" blockId="22.[189,334,613,639]" box="[189,334,613,639]" pageId="22" pageNumber="23">
<heading id="4F6281CEBB2DE422FF33FDF7FEB0FDCD" bold="true" box="[189,334,613,639]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="26E1EAB0BB2DE422FF33FDF7FEB0FDCD" bold="true" box="[189,334,613,639]" pageId="22" pageNumber="23">Description</emphasis>
</heading>
</paragraph>
<paragraph id="142A36A2BB2DE422FF33FD07FDB4FB0D" blockId="22.[189,1399,661,1215]" pageId="22" pageNumber="23">
Colony encrusting, multiserial, unilamellar. Ancestrula and early astogeny not observed. Autozooids subhexagonal, small, about
<quantity id="D36D9B47BB2DE422FD82FD2AFD47FD60" box="[524,697,696,722]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="22" pageNumber="23" unit="mm" value="0.31" valueMax="0.32" valueMin="0.3">0.300.32 mm</quantity>
long by
<quantity id="D36D9B47BB2DE422FCA7FD2AFC28FD60" box="[809,982,696,722]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="22" pageNumber="23" unit="mm" value="0.265" valueMax="0.28" valueMin="0.25">0.250.28 mm</quantity>
wide; boundary walls salient with a fine fissure; frontal shield convex, coarsely pustulose, marginal areolar pores large, no pseudopores; secondary orifice elliptical, about
<quantity id="D36D9B47BB2DE422FDC7FD6CFD0DFCAA" box="[585,755,766,792]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="22" pageNumber="23" unit="mm" value="0.11" valueMax="0.12" valueMin="0.1">0.100.12 mm</quantity>
wide, lappets highest proximolaterally, lyrula variable, typically narrow, 1318 μm wide at tip; oral spine bases not observed; ovicell hyperstomial, broad, about
<quantity id="D36D9B47BB2DE422FE8BFCD7FE50FCED" box="[261,430,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="22" pageNumber="23" unit="mm" value="0.14" valueMax="0.16" valueMin="0.12">0.120.16 mm</quantity>
long by
<quantity id="D36D9B47BB2DE422FD9DFCD7FD43FCED" box="[531,701,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="22" pageNumber="23" unit="mm" value="0.175" valueMax="0.2" valueMin="0.15">0.150.20 mm</quantity>
wide, ectooecium with about 30 ridged pores, proximal and distal edges of ovicell overgrown by cryptocystal calcification. Avicularia of three
<typeStatus id="CB2E8800BB2DE422FBFDFCFAFB4BFC30" box="[1139,1205,872,898]" pageId="22" pageNumber="23">types</typeStatus>
: (1) adventitious acuminate avicularia present on some proximal frontal shields, about
<quantity id="D36D9B47BB2DE422FC7AFC19FBA2FC17" box="[1012,1116,907,933]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="mm" value="0.11">0.11 mm</quantity>
long by
<quantity id="D36D9B47BB2DE422FB4BFC19FAD1FC17" box="[1221,1327,907,933]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="22" pageNumber="23" unit="mm" value="0.07">0.07 mm</quantity>
wide, usually distally or distolaterally directed, rostrum an acute triangle, tip raised, rostral shelf shallow, cross-bar calcified, opesia semicircular; (2) adventitious spatulate avicularia present on some frontal shields, about
<quantity id="D36D9B47BB2DE422FEE0FC67FE25FBBD" box="[366,475,1013,1039]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="23" unit="mm" value="0.14">0.14 mm</quantity>
long by
<quantity id="D36D9B47BB2DE422FDC1FC67FD43FBBD" box="[591,701,1013,1039]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="0.05">0.05 mm</quantity>
wide, directed proximally or laterally towards autozooid median line, rostrum rounded with deep shelf, cross-bar calcified, opesia semicircular; (3) vicarious avicularia large,
<quantity id="D36D9B47BB2DE422FE09FBA9FDCBFBE7" box="[391,565,1083,1109]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" metricValueMax="3.8" metricValueMin="2.3" pageId="22" pageNumber="23" unit="mm" value="0.305" valueMax="0.38" valueMin="0.23">0.230.38 mm</quantity>
long by
<quantity id="D36D9B47BB2DE422FD2BFBA9FCADFBE7" box="[677,851,1083,1109]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="22" pageNumber="23" unit="mm" value="0.14" valueMax="0.15" valueMin="0.13">0.130.15 mm</quantity>
wide, scattered, directed proximally, rostrum initially tapering before becoming parallel-sided, tip well rounded, distal edge slightly serrated, rostral shelf deep, occupying half or more of rostrum length, cross-bar calcified, concave, opesia relatively small, crescent shaped (
<figureCitation id="8CAE2A27BB2DE422FE5AFB37FDC3FB0D" box="[468,573,1189,1215]" captionStart="Fig" captionStartId="21.[189,232,1677,1703]" captionTargetBox="[188,1399,702,1642]" captionTargetId="figure@21.[173,1411,696,1655]" captionTargetPageId="21" captionText="Fig. 12. Parasmittina spp. — AC. P. raigioidea Liu et al., 2001. Pulau Betong, Penang, MSL BRY020b. A. Part of colony. B. Autozooids and avicularia. C. Orifice with lyrula and proximally directed adventitious avicularium. — DF. Parasmittina sp. 1. Kuah jetty, Penang, MSL BRY021b. D. Colony (bottom) becoming overgrown by a colony of P. winstonae (top). E. Ovicellate zooids and proximally directed vicarious avicularia. F. Detail showing a proximally directed vicarious avicularium (upper left of centre) with a proximally directed spatulate adventitious avicularium to its lower right and a distally directed acuminate adventitious avicularium to the lower right of that. — GI. Parasmittina sp. 2. Kampung Kuala Temoyong, Langkawi, MSL BRY012b; encrusting a large gastropod shell. G. Part of colony. H. Group of zooids. I. Autozooids and an adventitious avicularium. Scale bars: A, H = 500 μm; B, EF = 200 μm; C, I = 100 μm; D, G = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787578" httpUri="https://zenodo.org/record/3787578/files/figure.png" pageId="22" pageNumber="23">Fig. 12F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5C8F6529BB2DE422FF33FB77FC9FFA2A" pageId="22" pageNumber="23" type="discussion">
<paragraph id="142A36A2BB2DE422FF33FB77FED1FB4D" blockId="22.[189,303,1253,1279]" box="[189,303,1253,1279]" pageId="22" pageNumber="23">
<heading id="4F6281CEBB2DE422FF33FB77FED1FB4D" bold="true" box="[189,303,1253,1279]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="26E1EAB0BB2DE422FF33FB77FED1FB4D" bold="true" box="[189,303,1253,1279]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="142A36A2BB2DE422FF33FA87FC9FFA2A" blockId="22.[189,1399,1301,1433]" pageId="22" pageNumber="23">
It has not been possible to identify this species of
<taxonomicName id="D3954D21BB2DE422FC9BFA84FC50FA9D" authorityName="Osburn" authorityYear="1952" box="[789,942,1302,1327]" class="Gymnolaemata" family="Smittinidae" genus="Parasmittina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="23" phylum="Bryozoa" rank="genus">
<emphasis id="26E1EAB0BB2DE422FC9BFA84FC50FA9D" box="[789,942,1302,1327]" italics="true" pageId="22" pageNumber="23">Parasmittina</emphasis>
</taxonomicName>
despite the combination of distinctive large vicarious avicularia and small adventitious avicularia of spatulate and acuminate
<typeStatus id="CB2E8800BB2DE422FB38FAAAFB06FAE0" box="[1206,1272,1336,1362]" pageId="22" pageNumber="23">types</typeStatus>
. Pending a more detailed study and a comprehensive comparison with the myriad of species of
<taxonomicName id="D3954D21BB2DE422FB2CFACEFAC5FAC7" authorityName="Osburn" authorityYear="1952" box="[1186,1339,1372,1397]" class="Gymnolaemata" family="Smittinidae" genus="Parasmittina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="23" phylum="Bryozoa" rank="genus">
<emphasis id="26E1EAB0BB2DE422FB2CFACEFAC5FAC7" box="[1186,1339,1372,1397]" italics="true" pageId="22" pageNumber="23">Parasmittina</emphasis>
</taxonomicName>
, it is therefore left in open nomenclature as
<taxonomicName id="D3954D21BB2DE422FD0CFA12FCE5FA2B" authorityName="Osburn" authorityYear="1952" box="[642,795,1408,1433]" class="Gymnolaemata" family="Smittinidae" genus="Parasmittina" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="23" phylum="Bryozoa" rank="genus">
<emphasis id="26E1EAB0BB2DE422FD0CFA12FCE5FA2B" box="[642,795,1408,1433]" italics="true" pageId="22" pageNumber="23">Parasmittina</emphasis>
</taxonomicName>
sp. 1.
</paragraph>
</subSubSection>
</treatment>
</document>