treatments-xml/data/46/08/87/4608879FFFDCFFE86D8AFCD2FB6CFC79.xml
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<document id="94CA5D758D65BF26D0F7685883EFFF9F" ID-DOI="10.5281/zenodo.11404264" ID-ISSN="1525-9153" ID-Zenodo-Dep="11404264" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1716907766958" checkinUser="felipe" docAuthor="Barrio-Amorós, César L., Rojas-Runjaic, Fernando J. M. &amp; Señaris, J. Celsa" docDate="2019" docId="4608879FFFDCFFE86D8AFCD2FB6CFC79" docLanguage="en" docName="AmphibReptileCons.13.1.e180.pdf" docOrigin="Amphibian &amp; Reptile Conservation (e 180) 13 (1)" docStyle="DocumentStyle:2DCF38F802BD4DFFCCCB5916C58E77B3.2:AmphibReptileConse.2015-.journal_article.2cover" docStyleId="2DCF38F802BD4DFFCCCB5916C58E77B3" docStyleName="AmphibReptileConse.2015-.journal_article.2cover" docStyleVersion="2" docTitle="Pristimantis Jimenez de la Espada 1870" docType="treatment" docVersion="5" lastPageNumber="111" masterDocId="BA31FFE7FFACFF986D36FA1CFFB9FFDF" masterDocTitle="Catalogue of the amphibians of Venezuela: Illustrated and annotated species list, distribution, and conservation" masterLastPageNumber="198" masterPageNumber="1" pageNumber="111" updateTime="1717186961153" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7279BD7B1DBD83A716145FBA3B4681DB">Catalogue of the amphibians of Venezuela: Illustrated and annotated species list, distribution, and conservation</mods:title>
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<mods:namePart id="C91EE068E0ADC82397CAA17F41692454">Rojas-Runjaic, Fernando J. M.</mods:namePart>
<mods:affiliation id="2559D51BE363CD13EE9B7005DD9FBA43">Fundación La Salle de Ciencias Naturales, Museo de Historia Natural La Salle, Apartado Postal 1930, Caracas 1010 - A, VENEZUELA &amp; Current address: Pontifícia Universidade Católica do Río Grande do Sul (PUCRS), Laboratório de Sistemática de Vertebrados, Av. Ipiranga 6681, Porto Alegre, RS 90619 - 900, BRAZIL</mods:affiliation>
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<mods:namePart id="6B067E54C7AC8A166DC1EAC3AD1B7764">Señaris, J. Celsa</mods:namePart>
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<paragraph id="CE1E3689FFDCFFE86D8AFCD2FD03F939" blockId="112.[188,698,1742,1766]" box="[188,698,1742,1766]" pageId="112" pageNumber="111">
<heading id="955681E5FFDCFFE86D8AFCD2FD03F939" box="[188,698,1742,1766]" centered="true" fontSize="10" level="2" pageId="112" pageNumber="111" reason="7">
Genus
<taxonomicName id="09A14D0AFFDCFFE86C31FCD2FD03F939" ID-CoL="6VX3" authority="Jimenez de la Espada, 1870" authorityName="Jimenez de la Espada" authorityYear="1870" box="[263,698,1742,1766]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE86C31FCD2FE35F939" bold="true" box="[263,396,1742,1766]" italics="true" pageId="112" pageNumber="111">Pristimantis</emphasis>
Jiménez de la Espada, 1870
</taxonomicName>
</heading>
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<paragraph id="CE1E3689FFDCFFE86DBBFD12FEDAF899" blockId="112.[141,746,1805,1958]" pageId="112" pageNumber="111">
<emphasis id="FCD5EA9BFFDCFFE86DBBFD12FE9CF8FA" bold="true" box="[141,293,1805,1830]" pageId="112" pageNumber="111">
<typeStatus id="111A882BFFDCFFE86DBBFD12FF7CF8F9" box="[141,197,1806,1830]" pageId="112" pageNumber="111">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="09A14D0AFFDCFFE86C18FD12FF7CF899" ID-CoL="77WHP" authority="Jimenez de la Espada, 1870" authorityName="Jimenez de la Espada" authorityYear="1870" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="species" species="galdi">
<emphasis id="FCD5EA9BFFDCFFE86C18FD12FE49F8F9" box="[302,496,1806,1830]" italics="true" pageId="112" pageNumber="111">Pristimantis galdi</emphasis>
Jiménez de la Espada, 1870
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="86BB6502FFDCFFE86DBBFD52FB6CFC79" pageId="112" pageNumber="111" type="discussion">
<paragraph id="CE1E3689FFDCFFE86DBBFD52FAB1FCF9" blockId="112.[141,746,1805,1958]" lastBlockId="112.[791,1396,142,934]" pageId="112" pageNumber="111">
<emphasis id="FCD5EA9BFFDCFFE86DBBFD52FF47F8B9" bold="true" box="[141,254,1870,1894]" pageId="112" pageNumber="111">Remarks:</emphasis>
<bibRefCitation id="AA304B78FFDCFFE86C3EFD52FE40F8B9" author="Heinicke MP &amp; Duellman WE &amp; Hedges SB" box="[264,505,1870,1894]" pageId="112" pageNumber="111" refId="ref94231" refString="Heinicke MP, Duellman WE, Hedges SB. 2007. Major Caribbean and Central American frog faunas originated by ancient oceanic dispersal. Proceedings of the National Academy of Sciences of the United States of America 104: 10,092 - 10,097." type="journal volume" year="2007">Heinicke et al. (2007)</bibRefCitation>
split long-established genus
<taxonomicName id="09A14D0AFFDCFFE86DC5FD72FE0DF859" authorityName="Dumeril and Bibron" authorityYear="1841" box="[243,436,1902,1926]" class="Amphibia" family="Craugastoridae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE86DC5FD72FE0DF859" box="[243,436,1902,1926]" italics="true" pageId="112" pageNumber="111">Eleutherodactylus</emphasis>
</taxonomicName>
, formerly in family
<taxonomicName id="09A14D0AFFDCFFE86DBBFD92FE80F879" authorityName="Werner" authorityYear="1896" box="[141,313,1934,1958]" class="Amphibia" family="Leptodactylidae" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="family">Leptodactylidae</taxonomicName>
(Lynch 1971, 1981;
<bibRefCitation id="AA304B78FFDCFFE86F68FD92FD5CF879" author="Frost DR &amp; Kansas, USA" box="[606,741,1934,1958]" pageId="112" pageNumber="111" refId="ref92135" refString="Frost DR (ed.) 1985. Amphibian Species of the World. Allen Press Inc. and Association of Systematic Collections, Lawrence, Kansas, USA. 732 p." type="book" year="1985">Frost 1985</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86E21FA92FC79FF79" author="Duellman WE" box="[791,960,142,166]" pageId="112" pageNumber="111" pagination="1 - 372" refId="ref90241" refString="Duellman WE. 1993. Amphibian species of the world. Additions and corrections. University of Kansas, Museum of Natural History, Special Publications 21: 1 - 372." type="journal article" year="1993">Duellman 1993</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86EFBFA92FB56FF79" author="Lynch JD &amp; Duellman WE" box="[973,1263,142,166]" pageId="112" pageNumber="111" pagination="1 - 236" refId="ref99962" refString="Lynch JD, Duellman WE. 1997. Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador: systematics, ecology and biogeography. University of Kansas Natural History Museum, Special Publications 23: 1 - 236." type="journal article" year="1997">Lynch and Duellman 1997</bibRefCitation>
) and placed afterwards in family
<taxonomicName id="09A14D0AFFDCFFE86933FAB2FB7DFF19" authorityName="Gunther" authorityYear="1858" box="[1029,1220,174,198]" class="Amphibia" family="Brachycephalidae" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="family">Brachycephalidae</taxonomicName>
by
<bibRefCitation id="AA304B78FFDCFFE869C1FAB2FCE2FF39" author="Frost DR &amp; Grant T &amp; Faivovich J &amp; Bain RH &amp; Haas A &amp; Haddad CFB &amp; De Sa RO &amp; Channing A &amp; Wilkinson M &amp; Donnellan SC" pageId="112" pageNumber="111" pagination="1 - 370" refId="ref92213" refString="Frost DR, Grant T, Faivovich J, Bain RH, Haas A, Haddad CFB, De Sa RO, Channing A, Wilkinson M, Donnellan SC, et al. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1 - 370." type="journal article" year="2006">Frost et al. (2006)</bibRefCitation>
containing three major genera:
<taxonomicName id="09A14D0AFFDCFFE8698CFAD2FACAFF39" authorityName="Dumeril and Bibron" authorityYear="1841" box="[1210,1395,206,230]" class="Amphibia" family="Craugastoridae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE8698CFAD2FACAFF39" box="[1210,1395,206,230]" italics="true" pageId="112" pageNumber="111">Eleutherodactylus</emphasis>
</taxonomicName>
for mainly Caribbean species,
<taxonomicName id="09A14D0AFFDCFFE86960FAF2FAF4FED9" authority="Cope, 1862" authorityName="Cope" authorityYear="1862" box="[1110,1357,238,262]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE86960FAF2FB75FEDA" box="[1110,1228,238,261]" italics="true" pageId="112" pageNumber="111">Craugastor</emphasis>
Cope, 1862
</taxonomicName>
for mainly Central American species, and
<taxonomicName id="09A14D0AFFDCFFE869FCFB12FAFEFEFA" authorityName="Jimenez de la Espada" authorityYear="1870" box="[1226,1351,270,293]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE869FCFB12FAFEFEFA" box="[1226,1351,270,293]" italics="true" pageId="112" pageNumber="111">Pristimantis</emphasis>
</taxonomicName>
for almost all South American (and some Central American) species.
<bibRefCitation id="AA304B78FFDCFFE86E5AFB52FB81FEB9" author="Hedges SB &amp; Duellman WE &amp; Heinicke MP" box="[876,1080,334,358]" pageId="112" pageNumber="111" pagination="1 - 182" refId="ref94193" refString="Hedges SB, Duellman WE, Heinicke MP. 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1 - 182." type="journal article" year="2008">Hedges et al. (2008)</bibRefCitation>
offered a more comprehensive view of the so-called “eleutherodactylines” and erected the new unranked taxon Terrarana (emended as Terraranae by
<bibRefCitation id="AA304B78FFDCFFE86E21FBB2FC52FE19" author="Heinicke MP &amp; Lemmon AR &amp; Lemmon EM &amp; McGrath K &amp; Hedges SB" box="[791,1003,430,454]" pageId="112" pageNumber="111" pagination="145 - 155" refId="ref94320" refString="Heinicke MP, Lemmon AR, Lemmon EM, McGrath K, Hedges SB. 2018. Phylogenomic support for evolutionary relationships of New World direct-developing frogs (Anura: Terraranae). Molecular Phylogenetics and Evolution 118: 145 - 155." type="journal article" year="2018">Heinicke et al. 2018</bibRefCitation>
) for four families (
<taxonomicName id="09A14D0AFFDCFFE86982FBB2FAD6FE19" authorityName="Gunther" authorityYear="1858" box="[1204,1391,430,454]" class="Amphibia" family="Brachycephalidae" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="family">Brachycephalidae</taxonomicName>
,
<taxonomicName id="09A14D0AFFDCFFE86E21FBD2FC0DFE39" authorityName="Hedges, Duellman and Heinicke" authorityYear="2008" box="[791,948,462,486]" class="Amphibia" family="Craugastoridae" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="family">Craugastoridae</taxonomicName>
,
<taxonomicName id="09A14D0AFFDCFFE86E88FBD2FB35FE39" authorityName="Lutz" authorityYear="1954" box="[958,1164,462,486]" class="Amphibia" family="Eleutherodactylidae" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="family">Eleutherodactylidae</taxonomicName>
, and
<taxonomicName id="09A14D0AFFDCFFE869F2FBD2FAD3FE39" authorityName="Hedges, Duellman and Heinicke" authorityYear="2008" box="[1220,1386,462,486]" family="Strabomantidae" pageId="112" pageNumber="111" rank="family">Strabomantidae</taxonomicName>
). As of this writing, the phylogenetic arrangement remains in flux. Most South American species formerly in
<taxonomicName id="09A14D0AFFDCFFE86E21F832FC69FD99" authorityName="Dumeril and Bibron" authorityYear="1841" box="[791,976,558,582]" class="Amphibia" family="Craugastoridae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE86E21F832FC69FD99" box="[791,976,558,582]" italics="true" pageId="112" pageNumber="111">Eleutherodactylus</emphasis>
</taxonomicName>
are now members of
<taxonomicName id="09A14D0AFFDCFFE869C6F832FAD5FD9A" authorityName="Jimenez de la Espada" authorityYear="1870" box="[1264,1388,558,581]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE869C6F832FAD5FD9A" box="[1264,1388,558,581]" italics="true" pageId="112" pageNumber="111">Pristimantis</emphasis>
</taxonomicName>
, within
<taxonomicName id="09A14D0AFFDCFFE86E5FF852FBBDFDB9" authorityName="Hedges, Duellman and Heinicke" authorityYear="2008" box="[873,1028,590,614]" class="Amphibia" family="Craugastoridae" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="family">Craugastoridae</taxonomicName>
(after
<bibRefCitation id="AA304B78FFDCFFE8696DF852FA89FDB9" author="Padial JM &amp; Grant T &amp; Frost D." box="[1115,1328,590,614]" pageId="112" pageNumber="111" pagination="1 - 132" refId="ref103173" refString="Padial JM, Grant T, Frost D. 2014. Molecular systematics of terraranas (Anura; Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa 3825: 1 - 132." type="journal article" year="2014">Padial et al. 2014</bibRefCitation>
). The most recent approach (
<bibRefCitation id="AA304B78FFDCFFE8692CF872FABFFD59" author="Heinicke MP &amp; Lemmon AR &amp; Lemmon EM &amp; McGrath K &amp; Hedges SB" box="[1050,1286,622,646]" pageId="112" pageNumber="111" pagination="145 - 155" refId="ref94320" refString="Heinicke MP, Lemmon AR, Lemmon EM, McGrath K, Hedges SB. 2018. Phylogenomic support for evolutionary relationships of New World direct-developing frogs (Anura: Terraranae). Molecular Phylogenetics and Evolution 118: 145 - 155." type="journal article" year="2018">Heinicke et al. 2018</bibRefCitation>
) recovers
<taxonomicName id="09A14D0AFFDCFFE86E21F892FC13FD79" box="[791,938,654,678]" class="Amphibia" family="Craugastoridae" genus="Tachiramantis" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE86E21F892FC13FD79" box="[791,938,654,678]" italics="true" pageId="112" pageNumber="111">Tachiramantis</emphasis>
</taxonomicName>
as part of
<taxonomicName id="09A14D0AFFDCFFE86914F892FB06FD79" authorityName="Hedges, Duellman and Heinicke" authorityYear="2008" box="[1058,1215,654,678]" class="Amphibia" family="Craugastoridae" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="family">Craugastoridae</taxonomicName>
, and
<taxonomicName id="09A14D0AFFDCFFE869C0F892FACAFD7A" authorityName="Jimenez de la Espada" authorityYear="1870" box="[1270,1395,654,677]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE869C0F892FACAFD7A" box="[1270,1395,654,677]" italics="true" pageId="112" pageNumber="111">Pristimantis</emphasis>
</taxonomicName>
into
<taxonomicName id="09A14D0AFFDCFFE86E66F8B2FC4DFD19" authorityName="Hedges, Duellman and Heinicke" authorityYear="2008" box="[848,1012,686,710]" family="Strabomantidae" pageId="112" pageNumber="111" rank="family">Strabomantidae</taxonomicName>
.
<taxonomicName id="09A14D0AFFDCFFE86931F8B2FACAFD19" authority="La Marca, 2007" authorityName="La Marca" authorityYear="2007" box="[1031,1395,686,710]" class="Amphibia" family="Craugastoridae" genus="Mucubatrachus" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE86931F8B2FB11FD19" box="[1031,1192,686,710]" italics="true" pageId="112" pageNumber="111">Mucubatrachus</emphasis>
<bibRefCitation id="AA304B78FFDCFFE8698FF8B2FACAFD19" author="La Marca E." box="[1209,1395,686,710]" pageId="112" pageNumber="111" pagination="67 - 87" refId="ref98095" refString="La Marca E. 2007. Sinopsis taxonomica de dos nuevos generos de anfibios (Anura: Leptodactylidae) de los Andes de Venezuela. Herpetotopicos 3: 67 - 87." type="journal article" year="2007">La Marca, 2007</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="09A14D0AFFDCFFE86E7DF8D2FB7AFD39" authority="La Marca, 2007" authorityName="La Marca" authorityYear="2007" box="[843,1219,718,742]" class="Amphibia" family="Craugastoridae" genus="Paramophrynella" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="genus">
<emphasis id="FCD5EA9BFFDCFFE86E7DF8D2FBB9FD39" box="[843,1024,718,742]" italics="true" pageId="112" pageNumber="111">Paramophrynella</emphasis>
<bibRefCitation id="AA304B78FFDCFFE86938F8D2FB7AFD39" author="La Marca E." box="[1038,1219,718,742]" pageId="112" pageNumber="111" pagination="67 - 87" refId="ref98095" refString="La Marca E. 2007. Sinopsis taxonomica de dos nuevos generos de anfibios (Anura: Leptodactylidae) de los Andes de Venezuela. Herpetotopicos 3: 67 - 87." type="journal article" year="2007">La Marca, 2007</bibRefCitation>
</taxonomicName>
are considered synonyms of
<taxonomicName id="09A14D0AFFDCFFE86E85F8F2FBC4FCDA" box="[947,1149,750,773]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="112" pageNumber="111" phylum="Chordata" rank="species" species="sensu">
<emphasis id="FCD5EA9BFFDCFFE86E85F8F2FBC4FCDA" box="[947,1149,750,773]" italics="true" pageId="112" pageNumber="111">Pristimantis sensu</emphasis>
</taxonomicName>
(
<bibRefCitation id="AA304B78FFDCFFE869A7F8F2FAD6FCD9" author="Hedges SB &amp; Duellman WE &amp; Heinicke MP" box="[1169,1391,750,774]" pageId="112" pageNumber="111" pagination="1 - 182" refId="ref94193" refString="Hedges SB, Duellman WE, Heinicke MP. 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1 - 182." type="journal article" year="2008">Hedges et al. 2008</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86E21F912FB8DFCF9" author="Barrio-Amoros CL &amp; Heinicke M &amp; Hedges SB" box="[791,1076,782,806]" pageId="112" pageNumber="111" pagination="43 - 62" refId="ref86759" refString="Barrio-Amoros CL, Heinicke M, Hedges SB. 2013. A new tuberculated Pristimantis (Anura, Terrarana, Strabomantidae) from the Venezuelan Andes, redescription of Pristimantis pleurostriatus and variation within Pristimantis vanadisae. Zootaxa 3647: 43 - 62." type="journal article" year="2013">Barrio-Amorós et al. 2013</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86908F912FB45FCF9" author="Padial JM &amp; Grant T &amp; Frost D." box="[1086,1276,782,806]" pageId="112" pageNumber="111" pagination="1 - 132" refId="ref103173" refString="Padial JM, Grant T, Frost D. 2014. Molecular systematics of terraranas (Anura; Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa 3825: 1 - 132." type="journal article" year="2014">Padial et al. 2014</bibRefCitation>
).
</paragraph>
<paragraph id="CE1E3689FFDCFFE86E21F931FB6CFC79" blockId="112.[791,1396,142,934]" pageId="112" pageNumber="111">
<emphasis id="FCD5EA9BFFDCFFE86E21F931FC46FC9A" bold="true" box="[791,1023,813,837]" pageId="112" pageNumber="111">Selected references:</emphasis>
<bibRefCitation id="AA304B78FFDCFFE8693BF932FB2FFC99" author="Lutz A." box="[1037,1174,814,838]" pageId="112" pageNumber="111" pagination="35 - 65" refId="ref99402" refString="Lutz A. 1927. Notas sobre batrachios da Venezuela e da Ilha de trinidad. Memorias do Instituto Oswaldo Cruz 20: 35 - 65." type="journal article" year="1927">Lutz (1927)</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE8699FF932FAFDFC99" author="Lynch JD" box="[1193,1348,814,838]" pageId="112" pageNumber="111" pagination="1 - 24" refId="ref99492" refString="Lynch JD. 1976. The species groups of the South American frogs of the genus Eleutherodactylus (Leptodactylidae). Occasional Papers of the Museum of Natural History, University of Kansas 61: 1 - 24." type="journal article" year="1976">Lynch (1976)</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86861F932FC01FCB9" author="La Marca E." pageId="112" pageNumber="111" refId="ref97345" refString="La Marca E. 1992. Catalogo Taxonomico, Biogeografico y Bibliografico de las Ranas de Venezuela. Cuadernos Geograficos ULA 9. Universidad de Los Andes, Merida, Merida, Venezuela. 197 p." type="book" year="1992">La Marca (1992)</bibRefCitation>
; Barrio-Amorós (1998, 2004, 2009);
<bibRefCitation id="AA304B78FFDCFFE86E21F972FBACFC59" author="Heinicke MP &amp; Duellman WE &amp; Hedges SB" box="[791,1045,878,902]" pageId="112" pageNumber="111" refId="ref94231" refString="Heinicke MP, Duellman WE, Hedges SB. 2007. Major Caribbean and Central American frog faunas originated by ancient oceanic dispersal. Proceedings of the National Academy of Sciences of the United States of America 104: 10,092 - 10,097." type="journal volume" year="2007">Heinicke et al. (2007)</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86910F972FAACFC59" author="Hedges SB &amp; Duellman WE &amp; Heinicke MP" box="[1062,1301,878,902]" pageId="112" pageNumber="111" pagination="1 - 182" refId="ref94193" refString="Hedges SB, Duellman WE, Heinicke MP. 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1 - 182." type="journal article" year="2008">Hedges et al. (2008)</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86810F972FC41FC79" author="Barrio-Amoros CL &amp; Heinicke M &amp; Hedges SB" pageId="112" pageNumber="111" pagination="43 - 62" refId="ref86759" refString="Barrio-Amoros CL, Heinicke M, Hedges SB. 2013. A new tuberculated Pristimantis (Anura, Terrarana, Strabomantidae) from the Venezuelan Andes, redescription of Pristimantis pleurostriatus and variation within Pristimantis vanadisae. Zootaxa 3647: 43 - 62." type="journal article" year="2013">Barrio-Amorós et al. (2013)</bibRefCitation>
;
<bibRefCitation id="AA304B78FFDCFFE86935F992FB68FC79" author="Padial JM &amp; Grant T &amp; Frost D." box="[1027,1233,910,934]" pageId="112" pageNumber="111" pagination="1 - 132" refId="ref103173" refString="Padial JM, Grant T, Frost D. 2014. Molecular systematics of terraranas (Anura; Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa 3825: 1 - 132." type="journal article" year="2014">Padial et al. (2014)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>