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<document id="8F204D70CE98F55F302B42FAA84A7E84" ID="10.11646/zootaxa.4276.2.7SLASH11313" ID-CLB-Dataset="32860" ID-DOI="10.11646/zootaxa.4276.2.7" ID-GBIF-Dataset="1cb0e227-6a7f-41f2-8ac7-4a98a9641ad2" ID-ISSN="1175-5326" ID-Zenodo-Dep="806115" ID-ZooBank="72DC4B80-2CFE-414F-AA86-DAA73618D977" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497253899257" checkinUser="plazi" docAuthor="Ronderos, Maria M., Spinelli, Gustavo R. &amp; Grogan, Willliam L." docDate="2017" docId="D97387A89D38FF9AFF448AA1FCC38960" docLanguage="en" docName="zootaxa.4276.2.7SLASH11313.pdf" docOrigin="Zootaxa 4276 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austrohelea spinosa Ronderos &amp; Spinelli &amp; Grogan 2017, New Species" docType="treatment" docVersion="8" lastPageNumber="268" masterDocId="254AFFD09D30FF97FFD38C26FFED8F28" masterDocTitle="The Neotropical species of the predaceous midge genus Austrohelea Wirth &amp; Grogan (Diptera: Ceratopogonidae)" masterLastPageNumber="269" masterPageNumber="255" pageNumber="263" updateTime="1698440399754" updateUser="ExternalLinkService">
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<mods:title id="0F3896ED4F0BD0C44B2520280371B38E">The Neotropical species of the predaceous midge genus Austrohelea Wirth &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="82042712A38C6E00C3DD445F8641ED67">Ronderos, Maria M.</mods:namePart>
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<mods:namePart id="347AA29EBC3B8D630D6014EDAA55D78E">Spinelli, Gustavo R.</mods:namePart>
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<mods:namePart id="340025411AE448F95F1A3C24258251EA">Grogan, Willliam L.</mods:namePart>
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<mods:typeOfResource id="C06E8ACF75C9E84505F32B6AA08A8027">text</mods:typeOfResource>
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<mods:title id="9136D9EE4B472002E7FD5FDEBBB129A8">Zootaxa</mods:title>
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<mods:date id="C64FE3AD8065FF91D4E82192BE3509AC">2017</mods:date>
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<mods:number id="DFA745A7371287CA574A0EE290FD9DB9">4276</mods:number>
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<treatment id="D97387A89D38FF9AFF448AA1FCC38960" ID-DOI="http://doi.org/10.5281/zenodo.6017421" ID-GBIF-Taxon="131876629" ID-Zenodo-Dep="6017421" LSID="urn:lsid:plazi:treatment:D97387A89D38FF9AFF448AA1FCC38960" httpUri="http://treatment.plazi.org/id/D97387A89D38FF9AFF448AA1FCC38960" lastPageId="13" lastPageNumber="268" pageId="8" pageNumber="263">
<subSubSection id="19C065359D38FF9FFF448AA1FE6889EB" pageId="8" pageNumber="263" type="nomenclature">
<paragraph id="516536BE9D38FF9FFF448AA1FDC0898A" blockId="8.[151,557,1671,1732]" box="[151,557,1671,1698]" pageId="8" pageNumber="263">
<heading id="0A2D81D29D38FF9FFF448AA1FDC0898A" bold="true" box="[151,557,1671,1698]" fontSize="11" level="1" pageId="8" pageNumber="263" reason="1">
<emphasis id="63AEEAAC9D38FF9FFF448AA1FDC0898A" bold="true" box="[151,557,1671,1698]" pageId="8" pageNumber="263">
<taxonomicName id="96DA4D3D9D38FF9FFF448AA1FE618989" ID-CoL="9HWY2" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[151,396,1671,1697]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D38FF9FFF448AA1FE618989" bold="true" box="[151,396,1671,1697]" italics="true" pageId="8" pageNumber="263">Austrohelea spinosa</emphasis>
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<taxonomicNameLabel id="789D57D79D38FF9FFE408AAEFDC0898A" box="[403,557,1672,1698]" pageId="8" pageNumber="263" rank="species">New Species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="516536BE9D38FF9FFF448A8DFE6889EB" blockId="8.[151,557,1671,1732]" box="[151,389,1707,1732]" pageId="8" pageNumber="263">
(
<figureCitation id="C9E12A3B9D38FF9FFF4C8A8DFF1389EB" box="[159,254,1707,1732]" captionStart="FIGURES 20 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="8" pageNumber="263">Figs. 20</figureCitation>
̄28, 33̄34)
</paragraph>
</subSubSection>
<subSubSection id="19C065359D38FF9FFF448AD5FCEF88CC" pageId="8" pageNumber="263" type="diagnosis">
<paragraph id="516536BE9D38FF9FFF448AD5FCEF88CC" blockId="8.[151,1437,1779,2020]" pageId="8" pageNumber="263">
<emphasis id="63AEEAAC9D38FF9FFF448AD5FEFC8824" bold="true" box="[151,273,1779,1804]" pageId="8" pageNumber="263">Diagnosis.</emphasis>
A large (male wing length 1.50̄
<quantity id="96229B5B9D38FF9FFD478AD5FD128823" box="[660,767,1779,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="263" unit="mm" value="1.6">1.60 mm</quantity>
, female wing length 1.57̄
<quantity id="96229B5B9D38FF9FFBE78AD5FB4C8823" box="[1076,1185,1779,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="8" pageNumber="263" unit="mm" value="1.68">1.68 mm</quantity>
), dark brown species. Males with hind tibia with 3 long, stout, black subapical spines; tergite 9 with narrow shallow posterior excavation; gonocoxite short, stout, tapering slightly distally with basomesal tubercle; gonostylus longer than gonocoxite; parameres with very broad basal apodemes that form an obtuse angle between the anterior and posterior arms; and the aedeagus is very large, heavily sclerotized with long basal arms, a U-shaped basal arch and lightly sclerotized apical extension. Females with palpal segment 3 lacking sensory pit, hind femur greatly swollen, and subequal, ovoid spermathecae with moderately long broad necks.
</paragraph>
</subSubSection>
<caption id="05A566369D39FF9EFF448B9CFF1888D8" httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="9" pageNumber="264" targetBox="[260,1331,213,1923]" targetPageId="9">
<paragraph id="516536BE9D39FF9EFF448B9CFF1888D8" blockId="9.[151,1436,1978,2032]" pageId="9" pageNumber="264">
<emphasis id="63AEEAAC9D39FF9EFF448B9CFEBB88F8" bold="true" box="[151,342,1978,2001]" pageId="9" pageNumber="264">FIGURES 2024.</emphasis>
<taxonomicName id="96DA4D3D9D39FF9EFE8D8B9DFDC188F9" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[350,556,1979,2001]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D39FF9EFE8D8B9DFDC188F9" box="[350,556,1979,2001]" italics="true" pageId="9" pageNumber="264">Austrohelea spinosa</emphasis>
</taxonomicName>
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<emphasis id="63AEEAAC9D39FF9EFDE58B9DFD9C88F8" bold="true" box="[566,625,1979,2000]" pageId="9" pageNumber="264">n. sp.</emphasis>
</taxonomicNameLabel>
Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia.
</paragraph>
</caption>
<caption id="05A566369D3AFF9DFF448B95FE6188C0" httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="10" pageNumber="265" targetBox="[203,1371,219,1911]" targetPageId="10">
<paragraph id="516536BE9D3AFF9DFF448B95FE6188C0" blockId="10.[151,1436,1971,2024]" pageId="10" pageNumber="265">
<emphasis id="63AEEAAC9D3AFF9DFF448B95FEB888E1" bold="true" box="[151,341,1971,1994]" pageId="10" pageNumber="265">FIGURES 2528.</emphasis>
<taxonomicName id="96DA4D3D9D3AFF9DFE8F8B92FDC488E2" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[348,553,1972,1994]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D3AFF9DFE8F8B92FDC488E2" box="[348,553,1972,1994]" italics="true" pageId="10" pageNumber="265">Austrohelea spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3AFF9DFDE28B92FD8688E1" box="[561,619,1972,1993]" pageId="10" pageNumber="265" rank="species">
<emphasis id="63AEEAAC9D3AFF9DFDE28B92FD8688E1" bold="true" box="[561,619,1972,1993]" pageId="10" pageNumber="265">n. sp.</emphasis>
</taxonomicNameLabel>
Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 710 and spermathecae.
</paragraph>
</caption>
<subSubSection id="19C065359D3BFF9CFF148CB1FE6F883A" pageId="11" pageNumber="266" type="description">
<paragraph id="516536BE9D3BFF9CFF148CB1FE678B31" blockId="11.[151,1437,151,1990]" pageId="11" pageNumber="266">
<emphasis id="63AEEAAC9D3BFF9CFF148CB1FEE68F98" bold="true" box="[199,267,151,176]" pageId="11" pageNumber="266">Male.</emphasis>
Head (
<figureCitation id="C9E12A3B9D3BFF9CFE888CB1FE468F98" box="[347,427,151,176]" captionStart="FIGURES 20 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 20</figureCitation>
) dark brown. Eyes barely contiguous with fine interommatidial spicules. Antennal pedicel very dark brown; flagellomeres 2̄8 globular, 9̄10 slightly elongate, cylindrical, 11̄13 elongate, 11 longest; plume sparse, well developed, extending beyond apex of flagellomere 11; flagellomere 12 with single subapical sensillum coeloconicum; antennal ratio 0.72̄0.77 (0.75, n = 3). Clypeus with 2̄3 pairs of setae. Palpus slender, extending beyond apex of proboscis; segment 3 without apical pit; palpal ratio 3.64̄4.02 (3.78, n = 3). Thorax. Dark brown; scutellum slightly paler with 9̄11 large, 18̄25 smaller setae. Legs (
<figureCitation id="C9E12A3B9D3BFF9CFBE08D6AFB688E4C" box="[1075,1157,332,357]" captionStart="FIGURES 20 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 21</figureCitation>
) dark brown, fore femur slightly paler; fore, mid femora stout, hind femur greatly swollen; hind tibia broader than fore, mid tibiae, hind tibia (
<figureCitation id="C9E12A3B9D3BFF9CFF4C8DB2FF188E84" box="[159,245,404,429]" captionStart="FIGURES 20 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 22</figureCitation>
) with 3 long, black subapical spines and moderately numerous coarse setae on ventral surface (only insertions remain on most specimens); hind tibial comb with 5 long spines; hind tarsomere 1 with large basal, apical spines; hind tibial ratio 7.90̄8.57 (8.16, n = 3); hind tarsal ratio 1.82̄2.00 (1.89, n = 3); tarsomeres 4 cylindrical; claws short, equal size, without basal inner teeth, apices bifid. Wing (
<figureCitation id="C9E12A3B9D3BFF9CFBE18DD9FB698D30" box="[1074,1156,511,536]" captionStart="FIGURES 20 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Fig. 23</figureCitation>
) membrane mostly pale, lightly infuscated on costa, radius and base of medius and on membrane between these veins; veins M1, M2, CuA1, CuA2 and A infuscated; base of M2 obsolete; macrotrichia present on costa, radius and a few along anterior margin of cell r3; wing length 1.50̄1.60 (1.65, n = 3) mm, width 0.50̄0.54 (0.53, n = 3) mm; costal ratio 0.58̄0.62 (0.60, n = 3). Halter pale. Abdomen. Brown. Genitalia (
<figureCitation id="C9E12A3B9D3BFF9CFD1A8EBEFCC58D99" box="[713,808,664,689]" captionStart="FIGURES 20 24" captionStartId="9.[151,264,1978,2001]" captionTargetBox="[260,1331,213,1923]" captionTargetId="figure@9.[234,1347,185,1970]" captionTargetPageId="9" captionText="FIGURES 20 24. Austrohelea spinosa n. sp. Male. 20. head. 21. legs. 22. hind tibial comb and tarsomere 1. 23. wing. 24. genitalia." httpUri="https://zenodo.org/record/806125/files/figure.png" pageId="11" pageNumber="266">Figs. 24</figureCitation>
,
<figureCitation id="C9E12A3B9D3BFF9CFCE68EBFFCB98D99" box="[821,852,665,689]" captionStart="FIGURES 29 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="11" pageNumber="266">33</figureCitation>
̄34). Tergite 9 tapering gradually distally, posterior margin straight, apicolateral processes cylindrical, moderately long with a single apical seta, cercus hyaline, slender with 3-4 setae; sternite 9 2.2 x broader than long with narrow shallow, rounded posteromedian excavation. Gonocoxite short, stout, tapering slightly distally, with a small basomesal tubercle; gonostylus 1.3 x longer than gonocoxite, nearly straight, apex broad, spoon-shaped, tip pointed. Parameres (
<figureCitation id="C9E12A3B9D3BFF9CFBE88F0EFB7C8C69" box="[1083,1169,808,833]" captionStart="FIGURES 29 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="11" pageNumber="266">Fig. 33</figureCitation>
) separate, boomerangshaped; basal apodeme very broad, forming an obtuse angle between anterior and posterior arms; distal portion broadest proximally, curved, tapered distally to mid length, apices recurved, with widely divergent pointed tips. Aedeagus (
<figureCitation id="C9E12A3B9D3BFF9CFEC48FB2FE868C85" box="[279,363,916,941]" captionStart="FIGURES 29 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 34. Austrohelea shannoni (Wirth &amp; Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="11" pageNumber="266">Fig. 34</figureCitation>
) triangular, heavily sclerotized; basal arch U-shaped, extending 1/3 of total aedeagus length; basal arm broad, recurved 90˚ near base, distal portion long, tapered apically, apex rounded; distal portion tapering abruptly distally to a narrow sharply pointed tip, with a lightly sclerotized underlying apical process with a bulbous apex and rounded tip.
</paragraph>
<paragraph id="516536BE9D3BFF9CFF148802FE6F883A" blockId="11.[151,1437,151,1990]" pageId="11" pageNumber="266">
<emphasis id="63AEEAAC9D3BFF9CFF148802FEC88B15" bold="true" box="[199,293,1060,1085]" pageId="11" pageNumber="266">Female.</emphasis>
Head (
<figureCitation id="C9E12A3B9D3BFF9CFEAD8802FE3E8B15" box="[382,467,1060,1085]" captionStart="FIGURES 25 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 25</figureCitation>
) dark brown. Eyes barely contiguous ventrally, with fine interommatidial spicules. Antennal pedicel very dark brown; flagellum dark brown, flagellomeres 2̄8 (
<figureCitation id="C9E12A3B9D3BFF9CFBC8886EFB828B49" box="[1051,1135,1096,1121]" captionStart="FIGURES 25 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 26</figureCitation>
) moderately short, barrelshaped, 9̄13 elongate, cylindrical; flagellomeres 8, 12 with single subapical sensillum coeloconicum; antennal ratio 1.36̄1.47 (1.39, n = 5). Clypeus with 2̄4 pairs of setae. Palpus (
<figureCitation id="C9E12A3B9D3BFF9CFC6088B6FBE98B81" box="[947,1028,1168,1193]" captionStart="FIGURES 25 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 26</figureCitation>
) dark brown; segment 3 moderately slender with a few capitate sensilla on distal 1/3 of mesal surface; palpal ratio 3.00 (n = 5). Mandible with 8̄11 (9, n = 5) large teeth. Thorax. Scutum dark brown, humeral areas and scutellum paler; scutellum with 6̄7 large, 10̄12 smaller setae. Legs dark brown, hind tibia and femur darkest; fore, mid femora moderately swollen, hind femur greatly swollen; hind tibia without subapical spines, stouter than fore, mid tibiae; hind tibial comb with 5 large spines; hind tibial ratio 5.93̄7.17 (6.42, n = 5); tarsi dark brown, darkest on hind leg; tarsomeres 1 with large basal, apical ventral spines, 2 additional similar spines along ventral surface of mid leg, tarsomeres 2̄3 of fore, mid legs with apical spine; tarsomeres 1̄2 of hind leg with two rows of ventral palisade setae; hind tarsal ratio 1.76̄1.88 (1.83, n = 5); tarsomere 4 of fore, mid legs short, cordiform, tarsomere 4 of hind leg elongate, cylindrical; claws small, equal on fore, mid legs, hind leg with single long claw that is slightly longer than tarsomere 5 with a stout basal tooth about 1/3 as long as claw. Wing (
<figureCitation id="C9E12A3B9D3BFF9CFD5589DEFD3A8939" box="[646,727,1528,1553]" captionStart="FIGURES 25 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 27</figureCitation>
) membrane infuscated, particularly dark along veins and in and around radial cells; two well developed radial cells, 2nd broader, twice as long as1st; macrotrichia fairly numerous on distal ½ of cell r3, apices of m1, m2, a few in cua1 and anal cell; wing length 1.57̄1.68 (1.63, n = 5) mm, width 0.65̄0.68 (0.67, n = 5) mm; costal ratio 0.64̄0.66 (0.65, n = 5). Halter pale. Abdomen (
<figureCitation id="C9E12A3B9D3BFF9CFB788A4CFB1289AA" box="[1195,1279,1642,1667]" captionStart="FIGURES 25 28" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[203,1371,219,1911]" captionTargetId="figure@10.[173,1408,190,1946]" captionTargetPageId="10" captionText="FIGURES 25 28. Austrohelea spinosa n. sp. Female. 25. head. 26. palpi and mouthparts. 27. wing. 28. abdominal segments 7 10 and spermathecae." httpUri="https://zenodo.org/record/806127/files/figure.png" pageId="11" pageNumber="266">Fig. 28</figureCitation>
) dark brown. Sternite 8 with moderately deep V-shaped posteromedian excavation; sternite 9 separated, each half tapered distally with hook-like apices; sternite 10 with two pairs of setae. Two ovoid, subequal size spermathecae with moderately long, broad necks, measuring 55̄62 (59, n = 5) µm by 35̄42 (39, n = 5) µm; a small rudimentary third spermatheca present.
</paragraph>
</subSubSection>
<subSubSection id="19C065359D3BFF9CFF148B3BFA9188EE" pageId="11" pageNumber="266" type="materials_examined">
<paragraph id="516536BE9D3BFF9CFF148B3BFA9188EE" blockId="11.[151,1437,151,1990]" pageId="11" pageNumber="266">
<materialsCitation id="E1B23CE39D3BFF9CFF148B3BFED48872" ID-GBIF-Occurrence="1563642360" collectingDate="2004-01-15" collectionCode="MLPA" collectorName="Villa La Angostura &amp; G. Spinelli" country="Argentina" location="Neuquen" pageId="11" pageNumber="266" specimenCount="2" stateProvince="Neuquen" typeStatus="holotype">
<emphasis id="63AEEAAC9D3BFF9CFF148B3BFE95881E" bold="true" box="[199,376,1821,1846]" pageId="11" pageNumber="266">
<typeStatus id="8E61881C9D3BFF9CFF148B3BFEEF881E" box="[199,258,1821,1846]" pageId="11" pageNumber="266">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8E61881C9D3BFF9CFE508B3BFE06881E" box="[387,491,1821,1846]" pageId="11" pageNumber="266" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="29CD762E9D3BFF9CFDED8B38FD09881E" box="[574,740,1822,1846]" name="Argentina" pageId="11" pageNumber="266">ARGENTINA</collectingCountry>
,
<collectingRegion id="931EF85C9D3BFF9CFD218B39FCB7881E" box="[754,858,1823,1846]" country="Argentina" name="Neuquen" pageId="11" pageNumber="266">Neuquen</collectingRegion>
,
<collectorName id="FC2F53689D3BFF9CFCBA8B3BFBA7881E" box="[873,1098,1821,1846]" pageId="11" pageNumber="266">Villa La Angostura</collectorName>
, 11/
<collectingDate id="3520E9969D3BFF9CFBA88B3BFB03881E" box="[1147,1262,1821,1846]" pageId="11" pageNumber="266" value="2004-01-15">
<date id="2564107E9D3BFF9CFBA88B3BFB03881E" box="[1147,1262,1821,1846]" pageId="11" pageNumber="266" value="2004-01-15">15-I-2004</date>
</collectingDate>
,
<collectorName id="FC2F53689D3BFF9CFB2E8B38FA94881E" box="[1277,1401,1821,1846]" pageId="11" pageNumber="266">G. Spinelli</collectorName>
, at light (
<collectionCode id="37CBAE7B9D3BFF9CFF0C8B64FEDC8872" box="[223,305,1858,1882]" pageId="11" pageNumber="266">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1B23CE39D3BFF9CFE998B64FA748856" ID-GBIF-Occurrence="1563642355" collectingDate="2006-12-14" collectingDateMax="2007-01-08" collectingDateMin="2006-12-14" collectionCode="MLPA" collectorName="La Cortadera &amp; A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="769" latitude="-41.086945" location="La Cortadera" longLatPrecision="19" longitude="-71.80723" municipality="Parque Nacional Nahuel Huapi" pageId="11" pageNumber="266" specimenCount="1" stateProvince="Rio Negro" typeStatus="allotype">
<typeStatus id="8E61881C9D3BFF9CFE998B64FE498873" box="[330,420,1858,1883]" pageId="11" pageNumber="266" type="allotype">allotype</typeStatus>
female,
<collectingCountry id="29CD762E9D3BFF9CFDC28B64FD6E8873" box="[529,643,1858,1883]" name="Argentina" pageId="11" pageNumber="266">Argentina</collectingCountry>
,
<collectingRegion id="931EF85C9D3BFF9CFD468B64FCFE8873" box="[661,787,1858,1883]" country="Argentina" name="Rio Negro" pageId="11" pageNumber="266">Río Negro</collectingRegion>
,
<collectingMunicipality id="B101ACC49D3BFF9CFCF78B65FB7B8873" box="[804,1174,1858,1883]" pageId="11" pageNumber="266">Parque Nacional Nahuel Huapi</collectingMunicipality>
, mallín
<collectorName id="FC2F53689D3BFF9CFB2F8B65FA758873" box="[1276,1432,1858,1883]" pageId="11" pageNumber="266">
<location id="540560659D3BFF9CFB2F8B65FA758873" LSID="urn:lsid:plazi:treatment:D97387A89D38FF9AFF448AA1FCC38960:540560659D3BFF9CFB2F8B65FA758873" box="[1276,1432,1858,1883]" country="Argentina" latitude="-41.086945" longLatPrecision="19" longitude="-71.80723" municipality="Parque Nacional Nahuel Huapi" name="La Cortadera" pageId="11" pageNumber="266" stateProvince="Rio Negro">La Cortadera</location>
</collectorName>
,
<geoCoordinate id="34EE50799D3BFF9CFF448B43FEF48856" box="[151,281,1893,1918]" direction="south" orientation="latitude" pageId="11" pageNumber="266" precision="15" value="-41.086945">41°05'13&quot;S</geoCoordinate>
,
<geoCoordinate id="34EE50799D3BFF9CFEFF8B43FE5B8856" box="[300,438,1893,1918]" direction="west" orientation="longitude" pageId="11" pageNumber="266" precision="15" value="-71.80723">71°48'26&quot;W</geoCoordinate>
,
<elevation id="DAF7D18D9D3BFF9CFE1A8B43FDE68856" box="[457,523,1893,1918]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="11" pageNumber="266" unit="m" value="769.0">
<quantity id="96229B5B9D3BFF9CFE1A8B43FDE68856" box="[457,523,1893,1918]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="11" pageNumber="266" unit="m" value="769.0">769m</quantity>
</elevation>
,
<collectingDate id="3520E9969D3BFF9CFDCD8B43FD428856" box="[542,687,1893,1918]" pageId="11" pageNumber="266" value="2006-12-14">
<date id="2564107E9D3BFF9CFDCD8B43FD428856" box="[542,687,1893,1918]" pageId="11" pageNumber="266" value="2006-12-14">14-XII-2006</date>
</collectingDate>
/
<collectingDate id="3520E9969D3BFF9CFD648B43FCF58856" box="[695,792,1893,1918]" pageId="11" pageNumber="266" value="2007-01-08">
<date id="2564107E9D3BFF9CFD648B43FCF58856" box="[695,792,1893,1918]" pageId="11" pageNumber="266" value="2007-01-08">8-I-2007</date>
</collectingDate>
,
<collectorName id="FC2F53689D3BFF9CFCF88B40FC768856" box="[811,923,1894,1918]" pageId="11" pageNumber="266">A. Garré</collectorName>
̄
<collectorName id="FC2F53689D3BFF9CFC778B40FB6E8856" box="[932,1155,1893,1918]" pageId="11" pageNumber="266">F. Montes de Oca</collectorName>
,
<collectingMethod id="889B4EA99D3BFF9CFB458B43FAC48856" box="[1174,1321,1893,1918]" pageId="11" pageNumber="266">Malaise trap</collectingMethod>
(
<collectionCode id="37CBAE7B9D3BFF9CFAEC8B40FA7C8856" box="[1343,1425,1894,1918]" pageId="11" pageNumber="266">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E1B23CE39D3BFF9CFF448BADFB6A888A" ID-GBIF-Occurrence="1563642356" box="[151,1159,1930,1955]" collectingDate="2004-01-15" collectionCode="MLPA" collectorName="Villa La Angostura &amp; G. Spinelli" country="Argentina" location="Neuquen" pageId="11" pageNumber="266" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Neuquen" typeStatus="paratype">
<typeStatus id="8E61881C9D3BFF9CFF448BADFEEA888B" box="[151,263,1931,1955]" pageId="11" pageNumber="266" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="47DCFD379D3BFF9CFEC78BACFE9D888B" box="[276,368,1930,1955]" pageId="11" pageNumber="266" type="male">2 males</specimenCount>
,
<specimenCount id="47DCFD379D3BFF9CFEAE8BACFE02888B" box="[381,495,1930,1955]" pageId="11" pageNumber="266" type="female">4 females</specimenCount>
, as follows:
<specimenCount id="47DCFD379D3BFF9CFD518BACFD22888B" box="[642,719,1930,1955]" pageId="11" pageNumber="266" type="male">1 male</specimenCount>
with same data as holotype (
<collectionCode id="37CBAE7B9D3BFF9CFBFE8BACFB92888A" box="[1069,1151,1930,1954]" pageId="11" pageNumber="266">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E1B23CE39D3BFF9CFB478BACFA9588EE" ID-GBIF-Occurrence="1563642363" collectingDate="2006-12-14" collectingDateMax="2007-01-08" collectingDateMin="2006-12-14" collectionCode="MLPA, BMNH, USNM, FSCA" collectorName="La Cortadera &amp; A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="769" latitude="-41.086945" location="La Cortadera" longLatPrecision="19" longitude="-71.80723" municipality="Parque Nacional Nahuel Huapi" pageId="11" pageNumber="266" specimenCount="10" specimenCount-female="8" specimenCount-male="2" stateProvince="Rio Negro" typeStatus="paratype">
<specimenCount id="47DCFD379D3BFF9CFB478BACFB08888B" box="[1172,1253,1930,1955]" pageId="11" pageNumber="266" type="male">1 male</specimenCount>
,
<specimenCount id="47DCFD379D3BFF9CFB208BACFA8C888B" box="[1267,1377,1930,1955]" pageId="11" pageNumber="266" type="female">4 females</specimenCount>
with same data as allotype (
<specimenCount id="47DCFD379D3BFF9CFE4B8B88FE0988EE" box="[408,484,1965,1990]" pageId="11" pageNumber="266" type="male">1 male</specimenCount>
,
<specimenCount id="47DCFD379D3BFF9CFE258B88FDBA88EE" box="[502,599,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFDA88B88FD2488EE" box="[635,713,1966,1990]" pageId="11" pageNumber="266">MLPA</collectionCode>
,
<specimenCount id="47DCFD379D3BFF9CFD068B88FCD888EE" box="[725,821,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFC898B89FC5C88EE" box="[858,945,1967,1990]" pageId="11" pageNumber="266">BMNH</collectionCode>
,
<specimenCount id="47DCFD379D3BFF9CFC688B88FBF088EE" box="[955,1053,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFB928B88FB7888EE" box="[1089,1173,1966,1990]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="266">USNM</collectionCode>
,
<specimenCount id="47DCFD379D3BFF9CFB4C8B88FAED88EE" box="[1183,1280,1965,1990]" pageId="11" pageNumber="266" type="female">1 female</specimenCount>
in
<collectionCode id="37CBAE7B9D3BFF9CFAF68B88FA9D88EE" box="[1317,1392,1966,1990]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="11" pageNumber="266">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="05A566369D3CFF9BFF448ADBFBD4881A" httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="12" pageNumber="267" targetBox="[175,1421,225,1689]" targetPageId="12">
<paragraph id="516536BE9D3CFF9BFF448ADBFBD4881A" blockId="12.[151,1437,1789,1842]" pageId="12" pageNumber="267">
<emphasis id="63AEEAAC9D3CFF9BFF448ADBFEB9883B" bold="true" box="[151,340,1789,1812]" pageId="12" pageNumber="267">FIGURES 2934.</emphasis>
<taxonomicName id="96DA4D3D9D3CFF9BFE898ADBFD11883B" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[346,764,1789,1811]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3CFF9BFE898ADBFDDA883B" box="[346,567,1789,1811]" italics="true" pageId="12" pageNumber="267">Austrohelea shannoni</emphasis>
(Wirth &amp; Blanton)
</taxonomicName>
. Male. 29. parameres. 30. aedeagus.
<taxonomicName id="96DA4D3D9D3CFF9BFBA18ADBFAF6883C" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[1138,1307,1789,1812]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D3CFF9BFBA18ADBFAF6883C" box="[1138,1307,1789,1812]" italics="true" pageId="12" pageNumber="267">Austrohelea sirii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3CFF9BFAF18AD8FAB6883B" box="[1314,1371,1790,1811]" pageId="12" pageNumber="267" rank="species">
<emphasis id="63AEEAAC9D3CFF9BFAF18AD8FAB6883B" bold="true" box="[1314,1371,1790,1811]" pageId="12" pageNumber="267">n. sp.</emphasis>
</taxonomicNameLabel>
Male. 31. parameres. 32. aedeagus.
<taxonomicName id="96DA4D3D9D3CFF9BFE128B3AFD63881A" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[449,654,1820,1842]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D3CFF9BFE128B3AFD63881A" box="[449,654,1820,1842]" italics="true" pageId="12" pageNumber="267">Austrohelea spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3CFF9BFD478B3BFD20881A" box="[660,717,1821,1842]" pageId="12" pageNumber="267" rank="species">
<emphasis id="63AEEAAC9D3CFF9BFD478B3BFD20881A" bold="true" box="[660,717,1821,1842]" pageId="12" pageNumber="267">n. sp.</emphasis>
</taxonomicNameLabel>
Male. 33. parameres. 34. aedeagus.
</paragraph>
</caption>
<subSubSection id="19C065359D3CFF9BFF148B78FCE488CC" pageId="12" pageNumber="267" type="distribution">
<paragraph id="516536BE9D3CFF9BFF148B78FE1C88B4" blockId="12.[151,1436,1886,2020]" pageId="12" pageNumber="267">
<emphasis id="63AEEAAC9D3CFF9BFF148B78FE8D885F" bold="true" box="[199,352,1886,1911]" pageId="12" pageNumber="267">Distribution.</emphasis>
Known only from
<taxonomicName id="96DA4D3D9D3CFF9BFDE68B79FD5B885F" authorityName="Blume" authorityYear="1850" box="[565,694,1887,1911]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="267" phylum="Tracheophyta" rank="genus">
<emphasis id="63AEEAAC9D3CFF9BFDE68B79FD5B885F" box="[565,694,1887,1911]" italics="true" pageId="12" pageNumber="267">Nothofagus</emphasis>
</taxonomicName>
forests of Nahuel Huapi National Park in
<collectingRegion id="931EF85C9D3CFF9BFB5F8B46FB1C885F" box="[1164,1265,1888,1911]" country="Argentina" name="Neuquen" pageId="12" pageNumber="267">Neuquen</collectingRegion>
and
<collectingRegion id="931EF85C9D3CFF9BFAF48B79FA718850" box="[1319,1436,1887,1912]" country="Argentina" name="Rio Negro" pageId="12" pageNumber="267">Río Negro</collectingRegion>
provinces,
<collectingCountry id="29CD762E9D3CFF9BFEC18BA5FE6C88B4" box="[274,385,1923,1948]" name="Argentina" pageId="12" pageNumber="267">Argentina</collectingCountry>
(
<figureCitation id="C9E12A3B9D3CFF9BFE438BA5FE0988B4" box="[400,484,1923,1948]" captionStart="FIGURE 35" captionStartId="13.[151,250,1234,1256]" captionTargetBox="[267,1325,252,1118]" captionTargetId="figure@13.[205,1396,196,1194]" captionTargetPageId="13" captionText="FIGURE 35. Map of Argentine and Chilean Patagonia with distribution records of: Austrohelea shannoni (Wirth &amp; Blanton), green circles. Austrohelea sirii n. sp., yellow triangles. Austrohelea spinosa n. sp., red diamonds." httpUri="https://zenodo.org/record/806131/files/figure.png" pageId="12" pageNumber="267">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="516536BE9D3CFF9BFF148B80FCE488CC" blockId="12.[151,1436,1886,2020]" pageId="12" pageNumber="267">
<emphasis id="63AEEAAC9D3CFF9BFF148B80FDB78897" bold="true" box="[199,602,1958,1983]" pageId="12" pageNumber="267">Derivation of the specific epithet.</emphasis>
The specific name of this new species is Latin, in reference to the long, coarse, subapical spine-like setae of the male hind tibia.
</paragraph>
</subSubSection>
<caption id="05A566369D3DFF9AFF4488F4FB9C8A2F" httpUri="https://zenodo.org/record/806131/files/figure.png" pageId="13" pageNumber="268" targetBox="[267,1325,252,1118]" targetPageId="13">
<paragraph id="516536BE9D3DFF9AFF4488F4FB9C8A2F" blockId="13.[151,1436,1234,1287]" pageId="13" pageNumber="268">
<emphasis id="63AEEAAC9D3DFF9AFF4488F4FECD8BCF" bold="true" box="[151,288,1234,1256]" pageId="13" pageNumber="268">FIGURE 35.</emphasis>
Map of Argentine and Chilean Patagonia with distribution records of:
<taxonomicName id="96DA4D3D9D3DFF9AFC2788F4FA758BC0" authority="(Wirth &amp; Blanton)" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[1012,1432,1234,1256]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3DFF9AFC2788F4FB3F8BC0" box="[1012,1234,1234,1256]" italics="true" pageId="13" pageNumber="268">Austrohelea shannoni</emphasis>
(Wirth &amp; Blanton)
</taxonomicName>
, green circles.
<taxonomicName id="96DA4D3D9D3DFF9AFEF788D7FE208A2F" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[292,461,1265,1287]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="63AEEAAC9D3DFF9AFEF788D7FE208A2F" box="[292,461,1265,1287]" italics="true" pageId="13" pageNumber="268">Austrohelea sirii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3DFF9AFE0088D7FDE18A2E" box="[467,524,1265,1286]" pageId="13" pageNumber="268" rank="species">
<emphasis id="63AEEAAC9D3DFF9AFE0088D7FDE18A2E" bold="true" box="[467,524,1265,1286]" pageId="13" pageNumber="268">n. sp.</emphasis>
</taxonomicNameLabel>
, yellow triangles.
<taxonomicName id="96DA4D3D9D3DFF9AFD1488D7FC798A2F" authority="Ronderos &amp; Spinelli &amp; Grogan, 2017" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[711,916,1265,1287]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="63AEEAAC9D3DFF9AFD1488D7FC798A2F" box="[711,916,1265,1287]" italics="true" pageId="13" pageNumber="268">Austrohelea spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789D57D79D3DFF9AFC4888D7FC398A2E" box="[923,980,1265,1286]" pageId="13" pageNumber="268" rank="species">
<emphasis id="63AEEAAC9D3DFF9AFC4888D7FC398A2E" bold="true" box="[923,980,1265,1286]" pageId="13" pageNumber="268">n. sp.</emphasis>
</taxonomicNameLabel>
, red diamonds.
</paragraph>
</caption>
<subSubSection id="19C065359D3DFF9AFF148915FCC38960" pageId="13" pageNumber="268" type="discussion">
<paragraph id="516536BE9D3DFF9AFF148915FE428928" blockId="13.[151,1436,1331,1608]" pageId="13" pageNumber="268">
<emphasis id="63AEEAAC9D3DFF9AFF148915FEA18A64" bold="true" box="[199,332,1331,1356]" pageId="13" pageNumber="268">Discussion.</emphasis>
Males of
<taxonomicName id="96DA4D3D9D3DFF9AFE188912FDA98A63" authorityName="Ronderos &amp; Spinelli &amp; Grogan" authorityYear="2017" box="[459,580,1332,1355]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="63AEEAAC9D3DFF9AFE188912FE318A63" box="[459,476,1332,1355]" italics="true" pageId="13" pageNumber="268">A</emphasis>
.
<emphasis id="63AEEAAC9D3DFF9AFE3C8912FDA98A63" box="[495,580,1332,1355]" italics="true" pageId="13" pageNumber="268">spinosa</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="96DA4D3D9D3DFF9AFC768912FBDD8A64" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[933,1072,1332,1356]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3DFF9AFC768912FC5B8A63" box="[933,950,1332,1355]" italics="true" pageId="13" pageNumber="268">A</emphasis>
.
<emphasis id="63AEEAAC9D3DFF9AFC1A8912FBDD8A64" box="[969,1072,1332,1356]" italics="true" pageId="13" pageNumber="268">shannoni</emphasis>
</taxonomicName>
as they have similar genitalia, however; this new species differs from this previously described congener by the very long stout, black, subapical spines on the hind tibia of males; the posterior margin of tergite 9 is straight and the apicolateral processes are slender and much longer; the gonostylus is longer than the gonocoxite; the parameres are very broad and their basal apodemes form a shallow obtuse angle between the anterior and posterior arms; and the basal arch of the aedeagus is broader and U-shaped.
</paragraph>
<paragraph id="516536BE9D3DFF9AFF148A2DFCC38960" blockId="13.[151,1436,1331,1608]" pageId="13" pageNumber="268">
Females of this new species can be distinguished from those of
<taxonomicName id="96DA4D3D9D3DFF9AFC648A2AFBAD890C" authorityName="Wirth &amp; Grogan" authorityYear="1988" baseAuthorityName="Wirth &amp; Blanton" baseAuthorityYear="1972" box="[951,1088,1548,1572]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="63AEEAAC9D3DFF9AFC648A2AFC25890B" box="[951,968,1548,1571]" italics="true" pageId="13" pageNumber="268">A</emphasis>
.
<emphasis id="63AEEAAC9D3DFF9AFC098A2AFBAD890C" box="[986,1088,1548,1572]" italics="true" pageId="13" pageNumber="268">shannoni</emphasis>
</taxonomicName>
by their greatly swollen hind femora and subequal size spermathecae with shorter necks.
</paragraph>
</subSubSection>
</treatment>
</document>