287 lines
37 KiB
XML
287 lines
37 KiB
XML
<document id="2BE4C1AEB62800143476E5FE7E1A9676" ID="10.11646/zootaxa.4276.2.7SLASH11313" ID-CLB-Dataset="32860" ID-DOI="10.11646/zootaxa.4276.2.7" ID-GBIF-Dataset="1cb0e227-6a7f-41f2-8ac7-4a98a9641ad2" ID-ISSN="1175-5326" ID-Zenodo-Dep="806115" ID-ZooBank="72DC4B80-2CFE-414F-AA86-DAA73618D977" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497253899257" checkinUser="plazi" docAuthor="Ronderos, Maria M., Spinelli, Gustavo R. & Grogan, Willliam L." docDate="2017" docId="D97387A89D35FF9FFF448805FB8F891D" docLanguage="en" docName="zootaxa.4276.2.7SLASH11313.pdf" docOrigin="Zootaxa 4276 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austrohelea sirii Ronderos & Spinelli & Grogan 2017, New Species" docType="treatment" docVersion="8" lastPageNumber="263" masterDocId="254AFFD09D30FF97FFD38C26FFED8F28" masterDocTitle="The Neotropical species of the predaceous midge genus Austrohelea Wirth & Grogan (Diptera: Ceratopogonidae)" masterLastPageNumber="269" masterPageNumber="255" pageNumber="260" updateTime="1698440399754" updateUser="ExternalLinkService">
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<mods:mods id="27008F062710DB3BB003FD594C7FAEB1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E4556578201DEF3B66AFF5C6406AEC92">
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<mods:title id="16B1E37253974670D31DBF08BD29B38E">The Neotropical species of the predaceous midge genus Austrohelea Wirth & Grogan (Diptera: Ceratopogonidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="EA45C96730BCFEAA545A8D5735D43FF8" type="personal">
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<mods:role id="B7E51FAC9541AC2279264B6A7308C326">
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<mods:roleTerm id="EDECBFECBE26D2D200CA6FA1FBB78F6D">Author</mods:roleTerm>
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<mods:namePart id="9CA57E1B040A7A58AC91E7056F843FA1">Ronderos, Maria M.</mods:namePart>
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<mods:name id="208AE452758C89BB8D35B3FA4B3EC31C" type="personal">
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<mods:roleTerm id="7402667AF0C8E4DB2E5129B812B46EC3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C4A315556AA8A06C0EE5487779922902">Spinelli, Gustavo R.</mods:namePart>
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</mods:name>
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<mods:name id="33F48BD5AB1FCD3B0FC0103DBC31BF79" type="personal">
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<mods:role id="27687FF868FD674532B7945B07176476">
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<mods:roleTerm id="61649A8DE6276EF11E352874C2B3A202">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EE4BC4163F7A8ACEF0502FD3DD8889D9">Grogan, Willliam L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="79DB23E5CFC968EFF16964C6F6916EBF">text</mods:typeOfResource>
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<mods:titleInfo id="9310E2C87A74CE284EC8BC80125CA82C">
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<mods:title id="E0919618618AE3ECE953FF89A6F0971A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F41CA3957867F503FDE9E7B1812298F4">
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<mods:date id="676B5E6058373F1062D81D22D8A5CDC4">2017</mods:date>
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<mods:number id="72F07E1F306716D7FF0E00E51A4C7484">4276</mods:number>
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<mods:number id="E03185347CABFCAEDF2EAF7656A5F2DB">2</mods:number>
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<mods:end id="BD80E3F3513C079A1622887C42BF024F">269</mods:end>
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<mods:classification id="8A54F2EEBEF39C189C7D51E61DA45FF1">journal article</mods:classification>
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<mods:identifier id="3B486BEEB0D291B4F4D5D9234DE23490" type="CLB-Dataset">32860</mods:identifier>
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<mods:identifier id="26A89F53CD73DEA5E92ED09E1A85F396" type="DOI">10.11646/zootaxa.4276.2.7</mods:identifier>
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<treatment id="D97387A89D35FF9FFF448805FB8F891D" ID-DOI="http://doi.org/10.5281/zenodo.6017419" ID-GBIF-Taxon="131876630" ID-Zenodo-Dep="6017419" LSID="urn:lsid:plazi:treatment:D97387A89D35FF9FFF448805FB8F891D" httpUri="http://treatment.plazi.org/id/D97387A89D35FF9FFF448805FB8F891D" lastPageId="8" lastPageNumber="263" pageId="5" pageNumber="260">
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<subSubSection id="19C065359D35FF92FF448805FE688B77" pageId="5" pageNumber="260" type="nomenclature">
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<paragraph id="516536BE9D35FF92FF448805FDEC8B16" blockId="5.[151,513,1059,1119]" box="[151,513,1059,1086]" pageId="5" pageNumber="260">
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<heading id="0A2D81D29D35FF92FF448805FDEC8B16" bold="true" box="[151,513,1059,1086]" fontSize="11" level="1" pageId="5" pageNumber="260" reason="1">
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<emphasis id="63AEEAAC9D35FF92FF448805FDEC8B16" bold="true" box="[151,513,1059,1086]" pageId="5" pageNumber="260">
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<taxonomicName id="96DA4D3D9D35FF92FF448805FE8D8B15" ID-CoL="9HTKS" authority="Ronderos & Spinelli & Grogan, 2017" authorityName="Ronderos & Spinelli & Grogan" authorityYear="2017" box="[151,352,1059,1085]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
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<emphasis id="63AEEAAC9D35FF92FF448805FE8D8B15" bold="true" box="[151,352,1059,1085]" italics="true" pageId="5" pageNumber="260">Austrohelea sirii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="789D57D79D35FF92FEB48802FDEC8B16" box="[359,513,1060,1086]" pageId="5" pageNumber="260" rank="species">New Species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="516536BE9D35FF92FF448860FE688B77" blockId="5.[151,513,1059,1119]" box="[151,389,1094,1119]" pageId="5" pageNumber="260">
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(
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<figureCitation id="C9E12A3B9D35FF92FF4C8860FF118B77" box="[159,252,1094,1119]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="5" pageNumber="260">Figs. 13</figureCitation>
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̄19, 31̄32)
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</paragraph>
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</subSubSection>
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<subSubSection id="19C065359D35FF92FF4488A8FAB78AA8" pageId="5" pageNumber="260" type="diagnosis">
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<paragraph id="516536BE9D35FF92FF4488A8FAB78AA8" blockId="5.[151,1437,1166,2028]" pageId="5" pageNumber="260">
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<emphasis id="63AEEAAC9D35FF92FF4488A8FEFC8B8F" bold="true" box="[151,273,1166,1191]" pageId="5" pageNumber="260">Diagnosis.</emphasis>
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A small (male wing length
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<quantity id="96229B5B9D35FF92FD8488A8FD528B8F" box="[599,703,1166,1191]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" pageId="5" pageNumber="260" unit="mm" value="1.06">1.06 mm</quantity>
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, female wing length
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<quantity id="96229B5B9D35FF92FC6088A8FBF38B8F" box="[947,1054,1166,1191]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="260" unit="mm" value="1.16">1.16 mm</quantity>
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), brown species. Both sexes with sensory pit on third palpal segment and slender legs. Males with posterior margin of tergite 9 deeply concave with elongate, cylindrical, slightly divergent apicolateral processes; gonocoxite short, stout, without basomesal tubercle; gonostylus longer than gonocoxite; parameres with moderately slender basal apodeme with an elongate, curved anterior arm; aedeagus triangular with slender, slightly curved basal arms, very low basal arch, and lightly sclerotized distal portion with rounded apex. Females with wing darkly infuscated with dense, coarse macrotrichia in all cells beyond r-m crossvein; and unequal size, pyriform spermathecae with short, broad, truncate necks.
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</paragraph>
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</subSubSection>
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<subSubSection id="19C065359D35FF9FFF1489ADFBC98B31" lastPageId="8" lastPageNumber="263" pageId="5" pageNumber="260" type="materials_examined">
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<paragraph id="516536BE9D35FF9FFF1489ADFD9B8FD0" blockId="5.[151,1437,1166,2028]" lastBlockId="8.[151,1437,151,1589]" lastPageId="8" lastPageNumber="263" pageId="5" pageNumber="260">
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<emphasis id="63AEEAAC9D35FF92FF1489ADFE6F8A8C" bold="true" box="[199,386,1419,1444]" pageId="5" pageNumber="260">
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<typeStatus id="8E61881C9D35FF92FF1489ADFED98A8C" box="[199,308,1419,1444]" pageId="5" pageNumber="260" type="holotype">Holotype</typeStatus>
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male.
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</emphasis>
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Head (
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<figureCitation id="C9E12A3B9D35FF92FE3389ADFDD58A8B" box="[480,568,1419,1444]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="5" pageNumber="260">Fig. 13</figureCitation>
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) brown. Eyes narrowly contiguous with sparse interommatidial spicules. Antennal pedicel dark brown; flagellomeres 2̄8 globular, 9̄10 vasiform, 11̄13 elongate, 11 longest; plume sparse, well developed, extending beyond flagellomere 11; flagellomere 12 with single subapical sensillum coeloconicum; antennal ratio 0.81. Clypeus with 3 pairs of setae. Palpus moderately long, segments 4̄5 extending beyond proboscis; segment 3 moderately broad with small apicomesal pit; palpal ratio 2.33. Thorax brown except for pale scutellum; scutellum with 6 large, 8 smaller setae. Legs similar to those of female except for tarsal claws, which are shorter, equal size, without basal inner teeth, apices bifid; hind tibial ratio 9.00; hind tarsal ratio 1.68. Wing (
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<figureCitation id="C9E12A3B9D35FF92FF308AA1FED889B7" box="[227,309,1671,1696]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="5" pageNumber="260">Fig. 14</figureCitation>
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) membrane slightly infuscated, surface with dense macrotrichia along distal margin of cell r3, sparse above and below distal portion of vein M1 and apex of cell m1; veins brown, base of M2 thin, joined to M1; radial cells well developed, 2nd broader, nearly twice as long as 1st; wing length
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<quantity id="96229B5B9D35FF92FC3B8AF1FBBD89C7" box="[1000,1104,1751,1776]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" pageId="5" pageNumber="260" unit="mm" value="1.06">1.06 mm</quantity>
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, width
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<quantity id="96229B5B9D35FF92FB748AFEFAFD89C7" box="[1191,1296,1752,1776]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="260" unit="mm" value="0.38">0.38 mm</quantity>
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; costal ratio 0.57. Halter pale. Genitalia (
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<figureCitation id="C9E12A3B9D35FF92FE078ADDFDDC883C" box="[468,561,1787,1812]" captionStart="FIGURES 13 – 15" captionStartId="6.[151,264,2006,2029]" captionTargetBox="[201,1416,202,1972]" captionTargetId="figure@6.[176,1424,185,1999]" captionTargetPageId="6" captionText="FIGURES 13 – 15. Austrohelea sirii n. sp. Male. 13. head. 14. wing. 15. genitalia." httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="5" pageNumber="260">Figs. 15</figureCitation>
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,
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<figureCitation id="C9E12A3B9D35FF92FDEE8ADAFDB6883C" box="[573,603,1788,1812]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth & Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="5" pageNumber="260">31</figureCitation>
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̄32). Tergite 9 tapering gradually distally, posterior margin narrow, deeply concave, apicolateral processes elongate, cylindrical, slightly divergent with a single apical seta; cercus slender; sternite 9 3.5 x broader than long with narrow, moderately deep, concave posteromedian excavation. Gonocoxite short, stout, 1.25 x longer than broad, apex broad, sinuate; gonostylus 1.2 x longer than gonocoxite, broadest on proximal 1/5, tapering gradually distally, distal 2/3 slightly curved, apex rounded. Parameres (
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<figureCitation id="C9E12A3B9D35FF92FB008BADFACA888C" box="[1235,1319,1931,1956]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth & Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="5" pageNumber="260">Fig. 31</figureCitation>
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) separate, moderately slender, somewhat boomerang-shaped; basal apodeme with elongate anterior arm, apical 1/3 curved, posterior arm shorter with slightly expanded blunt apex; distal portions moderately short, curved distally, apices pointed, divergent. Aedeagus (
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<figureCitation id="C9E12A3B9D38FF9FFDD18CB1FDB68F98" box="[514,603,151,176]" captionStart="FIGURES 29 – 34" captionStartId="12.[151,264,1789,1812]" captionTargetBox="[175,1421,225,1689]" captionTargetId="figure@12.[151,1436,193,1768]" captionTargetPageId="12" captionText="FIGURES 29 – 34. Austrohelea shannoni (Wirth & Blanton). Male. 29. parameres. 30. aedeagus. Austrohelea sirii n. sp. Male. 31. parameres. 32. aedeagus. Austrohelea spinosa n. sp. Male. 33. parameres. 34. aedeagus." httpUri="https://zenodo.org/record/806129/files/figure.png" pageId="8" pageNumber="263">Fig. 32</figureCitation>
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) triangular; basal arch very low, slightly concave; basal arm heavily sclerotized, slender, nearly straight, laterally directed with blunt apex; distal portion barely discernible, lightly sclerotized, tapered distally, apex rounded.
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</paragraph>
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<caption id="05A566369D36FF91FF448BF0FC3988C5" box="[151,980,2006,2029]" httpUri="https://zenodo.org/record/806121/files/figure.png" pageId="6" pageNumber="261" targetBox="[201,1416,202,1972]" targetPageId="6">
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<paragraph id="516536BE9D36FF91FF448BF0FC3988C5" blockId="6.[151,980,2006,2029]" box="[151,980,2006,2029]" pageId="6" pageNumber="261">
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<emphasis id="63AEEAAC9D36FF91FF448BF0FEBE88C4" bold="true" box="[151,339,2006,2029]" pageId="6" pageNumber="261">FIGURES 13–15.</emphasis>
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<taxonomicName id="96DA4D3D9D36FF91FE898BF1FDEF88C5" authority="Ronderos & Spinelli & Grogan, 2017" authorityName="Ronderos & Spinelli & Grogan" authorityYear="2017" box="[346,514,2007,2029]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
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<emphasis id="63AEEAAC9D36FF91FE898BF1FDEF88C5" box="[346,514,2007,2029]" italics="true" pageId="6" pageNumber="261">Austrohelea sirii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="789D57D79D36FF91FDDB8BF1FDAC88C4" box="[520,577,2007,2028]" pageId="6" pageNumber="261" rank="species">
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<emphasis id="63AEEAAC9D36FF91FDDB8BF1FDAC88C4" bold="true" box="[520,577,2007,2028]" pageId="6" pageNumber="261">n. sp.</emphasis>
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</taxonomicNameLabel>
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Male. 13. head. 14. wing. 15. genitalia.
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</paragraph>
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</caption>
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<caption id="05A566369D37FF90FF448B9AFEBE88DA" httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="7" pageNumber="262" targetBox="[205,1391,209,1933]" targetPageId="7">
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<paragraph id="516536BE9D37FF90FF448B9AFEBE88DA" blockId="7.[151,1436,1980,2034]" pageId="7" pageNumber="262">
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<emphasis id="63AEEAAC9D37FF90FF448B9AFEB988FA" bold="true" box="[151,340,1980,2003]" pageId="7" pageNumber="262">FIGURES 16–19.</emphasis>
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<taxonomicName id="96DA4D3D9D37FF90FE888B9BFDE988FB" authority="Ronderos & Spinelli & Grogan, 2017" authorityName="Ronderos & Spinelli & Grogan" authorityYear="2017" box="[347,516,1981,2003]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
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<emphasis id="63AEEAAC9D37FF90FE888B9BFDE988FB" box="[347,516,1981,2003]" italics="true" pageId="7" pageNumber="262">Austrohelea sirii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="789D57D79D37FF90FDD88B98FDA888FB" box="[523,581,1982,2003]" pageId="7" pageNumber="262" rank="species">
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<emphasis id="63AEEAAC9D37FF90FDD88B98FDA888FB" bold="true" box="[523,581,1982,2003]" pageId="7" pageNumber="262">n. sp.</emphasis>
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</taxonomicNameLabel>
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Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7–9 and spermathecae.
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</paragraph>
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</caption>
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<paragraph id="516536BE9D38FF9FFF148D22FB128CA0" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
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<emphasis id="63AEEAAC9D38FF9FFF148D22FE678E35" bold="true" box="[199,394,260,285]" pageId="8" pageNumber="263">
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<typeStatus id="8E61881C9D38FF9FFF148D22FEC18E35" box="[199,300,260,285]" pageId="8" pageNumber="263" type="allotype">Allotype</typeStatus>
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female.
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</emphasis>
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Head (
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<figureCitation id="C9E12A3B9D38FF9FFE0E8D22FDC28E35" box="[477,559,260,285]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 16</figureCitation>
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) mostly dark brown. Eyes barely contiguous on extreme lower portions, with sparse interommatidial spicules. Antennal pedicel dark brown, flagellum light brown; flagellomeres 2-8 (
|
||
<figureCitation id="C9E12A3B9D38FF9FFAE88D01FA638E68" box="[1339,1422,295,320]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 17</figureCitation>
|
||
), moderately short, barrel-shaped, 9-13 elongate, cylindrical, 13 longest; flagellomeres 8, 12 with single subapical sensillum coeloconicum; antennal ratio 1.19. Clypeus with 3 pairs of setae. Palpus (
|
||
<figureCitation id="C9E12A3B9D38FF9FFBE88D49FB618EA0" box="[1083,1164,367,392]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 17</figureCitation>
|
||
) light brown; segment 3 moderately stout, with broad, shallow, subapicomesal sensory pit; palpal ratio 2.16. Mandible with 8 coarse teeth. Thorax. Scutum uniformly brown; scutellum pale with 6-7 large, 8 smaller setae. Femora, tibiae slender, uniformly brown (mid tibiae, tarsi missing), tarsi slightly paler; tarsomere 1 of fore, hind legs with ventral basal, apical spines; hind tarsomeres 1̄2 with two rows of ventral palisade setae, apparently with moderately numerous large setae (only insertions remain); hind tibial comb with 6 spines; hind tibial ratio 7.60; hind tarsal ratio 1.80; tarsomere 4 of fore leg short, cordiform, tarsomere 4 of hind leg elongate, cylindrical; claws small, equal size on fore leg, hind leg with single long claw that is slightly longer than tarsomere 5 with a stout basal tooth about 1/3 as long as claw. Wing (
|
||
<figureCitation id="C9E12A3B9D38FF9FFEA88EA9FE218D80" box="[379,460,655,680]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 18</figureCitation>
|
||
) membrane infuscated, particularly in radial sector, distal 2/3 of cell r3 and along veins; two well developed radial cells, 2nd broader, slightly longer than 1st; macrotrichia numerous on distal 2/3 of membrane, particularly dense on distal half of cell r3, and margins of m1, m2, about 40 on distal ½ of anal cell; wing length
|
||
<quantity id="96229B5B9D38FF9FFF378F22FEA68C34" box="[228,331,772,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="8" pageNumber="263" unit="mm" value="1.16">1.16 mm</quantity>
|
||
, width
|
||
<quantity id="96229B5B9D38FF9FFE4D8F23FDEB8C34" box="[414,518,773,797]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="8" pageNumber="263" unit="mm" value="0.47">0.47 mm</quantity>
|
||
; costal ratio 0.60. Halter missing. Abdomen (
|
||
<figureCitation id="C9E12A3B9D38FF9FFBD58F22FBB58C35" box="[1030,1112,772,797]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,264,1980,2003]" captionTargetBox="[205,1391,209,1933]" captionTargetId="figure@7.[182,1401,183,1968]" captionTargetPageId="7" captionText="FIGURES 16 – 19. Austrohelea sirii n. sp. Female. 16. head. 17. palpi and mouthparts. 18. wing. 19. abdominal segments 7 – 9 and spermathecae." httpUri="https://zenodo.org/record/806123/files/figure.png" pageId="8" pageNumber="263">Fig. 19</figureCitation>
|
||
) dark brown. Sternite 8 with V-shaped posteromedian cleft; sternite 9 halves moderately broad with rounded apices. Sternite 10 with two pairs of setae. Two unequal spermathecae; the larger pyriform with short, broad truncate neck, measuring 75 µm by 42 µm; the smaller partially collapsed with short narrow neck; a rudimentary third spermatheca present.
|
||
</paragraph>
|
||
<paragraph id="516536BE9D38FF9FFF148FB2FBC98B31" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
|
||
<materialsCitation id="E1B23CE39D38FF9FFF148FB2FA758CF9" ID-GBIF-Occurrence="1563642365" collectingDate="2007-01-07" collectionCode="MLPA" collectorName="La Heladera & A. Garre & F. Montes de Oca" country="Argentina" elevation="878" latitude="-41.015556" location="La Heladera" longLatPrecision="1" longitude="-71.82928" municipality="Parque Nacional Nahuel Huapi" pageId="8" pageNumber="263" specimenCount="2" stateProvince="Neuquen" typeStatus="holotype">
|
||
<emphasis id="63AEEAAC9D38FF9FFF148FB2FE988C85" bold="true" box="[199,373,916,941]" pageId="8" pageNumber="263">
|
||
<typeStatus id="8E61881C9D38FF9FFF148FB2FEEF8C85" box="[199,258,916,941]" pageId="8" pageNumber="263">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="8E61881C9D38FF9FFEA88FB2FE0E8C85" box="[379,483,916,941]" pageId="8" pageNumber="263" type="holotype">Holotype</typeStatus>
|
||
male,
|
||
<collectingCountry id="29CD762E9D38FF9FFDFD8FB3FD398C85" box="[558,724,917,941]" name="Argentina" pageId="8" pageNumber="263">ARGENTINA</collectingCountry>
|
||
,
|
||
<collectingRegion id="931EF85C9D38FF9FFD338FB3FCA58C84" box="[736,840,917,940]" country="Argentina" name="Neuquen" pageId="8" pageNumber="263">Neuquen</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="B101ACC49D38FF9FFC808FB3FB5F8C85" box="[851,1202,916,941]" pageId="8" pageNumber="263">Parque Nacional Nahuel Huapi</collectingMunicipality>
|
||
, mallin
|
||
<collectorName id="FC2F53689D38FF9FFADF8FB3FA758C85" box="[1292,1432,916,941]" pageId="8" pageNumber="263">
|
||
<location id="540560659D38FF9FFADF8FB3FA758C85" LSID="urn:lsid:plazi:treatment:D97387A89D35FF9FFF448805FB8F891D:540560659D38FF9FFADF8FB3FA758C85" box="[1292,1432,916,941]" country="Argentina" latitude="-41.015556" longLatPrecision="1" longitude="-71.82928" municipality="Parque Nacional Nahuel Huapi" name="La Heladera" pageId="8" pageNumber="263" stateProvince="Neuquen">La Heladera</location>
|
||
</collectorName>
|
||
,
|
||
<geoCoordinate id="34EE50799D38FF9FFF448F9EFEC38CF9" box="[151,302,952,977]" direction="south" orientation="latitude" pageId="8" pageNumber="263" precision="1" value="-41.015556">41°00'56.0"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="34EE50799D38FF9FFEEC8F9EFE018CF9" box="[319,492,952,977]" direction="west" orientation="longitude" pageId="8" pageNumber="263" precision="1" value="-71.82928">71°49'45.40"W</geoCoordinate>
|
||
,
|
||
<elevation id="DAF7D18D9D38FF9FFE2E8F9FFDAD8CF9" box="[509,576,953,977]" metricMagnitude="2" metricUnit="m" metricValue="8.78" pageId="8" pageNumber="263" unit="m" value="878.0">
|
||
<quantity id="96229B5B9D38FF9FFE2E8F9FFDAD8CF9" box="[509,576,953,977]" metricMagnitude="2" metricUnit="m" metricValue="8.78" pageId="8" pageNumber="263" unit="m" value="878.0">878m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="3520E9969D38FF9FFD838F9FFD5E8CF9" box="[592,691,953,977]" pageId="8" pageNumber="263" value="2007-01-07">
|
||
<date id="2564107E9D38FF9FFD838F9FFD5E8CF9" box="[592,691,953,977]" pageId="8" pageNumber="263" value="2007-01-07">7-I-2007</date>
|
||
</collectingDate>
|
||
/
|
||
<collectingDate id="3520E9969D38FF9FFD688F9FFD008CF9" box="[699,749,953,977]" bridgedPair="-" pageId="8" pageNumber="263">
|
||
<date id="2564107E9D38FF9FFD688F9FFD008CF9" box="[699,749,953,977]" bridgedPair="-" pageId="8" pageNumber="263">4-II-</date>
|
||
</collectingDate>
|
||
2 0 0 7,
|
||
<collectorName id="FC2F53689D38FF9FFCEA8F9FFC4B8CF9" box="[825,934,953,977]" pageId="8" pageNumber="263">A. Garré</collectorName>
|
||
̄
|
||
<collectorName id="FC2F53689D38FF9FFC638F9FFB658CF9" box="[944,1160,952,977]" pageId="8" pageNumber="263">F. Montes de Oca</collectorName>
|
||
,
|
||
<collectingMethod id="889B4EA99D38FF9FFB4A8F9EFAC78CF8" box="[1177,1322,952,977]" pageId="8" pageNumber="263">Malaise trap</collectingMethod>
|
||
(
|
||
<collectionCode id="37CBAE7B9D38FF9FFAED8F9FFA7D8CF9" box="[1342,1424,953,977]" pageId="8" pageNumber="263">MLPA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E1B23CE39D38FF9FFF448FFAFBCD8B31" ID-GBIF-Occurrence="1563642364" collectingDate="2007-02-05" collectionCode="MLPA" collectorName="A. Garre & F. Montes de Oca" country="Argentina" elevation="1194" latitude="-41.080166" location="Los Clavos" longLatPrecision="1" longitude="-71.82608" municipality="Parque Nacional Nahuel Huapi" pageId="8" pageNumber="263" specimenCount="1" stateProvince="Neuquen" typeStatus="allotype">
|
||
<typeStatus id="8E61881C9D38FF9FFF448FFAFF1C8CDD" box="[151,241,988,1013]" pageId="8" pageNumber="263" type="allotype">allotype</typeStatus>
|
||
female,
|
||
<collectingRegion id="931EF85C9D38FF9FFE8C8FFBFE2A8CDC" box="[351,455,989,1012]" country="Argentina" name="Neuquen" pageId="8" pageNumber="263">Neuquen</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="B101ACC49D38FF9FFE0A8FFBFCA18CDD" box="[473,844,988,1013]" pageId="8" pageNumber="263">Parque Nacional Nahuel Huapi</collectingMunicipality>
|
||
, laguna
|
||
<location id="540560659D38FF9FFC658FFBFBD28CDD" LSID="urn:lsid:plazi:treatment:D97387A89D35FF9FFF448805FB8F891D:540560659D38FF9FFC658FFBFBD28CDD" box="[950,1087,988,1013]" country="Argentina" latitude="-41.080166" longLatPrecision="1" longitude="-71.82608" municipality="Parque Nacional Nahuel Huapi" name="Los Clavos" pageId="8" pageNumber="263" stateProvince="Neuquen">Los Clavos</location>
|
||
,
|
||
<geoCoordinate id="34EE50799D38FF9FFB838FFAFB0A8CDD" box="[1104,1255,988,1013]" direction="south" orientation="latitude" pageId="8" pageNumber="263" precision="1" value="-41.080166">41°04'48.6"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="34EE50799D38FF9FFB2A8FFBFA758CDD" box="[1273,1432,989,1013]" direction="west" orientation="longitude" pageId="8" pageNumber="263" precision="1" value="-71.82608">71°49'33.9"W</geoCoordinate>
|
||
,
|
||
<elevation id="DAF7D18D9D38FF9FFF448827FF0B8B31" box="[151,230,1025,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.194" pageId="8" pageNumber="263" unit="m" value="1194.0">
|
||
<quantity id="96229B5B9D38FF9FFF448827FF0B8B31" box="[151,230,1025,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.194" pageId="8" pageNumber="263" unit="m" value="1194.0">1194m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="3520E9969D38FF9FFF218826FEB08B31" box="[242,349,1024,1049]" pageId="8" pageNumber="263" value="2007-02-05">
|
||
<date id="2564107E9D38FF9FFF218826FEB08B31" box="[242,349,1024,1049]" pageId="8" pageNumber="263" value="2007-02-05">5-II-2007</date>
|
||
</collectingDate>
|
||
/
|
||
<collectingDate id="3520E9969D38FF9FFEB48826FE4F8B31" box="[359,418,1024,1049]" bridgedPair="-" pageId="8" pageNumber="263">
|
||
<date id="2564107E9D38FF9FFEB48826FE4F8B31" box="[359,418,1024,1049]" bridgedPair="-" pageId="8" pageNumber="263">3-III-</date>
|
||
</collectingDate>
|
||
2 0 0 7,
|
||
<collectorName id="FC2F53689D38FF9FFE348827FDA28B31" box="[487,591,1025,1049]" pageId="8" pageNumber="263">A. Garré</collectorName>
|
||
̄
|
||
<collectorName id="FC2F53689D38FF9FFD8A8827FCCC8B31" box="[601,801,1024,1049]" pageId="8" pageNumber="263">F. Montes de Oca</collectorName>
|
||
,
|
||
<collectingMethod id="889B4EA99D38FF9FFCFF8826FC5A8B30" box="[812,951,1024,1049]" pageId="8" pageNumber="263">Malaise trap</collectingMethod>
|
||
(
|
||
<collectionCode id="37CBAE7B9D38FF9FFC158827FBF58B31" box="[966,1048,1025,1049]" pageId="8" pageNumber="263">MLPA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="19C065359D38FF9FFF148802FCC98BD9" pageId="8" pageNumber="263" type="distribution">
|
||
<paragraph id="516536BE9D38FF9FFF148802FDBF8B49" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
|
||
<emphasis id="63AEEAAC9D38FF9FFF148802FE8D8B15" bold="true" box="[199,352,1060,1085]" pageId="8" pageNumber="263">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="8E61881C9D38FF9FFD8C8803FD628B15" box="[607,655,1061,1085]" pageId="8" pageNumber="263">type</typeStatus>
|
||
locality, the
|
||
<taxonomicName id="96DA4D3D9D38FF9FFCCC8802FC4D8B14" authorityName="Blume" authorityYear="1850" box="[799,928,1060,1084]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="263" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="63AEEAAC9D38FF9FFCCC8802FC4D8B14" box="[799,928,1060,1084]" italics="true" pageId="8" pageNumber="263">Nothofagus</emphasis>
|
||
</taxonomicName>
|
||
forests of the Nahuel Huapi National Park of
|
||
<collectingRegion id="931EF85C9D38FF9FFF44886FFE858B49" box="[151,360,1096,1121]" country="Argentina" name="Neuquen" pageId="8" pageNumber="263">Neuquen province</collectingRegion>
|
||
,
|
||
<collectingCountry id="29CD762E9D38FF9FFEA0886EFE0F8B49" box="[371,482,1096,1121]" name="Argentina" pageId="8" pageNumber="263">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="C9E12A3B9D38FF9FFE21886EFDA88B49" box="[498,581,1096,1121]" captionStart="FIGURE 35" captionStartId="13.[151,250,1234,1256]" captionTargetBox="[267,1325,252,1118]" captionTargetId="figure@13.[205,1396,196,1194]" captionTargetPageId="13" captionText="FIGURE 35. Map of Argentine and Chilean Patagonia with distribution records of: Austrohelea shannoni (Wirth & Blanton), green circles. Austrohelea sirii n. sp., yellow triangles. Austrohelea spinosa n. sp., red diamonds." httpUri="https://zenodo.org/record/806131/files/figure.png" pageId="8" pageNumber="263">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="516536BE9D38FF9FFF14884AFCC98BD9" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
|
||
<emphasis id="63AEEAAC9D38FF9FFF14884AFDBE8BAD" bold="true" box="[199,595,1132,1157]" pageId="8" pageNumber="263">Derivation of the specific epithet.</emphasis>
|
||
We are pleased to name this new species after our colleague Augusto Siri, at the Instituto de Limnología “Dr. Raúl A. Ringuelet”, La Plata,
|
||
<collectingCountry id="29CD762E9D38FF9FFC7E88B6FBF28B81" box="[941,1055,1168,1193]" name="Argentina" pageId="8" pageNumber="263">Argentina</collectingCountry>
|
||
, in recognition of his important contributions to the taxonomy of
|
||
<taxonomicName id="96DA4D3D9D38FF9FFDF28892FD2C8BE5" authorityName="Newman" authorityYear="1834" box="[545,705,1204,1229]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
|
||
in the Neotropical Region, and for his valuable collections of
|
||
<taxonomicName id="96DA4D3D9D38FF9FFF4488FEFEBB8BD9" authorityName="Newman" authorityYear="1834" box="[151,342,1240,1265]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
|
||
from
|
||
<collectingCountry id="29CD762E9D38FF9FFE4888FEFDE78BD9" box="[411,522,1240,1265]" name="Argentina" pageId="8" pageNumber="263">Argentina</collectingCountry>
|
||
during the past 10 years.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="19C065359D38FF9FFF1488DAFB8F891D" pageId="8" pageNumber="263" type="discussion">
|
||
<paragraph id="516536BE9D38FF9FFF1488DAFAB68A8D" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
|
||
<emphasis id="63AEEAAC9D38FF9FFF1488DAFEA18A3D" bold="true" box="[199,332,1276,1301]" pageId="8" pageNumber="263">Discussion.</emphasis>
|
||
This new species is readily distinguished from its Neotropical congeners by its small size and the well-defined sensory pit on palpal segment 3. The male gonocoxite is very short, stout and lacks a basomesal tubercle; the gonostylus is much longer than the gonocoxite; and, the aedeagus is small, lightly sclerotized with a very low slightly concave basal arch, the apex is rounded and lacks a distal extension. The female wing membrane has dense, coarse macrotrichia and the spermathecae are pyriform, unequal, with short, broad truncate necks.
|
||
</paragraph>
|
||
<paragraph id="516536BE9D38FF9FFF148997FB8F891D" blockId="8.[151,1437,151,1589]" pageId="8" pageNumber="263">
|
||
Two other species from Australasia also have a sensory pit on palpal segment 3. In his original description of
|
||
<taxonomicName id="96DA4D3D9D38FF9FFF4489F3FEFC8AC4" authorityName="Macfie" authorityYear="1932" baseAuthorityName="Macfie" baseAuthorityYear="1932" box="[151,273,1493,1516]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="tonnoiri">
|
||
<emphasis id="63AEEAAC9D38FF9FFF4489F3FF458AC4" box="[151,168,1493,1516]" italics="true" pageId="8" pageNumber="263">A</emphasis>
|
||
.
|
||
<emphasis id="63AEEAAC9D38FF9FFF6489F3FEFC8AC4" box="[183,273,1493,1516]" italics="true" pageId="8" pageNumber="263">tonnoiri</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="29CD762E9D38FF9FFE8B89F3FE1F8AC5" box="[344,498,1492,1517]" name="New Zealand" pageId="8" pageNumber="263">New Zealand</collectingCountry>
|
||
,
|
||
<bibRefCitation id="354B4B4F9D38FF9FFE2D89F2FD4D8AC5" author="Macfie" box="[510,672,1492,1517]" pageId="8" pageNumber="263" refString="Macfie, J. W. S. (1932) New Zealand biting midges (Diptera, Ceratopogonidae). Annals of Tropical Medicine and Parasitology, 26, 23 - 53." type="journal article" year="1932">Macfie (1932)</bibRefCitation>
|
||
stated “…third (palpal segment) not inflated, bearing sensory hairs on a depressed area on apical third” and
|
||
<bibRefCitation id="354B4B4F9D38FF9FFDBA89DEFCC78939" author="Tokunaga" box="[617,810,1528,1553]" pageId="8" pageNumber="263" refString="Tokunaga, M. (1964) Insects of Campbell Island. Diptera: Ceratopogonidae. Pacific Insects Monograph, 7, 289 - 291." type="journal article" year="1964">Tokunaga (1964)</bibRefCitation>
|
||
noted for
|
||
<taxonomicName id="96DA4D3D9D38FF9FFC7189DFFB8F8938" authorityName="Tokunaga" authorityYear="1964" baseAuthorityName="Tokunaga" baseAuthorityYear="1964" box="[930,1122,1528,1552]" class="Insecta" family="Ceratopogonidae" genus="Austrohelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="campbellensis">
|
||
<emphasis id="63AEEAAC9D38FF9FFC7189DFFC5E8938" box="[930,947,1529,1552]" italics="true" pageId="8" pageNumber="263">A</emphasis>
|
||
.
|
||
<emphasis id="63AEEAAC9D38FF9FFC1089DEFB8F8938" box="[963,1122,1528,1552]" italics="true" pageId="8" pageNumber="263">campbellensis</emphasis>
|
||
</taxonomicName>
|
||
from Campbell Island, that palpal segment 3 is “…slightly thickened, with very small sensory pore on apical 1/3.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |