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<document id="2BDB88516D1A8DA93742AC3E8864884E" ID-DOI="10.1206/3928.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5369854" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268040215" checkinUser="carolina" docAuthor="Hopkins, Melanie J." docDate="2019" docId="B10287DEFF90B05FFE2EFCECFD43FAF0" docLanguage="en" docName="N3928 highres.pdf" docOrigin="American Museum Novitates 2019 (3928)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2019/issue-3928/3928.1/Phylogenetic-Analysis-and-Revision-of-the-Trilobite-Subfamily-Balnibarbiinae-Olenidae/10.1206/3928.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Cloacaspis ceryx subsp. anataphra" docType="treatment" docVersion="4" masterDocId="4D3BFFA6FF9CB053FF99FFE8FF9AFFCB" masterDocTitle="Phylogenetic analysis and revision of the trilobite subfamily Balnibarbiinae (Olenidae)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1699205220592" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6DB7E30DECC2E92F522270E2E50E2855">Phylogenetic analysis and revision of the trilobite subfamily Balnibarbiinae (Olenidae)</mods:title>
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<mods:namePart id="A38DC923C3284B6A3B3328165FB93C30">Hopkins, Melanie J.</mods:namePart>
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<treatment id="B10287DEFF90B05FFE2EFCECFD43FAF0" ID-DOI="http://doi.org/10.5281/zenodo.5461578" ID-Zenodo-Dep="5461578" LSID="urn:lsid:plazi:treatment:B10287DEFF90B05FFE2EFCECFD43FAF0" httpUri="http://treatment.plazi.org/id/B10287DEFF90B05FFE2EFCECFD43FAF0" pageId="12">
<subSubSection id="71B16543FF90B05FFE2EFCECFC3BFCD5" box="[439,929,772,798]" pageId="12" type="nomenclature">
<paragraph id="391436C8FF90B05FFE2EFCECFC3BFCD5" blockId="12.[439,929,772,798]" box="[439,929,772,798]" pageId="12">
<heading id="625C81A4FF90B05FFE2EFCECFC3BFCD5" box="[439,929,772,798]" centered="true" fontSize="11" level="2" pageId="12" reason="5">
<taxonomicName id="FEAB4D4BFF90B05FFE2EFCECFC3BFCD5" authority="(Fortey, 1974)" baseAuthorityName="Fortey" baseAuthorityYear="1974" box="[439,929,772,798]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="12" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="anataphra">
<emphasis id="0BDFEADAFF90B05FFE2EFCECFD6EFCD5" box="[439,756,772,798]" italics="true" pageId="12">Cloacaspis ceryx anataphra</emphasis>
(
<bibRefCitation id="5D3A4B39FF90B05FFC9DFCECFC02FCD5" author="Fortey, R. A." box="[772,920,772,798]" pageId="12" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey, 1974</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="71B16543FF90B05FFDFBFCD4FD6DFC9D" box="[610,759,828,854]" pageId="12" type="description">
<paragraph id="391436C8FF90B05FFDFBFCD4FD6DFC9D" blockId="12.[610,759,828,854]" box="[610,759,828,854]" pageId="12">
<heading id="625C81A4FF90B05FFDFBFCD4FD6DFC9D" box="[610,759,828,854]" centered="true" fontSize="36" level="3" pageId="12" reason="7">
<figureCitation id="A1902A4DFF90B05FFDFBFCD4FD6DFC9D" box="[610,759,828,854]" captionStart="FIGURE 4" captionStartId="10.[132,218,1492,1513]" captionTargetBox="[163,1205,248,1465]" captionTargetId="figure-41@10.[163,719,685,1129]" captionTargetPageId="10" captionText="FIGURE 4. Cloacaspis spp. A. Cloacaspis ekphymosa Fortey, 1974, USNM 144232. Collected from the Lost River Area on Seward Peninsula, Alaska, and previously identified as Triarthrus sp. (Ross, 1965). Arrow pointing to postpalpebral furrow. B. Anterior border of Cloacaspis ceryx anataphra (Fortey, 1974), AMNH- FI-101655. Black arrows pointing to faint anterior border pits on internal mold. C. Cloacaspis ceryx anataphra (Fortey, 1974), AMNH-FI-101655. Collected from PO-18 of the Olenidsletta Member of the Valhallfonna Formation (Kröger et al., 2017). D) Cloacaspis ceryx ceryx (Fortey, 1974), AMNH-FI-101657. Close-up of E, anterior glabellar lobe and F,posterior wing, showing presence of fine granulation of recently prepared surface (white arrow) and lack of granulation on exposed and slightly eroded surface (black arrow). Collected from bed PO-18 of the Olenidsletta Member of the Valhallfonna Formation (Kröger et al., 2017). Scale bars = 1 mm except A and D, where scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5369862" httpUri="https://zenodo.org/record/5369862/files/figure.png" pageId="12">Figure 4B, C</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="71B16543FF90B05FFF1DFC9EFD6AFC45" box="[132,752,886,910]" pageId="12" type="reference_group">
<paragraph id="391436C8FF90B05FFF1DFC9EFD6AFC45" blockId="12.[132,752,886,910]" box="[132,752,886,910]" pageId="12">
<treatmentCitationGroup id="19BB11E6FF90B05FFF1DFC9EFD6AFC45" box="[132,752,886,910]" pageId="12">
<treatmentCitation id="B80A10D9FF90B05FFF1DFC9EFDC2FC45" author="Fortey, R. A." box="[132,600,886,910]" page="30" pageId="12" year="1974">
<taxonomicName id="FEAB4D4BFF90B05FFF1DFC9EFDC2FC45" authority="Fortey, 1974: 30" authorityName="Fortey" authorityPageNumber="30" authorityYear="1974" box="[132,600,886,910]" class="Trilobita" family="Olenidae" genus="Balnibarbi" kingdom="Animalia" order="Ptychopariida" pageId="12" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="anataphra">
<emphasis id="0BDFEADAFF90B05FFF1DFC9EFE3EFC45" box="[132,420,886,910]" italics="true" pageId="12">Balnibarbi ceryx anataphra</emphasis>
<bibRefCitation id="5D3A4B39FF90B05FFE32FC9EFDC2FC45" author="Fortey, R. A." box="[427,600,886,910]" pageId="12" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey, 1974: 30</bibRefCitation>
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, pl. 9, fig. 5-7.
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</paragraph>
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<subSubSection id="71B16543FF90B05FFF2DFC44FE5EFB3A" pageId="12" type="materials_examined">
<paragraph id="391436C8FF90B05FFF2DFC44FBB6FC0C" blockId="12.[132,1236,940,1340]" box="[180,1068,940,967]" pageId="12">
<typeStatus id="E610886AFF90B05FFF2DFC44FEA2FC0D" box="[180,312,940,966]" pageId="12" type="holotype">HOLOTYPE</typeStatus>
: Cranidium, PMO NF 651 (figured in
<bibRefCitation id="5D3A4B39FF90B05FFC99FC45FC09FC0D" author="Fortey, R. A." box="[768,915,940,967]" pageId="12" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey, 1974</bibRefCitation>
: pl. 9, fig. 5).
</paragraph>
<paragraph id="391436C8FF90B05FFF2DFC3AFE1DFBDA" blockId="12.[132,1236,940,1340]" pageId="12">
<collectionCode id="5FBAAE0DFF90B05FFF2DFC3AFEF6FC20" box="[180,364,977,1004]" pageId="12">TYPE LOCALITY</collectionCode>
: Profilstranda, Olenidsletta Member, Valhallfonna Formation, Ny Friesland, Spitsbergen,
<collectingCountry id="41BC7658FF90B05FFE82FC1FFE19FBDA" box="[283,387,1015,1041]" name="Svalbard and Jan Mayen" pageId="12">Svalbard</collectingCountry>
.
</paragraph>
<paragraph id="391436C8FF90B05FFF2DFBF4FE5EFB3A" blockId="12.[132,1236,940,1340]" pageId="12">
<materialsCitation id="89C33C95FF90B05FFF2DFBF4FE24FB3A" collectionCode="STRATIGRAPHIC, RANGE" elevation="8" pageId="12" specimenCount="1">
<collectionCode id="5FBAAE0DFF90B05FFF2DFBF4FE4FFBFD" box="[180,469,1052,1078]" pageId="12">STRATIGRAPHIC RANGE</collectionCode>
:
<bibRefCitation id="5D3A4B39FF90B05FFE7BFBF5FD13FBFD" author="Fortey, R. A." box="[482,649,1052,1079]" pageId="12" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey (1974)</bibRefCitation>
reported this subspecies from a single horizon about
<quantity id="FE539B2DFF90B05FFF49FBAAFF64FB97" box="[208,254,1090,1116]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="12" unit="m" value="8.0">
<elevation id="B286D1FBFF90B05FFF49FBAAFF64FB97" box="[208,254,1090,1116]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="12" unit="m" value="8.0">8 m</elevation>
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above the base of the Olenidsletta Member, which is estimated to be around
<quantity id="FE539B2DFF90B05FFB3BFBA9FF06FB4A" metricMagnitude="1" metricUnit="m" metricValue="1.25" pageId="12" unit="m" value="12.5">
<elevation id="B286D1FBFF90B05FFB3BFBA9FF06FB4A" metricMagnitude="1" metricUnit="m" metricValue="1.25" pageId="12" unit="m" value="12.5">12.5 m</elevation>
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in the 2016 stratigraphic section. Additional specimens were recovered in 2016 from
<quantity id="FE539B2DFF90B05FFF1DFB64FF75FB6C" box="[132,239,1164,1191]" metricMagnitude="1" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="12" unit="m" value="18.0" valueMax="19.0" valueMin="17.0">
<elevation id="B286D1FBFF90B05FFF1DFB64FF75FB6C" box="[132,239,1164,1191]" metricMagnitude="1" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="12" unit="m" value="18.0" valueMax="19.0" valueMin="17.0">1719 m</elevation>
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in section, which extends the stratigraphic range of this subspecies, but the total range remains within the early half of the stratigraphic range of the other subspecies,
<taxonomicName id="FEAB4D4BFF90B05FFB3BFB59FEFEFB3A" authority="ceryx" authorityName="ceryx (Fortey" authorityYear="1974" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="12" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="ceryx">
<emphasis id="0BDFEADAFF90B05FFB3BFB59FEFEFB3A" italics="true" pageId="12">Cloacaspis ceryx ceryx</emphasis>
</taxonomicName>
(fig. 1)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71B16543FF90B05FFF2DFB14FD43FAF0" pageId="12" type="diagnosis">
<paragraph id="391436C8FF90B05FFF2DFB14FD43FAF0" blockId="12.[132,1236,940,1340]" pageId="12">
DIAGNOSIS:
<taxonomicName id="FEAB4D4BFF90B05FFED1FB14FD93FADD" baseAuthorityName="Fortey" baseAuthorityYear="1974" box="[328,521,1276,1302]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="12" phylum="Arthropoda" rank="species" species="ceryx">
<emphasis id="0BDFEADAFF90B05FFED1FB14FD93FADD" box="[328,521,1276,1302]" italics="true" pageId="12">Cloacaspis ceryx</emphasis>
</taxonomicName>
species with relatively narrow preglabellar field (0.060.10 length of glabella) and smooth exoskeletal surface.
</paragraph>
</subSubSection>
</treatment>
</document>