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<document id="7B7B3A5FCB5E320E1A396C003E05D7A2" ID-CLB-Dataset="24216" ID-DOI="10.11646/zootaxa.4729.3.1" ID-GBIF-Dataset="f7aee5da-a7ce-4660-b00c-b272b6204a19" ID-ISSN="1175-5326" ID-Zenodo-Dep="3632662" ID-ZooBank="330533E0-25B8-4C66-89F2-B93F3E88DD5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580475428538" checkinUser="plazi" docAuthor="Fominykh, Dmitriy D., Zamotajlov, Alexandr S., Khomitskiy, Eugeniy E. &amp; Titarenko, Andrey Yu." docDate="2020" docId="E676063DFFF33866FF70B2A63A75E18D" docLanguage="en" docName="zootaxa.4729.3.1.pdf" docOrigin="Zootaxa 4729 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carabus (Lipaster) stjernvalli subsp. gvalijai Retezar, Mravaldzali 1992" docType="treatment" docVersion="8" lastPageNumber="316" masterDocId="1A4F7E45FFFC3869FFE7B23C3B0DE566" masterDocTitle="Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini)" masterLastPageNumber="328" masterPageNumber="301" pageNumber="316" updateTime="1698779826648" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D8EE8C943D8DD51BCDEB47F39E196D06">Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini)</mods:title>
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<mods:name id="7F3A819FCEC3A305568A107F53BF1B59" type="personal">
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<mods:namePart id="536BBD4EB5B8CD8DC83A82E0C0360403">Fominykh, Dmitriy D.</mods:namePart>
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<mods:namePart id="B1EC0FE9CCFB8AC712C482C678AC8A95">Zamotajlov, Alexandr S.</mods:namePart>
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<mods:namePart id="8732BB8643EDAE4AFEC7D3D3F1D6B234">Khomitskiy, Eugeniy E.</mods:namePart>
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<mods:namePart id="C1A8D65AC254BAFAFD77DDADC5707D59">Titarenko, Andrey Yu.</mods:namePart>
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<mods:title id="A006D7442E4DEE050442AC285C5B8482">Zootaxa</mods:title>
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<mods:date id="844658F62ED3176BFC1DE1B5AF0535CF">2020</mods:date>
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<mods:number id="CD1C1AB512217A6FB51EFA2AC2256543">4729</mods:number>
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<mods:number id="63502A8B351DA37DF13DF658FD3153F8">3</mods:number>
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<mods:classification id="96798A10FA634193E35B5AD7B8BDE66A">journal article</mods:classification>
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<mods:identifier id="5CAFF1738E13DE4A07456B0FAD5455C7" type="DOI">10.11646/zootaxa.4729.3.1</mods:identifier>
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<treatment id="E676063DFFF33866FF70B2A63A75E18D" ID-GBIF-Taxon="161926262" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D" httpUri="http://treatment.plazi.org/id/E676063DFFF33866FF70B2A63A75E18D" lastPageNumber="316" pageId="15" pageNumber="316">
<subSubSection id="26C5E4A0FFF33866FF70B2A63827E5D5" box="[151,810,153,180]" pageId="15" pageNumber="316" type="nomenclature">
<paragraph id="6E60B72BFFF33866FF70B2A63827E5D5" blockId="15.[151,810,153,216]" box="[151,810,153,180]" pageId="15" pageNumber="316">
<heading id="35280047FFF33866FF70B2A63827E5D5" bold="true" box="[151,810,153,180]" fontSize="11" level="1" pageId="15" pageNumber="316" reason="1">
<emphasis id="5CAB6B39FFF33866FF70B2A63827E5D5" bold="true" box="[151,810,153,180]" pageId="15" pageNumber="316">
7.
<taxonomicName id="A9DFCCA8FFF33866FF52B2A53827E5D5" ID-CoL="8MHWS" authority="Retezar, 1992" authorityName="Retezar, Mravaldzali" authorityYear="1992" box="[181,810,153,180]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subGenus="Lipaster" subSpecies="gvalijai">
<emphasis id="5CAB6B39FFF33866FF52B2A53976E5D2" bold="true" box="[181,635,153,180]" italics="true" pageId="15" pageNumber="316">Carabus (Lipaster) stjernvalli gvalijai</emphasis>
<bibRefCitation id="0A4ECADAFFF33866FD65B2A53827E5D5" author="Retezar, I." box="[642,810,153,180]" pageId="15" pageNumber="316" pagination="85 - 88" refId="ref10287" refString="Retezar, I. (1992) Two new Carabus species from the Caucasus (Coleoptera: Carabidae). Folia Entomologica Hungarica, 52, 85 - 88." type="journal article" year="1992">Retezár, 1992</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="26C5E4A0FFF33866FF70B2813924E5BE" box="[151,553,189,216]" pageId="15" pageNumber="316" type="description">
<paragraph id="6E60B72BFFF33866FF70B2813924E5BE" blockId="15.[151,810,153,216]" box="[151,553,189,216]" pageId="15" pageNumber="316">
(
<figureCitation id="F6E4ABAEFFF33866FF78B2813BE0E5B1" box="[159,237,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1031,1056]" captionTargetBox="[416,1172,414,1002]" captionTargetId="figure@2.[416,1172,414,1002]" captionTargetPageId="2" captionText="FIGURE 1. Collecting sites of Carabus (Lipaster) stjernvalli Mannerheim in the South Caucasus: red square—C. stjernvalli ssp., Gorelovka env.; green square—C. stjernvalli paravanicus subsp. nov.; pink rhombs—C. stjernvalli tabackuriensis Novotný &amp; Vořišek; red rhombus—C. stjernvalli tabackuriensis Novotný &amp; Vořišek, type locality; red triangles—C. stjernvalli kartalinicus subsp. nov.; blue circle—C. stjernvalli bogatshevi Zamotajlov, type locality; violet empty squares—C. stjernvalli stjernvalli Mannerheim; red crosses—C. stjernvalli gvalijai Retezár." figureDoi="http://doi.org/10.5281/zenodo.3632664" httpUri="https://zenodo.org/record/3632664/files/figure.png" pageId="15" pageNumber="316">Figs. 1</figureCitation>
,
<figureCitation id="F6E4ABAEFFF33866FF1CB2813A49E5B1" box="[251,324,189,215]" captionStart-0="FIGURES 68–75" captionStart-1="FIGURES 76–83" captionStart-2="FIGURES 84–91" captionStartId-0="16.[151,264,1942,1967]" captionStartId-1="17.[151,264,1930,1955]" captionStartId-2="18.[151,264,1929,1954]" captionTargetBox-0="[159,1426,209,1853]" captionTargetBox-1="[160,1424,210,1856]" captionTargetBox-2="[164,1414,194,1885]" captionTargetId-0="figure@16.[151,1436,199,1867]" captionTargetId-1="figure@17.[151,1436,196,1866]" captionTargetId-2="figure@18.[151,1436,181,1904]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURES 68–75. Dorsal habitus of Carabus (Lipaster) stjernvalli gvalijai Retezár: 68–71—Mravaldzali env., 2200 m; 72– 75—Tli env., 2000 m. Scale bar 10.0 mm." captionText-1="FIGURES 76–83. Dorsal habitus of Carabus (Lipaster) stjernvalli gvalijai Retezár: 76–79—Askhi Plateau, 2300 m; 80–83— Sasashi env., 2300 m. Scale bar 10.0 mm." captionText-2="FIGURES 84–91. Dorsal habitus of Carabus (Lipaster) stjernvalli gvalijai Retezár: 84–87—Goldash Mt., 2600 m; 88–91— Goldash Mt., 1500 m. Scale bar 10.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3632688" figureDoi-1="http://doi.org/10.5281/zenodo.3632690" figureDoi-2="http://doi.org/10.5281/zenodo.3632692" httpUri-0="https://zenodo.org/record/3632688/files/figure.png" httpUri-1="https://zenodo.org/record/3632690/files/figure.png" httpUri-2="https://zenodo.org/record/3632692/files/figure.png" pageId="15" pageNumber="316">68–91</figureCitation>
,
<figureCitation id="F6E4ABAEFFF33866FEA8B2813ABFE5B1" box="[335,434,189,215]" captionStart="FIGURES 102–114" captionStartId="19.[151,264,1829,1854]" captionTargetBox="[230,1355,1226,1796]" captionTargetId="figure@19.[225,1361,1220,1804]" captionTargetPageId="19" captionText="FIGURES 102–114. Apical portion of aedeagus and fully inflated endophallus preparation of Carabus (Lipaster) stjernvalli Mannerheim, right lateral view: 102—C. stjernvalli stjernvalli Mannerheim, Chidila Pass env., 2350 m; 103—idem, Dzegvila Mt., 1100 m; 104—idem, Gomismta env., 2100 m; 105—idem, Gomi env., 1500 m; 106—Goderdzi Pass env., 2300 m; 107— idem, Zekari Pass env., 2100 m; 108—idem, Sairme env., 1500 m; 109—C. stjernvalli gvalijai Retezár, Mravaldzali env., 2200 m; 110—idem, Tli env., 2000 m; 111—idem, Askhi Plateau, 2300 m; 112—idem, Sasashi env., 2300 m; 113—idem, Goldash Mt., 2600 m; 114 —idem, Goldash Mt., 1500 m. Scale bar 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3632696" httpUri="https://zenodo.org/record/3632696/files/figure.png" pageId="15" pageNumber="316">109–114</figureCitation>
,
<figureCitation id="F6E4ABAEFFF33866FE59B281392CE5BE" box="[446,545,189,216]" captionStart-0="FIGURE 125" captionStart-1="FIGURE 126" captionStart-2="FIGURE 127" captionStartId-0="25.[151,250,1119,1144]" captionStartId-1="25.[151,250,2004,2029]" captionStartId-2="26.[151,250,1157,1182]" captionTargetBox-0="[176,1410,523,1100]" captionTargetBox-1="[173,1415,1158,1984]" captionTargetBox-2="[151,1435,174,1138]" captionTargetId-0="figure@25.[176,1410,174,1100]" captionTargetId-1="figure@25.[173,1415,1158,1984]" captionTargetId-2="figure@26.[151,1435,174,1138]" captionTargetPageId-0="25" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURE 125. Habitat of Carabus stjernvalli gvalijai Retezár, Georgia, Mravaldzali env., 2100 m." captionText-1="FIGURE 126. Habitat of Carabus stjernvalli gvalijai Retezár, Georgia, Askhi Plateau, 2300 m." captionText-2="FIGURE 127. Habitat of Carabus stjernvalli gvalijai Retezár, Georgia, Sasashi env., Jvari Pass, 2300 m." figureDoi-0="http://doi.org/10.5281/zenodo.3632720" figureDoi-1="http://doi.org/10.5281/zenodo.3632722" figureDoi-2="http://doi.org/10.5281/zenodo.3632724" httpUri-0="https://zenodo.org/record/3632720/files/figure.png" httpUri-1="https://zenodo.org/record/3632722/files/figure.png" httpUri-2="https://zenodo.org/record/3632724/files/figure.png" pageId="15" pageNumber="316">125–127</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="26C5E4A0FFF33866FF70B33939C2E65D" pageId="15" pageNumber="316" type="materials_examined">
<paragraph id="6E60B72BFFF33866FF70B33939C2E65D" blockId="15.[151,1437,261,1260]" pageId="15" pageNumber="316">
<emphasis id="5CAB6B39FFF33866FF70B3393A8CE446" bold="true" box="[151,385,261,288]" pageId="15" pageNumber="316">Material examined.</emphasis>
<materialsCitation id="DEB7BD76FFF33866FE6EB3393FBBE425" ID-GBIF-Occurrence="2561484944" collectingDate="2016-09-06" collectionCode="CFDD" collectorName="A. Zubov" country="Georgia" county="Khikhata Mt. Range" elevation="2100" latitude="42.48712" location="Racha" longLatPrecision="1" longitude="43.353386" municipality="Racha Mt. Range" pageId="15" pageNumber="316" specimenCount="18" specimenCount-female="6" specimenCount-male="12">
<specimenCount id="78D97CA2FFF33866FE6EB3393ADFE479" box="[393,466,261,287]" pageId="15" pageNumber="316" type="male">12♂♂</specimenCount>
,
<specimenCount id="78D97CA2FFF33866FE39B339391AE479" box="[478,535,261,287]" pageId="15" pageNumber="316" type="female">6♀♀</specimenCount>
,
<collectingCountry id="16C8F7BBFFF33866FDC2B339398FE479" box="[549,642,261,287]" name="Georgia" pageId="15" pageNumber="316">Georgia</collectingCountry>
,
<location id="6B00E1F0FFF33866FD6AB33939DBE479" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FD6AB33939DBE479" box="[653,726,261,287]" country="Georgia" county="Khikhata Mt. Range" latitude="42.48712" longLatPrecision="1" longitude="43.353386" municipality="Racha Mt. Range" name="Racha" pageId="15" pageNumber="316">Racha</location>
,
<collectingMunicipality id="8E042D51FFF33866FD05B33938A4E446" box="[738,937,261,288]" pageId="15" pageNumber="316">Racha Mt. Range</collectingMunicipality>
, N slopes of
<collectingCounty id="8701CFA7FFF33866FBD8B3393E28E446" box="[1087,1317,261,288]" pageId="15" pageNumber="316">Khikhata Mt. Range</collectingCounty>
near Mra- valdzali vill.,
<quantity id="A9271ACEFFF33866FED5B3153A84E422" box="[306,393,297,324]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="316" unit="m" value="2100.0">
<elevation id="E5F25018FFF33866FED5B3153A84E422" box="[306,393,297,324]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="316" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<geoCoordinate id="0BEBD1ECFFF33866FE72B3163944E425" box="[405,585,297,324]" degrees="42" direction="north" minutes="29" orientation="latitude" pageId="15" pageNumber="316" precision="1" seconds="13,64" value="42.48712">N 42°29’13,64”</geoCoordinate>
/
<geoCoordinate id="0BEBD1ECFFF33866FDB8B316381EE425" box="[607,787,297,324]" degrees="43" direction="east" minutes="21" orientation="longitude" pageId="15" pageNumber="316" precision="1" seconds="12,19" value="43.353386">E 43°21’12,19”</geoCoordinate>
, 08.-
<date id="1A6191EBFFF33866FCA3B31538C5E425" box="[836,968,297,323]" pageId="15" pageNumber="316" value="2016-09-06">
<collectingDate id="0A256803FFF33866FCA3B31538C5E425" box="[836,968,297,323]" pageId="15" pageNumber="316" value="2016-09-06">09.06.2016</collectingDate>
</date>
(
<collectorName id="C32AD2FDFFF33866FC3EB3153F4AE422" box="[985,1095,297,324]" pageId="15" pageNumber="316">A. Zubov</collectorName>
) (
<collectionCode id="08CE2FEEFFF33866FBB9B3153FA2E425" box="[1118,1199,297,323]" pageId="15" pageNumber="316">CFDD</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DEB7BD76FFF33866FB25B3153953E4ED" ID-GBIF-Occurrence="2561484967" collectingDate="2015-12-07" collectionCode="CFDD, CTAY, CZAM" collectorName="D. Fominykh" country="Georgia" elevation="1800" latitude="42.469646" location="Racha Mt. Range" longLatPrecision="1" longitude="43.76598" municipality="Ertzo Pass" pageId="15" pageNumber="316" specimenCount="17" specimenCount-female="7" specimenCount-male="10">
<specimenCount id="78D97CA2FFF33866FB25B3153E06E425" box="[1218,1291,297,323]" pageId="15" pageNumber="316" type="male">10♂♂</specimenCount>
,
<specimenCount id="78D97CA2FFF33866FAF1B3163E5DE422" box="[1302,1360,298,324]" pageId="15" pageNumber="316" type="female">7♀♀</specimenCount>
, S Os- setia,
<location id="6B00E1F0FFF33866FF3FB3713A96E40E" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FF3FB3713A96E40E" box="[216,411,333,360]" country="Georgia" latitude="42.469646" longLatPrecision="1" longitude="43.76598" municipality="Ertzo Pass" name="Racha Mt. Range" pageId="15" pageNumber="316">Racha Mt. Range</location>
near
<collectingMunicipality id="8E042D51FFF33866FE3FB3723942E40E" box="[472,591,334,360]" pageId="15" pageNumber="316">Ertzo Pass</collectingMunicipality>
,
<quantity id="A9271ACEFFF33866FDBDB37139BCE40E" box="[602,689,333,360]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="316" unit="m" value="1800.0">
<elevation id="E5F25018FFF33866FDBDB37139BCE40E" box="[602,689,333,360]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="316" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<geoCoordinate id="0BEBD1ECFFF33866FD5BB3723862E401" box="[700,879,333,360]" degrees="42" direction="north" minutes="28" orientation="latitude" pageId="15" pageNumber="316" precision="1" seconds="10.72" value="42.469646">N 42°28’10.72”</geoCoordinate>
/
<geoCoordinate id="0BEBD1ECFFF33866FC63B3723F3BE40E" box="[900,1078,333,360]" degrees="43" direction="east" minutes="45" orientation="longitude" pageId="15" pageNumber="316" precision="1" seconds="57.53" value="43.76598">E 43°45’57.53”</geoCoordinate>
,
<collectingMethod id="B79ECF3CFFF33866FBA6B3713FC9E40E" box="[1089,1220,333,360]" pageId="15" pageNumber="316">pitfall traps</collectingMethod>
, 29.05.-
<date id="1A6191EBFFF33866FAFDB3713E91E40E" box="[1306,1436,333,360]" pageId="15" pageNumber="316" value="2015-12-07">
<collectingDate id="0A256803FFF33866FAFDB3713E91E40E" box="[1306,1436,333,360]" pageId="15" pageNumber="316" value="2015-12-07">12.07.2015</collectingDate>
</date>
(
<collectorName id="C32AD2FDFFF33866FF47B34D3A34E4EA" box="[160,313,369,396]" pageId="15" pageNumber="316">D. Fominykh</collectorName>
) (
<collectionCode id="08CE2FEEFFF33866FEB6B34D3AADE4ED" box="[337,416,369,395]" pageId="15" pageNumber="316">CFDD</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FE4AB34D3AF9E4ED" box="[429,500,369,395]" pageId="15" pageNumber="316">CTAY</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FDE7B34D395BE4ED" box="[512,598,369,395]" pageId="15" pageNumber="316">CZAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DEB7BD76FFF33866FD8CB34D390BE4B5" ID-GBIF-Occurrence="2561484946" collectingDate="2016-10-07" collectionCode="CFDD, CTAY, CZAM" collectorName="D. Fominykh &amp; A. Zubov" country="Georgia" county="Alkhashenda Mt." elevation="1800" latitude="42.46103" location="Ossetia" longLatPrecision="1" longitude="43.773167" municipality="Ertzo" pageId="15" pageNumber="316" specimenCount="38" specimenCount-female="23" specimenCount-male="15">
<specimenCount id="78D97CA2FFF33866FD8CB34D39BEE4EA" box="[619,691,369,396]" pageId="15" pageNumber="316" type="male">15♂♂</specimenCount>
,
<specimenCount id="78D97CA2FFF33866FD26B34D3805E4ED" box="[705,776,369,395]" pageId="15" pageNumber="316" type="female">23♀♀</specimenCount>
, S
<location id="6B00E1F0FFF33866FCC9B34D388EE4ED" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FCC9B34D388EE4ED" box="[814,899,369,395]" country="Georgia" county="Alkhashenda Mt." latitude="42.46103" longLatPrecision="1" longitude="43.773167" municipality="Ertzo" name="Ossetia" pageId="15" pageNumber="316">Ossetia</location>
,
<location id="6B00E1F0FFF33866FC77B34D3F5AE4EA" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FC77B34D3F5AE4EA" box="[912,1111,369,396]" country="Georgia" county="Alkhashenda Mt." latitude="42.46103" longLatPrecision="1" longitude="43.773167" municipality="Ertzo" name="Racha Mt. Range" pageId="15" pageNumber="316">Racha Mt. Range</location>
near
<collectingMunicipality id="8E042D51FFF33866FB7FB34E3FD8E4EA" box="[1176,1237,370,396]" pageId="15" pageNumber="316">Ertzo</collectingMunicipality>
vill., N slopes of
<collectingCounty id="8701CFA7FFF33866FF70B3A93A51E4D6" box="[151,348,405,432]" pageId="15" pageNumber="316">Alkhashenda Mt.</collectingCounty>
,
<quantity id="A9271ACEFFF33866FE8FB3A93ACDE4D6" box="[360,448,405,432]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="316" unit="m" value="1800.0">
<elevation id="E5F25018FFF33866FE8FB3A93ACDE4D6" box="[360,448,405,432]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="316" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<geoCoordinate id="0BEBD1ECFFF33866FE2BB3AA398DE4C9" box="[460,640,405,432]" degrees="42" direction="north" minutes="27" orientation="latitude" pageId="15" pageNumber="316" precision="1" seconds="39,70" value="42.46103">N 42°27’39,70”</geoCoordinate>
/
<geoCoordinate id="0BEBD1ECFFF33866FD7FB3AA3846E4C9" box="[664,843,405,432]" degrees="43" direction="east" minutes="46" orientation="longitude" pageId="15" pageNumber="316" precision="1" seconds="23,40" value="43.773167">E 43°46’23,40”</geoCoordinate>
,
<collectingMethod id="B79ECF3CFFF33866FCBFB3A938D1E4D6" box="[856,988,405,432]" pageId="15" pageNumber="316">pitfall traps</collectingMethod>
, 17.05.-
<date id="1A6191EBFFF33866FBD5B3A93FBBE4D6" box="[1074,1206,405,432]" pageId="15" pageNumber="316" value="2016-10-07">
<collectingDate id="0A256803FFF33866FBD5B3A93FBBE4D6" box="[1074,1206,405,432]" pageId="15" pageNumber="316" value="2016-10-07">10.07.2016</collectingDate>
</date>
(
<collectorName id="C32AD2FDFFF33866FB21B3A93E50E4C9" box="[1222,1373,405,431]" pageId="15" pageNumber="316">D. Fominykh</collectorName>
&amp;
<collectorName id="C32AD2FDFFF33866FA66B3AA3BEFE4B2" pageId="15" pageNumber="316">A. Zubov</collectorName>
) (
<collectionCode id="08CE2FEEFFF33866FF1EB3853A45E4B5" box="[249,328,441,467]" pageId="15" pageNumber="316">CFDD</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FEB2B3853A91E4B5" box="[341,412,441,467]" pageId="15" pageNumber="316">CTAY</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FE4FB3853AF3E4B5" box="[424,510,441,467]" pageId="15" pageNumber="316">CZAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DEB7BD76FFF33866FDF5B3853995E77D" ID-GBIF-Occurrence="2561484957" collectingDate="2016-09-07" collectionCode="ZISP, MPSU, CFDD, CTAY, CZAM, CKEE" collectorName="D. Fominykh &amp; A. Zubov" country="Georgia" county="Valtkatzi River" elevation="2100" latitude="42.495808" location="Ossetia" longLatPrecision="1" longitude="43.845417" municipality="Tli" pageId="15" pageNumber="316" specimenCount="164" specimenCount-female="85" specimenCount-male="79">
<specimenCount id="78D97CA2FFF33866FDF5B3853957E4B5" box="[530,602,441,467]" pageId="15" pageNumber="316" type="male">79♂♂</specimenCount>
,
<specimenCount id="78D97CA2FFF33866FD80B38539A2E4B2" box="[615,687,441,468]" pageId="15" pageNumber="316" type="female">85♀♀</specimenCount>
, S
<location id="6B00E1F0FFF33866FD34B3853825E4B5" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FD34B3853825E4B5" box="[723,808,441,467]" country="Georgia" county="Valtkatzi River" latitude="42.495808" longLatPrecision="1" longitude="43.845417" municipality="Tli" name="Ossetia" pageId="15" pageNumber="316">Ossetia</location>
,
<location id="6B00E1F0FFF33866FCD2B38538F6E4B2" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FCD2B38538F6E4B2" box="[821,1019,441,468]" country="Georgia" county="Valtkatzi River" latitude="42.495808" longLatPrecision="1" longitude="43.845417" municipality="Tli" name="Racha Mt. Range" pageId="15" pageNumber="316">Racha Mt. Range</location>
near
<collectingMunicipality id="8E042D51FFF33866FBDDB3853F56E4B5" box="[1082,1115,441,467]" pageId="15" pageNumber="316">Tli</collectingMunicipality>
vill., right bank of
<collectingCounty id="8701CFA7FFF33866FADEB3853BDAE491" pageId="15" pageNumber="316">Valtkatzi River</collectingCounty>
,
<quantity id="A9271ACEFFF33866FF05B3E13A74E49E" box="[226,377,477,504]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.3" metricValueMin="1.9" pageId="15" pageNumber="316" unit="m" value="2100.0" valueMax="2300.0" valueMin="1900.0">
<elevation id="E5F25018FFF33866FF05B3E13A74E49E" box="[226,377,477,504]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.3" metricValueMin="1.9" pageId="15" pageNumber="316" unit="m" value="2100.0" valueMax="2300.0" valueMin="1900.0">1900-2300 m</elevation>
</quantity>
,
<geoCoordinate id="0BEBD1ECFFF33866FE62B3E2393AE491" box="[389,567,477,504]" degrees="42" direction="north" minutes="29" orientation="latitude" pageId="15" pageNumber="316" precision="1" seconds="44,90" value="42.495808">N 42°29’44,90”</geoCoordinate>
/
<geoCoordinate id="0BEBD1ECFFF33866FDABB3E239F0E49E" box="[588,765,477,504]" degrees="43" direction="east" minutes="50" orientation="longitude" pageId="15" pageNumber="316" precision="1" seconds="43,50" value="43.845417">E 43°50’43,50”</geoCoordinate>
,
<collectingMethod id="B79ECF3CFFF33866FCEFB3E13886E49E" box="[776,907,477,504]" pageId="15" pageNumber="316">pitfall traps</collectingMethod>
, 17.05.-
<date id="1A6191EBFFF33866FC39B3E13F6EE49E" box="[990,1123,477,504]" pageId="15" pageNumber="316" value="2016-09-07">
<collectingDate id="0A256803FFF33866FC39B3E13F6EE49E" box="[990,1123,477,504]" pageId="15" pageNumber="316" value="2016-09-07">09.07.2016</collectingDate>
</date>
(
<collectorName id="C32AD2FDFFF33866FB95B3E13E0AE491" box="[1138,1287,477,503]" pageId="15" pageNumber="316">D. Fominykh</collectorName>
&amp;
<collectorName id="C32AD2FDFFF33866FACFB3E23E98E49E" box="[1320,1429,477,504]" pageId="15" pageNumber="316">A. Zubov</collectorName>
) (
<collectionCode id="08CE2FEEFFF33866FF78B03D3BD7E77D" box="[159,218,513,539]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="15" pageNumber="316">ZISP</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FF03B03D3A3EE77D" box="[228,307,513,539]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="15" pageNumber="316">MPSU</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FEDAB03D3A86E77D" box="[317,395,513,539]" pageId="15" pageNumber="316">CFDD</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FE73B03D3AD6E77D" box="[404,475,513,539]" pageId="15" pageNumber="316">CTAY</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FE03B03D3935E77D" box="[484,568,513,539]" pageId="15" pageNumber="316">CZAM</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FDA5B03D399DE77D" box="[578,656,513,539]" pageId="15" pageNumber="316">CKEE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DEB7BD76FFF33866FD45B03D3E5CE759" ID-GBIF-Occurrence="2561484932" collectingDate="2014-06-22" collectionCode="ZISP, MPSU, CFDD, CTAY, CZAM" collectorName="A. Zubov" country="Georgia" county="Lekhi Mt." elevation="2250" latitude="42.583332" location="Samegrelo" longLatPrecision="1149" longitude="42.533333" municipality="Askhi Plateau" pageId="15" pageNumber="316" specimenCount="60" specimenCount-female="27" specimenCount-male="33">
<specimenCount id="78D97CA2FFF33866FD45B03D39E7E77D" box="[674,746,513,539]" pageId="15" pageNumber="316" type="male">33♂♂</specimenCount>
,
<specimenCount id="78D97CA2FFF33866FD12B03D3831E77D" box="[757,828,513,539]" pageId="15" pageNumber="316" type="female">27♀♀</specimenCount>
,
<collectingCountry id="16C8F7BBFFF33866FCA0B03D38A9E77D" box="[839,932,513,539]" name="Georgia" pageId="15" pageNumber="316">Georgia</collectingCountry>
,
<location id="6B00E1F0FFF33866FC49B03D3F24E77D" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FC49B03D3F24E77D" box="[942,1065,513,539]" country="Georgia" county="Lekhi Mt." latitude="42.583332" longLatPrecision="1149" longitude="42.533333" municipality="Askhi Plateau" name="Samegrelo" pageId="15" pageNumber="316">Samegrelo</location>
,
<collectingMunicipality id="8E042D51FFF33866FBD6B03D3FC3E77D" box="[1073,1230,513,539]" pageId="15" pageNumber="316">Askhi Plateau</collectingMunicipality>
, N slopes of
<collectingCounty id="8701CFA7FFF33866FABCB03D3BCFE726" pageId="15" pageNumber="316">Lekhi Mt.</collectingCounty>
,
<quantity id="A9271ACEFFF33866FF29B0193A6AE726" box="[206,359,549,576]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="15" pageNumber="316" unit="m" value="2250.0" valueMax="2300.0" valueMin="2200.0">
<elevation id="E5F25018FFF33866FF29B0193A6AE726" box="[206,359,549,576]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="15" pageNumber="316" unit="m" value="2250.0" valueMax="2300.0" valueMin="2200.0">2200-2300 m</elevation>
</quantity>
,
<geoCoordinate id="0BEBD1ECFFF33866FE94B01A3AD7E726" box="[371,474,549,576]" degrees="42" direction="north" minutes="35" orientation="latitude" pageId="15" pageNumber="316" precision="925" value="42.583332">N 42°35’</geoCoordinate>
/
<geoCoordinate id="0BEBD1ECFFF33866FE16B01A3954E759" box="[497,601,549,576]" degrees="42" direction="east" minutes="32" orientation="longitude" pageId="15" pageNumber="316" precision="925" value="42.533333">E 42°32’</geoCoordinate>
, 21.-
<date id="1A6191EBFFF33866FD6DB019381DE759" box="[650,784,549,575]" pageId="15" pageNumber="316" value="2014-06-22">
<collectingDate id="0A256803FFF33866FD6DB019381DE759" box="[650,784,549,575]" pageId="15" pageNumber="316" value="2014-06-22">22.06.2014</collectingDate>
</date>
(
<collectorName id="C32AD2FDFFF33866FCC7B0193882E726" box="[800,911,549,576]" pageId="15" pageNumber="316">A. Zubov</collectorName>
) (
<collectionCode id="08CE2FEEFFF33866FC42B01938EDE759" box="[933,992,549,575]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="15" pageNumber="316">ZISP</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FC0BB0193F36E759" box="[1004,1083,549,575]" country="Thailand" httpUri="http://grbio.org/cool/w22j-vay4" name="Department of Microbiology, Songkla University" pageId="15" pageNumber="316">MPSU</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FBA0B0193F98E759" box="[1095,1173,549,575]" pageId="15" pageNumber="316">CFDD</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FB46B0193FE5E759" box="[1185,1256,549,575]" pageId="15" pageNumber="316">CTAY</collectionCode>
,
<collectionCode id="08CE2FEEFFF33866FB14B0193E44E759" box="[1267,1353,549,575]" pageId="15" pageNumber="316">CZAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DEB7BD76FFF33866FABAB0193E95E7E1" ID-GBIF-Occurrence="2561484934" collectingDate="2016-06-27" collectionCode="CFDD" collectorName="A. Zubov" country="Georgia" county="Jvari Pass" elevation="2450" latitude="42.75874" location="Kvemo Svaneti" longLatPrecision="1" longitude="42.14179" municipality="Tskhenis-Tskali River" pageId="15" pageNumber="316" specimenCount="7" specimenCount-female="3" specimenCount-male="4">
<specimenCount id="78D97CA2FFF33866FABAB0193E9AE759" box="[1373,1431,549,575]" pageId="15" pageNumber="316" type="male">4♂♂</specimenCount>
,
<specimenCount id="78D97CA2FFF33866FF70B0753BDDE705" box="[151,208,585,611]" pageId="15" pageNumber="316" type="female">3♀♀</specimenCount>
,
<collectingCountry id="16C8F7BBFFF33866FF39B0753A36E705" box="[222,315,585,611]" name="Georgia" pageId="15" pageNumber="316">Georgia</collectingCountry>
,
<location id="6B00E1F0FFF33866FEA0B0763AF5E705" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FEA0B0763AF5E705" box="[327,504,585,612]" country="Georgia" county="Jvari Pass" latitude="42.75874" longLatPrecision="1" longitude="42.14179" municipality="Tskhenis-Tskali River" name="Kvemo Svaneti" pageId="15" pageNumber="316">Kvemo Svaneti</location>
, N slopes of
<location id="6B00E1F0FFF33866FD69B0753882E702" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FD69B0753882E702" box="[654,911,585,612]" country="Georgia" county="Jvari Pass" latitude="42.75874" longLatPrecision="1" longitude="42.14179" municipality="Tskhenis-Tskali River" name="Lechkhumi Mt. Range" pageId="15" pageNumber="316">Lechkhumi Mt. Range</location>
, left bank of
<collectingMunicipality id="8E042D51FFF33866FBC0B0753E12E705" box="[1063,1311,585,611]" pageId="15" pageNumber="316">Tskhenis-Tskali River</collectingMunicipality>
,
<locationDeviation id="7FD2AE09FFF33866FACCB0753BE6E7E1" location="Sasashi" pageId="15" pageNumber="316">
<quantity id="A9271ACEFFF33866FACCB0753E68E705" box="[1323,1381,585,611]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="316" unit="km" value="4.0">4 km</quantity>
S of Sasashi
</locationDeviation>
vill., near
<collectingCounty id="8701CFA7FFF33866FE83B0513ADBE7EE" box="[356,470,621,648]" pageId="15" pageNumber="316">Jvari Pass</collectingCounty>
,
<quantity id="A9271ACEFFF33866FE05B0513977E7EE" box="[482,634,621,648]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="15" pageNumber="316" unit="m" value="2450.0" valueMax="2600.0" valueMin="2300.0">
<elevation id="E5F25018FFF33866FE05B0513977E7EE" box="[482,634,621,648]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="15" pageNumber="316" unit="m" value="2450.0" valueMax="2600.0" valueMin="2300.0">2300-2600 m</elevation>
</quantity>
,
<geoCoordinate id="0BEBD1ECFFF33866FD61B0523837E7EE" box="[646,826,621,648]" degrees="42" direction="north" minutes="45" orientation="latitude" pageId="15" pageNumber="316" precision="1" seconds="31,46" value="42.75874">N 42°45’31,46”</geoCoordinate>
/
<geoCoordinate id="0BEBD1ECFFF33866FCB7B05238F8E7E1" box="[848,1013,621,648]" degrees="42" direction="east" minutes="8" orientation="longitude" pageId="15" pageNumber="316" precision="1" seconds="30,43" value="42.14179">E 42°8’30,43”</geoCoordinate>
, 26.-
<date id="1A6191EBFFF33866FBC0B0513FA6E7E1" box="[1063,1195,621,647]" pageId="15" pageNumber="316" value="2016-06-27">
<collectingDate id="0A256803FFF33866FBC0B0513FA6E7E1" box="[1063,1195,621,647]" pageId="15" pageNumber="316" value="2016-06-27">27.06.2016</collectingDate>
</date>
(
<collectorName id="C32AD2FDFFF33866FB5CB0513E25E7EE" box="[1211,1320,621,648]" pageId="15" pageNumber="316">A. Zubov</collectorName>
) (
<collectionCode id="08CE2FEEFFF33866FAD8B0513E9DE7E1" box="[1343,1424,621,647]" pageId="15" pageNumber="316">CFDD</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DEB7BD76FFF33866FF70B0AD3AF5E795" ID-GBIF-Occurrence="2561484950" collectingDate="2017-07-13" collectionCode="ZISP, MPSU, CFDD, CTAY, CZAM, CKEE" collectorName="D. Fominykh" country="Georgia" county="Goldash Mt." elevation="2615" latitude="42.83336" location="Kvemo-Svaneti" longLatPrecision="1" longitude="42.754696" municipality="Lentekhi" pageId="15" pageNumber="316" specimenCount="270" specimenCount-female="146" specimenCount-male="124">
<specimenCount id="78D97CA2FFF33866FF70B0AD3BE0E7CD" box="[151,237,657,683]" pageId="15" pageNumber="316" type="male">124♂♂</specimenCount>
,
<specimenCount id="78D97CA2FFF33866FF1DB0AD3A5DE7CD" box="[250,336,657,683]" pageId="15" pageNumber="316" type="female">146♀♀</specimenCount>
,
<collectingCountry id="16C8F7BBFFF33866FEBBB0AD3AB4E7CD" box="[348,441,657,683]" name="Georgia" pageId="15" pageNumber="316">Georgia</collectingCountry>
,
<location id="6B00E1F0FFF33866FE23B0AD397AE7CD" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FE23B0AD397AE7CD" box="[452,631,657,683]" country="Georgia" county="Goldash Mt." latitude="42.83336" longLatPrecision="1" longitude="42.754696" municipality="Lentekhi" name="Kvemo-Svaneti" pageId="15" pageNumber="316">Kvemo-Svaneti</location>
, S slopes of
<location id="6B00E1F0FFF33866FCE2B0AD38D5E7CA" LSID="urn:lsid:plazi:treatment:E676063DFFF33866FF70B2A63A75E18D:6B00E1F0FFF33866FCE2B0AD38D5E7CA" box="[773,984,657,684]" country="Georgia" county="Goldash Mt." latitude="42.83336" longLatPrecision="1" longitude="42.754696" municipality="Lentekhi" name="Svaneti Mt. Range" pageId="15" pageNumber="316">Svaneti Mt. Range</location>
near
<collectingMunicipality id="8E042D51FFF33866FBF1B0AD3F77E7CD" box="[1046,1146,657,683]" pageId="15" pageNumber="316">Lentekhi</collectingMunicipality>
vill., S slopes of
<collectingCounty id="8701CFA7FFF33866FAD8B0AD3BCFE7B6" pageId="15" pageNumber="316">Goldash Mt.</collectingCounty>
,
<quantity id="A9271ACEFFF33866FF37B0893A26E7B6" box="[208,299,693,720]" metricMagnitude="3" metricUnit="m" metricValue="2.6149999999999998" pageId="15" pageNumber="316" unit="m" value="2615.0">
<elevation id="E5F25018FFF33866FF37B0893A26E7B6" box="[208,299,693,720]" metricMagnitude="3" metricUnit="m" metricValue="2.6149999999999998" pageId="15" pageNumber="316" unit="m" value="2615.0">2615 m</elevation>
</quantity>
,
<geoCoordinate id="0BEBD1ECFFF33866FEDCB08A3AFCE7B6" box="[315,497,693,720]" degrees="42" direction="north" minutes="50" orientation="latitude" pageId="15" pageNumber="316" precision="1" seconds="00,10" value="42.83336">N 42°50’00,10”</geoCoordinate>
/
<geoCoordinate id="0BEBD1ECFFF33866FDE9B08A39C9E7B6" box="[526,708,693,720]" degrees="42" direction="east" minutes="45" orientation="longitude" pageId="15" pageNumber="316" precision="1" seconds="16,90" value="42.754696">E 42°45’16,90”</geoCoordinate>
,
<collectingMethod id="B79ECF3CFFF33866FD33B0893857E7B6" box="[724,858,693,720]" pageId="15" pageNumber="316">pitfall traps</collectingMethod>
, 18.06.-
<date id="1A6191EBFFF33866FC53B0893F3AE7A9" box="[948,1079,693,719]" pageId="15" pageNumber="316" value="2017-07-13">
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(
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) (
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,
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,
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,
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,
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,
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;
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,
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,
<collectingCountry id="16C8F7BBFFF33866FD4BB0E53804E795" box="[684,777,729,755]" name="Georgia" pageId="15" pageNumber="316">Georgia</collectingCountry>
,
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, S slopes of
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near Len- tekhi vill., S slope of
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,
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,
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/
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,
<collectingMethod id="B79ECF3CFFF33866FBF4B0C13F9AE67E" box="[1043,1175,765,792]" pageId="15" pageNumber="316">pitfall traps</collectingMethod>
, 17.06.-
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(
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) (
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,
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,
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,
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,
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="26C5E4A0FFF33866FF20B17938E9E15E" pageId="15" pageNumber="316" type="distribution">
<paragraph id="6E60B72BFFF33866FF20B17938E9E15E" blockId="15.[151,1437,261,1260]" pageId="15" pageNumber="316">
<emphasis id="5CAB6B39FFF33866FF20B1793AE2E606" bold="true" box="[199,495,837,864]" pageId="15" pageNumber="316">Distribution and habitat.</emphasis>
Lebarde was pointed originally as the
<typeStatus id="B1640989FFF33866FC7FB17A38C5E606" box="[920,968,838,864]" pageId="15" pageNumber="316">type</typeStatus>
locality of this subspecies (
<bibRefCitation id="0A4ECADAFFF33866FB10B1793E9DE639" author="Retezar, I." box="[1271,1424,837,863]" pageId="15" pageNumber="316" pagination="85 - 88" refId="ref10287" refString="Retezar, I. (1992) Two new Carabus species from the Caucasus (Coleoptera: Carabidae). Folia Entomologica Hungarica, 52, 85 - 88." type="journal article" year="1992">Retezár 1992</bibRefCitation>
). Later
<bibRefCitation id="0A4ECADAFFF33866FF3FB1553A72E6E2" author="Retezar, I." box="[216,383,873,900]" pageId="15" pageNumber="316" refId="ref10317" refString="Retezar, I. (2015) Atlas of the Carabus of the Caucasus (Coleoptera, Carabidae). Iconography, genital morphology, systematics and faunistics. Szerzoi kiadas, Mondat Kft, Budapest, 134 pp., 79 maps, 238 color pls." type="book" year="2015">Retezár (2015)</bibRefCitation>
added a further locality, Plateau Askhi, located south-eastwards of Lebarde. We do not possess material from the
<typeStatus id="B1640989FFF33866FEBAB1B23A80E6CE" box="[349,397,910,936]" pageId="15" pageNumber="316">type</typeStatus>
locality, though examination of the available specimens from Plateau Askhi revealed its identity with populations inhabiting various mountains in the Greater Caucasus (namely Racha, Lechkhumi, and Svaneti ranges) and morphologically corresponding to the original description. Thus the distribution range of
<taxonomicName id="A9DFCCA8FFF33866FAEDB1EA3BE0E175" authorityName="Retezar, Mravaldzali" authorityYear="1992" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="gvalijai">
<emphasis id="5CAB6B39FFF33866FAEDB1EA3BE0E175" italics="true" pageId="15" pageNumber="316">C. stjernvalli gvalijai</emphasis>
</taxonomicName>
appeared to be actually much more widespread than accepted earlier. Nearly all populations were found on alpine meadows above
<quantity id="A9271ACEFFF33866FE7AB6213AF9E15E" box="[413,500,1053,1080]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="316" unit="m" value="2000.0">2000 m</quantity>
, only one was discovered in the forest zone.
</paragraph>
</subSubSection>
<subSubSection id="26C5E4A0FFF33866FF20B67E3A75E18D" pageId="15" pageNumber="316" type="discussion">
<paragraph id="6E60B72BFFF33866FF20B67E3A75E18D" blockId="15.[151,1437,261,1260]" pageId="15" pageNumber="316">
<emphasis id="5CAB6B39FFF33866FF20B67E3A36E13A" bold="true" box="[199,315,1090,1116]" pageId="15" pageNumber="316">Remarks.</emphasis>
Despite some minor differences in general appearance of individuals from different populations (
<figureCitation id="F6E4ABAEFFF33866FF78B6593A24E1E6" box="[159,297,1125,1152]" captionStart-0="FIGURES 68–75" captionStart-1="FIGURES 76–83" captionStart-2="FIGURES 84–91" captionStartId-0="16.[151,264,1942,1967]" captionStartId-1="17.[151,264,1930,1955]" captionStartId-2="18.[151,264,1929,1954]" captionTargetBox-0="[159,1426,209,1853]" captionTargetBox-1="[160,1424,210,1856]" captionTargetBox-2="[164,1414,194,1885]" captionTargetId-0="figure@16.[151,1436,199,1867]" captionTargetId-1="figure@17.[151,1436,196,1866]" captionTargetId-2="figure@18.[151,1436,181,1904]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURES 68–75. Dorsal habitus of Carabus (Lipaster) stjernvalli gvalijai Retezár: 68–71—Mravaldzali env., 2200 m; 72– 75—Tli env., 2000 m. Scale bar 10.0 mm." captionText-1="FIGURES 76–83. Dorsal habitus of Carabus (Lipaster) stjernvalli gvalijai Retezár: 76–79—Askhi Plateau, 2300 m; 80–83— Sasashi env., 2300 m. Scale bar 10.0 mm." captionText-2="FIGURES 84–91. Dorsal habitus of Carabus (Lipaster) stjernvalli gvalijai Retezár: 84–87—Goldash Mt., 2600 m; 88–91— Goldash Mt., 1500 m. Scale bar 10.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3632688" figureDoi-1="http://doi.org/10.5281/zenodo.3632690" figureDoi-2="http://doi.org/10.5281/zenodo.3632692" httpUri-0="https://zenodo.org/record/3632688/files/figure.png" httpUri-1="https://zenodo.org/record/3632690/files/figure.png" httpUri-2="https://zenodo.org/record/3632692/files/figure.png" pageId="15" pageNumber="316">Figs. 68–91</figureCitation>
), their body proportions are rather close (
<tableCitation id="235D8290FFF33866FD1EB6593841E119" box="[761,844,1125,1151]" captionStart="TABLE 1" captionStartId="3.[151,244,453,480]" captionTargetBox="[162,2021,538,1731]" captionText="TABLE 1. Morphometric data of Carabus (Lipaster) stjernvalli Mannerheim subspecies in the South Caucasus. Characters abbreviations as indicated in Material and Methods." httpUri="http://table.plazi.org/id/3AA0E7A3FFFF386AFF70B3F93A0CE762" pageId="15" pageNumber="316" tableUuid="3AA0E7A3FFFF386AFF70B3F93A0CE762">Table 1</tableCitation>
). Male genitalia structure is stable in all populations studied (
<figureCitation id="F6E4ABAEFFF33866FF10B6B53A91E1C5" box="[247,412,1161,1187]" captionStart="FIGURES 102–114" captionStartId="19.[151,264,1829,1854]" captionTargetBox="[230,1355,1226,1796]" captionTargetId="figure@19.[225,1361,1220,1804]" captionTargetPageId="19" captionText="FIGURES 102–114. Apical portion of aedeagus and fully inflated endophallus preparation of Carabus (Lipaster) stjernvalli Mannerheim, right lateral view: 102—C. stjernvalli stjernvalli Mannerheim, Chidila Pass env., 2350 m; 103—idem, Dzegvila Mt., 1100 m; 104—idem, Gomismta env., 2100 m; 105—idem, Gomi env., 1500 m; 106—Goderdzi Pass env., 2300 m; 107— idem, Zekari Pass env., 2100 m; 108—idem, Sairme env., 1500 m; 109—C. stjernvalli gvalijai Retezár, Mravaldzali env., 2200 m; 110—idem, Tli env., 2000 m; 111—idem, Askhi Plateau, 2300 m; 112—idem, Sasashi env., 2300 m; 113—idem, Goldash Mt., 2600 m; 114 —idem, Goldash Mt., 1500 m. Scale bar 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3632696" httpUri="https://zenodo.org/record/3632696/files/figure.png" pageId="15" pageNumber="316">Figs. 109–114</figureCitation>
). Only one population from Racha Mountain Range (Mravaldzali vicinity) comprises black individuals. The upper side is usually with metallic luster, but generally more matt than in the nominotypical subspecies, bronze to blue.
</paragraph>
</subSubSection>
</treatment>
</document>
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