treatments-xml/data/D9/5F/87/D95F87C6FF9AFFD44BF1FC6C508668BD.xml
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<document id="1BDC831F32910D5E123F0B9C543959A1" ID-CLB-Dataset="25274" ID-DOI="10.11646/zootaxa.4683.3.5" ID-GBIF-Dataset="39bcbc6b-37c9-416c-b655-46521260f336" ID-ISSN="1175-5326" ID-Zenodo-Dep="3479023" ID-ZooBank="2031FAC2-D64B-4315-B4D7-DEB69D80C095" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570615964789" checkinUser="plazi" docAuthor="Vilela, Diogo Silva, Guillermo-Ferreira, Rhainer, Encalada, Andrea C. &amp; Cordero-Rivera, Adolfo" docDate="2019" docId="D95F87C6FF9AFFD44BF1FC6C508668BD" docLanguage="en" docName="zootaxa.4683.3.5.pdf" docOrigin="Zootaxa 4683 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philogenia gaiae Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="419" masterDocId="2566FFBEFF9BFFD34B66FFDD511F6936" masterDocTitle="Philogenia gaiae sp. nov. (Zygoptera: Philogeniidae) and description of the female of P. macuma Dunkle, 1986, two species from the Ecuadorean lowland rainforest" masterLastPageNumber="420" masterPageNumber="412" pageNumber="413" updateTime="1698758593221" updateUser="ExternalLinkService">
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<mods:title id="1BF2384BED4971510396E2938EB575F9">Philogenia gaiae sp. nov. (Zygoptera: Philogeniidae) and description of the female of P. macuma Dunkle, 1986, two species from the Ecuadorean lowland rainforest</mods:title>
</mods:titleInfo>
<mods:name id="B646783C89808F7C0B97F84A6E37EA79" type="personal">
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<mods:namePart id="E1848F37821A0E09BC68B868479B82D7">Vilela, Diogo Silva</mods:namePart>
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<mods:namePart id="08FD9F2CAF81BD799E103B70A081E168">Guillermo-Ferreira, Rhainer</mods:namePart>
</mods:name>
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<mods:role id="2E62B69D9FD6F8E80EE90E085AB6E94C">
<mods:roleTerm id="00455892E61C7C9CD5D0697EEFDCC187">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4943166E32F1D2EF553D98D3E39E425">Encalada, Andrea C.</mods:namePart>
</mods:name>
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<mods:role id="5B288D9FF6595063BC861CF7F74A10EF">
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</mods:role>
<mods:namePart id="1FCA51A1CEFDAB672A4A8A45E5AD5698">Cordero-Rivera, Adolfo</mods:namePart>
</mods:name>
<mods:typeOfResource id="BA625D5C0B66B63D1C02C53D2D14296B">text</mods:typeOfResource>
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<mods:title id="5206F5AEEE1A35F302A21D79E9ADB857">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8191F90A23EE92DCA8A82B47962B1427">
<mods:date id="1527FFF2B08276898EE37A5852490E64">2019</mods:date>
<mods:detail id="667BA7D68738E3C9BF4CB350FE040473" type="pubDate">
<mods:number id="A62BD606E734621888D2CB1545B4210D">2019-10-09</mods:number>
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<mods:number id="E54DFC98074C15F7B19F7ACC2F9FF4CB">4683</mods:number>
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<mods:number id="4737F75A4B9DCD6F185683BFE5B74D7A">3</mods:number>
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<mods:identifier id="0C9C5370C6FFF07C05AA7E69F21A8F57" type="DOI">10.11646/zootaxa.4683.3.5</mods:identifier>
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<treatment id="D95F87C6FF9AFFD44BF1FC6C508668BD" ID-GBIF-Taxon="159673613" LSID="urn:lsid:plazi:treatment:D95F87C6FF9AFFD44BF1FC6C508668BD" httpUri="http://treatment.plazi.org/id/D95F87C6FF9AFFD44BF1FC6C508668BD" lastPageId="7" lastPageNumber="419" pageId="1" pageNumber="413">
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<heading id="0A0181BCFF9AFFD24BF1FC6C52186AFD" bold="true" box="[151,775,945,972]" fontSize="11" level="1" pageId="1" pageNumber="413" reason="1">
<emphasis id="6382EAC2FF9AFFD24BF1FC6C52186AFD" bold="true" box="[151,775,945,972]" pageId="1" pageNumber="413">
<taxonomicName id="96F64D53FF9AFFD24BF1FC6C507B6AFD" authority="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera, 2019" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[151,356,945,972]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="gaiae" status="sp. nov.">
<emphasis id="6382EAC2FF9AFFD24BF1FC6C507B6AFD" bold="true" box="[151,356,945,972]" italics="true" pageId="1" pageNumber="413">Philogenia gaiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78B157B9FF9AFFD24A0DFC6C50D66AFD" box="[363,457,945,971]" pageId="1" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
Vilela &amp; Cordero-Rivera
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="19EC655BFF9AFFD24BF1FC0853156AC6" box="[151,522,980,1008]" pageId="1" pageNumber="413" type="description">
<paragraph id="514936D0FF9AFFD24BF1FC0853156AC6" blockId="1.[151,775,945,1008]" box="[151,522,980,1008]" pageId="1" pageNumber="413">
(
<figureCitation id="C9CD2A55FF9AFFD24BF9FC0851F26AD9" box="[159,237,981,1007]" captionStart="FIGURE 1" captionStartId="2.[151,250,1062,1087]" captionTargetBox="[156,1429,375,1030]" captionTargetId="figure@2.[151,1436,371,1040]" captionTargetPageId="2" captionText="FIGURE 1. Philogenia gaiae sp. nov., holotype ♂; (1a) habitus; (1b) head; (1c) wings." figureDoi="http://doi.org/10.5281/zenodo.3479025" httpUri="https://zenodo.org/record/3479025/files/figure.png" pageId="1" pageNumber="413">Figs. 1</figureCitation>
,
<figureCitation id="C9CD2A55FF9AFFD24B9DFC0850146AD9" box="[251,267,981,1007]" captionStart="FIGURE 2" captionStartId="2.[151,250,1979,2004]" captionTargetBox="[162,1425,1101,1960]" captionTargetId="figure@2.[162,1425,1101,1960]" captionTargetPageId="2" captionText="FIGURE 2. Philogenia gaiae sp. nov., paratype ♂ ACR00579; (2a) lateral view of genital ligula; (2b) detail of genital ligula distal flagella; (2c) detail of genital ligula flagellum apex; (2d) detail of lateral first segment of genital ligula." figureDoi="http://doi.org/10.5281/zenodo.3479027" httpUri="https://zenodo.org/record/3479027/files/figure.png" pageId="1" pageNumber="413">2</figureCitation>
,
<figureCitation id="C9CD2A55FF9AFFD24A71FC08503B6AD9" box="[279,292,981,1007]" captionStart="FIGURE 3" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[186,1400,352,1861]" captionTargetId="figure@3.[174,1413,337,1882]" captionTargetPageId="3" captionText="FIGURE 3. Philogenia gaiae sp. nov., paratype ♂ from Ecuador, Tiputini at RWG; (3a) appendages in lateral view; (3b) detail of right paraproct in lateral view; (3c) appendages in dorsal view. Philogenia minteri, ♂ from Ecuador, Limoncocha at RWG; (3d) appendages in lateral view; (3e) detail of right paraproct in lateral view; (3f) appendages in dorsal view. (Photos 3a,c, d, f by RWG)" figureDoi="http://doi.org/10.5281/zenodo.3479029" httpUri="https://zenodo.org/record/3479029/files/figure.png" pageId="1" pageNumber="413">3</figureCitation>
ac, 5ae and 6ab)
</paragraph>
</subSubSection>
<subSubSection id="19EC655BFF9AFFD24BF1FBC055DC6F65" pageId="1" pageNumber="413" type="materials_examined">
<paragraph id="514936D0FF9AFFD24BF1FBC0558E6D6D" blockId="1.[151,1437,1052,2015]" pageId="1" pageNumber="413">
<materialsCitation id="E19E3C8DFF9AFFD24BF1FBC0558E6D6D" ID-GBIF-Occurrence="2425441296" collectingDate="2012-12-13" collectionCode="ECOEVO, ACR" collectorName="A. Cordero-Rivera &amp; M. Sanchez-Herrera" country="Ecuador" elevation="241" latitude="-0.6349" location="Tiputini Biodiversity Station" longLatPrecision="7" longitude="-76.1501" pageId="1" pageNumber="413" specimenCount="1" specimenCount-male="1" stateProvince="Orellana" typeStatus="holotype">
<emphasis id="6382EAC2FF9AFFD24BF1FBC050146D01" bold="true" box="[151,267,1053,1079]" pageId="1" pageNumber="413">
<typeStatus id="8E4D8872FF9AFFD24BF1FBC050196D01" box="[151,262,1053,1079]" pageId="1" pageNumber="413" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="47F0FD59FF9AFFD24A74FBC050386D01" box="[274,295,1053,1079]" pageId="1" pageNumber="413" type="male"></specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD24A50FBC050B36D01" box="[310,428,1053,1079]" pageId="1" pageNumber="413">ECOEVO</collectionCode>
,
<collectionCode id="37E7AE15FF9AFFD24AD0FBC050EE6D01" box="[438,497,1053,1079]" pageId="1" pageNumber="413">ACR</collectionCode>
00636, in copula with
<collectionCode id="37E7AE15FF9AFFD2498DFBC152386D01" box="[747,807,1052,1079]" pageId="1" pageNumber="413">ACR</collectionCode>
00635),
<collectingCountry id="29E17640FF9AFFD248E4FBC052FE6D01" box="[898,993,1053,1079]" name="Ecuador" pageId="1" pageNumber="413">Ecuador</collectingCountry>
,
<collectingRegion id="9332F832FF9AFFD2488AFBC055506D01" box="[1004,1103,1053,1079]" country="Ecuador" name="Orellana" pageId="1" pageNumber="413">Orellana</collectingRegion>
,
<location id="5429600BFF9AFFD24F3DFBC054836D01" LSID="urn:lsid:plazi:treatment:D95F87C6FF9AFFD44BF1FC6C508668BD:5429600BFF9AFFD24F3DFBC054836D01" box="[1115,1436,1053,1079]" country="Ecuador" latitude="-0.6349" longLatPrecision="7" longitude="-76.1501" name="Tiputini Biodiversity Station" pageId="1" pageNumber="413" stateProvince="Orellana">Tiputini Biodiversity Station</location>
(
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,
<geoCoordinate id="34C25017FF9AFFD24A63FB9D50746D6D" box="[261,363,1088,1115]" degrees="76.1501" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1501">-76.1501</geoCoordinate>
,
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<quantity id="960E9B35FF9AFFD24A11FB9C50A16D6D" box="[375,446,1089,1115]" metricMagnitude="2" metricUnit="m" metricValue="2.41" pageId="1" pageNumber="413" unit="m" value="241.0">241 m</quantity>
asl
</elevation>
),
<date id="25481010FF9AFFD24A9DFB9C53626D6D" box="[507,637,1089,1115]" pageId="1" pageNumber="413" value="2012-12-13">
<collectingDate id="350CE9F8FF9AFFD24A9DFB9C53626D6D" box="[507,637,1089,1115]" pageId="1" pageNumber="413" value="2012-12-13">13.xii.2012</collectingDate>
</date>
,
<collectorName id="FC035306FF9AFFD249E1FB9C52456D6D" box="[647,858,1089,1115]" pageId="1" pageNumber="413">A. Cordero-Rivera</collectorName>
&amp;
<collectorName id="FC035306FF9AFFD2481BFB9C557F6D6D" box="[893,1120,1089,1115]" pageId="1" pageNumber="413">M. Sánchez-Herrera</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="514936D0FF9AFFD24BA1FBB851DE6F3D" blockId="1.[151,1437,1052,2015]" pageId="1" pageNumber="413">
<materialsCitation id="E19E3C8DFF9AFFD24BA1FBB8549F6DB6" ID-GBIF-Occurrence="2425441273" box="[199,1408,1124,1152]" collectionCode="ECOEVO" pageId="1" pageNumber="413" specimenCount="6" specimenCount-male="6" typeStatus="paratype">
<emphasis id="6382EAC2FF9AFFD24BA1FBB850596D49" bold="true" box="[199,326,1125,1151]" pageId="1" pageNumber="413">
<typeStatus id="8E4D8872FF9AFFD24BA1FBB8505E6D49" box="[199,321,1125,1151]" pageId="1" pageNumber="413" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="47F0FD59FF9AFFD24A36FBB8508C6D49" box="[336,403,1125,1151]" pageId="1" pageNumber="413" type="male">6 ♂♂</specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD24AC2FBB853056D49" box="[420,538,1125,1151]" pageId="1" pageNumber="413">ECOEVO</collectionCode>
, ACR00477, ACR00501, ACR00579, ACR00580, ACR00582, ACR00624)
</materialsCitation>
,
<materialsCitation id="E19E3C8DFF9AFFD24EE8FBB8533A6DDD" ID-GBIF-Occurrence="2425441219" collectingDate="2012-12-04" collectingDateMax="2012-12-12" collectingDateMin="2012-12-04" collectionCode="ECOEVO, ACR" collectorName="A. Cordero-Rivera &amp; M. Sanchez-Herrera" country="Ecuador" elevation="241" latitude="-0.6349" location="Tiputini Biodiversity Station" longLatPrecision="7" longitude="-76.1501" pageId="1" pageNumber="413" specimenCount="1" specimenCount-female="1" stateProvince="Orellana" typeStatus="paratype">
<specimenCount id="47F0FD59FF9AFFD24EE8FBB851B36D95" pageId="1" pageNumber="413" type="female">1 ♀</specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD24BD9FB54502A6D95" box="[191,309,1161,1187]" pageId="1" pageNumber="413">ECOEVO</collectionCode>
,
<collectionCode id="37E7AE15FF9AFFD24A24FB5550606D95" box="[322,383,1160,1187]" pageId="1" pageNumber="413">ACR</collectionCode>
00581), locality data same as for holotype, except for
<collectionCode id="37E7AE15FF9AFFD2489CFB54552B6D95" box="[1018,1076,1161,1187]" pageId="1" pageNumber="413">ACR</collectionCode>
00477 collected in
<date id="25481010FF9AFFD24E73FB5454876D95" box="[1301,1432,1161,1187]" pageId="1" pageNumber="413" value="2012-12-04">
<collectingDate id="350CE9F8FF9AFFD24E73FB5454876D95" box="[1301,1432,1161,1187]" pageId="1" pageNumber="413" value="2012-12-04">04.xii.2012</collectingDate>
</date>
;
<collectionCode id="37E7AE15FF9AFFD24BF1FB7151CD6DF1" box="[151,210,1196,1223]" pageId="1" pageNumber="413">ACR</collectionCode>
00501 collected in
<date id="25481010FF9AFFD24ACEFB7153346DF1" box="[424,555,1196,1223]" pageId="1" pageNumber="413" value="2012-12-05">
<collectingDate id="350CE9F8FF9AFFD24ACEFB7153346DF1" box="[424,555,1196,1223]" pageId="1" pageNumber="413" value="2012-12-05">05.xii.2012</collectingDate>
</date>
; ACR00579, ACR00580, ACR00581 and ACR00582 collected in
<date id="25481010FF9AFFD24E70FB7054876DF1" box="[1302,1432,1197,1223]" pageId="1" pageNumber="413" value="2012-12-09">
<collectingDate id="350CE9F8FF9AFFD24E70FB7054876DF1" box="[1302,1432,1197,1223]" pageId="1" pageNumber="413" value="2012-12-09">09.xii.2012</collectingDate>
</date>
,
<collectionCode id="37E7AE15FF9AFFD24BF1FB0C51CE6DDD" box="[151,209,1233,1259]" pageId="1" pageNumber="413">ACR</collectionCode>
00624 collected in
<date id="25481010FF9AFFD24AC5FB0C533A6DDD" box="[419,549,1233,1259]" pageId="1" pageNumber="413" value="2012-12-12">
<collectingDate id="350CE9F8FF9AFFD24AC5FB0C533A6DDD" box="[419,549,1233,1259]" pageId="1" pageNumber="413" value="2012-12-12">12.xii.2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="E19E3C8DFF9AFFD24956FB0C541B6DDD" ID-GBIF-Occurrence="2425441320" box="[560,1284,1233,1259]" collectingDate="2012-12-04" collectingDateMax="2012-12-12" collectingDateMin="2012-12-04" collectionCode="RWG" collectorName="A. Cordero-Rivera &amp; M. Sanchez-Herrera" country="Ecuador" elevation="241" latitude="-0.6349" location="Tiputini Biodiversity Station" longLatPrecision="7" longitude="-76.1501" pageId="1" pageNumber="413" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Orellana" typeStatus="paratype">
<specimenCount id="47F0FD59FF9AFFD24956FB0C53786DDD" box="[560,615,1233,1259]" pageId="1" pageNumber="413" type="male">3♂♂</specimenCount>
and
<specimenCount id="47F0FD59FF9AFFD249FDFB0C53A16DDD" box="[667,702,1233,1259]" pageId="1" pageNumber="413" type="female">1♀</specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD249AAFB0C520D6DDD" box="[716,786,1233,1259]" pageId="1" pageNumber="413">RWG</collectionCode>
), same locality as other
<typeStatus id="8E4D8872FF9AFFD24F7EFB0C55996DDD" box="[1048,1158,1233,1259]" pageId="1" pageNumber="413" type="paratype">paratypes</typeStatus>
, except for
</materialsCitation>
:
<specimenCount id="47F0FD59FF9AFFD24E6BFB0C542F6DDD" box="[1293,1328,1233,1259]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(
<geoCoordinate id="34C25017FF9AFFD24E5BFB0C54876DDD" box="[1341,1432,1233,1259]" degrees="0.6378" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6378">-0.6378</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD24BF1FB2851E16C26" box="[151,254,1269,1296]" degrees="76.1486" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1486">-76.1486</geoCoordinate>
) and
<specimenCount id="47F0FD59FF9AFFD24A59FB28507E6C39" box="[319,353,1269,1295]" pageId="1" pageNumber="413" type="female">1♀</specimenCount>
(
<geoCoordinate id="34C25017FF9AFFD24A14FB2950D26C39" box="[370,461,1268,1295]" degrees="0.6256" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6256">-0.6256</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD24ABCFB28535F6C26" box="[474,576,1269,1296]" degrees="76.1494" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1494">-76.1494</geoCoordinate>
) collected in
<date id="25481010FF9AFFD249BAFB2852586C39" box="[732,839,1269,1295]" pageId="1" pageNumber="413" value="2009-01-11">11.i.2009</date>
,
<specimenCount id="47F0FD59FF9AFFD24832FB2852686C39" box="[852,887,1269,1295]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(-0.63, -76.1525) collected in
<date id="25481010FF9AFFD24FB0FB28545F6C39" box="[1238,1344,1269,1295]" pageId="1" pageNumber="413" value="2009-01-12">12.i.2009</date>
and
<specimenCount id="47F0FD59FF9AFFD24E1FFB2854836C39" box="[1401,1436,1269,1295]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(-0.635, -76.15) collected in
<date id="25481010FF9AFFD24ABFFAC4535C6C05" box="[473,579,1305,1331]" pageId="1" pageNumber="413" value="2009-01-16">16.i.2009</date>
all by R.W. Garrison &amp; N. von Ellenrieder leg;
<materialsCitation id="E19E3C8DFF9AFFD24F39FAC4535B6C61" ID-GBIF-Occurrence="2425441246" collectingDate="2012-12-04" collectingDateMax="2012-12-12" collectingDateMin="2012-12-04" collectionCode="CSCA" collectorName="A. Cordero-Rivera &amp; M. Sanchez-Herrera" country="Ecuador" elevation="241" latitude="-0.6349" location="Tiputini Biodiversity Station" longLatPrecision="7" longitude="-76.1501" pageId="1" pageNumber="413" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Orellana" typeStatus="paratype">
<specimenCount id="47F0FD59FF9AFFD24F39FAC455876C05" box="[1119,1176,1305,1331]" pageId="1" pageNumber="413" type="male">2♂♂</specimenCount>
and
<specimenCount id="47F0FD59FF9AFFD24FA8FAC455EE6C05" box="[1230,1265,1305,1331]" pageId="1" pageNumber="413" type="female">1♀</specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD24E67FAC4544F6C05" box="[1281,1360,1305,1331]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="1" pageNumber="413" type="Museum">CSCA</collectionCode>
), same locality as other
<typeStatus id="8E4D8872FF9AFFD24A32FAE050DD6C61" box="[340,450,1341,1367]" pageId="1" pageNumber="413" type="paratype">paratypes</typeStatus>
, except for
</materialsCitation>
:
<specimenCount id="47F0FD59FF9AFFD24929FAE0536D6C61" box="[591,626,1341,1367]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(
<geoCoordinate id="34C25017FF9AFFD249E4FAE053C26C61" box="[642,733,1341,1367]" degrees="0.6383" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6383">-0.6383</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD2498FFAE052516C6E" box="[745,846,1341,1368]" degrees="76.1491" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1491">-76.1491</geoCoordinate>
) collected in
<date id="25481010FF9AFFD2488FFAE0554A6C61" box="[1001,1109,1341,1367]" pageId="1" pageNumber="413" value="2009-01-16">16.i.2009</date>
,
<specimenCount id="47F0FD59FF9AFFD24F04FAE0559A6C61" box="[1122,1157,1341,1367]" pageId="1" pageNumber="413" type="female">1♀</specimenCount>
(
<geoCoordinate id="34C25017FF9AFFD24FF3FAE055F06C61" box="[1173,1263,1341,1367]" degrees="0.6383" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6383">-0.6383</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD24F9DFAE0547E6C6E" box="[1275,1377,1341,1368]" degrees="76.1491" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1491">-76.1491</geoCoordinate>
) col- lected in
<date id="25481010FF9AFFD24A66FABC50756C4D" box="[256,362,1377,1403]" pageId="1" pageNumber="413" value="2009-01-18">18.i.2009</date>
and
<specimenCount id="47F0FD59FF9AFFD24AC5FABC50D96C4D" box="[419,454,1377,1403]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(-0.635, -76.15) collected in
<date id="25481010FF9AFFD24872FABC52616C4D" box="[788,894,1377,1403]" pageId="1" pageNumber="413" value="2009-01-16">16.i.2009</date>
all by R.W. Garrison &amp; N. von Ellenrieder leg;
<materialsCitation id="E19E3C8DFF9AFFD24BF1FA5852356CA9" ID-GBIF-Occurrence="2425441229" box="[151,810,1413,1439]" collectingDate="2012-12-04" collectingDateMax="2012-12-12" collectingDateMin="2012-12-04" collectionCode="NVE" collectorName="A. Cordero-Rivera &amp; M. Sanchez-Herrera" country="Ecuador" elevation="241" latitude="-0.6349" location="Tiputini Biodiversity Station" longLatPrecision="7" longitude="-76.1501" pageId="1" pageNumber="413" specimenCount="2" specimenCount-male="2" stateProvince="Orellana" typeStatus="paratype">
<specimenCount id="47F0FD59FF9AFFD24BF1FA5851D06CA9" box="[151,207,1413,1439]" pageId="1" pageNumber="413" type="male">2♂♂</specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD24B86FA5850006CA9" box="[224,287,1413,1439]" pageId="1" pageNumber="413">NVE</collectionCode>
), same locality as other
<typeStatus id="8E4D8872FF9AFFD24951FA5853BA6CA9" box="[567,677,1413,1439]" pageId="1" pageNumber="413" type="paratype">paratypes</typeStatus>
, except for
</materialsCitation>
:
<specimenCount id="47F0FD59FF9AFFD24850FA5852466CA9" box="[822,857,1413,1439]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(
<geoCoordinate id="34C25017FF9AFFD2480CFA5852DA6CA9" box="[874,965,1413,1439]" degrees="0.6378" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6378">-0.6378</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD248B4FA5855276C96" box="[978,1080,1413,1440]" degrees="76.1486" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1486">-76.1486</geoCoordinate>
) collected in
<date id="25481010FF9AFFD24FB0FA5854206CA9" box="[1238,1343,1413,1439]" pageId="1" pageNumber="413" value="2009-01-11">11.i.2009</date>
and
<specimenCount id="47F0FD59FF9AFFD24E1FFA5854836CA9" box="[1401,1436,1413,1439]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(
<geoCoordinate id="34C25017FF9AFFD24BF9FA7451E56CF5" box="[159,250,1449,1475]" degrees="0.6369" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6369">-0.6369</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD24A63FA7550746CF5" box="[261,363,1448,1475]" degrees="76.1457" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1457">-76.1457</geoCoordinate>
) collected in
<date id="25481010FF9AFFD24965FA7453726CF5" box="[515,621,1449,1475]" pageId="1" pageNumber="413" value="2009-01-14">14.i.2009</date>
all by R.W. Garrison &amp; N. von Ellenrieder leg;
<materialsCitation id="E19E3C8DFF9AFFD24FEFFA7451DE6F3D" ID-GBIF-Occurrence="2425441248" collectingDate="2009-01-10" collectionCode="USFQ" collectorName="R. W. Garrison &amp; N. von Ellenrieder" country="Ecuador" latitude="-0.6358" longLatPrecision="7" longitude="-76.1503" pageId="1" pageNumber="413" specimenCount="1" specimenCount-male="1" stateProvince="Orellana" typeStatus="paratype">
<specimenCount id="47F0FD59FF9AFFD24FEFFA7455B36CF5" box="[1161,1196,1449,1475]" pageId="1" pageNumber="413" type="male">1♂</specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD24FDDFA7454176CF5" box="[1211,1288,1449,1475]" pageId="1" pageNumber="413">USFQ</collectionCode>
), same local- ity as other
<typeStatus id="8E4D8872FF9AFFD24A7CFA1050976CD1" box="[282,392,1485,1511]" pageId="1" pageNumber="413" type="paratype">paratypes</typeStatus>
, except for (
<geoCoordinate id="34C25017FF9AFFD24974FA1153726CD1" box="[530,621,1484,1511]" degrees="0.6358" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6358">-0.6358</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD2491EFA1153C16CD1" box="[632,734,1484,1511]" degrees="76.1503" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1503">-76.1503</geoCoordinate>
) collected in
<date id="25481010FF9AFFD24813FA1052C06CD1" box="[885,991,1485,1511]" pageId="1" pageNumber="413" value="2009-01-10">
<collectingDate id="350CE9F8FF9AFFD24813FA1052C06CD1" box="[885,991,1485,1511]" pageId="1" pageNumber="413" value="2009-01-10">10.i.2009</collectingDate>
</date>
by
<collectorName id="FC035306FF9AFFD24F6EFA1055B56CD1" box="[1032,1194,1485,1511]" pageId="1" pageNumber="413">R.W. Garrison</collectorName>
&amp;
<collectorName id="FC035306FF9AFFD24FABFA1054836CD1" box="[1229,1436,1485,1511]" pageId="1" pageNumber="413">N. von Ellenrieder</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="514936D0FF9AFFD24BA1F9C955DC6F65" blockId="1.[151,1437,1052,2015]" pageId="1" pageNumber="413">
<materialsCitation id="E19E3C8DFF9AFFD24BA1F9C955DC6F65" ID-GBIF-Occurrence="2425441292" collectingDate="2012-12-13" collectionCode="ECOEVO, ACR" collectorName="A. Cordero-Rivera &amp; M. Sanchez-Herrera" country="Ecuador" elevation="241" latitude="-0.6349" location="Tiputini Biodiversity" longLatPrecision="7" longitude="-76.1501" pageId="1" pageNumber="413" specimenCount="1" specimenCount-female="1" stateProvince="Orellana" typeStatus="allotype">
<emphasis id="6382EAC2FF9AFFD24BA1F9C9502C6F19" bold="true" box="[199,307,1556,1583]" pageId="1" pageNumber="413">
<typeStatus id="8E4D8872FF9AFFD24BA1F9C950316F19" box="[199,302,1556,1583]" pageId="1" pageNumber="413" type="allotype">Allotype</typeStatus>
.
</emphasis>
<specimenCount id="47F0FD59FF9AFFD24A5CF9C850506F19" box="[314,335,1557,1583]" pageId="1" pageNumber="413" type="female"></specimenCount>
(
<collectionCode id="37E7AE15FF9AFFD24A3BF9C850CC6F19" box="[349,467,1557,1583]" pageId="1" pageNumber="413">ECOEVO</collectionCode>
,
<collectionCode id="37E7AE15FF9AFFD24ABBF9C953066F19" box="[477,537,1556,1583]" pageId="1" pageNumber="413">ACR</collectionCode>
00635, in copula with
<collectionCode id="37E7AE15FF9AFFD24869F9C852546F06" box="[783,843,1557,1584]" pageId="1" pageNumber="413">ACR</collectionCode>
00636),
<collectingCountry id="29E17640FF9AFFD248C0F9C8551A6F19" box="[934,1029,1557,1583]" name="Ecuador" pageId="1" pageNumber="413">Ecuador</collectingCountry>
,
<collectingRegion id="9332F832FF9AFFD24F76F9C8556C6F19" box="[1040,1139,1557,1583]" country="Ecuador" name="Orellana" pageId="1" pageNumber="413">Orellana</collectingRegion>
,
<location id="5429600BFF9AFFD24F18F9C854776F19" LSID="urn:lsid:plazi:treatment:D95F87C6FF9AFFD44BF1FC6C508668BD:5429600BFF9AFFD24F18F9C854776F19" box="[1150,1384,1557,1583]" country="Ecuador" latitude="-0.6349" longLatPrecision="7" longitude="-76.1501" name="Tiputini Biodiversity" pageId="1" pageNumber="413" stateProvince="Orellana">Tiputini Biodiversity</location>
Sta- tion (
<geoCoordinate id="34C25017FF9AFFD24BB4F9E450336F65" box="[210,300,1593,1619]" degrees="0.6349" direction="south" orientation="latitude" pageId="1" pageNumber="413" precision="5" value="-0.6349">-0.6349</geoCoordinate>
,
<geoCoordinate id="34C25017FF9AFFD24A51F9E550816F65" box="[311,414,1592,1619]" degrees="76.1501" direction="west" orientation="longitude" pageId="1" pageNumber="413" precision="5" value="-76.1501">-76.1501</geoCoordinate>
,
<elevation id="DADBD1E3FF9AFFD24ACCF9E453056F62" box="[426,538,1593,1620]" metricMagnitude="2" metricUnit="m" metricValue="2.41" pageId="1" pageNumber="413" unit="m" value="241.0">
<quantity id="960E9B35FF9AFFD24ACCF9E450EE6F65" box="[426,497,1593,1619]" metricMagnitude="2" metricUnit="m" metricValue="2.41" pageId="1" pageNumber="413" unit="m" value="241.0">241 m</quantity>
asl
</elevation>
),
<date id="25481010FF9AFFD24948F9E453AF6F65" box="[558,688,1593,1619]" pageId="1" pageNumber="413" value="2012-12-13">
<collectingDate id="350CE9F8FF9AFFD24948F9E453AF6F65" box="[558,688,1593,1619]" pageId="1" pageNumber="413" value="2012-12-13">13.xii.2012</collectingDate>
</date>
,
<collectorName id="FC035306FF9AFFD249DCF9E452936F65" box="[698,908,1593,1619]" pageId="1" pageNumber="413">A. Cordero-Rivera</collectorName>
&amp;
<collectorName id="FC035306FF9AFFD248D6F9E4558C6F65" box="[944,1171,1593,1619]" pageId="1" pageNumber="413">M. Sanchez-Herrera</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="19EC655BFF9AFFD24BA1F98054426FAD" pageId="1" pageNumber="413" type="etymology">
<paragraph id="514936D0FF9AFFD24BA1F98054426FAD" blockId="1.[151,1437,1052,2015]" pageId="1" pageNumber="413">
<emphasis id="6382EAC2FF9AFFD24BA1F98050516F41" bold="true" box="[199,334,1629,1655]" pageId="1" pageNumber="413">Etymology.</emphasis>
Named
<taxonomicName id="96F64D53FF9AFFD24ACDF98050F66F41" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[427,489,1629,1655]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9AFFD24ACDF98050F66F41" bold="true" box="[427,489,1629,1655]" italics="true" pageId="1" pageNumber="413">gaiae</emphasis>
</taxonomicName>
(noun in the genitive case) after the daughter of ACR, Gaia Cordero Santolamazza, a dragonfly enthusiast, who has suffered her fathers long absences while sampling odonates around the world.
</paragraph>
</subSubSection>
<subSubSection id="19EC655BFF9AFFD44BA1F979508668BD" lastPageId="7" lastPageNumber="419" pageId="1" pageNumber="413" type="description">
<paragraph id="514936D0FF9AFFD24BA1F97954306E31" blockId="1.[151,1437,1052,2015]" pageId="1" pageNumber="413">
<emphasis id="6382EAC2FF9AFFD24BA1F97950FC6F89" bold="true" box="[199,483,1700,1727]" pageId="1" pageNumber="413">
Description of
<typeStatus id="8E4D8872FF9AFFD24A13F97850C16F89" box="[373,478,1701,1727]" pageId="1" pageNumber="413" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="6382EAC2FF9AFFD24A8FF97853396F89" box="[489,550,1701,1727]" italics="true" pageId="1" pageNumber="413">Head</emphasis>
(
<figureCitation id="C9CD2A55FF9AFFD24952F97853646FF6" box="[564,635,1701,1728]" captionStart="FIGURE 1" captionStartId="2.[151,250,1062,1087]" captionTargetBox="[156,1429,375,1030]" captionTargetId="figure@2.[151,1436,371,1040]" captionTargetPageId="2" captionText="FIGURE 1. Philogenia gaiae sp. nov., holotype ♂; (1a) habitus; (1b) head; (1c) wings." figureDoi="http://doi.org/10.5281/zenodo.3479025" httpUri="https://zenodo.org/record/3479025/files/figure.png" pageId="1" pageNumber="413">Figs. 1</figureCitation>
ab). Dark brown with labrum, mandibular base and basal half of genae greenish, antennal socket pale, lighter brown tinge between base of antennae and lateral ocelli, postocular area pale, rear of head pale. Frons slightly rounded. Postocular lobes at the level of hind margin of compound eyes.
</paragraph>
<paragraph id="514936D0FF9AFFD24BA1F8CC52BD6EA1" blockId="1.[151,1437,1052,2015]" pageId="1" pageNumber="413">
<emphasis id="6382EAC2FF9AFFD24BA1F8CC50076E1D" box="[199,280,1809,1835]" italics="true" pageId="1" pageNumber="413">Thorax</emphasis>
(
<figureCitation id="C9CD2A55FF9AFFD24A4EF8CC50686E1D" box="[296,375,1809,1835]" captionStart="FIGURE 1" captionStartId="2.[151,250,1062,1087]" captionTargetBox="[156,1429,375,1030]" captionTargetId="figure@2.[151,1436,371,1040]" captionTargetPageId="2" captionText="FIGURE 1. Philogenia gaiae sp. nov., holotype ♂; (1a) habitus; (1b) head; (1c) wings." figureDoi="http://doi.org/10.5281/zenodo.3479025" httpUri="https://zenodo.org/record/3479025/files/figure.png" pageId="1" pageNumber="413">Fig. 1a</figureCitation>
). Prothorax dark brown with yellowish laterodorsal stripes each extending up to 1/2 of mese- pisternum; posterior prothoracic lobe rounded. Pterothorax dark brown except for black antehumeral stripe, mesin- fraepisternum black, first 1/3 of metepisternum, upper half of metepimeron and venter pale. Coxae and internal surface of legs yellowish, external surfaces and armature dark brown.
</paragraph>
<paragraph id="514936D0FF9AFFD24BA1F87F51F76EE9" blockId="1.[151,1437,1052,2015]" pageId="1" pageNumber="413">
<emphasis id="6382EAC2FF9AFFD24BA1F87F50146E8D" box="[199,267,1954,1979]" italics="true" pageId="1" pageNumber="413">Wings</emphasis>
(
<figureCitation id="C9CD2A55FF9AFFD24A7DF87C50756E8D" box="[283,362,1953,1979]" captionStart="FIGURE 1" captionStartId="2.[151,250,1062,1087]" captionTargetBox="[156,1429,375,1030]" captionTargetId="figure@2.[151,1436,371,1040]" captionTargetPageId="2" captionText="FIGURE 1. Philogenia gaiae sp. nov., holotype ♂; (1a) habitus; (1b) head; (1c) wings." figureDoi="http://doi.org/10.5281/zenodo.3479025" httpUri="https://zenodo.org/record/3479025/files/figure.png" pageId="1" pageNumber="413">Fig. 1c</figureCitation>
). Hyaline. Pt dark brown surmounting four cells in FW and six cells in HW. Px =
<quantity id="960E9B35FF9AFFD24E6CF87C545C6E8D" box="[1290,1347,1953,1979]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="1" pageNumber="413" unit="in" value="27.0">27 in</quantity>
FW,
<quantity id="960E9B35FF9AFFD24EE6F87C51B26EE9" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="1" pageNumber="413" unit="in" value="24.0">24 in</quantity>
HW.
</paragraph>
<paragraph id="514936D0FF99FFD14BA1FF4452C069CD" blockId="2.[151,1437,153,359]" pageId="2" pageNumber="414">
<emphasis id="6382EAC2FF99FFD14BA1FF44502E6985" box="[199,305,153,179]" italics="true" pageId="2" pageNumber="414">Abdomen</emphasis>
(
<figureCitation id="C9CD2A55FF99FFD14A26FF4450926985" box="[320,397,153,179]" captionStart="FIGURE 1" captionStartId="2.[151,250,1062,1087]" captionTargetBox="[156,1429,375,1030]" captionTargetId="figure@2.[151,1436,371,1040]" captionTargetPageId="2" captionText="FIGURE 1. Philogenia gaiae sp. nov., holotype ♂; (1a) habitus; (1b) head; (1c) wings." figureDoi="http://doi.org/10.5281/zenodo.3479025" httpUri="https://zenodo.org/record/3479025/files/figure.png" pageId="2" pageNumber="414">Fig. 1a</figureCitation>
). S1 and S2 dark brown, S1 with a pale rounded spot laterally; S2 with a thin slightly oblique pale stripe laterally; S3 with basal ¾ brown becoming darker apically; S4S5 with basal half brown becoming darker posteriorly; S6S8 black; S910 black with dorsal white pruinosity.
</paragraph>
<paragraph id="514936D0FF99FFD14BA1FED850556851" blockId="2.[151,1437,153,359]" pageId="2" pageNumber="414">
<emphasis id="6382EAC2FF99FFD14BA1FED8507C6829" box="[199,355,261,287]" italics="true" pageId="2" pageNumber="414">Genital ligula</emphasis>
(
<figureCitation id="C9CD2A55FF99FFD14A14FED850A16816" box="[370,446,261,288]" captionStart="FIGURE 2" captionStartId="2.[151,250,1979,2004]" captionTargetBox="[162,1425,1101,1960]" captionTargetId="figure@2.[162,1425,1101,1960]" captionTargetPageId="2" captionText="FIGURE 2. Philogenia gaiae sp. nov., paratype ♂ ACR00579; (2a) lateral view of genital ligula; (2b) detail of genital ligula distal flagella; (2c) detail of genital ligula flagellum apex; (2d) detail of lateral first segment of genital ligula." figureDoi="http://doi.org/10.5281/zenodo.3479027" httpUri="https://zenodo.org/record/3479027/files/figure.png" pageId="2" pageNumber="414">Figs. 2</figureCitation>
ad). In lateral view with paired long flagella on the distal segment with a coiled confor- mity, apex of each flagellum covered with numerous spines. First segment covered with small bristles and one row of several hairs.
</paragraph>
<caption id="05896658FF99FFD14BF1FBFB550E6D09" ID-DOI="http://doi.org/10.5281/zenodo.3479025" ID-Zenodo-Dep="3479025" box="[151,1041,1062,1087]" httpUri="https://zenodo.org/record/3479025/files/figure.png" pageId="2" pageNumber="414" startId="2.[151,250,1062,1087]" targetBox="[156,1429,375,1030]" targetPageId="2">
<paragraph id="514936D0FF99FFD14BF1FBFB550E6D09" blockId="2.[151,1041,1062,1087]" box="[151,1041,1062,1087]" pageId="2" pageNumber="414">
<emphasis id="6382EAC2FF99FFD14BF1FBFB500C6D09" bold="true" box="[151,275,1062,1087]" pageId="2" pageNumber="414">FIGURE 1.</emphasis>
<taxonomicName id="96F64D53FF99FFD14A7FFBFB50D86D08" authority="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera, 2019" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[281,455,1062,1086]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="414" phylum="Arthropoda" rank="species" species="gaiae" status="sp. nov.">
<emphasis id="6382EAC2FF99FFD14A7FFBFB50D86D08" box="[281,455,1062,1086]" italics="true" pageId="2" pageNumber="414">Philogenia gaiae</emphasis>
</taxonomicName>
<emphasis id="6382EAC2FF99FFD14AABFBFB53016D08" bold="true" box="[461,542,1062,1086]" pageId="2" pageNumber="414">
<taxonomicNameLabel id="78B157B9FF99FFD14AABFBFB53016D08" box="[461,542,1062,1086]" pageId="2" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂; (1a) habitus; (1b) head; (1c) wings.
</paragraph>
</caption>
<caption id="05896658FF99FFD14BF1F86655F66ECE" ID-DOI="http://doi.org/10.5281/zenodo.3479027" ID-Zenodo-Dep="3479027" httpUri="https://zenodo.org/record/3479027/files/figure.png" pageId="2" pageNumber="414" startId="2.[151,250,1979,2004]" targetBox="[162,1425,1101,1960]" targetPageId="2">
<paragraph id="514936D0FF99FFD14BF1F86655F66ECE" blockId="2.[151,1437,1979,2040]" pageId="2" pageNumber="414">
<emphasis id="6382EAC2FF99FFD14BF1F866500B6EE2" bold="true" box="[151,276,1979,2004]" pageId="2" pageNumber="414">FIGURE 2.</emphasis>
<taxonomicName id="96F64D53FF99FFD14A7DF86150D56EE2" authority="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera, 2019" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[283,458,1980,2004]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="414" phylum="Arthropoda" rank="species" species="gaiae" status="sp. nov.">
<emphasis id="6382EAC2FF99FFD14A7DF86150D56EE2" box="[283,458,1980,2004]" italics="true" pageId="2" pageNumber="414">Philogenia gaiae</emphasis>
</taxonomicName>
<emphasis id="6382EAC2FF99FFD14AB7F861533C6EE2" bold="true" box="[465,547,1980,2004]" pageId="2" pageNumber="414">
<taxonomicNameLabel id="78B157B9FF99FFD14AB7F861533C6EE2" box="[465,547,1980,2004]" pageId="2" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype ♂ ACR00579; (2a) lateral view of genital ligula; (2b) detail of genital ligula distal flagella; (2c) detail of genital ligula flagellum apex; (2d) detail of lateral first segment of genital ligula.
</paragraph>
</caption>
<paragraph id="514936D0FF98FFD04BA1FF4454636875" blockId="3.[151,1436,153,323]" pageId="3" pageNumber="415">
<emphasis id="6382EAC2FF98FFD04BA1FF4450966985" box="[199,393,153,179]" italics="true" pageId="3" pageNumber="415">Anal appendages</emphasis>
(
<figureCitation id="C9CD2A55FF98FFD04AF1FF4450FD6985" box="[407,482,153,179]" captionStart="FIGURE 3" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[186,1400,352,1861]" captionTargetId="figure@3.[174,1413,337,1882]" captionTargetPageId="3" captionText="FIGURE 3. Philogenia gaiae sp. nov., paratype ♂ from Ecuador, Tiputini at RWG; (3a) appendages in lateral view; (3b) detail of right paraproct in lateral view; (3c) appendages in dorsal view. Philogenia minteri, ♂ from Ecuador, Limoncocha at RWG; (3d) appendages in lateral view; (3e) detail of right paraproct in lateral view; (3f) appendages in dorsal view. (Photos 3a,c, d, f by RWG)" figureDoi="http://doi.org/10.5281/zenodo.3479029" httpUri="https://zenodo.org/record/3479029/files/figure.png" pageId="3" pageNumber="415">Figs. 3</figureCitation>
ac). Cerci in lateral view extending ventrally at approximately its basal half, extend- ing as long as width of appendage; its dorsal portion covered with a compliment of strong teeth gradually growing in size basally to broad apical portion; in dorsal view, cerci curved medially and of uniform width but expanding at apex. In lateral view, basal portion of paraprocts with dense tuft of hairs; apical portion roundly curved mediodor- sally terminating in a club-like structure; in dorsal view, paraprocts terminating as an acute arrow-like structure.
</paragraph>
<caption id="05896658FF98FFD04BF1F8B351E66EC4" ID-DOI="http://doi.org/10.5281/zenodo.3479029" ID-Zenodo-Dep="3479029" httpUri="https://zenodo.org/record/3479029/files/figure.png" pageId="3" pageNumber="415" startId="3.[151,250,1902,1927]" targetBox="[186,1400,352,1861]" targetPageId="3">
<paragraph id="514936D0FF98FFD04BF1F8B351E66EC4" blockId="3.[151,1437,1902,2035]" pageId="3" pageNumber="415">
<emphasis id="6382EAC2FF98FFD04BF1F8B3500C6EB1" bold="true" box="[151,275,1902,1927]" pageId="3" pageNumber="415">FIGURE 3.</emphasis>
<taxonomicName id="96F64D53FF98FFD04A7FF8B350D96EB0" authority="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera, 2019" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[281,454,1902,1926]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="415" phylum="Arthropoda" rank="species" species="gaiae" status="sp. nov.">
<emphasis id="6382EAC2FF98FFD04A7FF8B350D96EB0" box="[281,454,1902,1926]" italics="true" pageId="3" pageNumber="415">Philogenia gaiae</emphasis>
</taxonomicName>
<emphasis id="6382EAC2FF98FFD04AAAF8B3533C6EB0" bold="true" box="[460,547,1902,1926]" pageId="3" pageNumber="415">
<taxonomicNameLabel id="78B157B9FF98FFD04AAAF8B353016EB0" box="[460,542,1902,1926]" pageId="3" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
paratype ♂ from Ecuador, Tiputini at RWG; (3a) appendages in lateral view; (3b) detail of right paraproct in lateral view; (3c) appendages in dorsal view.
<taxonomicName id="96F64D53FF98FFD0485AF84F52E46E9C" authorityName="Dunkle" authorityYear="1986" box="[828,1019,1938,1962]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="415" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF98FFD0485AF84F52E46E9C" box="[828,1019,1938,1962]" italics="true" pageId="3" pageNumber="415">Philogenia minteri</emphasis>
</taxonomicName>
, ♂ from Ecuador, Limoncocha at RWG; (3d) appendages in lateral view; (3e) detail of right paraproct in lateral view; (3f) appendages in dorsal view. (Photos 3a,c, d, f by RWG)
</paragraph>
</caption>
<paragraph id="514936D0FF9FFFD74BA1FF4452DC6985" blockId="4.[151,1436,152,323]" box="[199,963,152,179]" pageId="4" pageNumber="416">Measurements. FW = 33.8; HW = 34.3; abdomen = 41; total = 52.3.</paragraph>
<paragraph id="514936D0FF9FFFD74BA1FF61542E6875" blockId="4.[151,1436,152,323]" pageId="4" pageNumber="416">
<emphasis id="6382EAC2FF9FFFD74BA1FF6150FA69E1" bold="true" box="[199,485,188,215]" pageId="4" pageNumber="416">
Variations in
<typeStatus id="8E4D8872FF9FFFD74A0EFF6050FF69E1" box="[360,480,189,215]" pageId="4" pageNumber="416" type="paratype">paratypes</typeStatus>
.
</emphasis>
<typeStatus id="8E4D8872FF9FFFD74A8AFF60535169E1" box="[492,590,189,215]" pageId="4" pageNumber="416" type="paratype">Paratype</typeStatus>
males do not vary in cerci morphology. Slight differences were observed in the amount of abdominal pruinosity, probably related to the age and/or postmortem differences. Male
<typeStatus id="8E4D8872FF9FFFD74F98FF3C547669CD" box="[1278,1385,225,251]" pageId="4" pageNumber="416" type="paratype">paratypes</typeStatus>
var- ied in size as follows: FW = 28.334, HW = 29.235, abdomen = 3641.2, total = 4552. Venation among
<typeStatus id="8E4D8872FF9FFFD74E57FED854836829" box="[1329,1436,261,287]" pageId="4" pageNumber="416" type="paratype">paratypes</typeStatus>
varied as follows: Pt surmounting 45 cells in FW and 56 cells in HW; Px in FW 2428, in HW 2226.
</paragraph>
<caption id="05896658FF9FFFD74BF1FA5153F86CDB" ID-DOI="http://doi.org/10.5281/zenodo.3479031" ID-Zenodo-Dep="3479031" httpUri="https://zenodo.org/record/3479031/files/figure.png" pageId="4" pageNumber="416" startId="4.[151,250,1420,1445]" targetBox="[164,1423,344,1389]" targetPageId="4">
<paragraph id="514936D0FF9FFFD74BF1FA5153F86CDB" blockId="4.[151,1436,1420,1517]" pageId="4" pageNumber="416">
<emphasis id="6382EAC2FF9FFFD74BF1FA51500D6C93" bold="true" box="[151,274,1420,1445]" pageId="4" pageNumber="416">FIGURE 4.</emphasis>
<taxonomicName id="96F64D53FF9FFFD74A7EFA5050F06C93" authorityName="Selys" authorityYear="1886" box="[280,495,1421,1445]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="416" phylum="Arthropoda" rank="species" species="raphaella">
<emphasis id="6382EAC2FF9FFFD74A7EFA5050F06C93" box="[280,495,1421,1445]" italics="true" pageId="4" pageNumber="416">Philogenia raphaella</emphasis>
</taxonomicName>
, holotype ♂ from Colombia, Bogota at BMNH; (4a) appendages in lateral view; (4b) append- ages in dorsal view.
<taxonomicName id="96F64D53FF9FFFD74A06FA6C53346CFF" authorityName="Dunkle" authorityYear="1986" box="[352,555,1457,1481]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="416" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9FFFD74A06FA6C53346CFF" box="[352,555,1457,1481]" italics="true" pageId="4" pageNumber="416">Philogenia macuma</emphasis>
</taxonomicName>
, ♂ from Ecuador, Jatun Yacu at RWG; (4c) appendages in lateral view; (4d) appendages in dorsal view. (Photos 4a, b by Ben Price; 4c, d by RWG)
</paragraph>
</caption>
<paragraph id="514936D0FF9FFFD74BA1F9C653BE6F6C" blockId="4.[151,1436,1563,2021]" pageId="4" pageNumber="416">
<emphasis id="6382EAC2FF9FFFD74BA1F9C6502C6F00" bold="true" box="[199,307,1563,1590]" pageId="4" pageNumber="416">
<typeStatus id="8E4D8872FF9FFFD74BA1F9C650316F00" box="[199,302,1563,1590]" pageId="4" pageNumber="416" type="allotype">Allotype</typeStatus>
.
</emphasis>
<emphasis id="6382EAC2FF9FFFD74A5CF9C650686F03" box="[314,375,1563,1589]" italics="true" pageId="4" pageNumber="416">Head</emphasis>
(
<figureCitation id="C9CD2A55FF9FFFD74AE0F9C650CC6F00" box="[390,467,1563,1590]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="4" pageNumber="416">Figs. 5</figureCitation>
ad). Uniformly dark brown except for base of the antennae light brown; genae with a greenish tinge; labrum with greenish contours.
</paragraph>
<paragraph id="514936D0FF9FFFD74BA1F9BE50DD6E3B" blockId="4.[151,1436,1563,2021]" pageId="4" pageNumber="416">
<emphasis id="6382EAC2FF9FFFD74BA1F9BE50076F4B" box="[199,280,1635,1661]" italics="true" pageId="4" pageNumber="416">Thorax</emphasis>
(
<figureCitation id="C9CD2A55FF9FFFD74A41F9BE506D6F48" box="[295,370,1635,1662]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="4" pageNumber="416">Figs. 5</figureCitation>
ab, de). Prothorax brown, similar to the
<typeStatus id="8E4D8872FF9FFFD7482BF9BE52B06F4B" box="[845,943,1635,1661]" pageId="4" pageNumber="416" type="holotype">holotype</typeStatus>
except with paler coloration; in dorsal view, posterior lobe rounded with a lateral pointed process; in lateral view, posterior lobe curved upwards. Intersternite (
<figureCitation id="C9CD2A55FF9FFFD74BC6F97651EF6FF0" box="[160,240,1707,1734]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="4" pageNumber="416">Fig. 5e</figureCitation>
) with an irregular contour on anterior portion; dorsal portion nearly quadrate; a notable cleft medially at level of setifer; setifer ovaloid, narrow, tuft of hairs decreasing in size ventrally. Coxae brown, internal surface of legs pale, armature brown.
</paragraph>
<paragraph id="514936D0FF9FFFD74BA1F8C5533A6E60" blockId="4.[151,1436,1563,2021]" pageId="4" pageNumber="416">
<emphasis id="6382EAC2FF9FFFD74BA1F8C550146E07" box="[199,267,1816,1841]" italics="true" pageId="4" pageNumber="416">Wings</emphasis>
(
<figureCitation id="C9CD2A55FF9FFFD74A7DF8CA50726E04" box="[283,365,1815,1842]" captionStart="FIGURE 6" captionStartId="6.[151,250,1952,1977]" captionTargetBox="[162,1429,1126,1864]" captionTargetId="figure@6.[151,1436,1114,1878]" captionTargetPageId="6" captionText="FIGURE 6. Philogenia gaiae sp. nov., allotype ♀; (6a) lateral view of S8S10; (6b) wings. Philogenia macuma, ♀ from Ecua- dor, Jatún Sacha at ECOEVO; (6c) lateral view of S8S10; (6d) wings." figureDoi="http://doi.org/10.5281/zenodo.3479035" httpUri="https://zenodo.org/record/3479035/files/figure.png" pageId="4" pageNumber="416">Fig. 6b</figureCitation>
). Hyaline (HW with a small rip on apical C). Pt dark brown surmounting four cells in both FW and HW. Px =
<quantity id="960E9B35FF9FFFD74A5AF8E1506A6E63" box="[316,373,1851,1878]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="4" pageNumber="416" unit="in" value="27.0">27 in</quantity>
FW,
<quantity id="960E9B35FF9FFFD74AD7F8E150F56E63" box="[433,490,1851,1878]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="4" pageNumber="416" unit="in" value="24.0">24 in</quantity>
HW.
</paragraph>
<paragraph id="514936D0FF9FFFD74BA1F882504A6ED3" blockId="4.[151,1436,1563,2021]" pageId="4" pageNumber="416">
<emphasis id="6382EAC2FF9FFFD74BA1F882502E6E4F" box="[199,305,1887,1913]" italics="true" pageId="4" pageNumber="416">Abdomen</emphasis>
(
<figureCitation id="C9CD2A55FF9FFFD74A26F88250836E4C" box="[320,412,1887,1914]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="4" pageNumber="416">Figs. 5a</figureCitation>
,
<figureCitation id="C9CD2A55FF9FFFD74ACEF88250DC6E4C" box="[424,451,1887,1914]" captionStart="FIGURE 6" captionStartId="6.[151,250,1952,1977]" captionTargetBox="[162,1429,1126,1864]" captionTargetId="figure@6.[151,1436,1114,1878]" captionTargetPageId="6" captionText="FIGURE 6. Philogenia gaiae sp. nov., allotype ♀; (6a) lateral view of S8S10; (6b) wings. Philogenia macuma, ♀ from Ecua- dor, Jatún Sacha at ECOEVO; (6c) lateral view of S8S10; (6d) wings." figureDoi="http://doi.org/10.5281/zenodo.3479035" httpUri="https://zenodo.org/record/3479035/files/figure.png" pageId="4" pageNumber="416">6a</figureCitation>
). Similar to the
<typeStatus id="8E4D8872FF9FFFD7491FF88253C16E4F" box="[633,734,1887,1913]" pageId="4" pageNumber="416" type="holotype">holotype</typeStatus>
, only with paler coloration and lacking pruinosity on the tip of abdomen. Genital valves dark brown, surpassing S10 for a distance slightly shorter than segment, with a pronounced process above base of styli (both missing). Ovipositor mostly red, serrulate at apical 1/3. Cerci black, conical with dense pruinosity.
</paragraph>
<caption id="05896658FF9EFFD64BF1F895500C6EC7" ID-DOI="http://doi.org/10.5281/zenodo.3479033" ID-Zenodo-Dep="3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="5" pageNumber="417" startId="5.[151,250,1864,1889]" targetBox="[157,1396,180,1833]" targetPageId="5">
<paragraph id="514936D0FF9EFFD64BF1F895500C6EC7" blockId="5.[151,1437,1864,2033]" pageId="5" pageNumber="417">
<emphasis id="6382EAC2FF9EFFD64BF1F895500C6E57" bold="true" box="[151,275,1864,1889]" pageId="5" pageNumber="417">FIGURE 5.</emphasis>
<taxonomicName id="96F64D53FF9EFFD64A7FF89550D86E56" authority="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera, 2019" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[281,455,1864,1888]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="417" phylum="Arthropoda" rank="species" species="gaiae" status="sp. nov.">
<emphasis id="6382EAC2FF9EFFD64A7FF89550D86E56" box="[281,455,1864,1888]" italics="true" pageId="5" pageNumber="417">Philogenia gaiae</emphasis>
</taxonomicName>
<emphasis id="6382EAC2FF9EFFD64AABF89453016E57" bold="true" box="[461,542,1865,1889]" pageId="5" pageNumber="417">
<taxonomicNameLabel id="78B157B9FF9EFFD64AABF89453016E57" box="[461,542,1865,1889]" pageId="5" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite.
<taxonomicName id="96F64D53FF9EFFD64BF1F84D507A6E9E" authorityName="Dunkle" authorityYear="1986" box="[151,357,1936,1960]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="417" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9EFFD64BF1F84D507A6E9E" box="[151,357,1936,1960]" italics="true" pageId="5" pageNumber="417">Philogenia macuma</emphasis>
</taxonomicName>
, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus.
</paragraph>
</caption>
<paragraph id="514936D0FF9DFFD54BA1FF4450C369E1" blockId="6.[151,1437,152,1079]" pageId="6" pageNumber="418">
Measurements. FW = 32.6; HW = 33.4; abdomen = 36.5; total = 47. Female
<typeStatus id="8E4D8872FF9DFFD54F44FF4455986985" box="[1058,1159,153,179]" pageId="6" pageNumber="418" type="paratype">paratype</typeStatus>
: FW = 30.2; HW = 30.7; abdomen = 33.1; total = 41.8
</paragraph>
<paragraph id="514936D0FF9DFFD54BA1FF3D54656B09" blockId="6.[151,1437,152,1079]" pageId="6" pageNumber="418">
<emphasis id="6382EAC2FF9DFFD54BA1FF3D50D069CD" bold="true" box="[199,463,224,251]" pageId="6" pageNumber="418">Differential diagnosis.</emphasis>
Male
<taxonomicName id="96F64D53FF9DFFD5497EFF3F536C69CD" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[536,627,226,251]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD5497EFF3F536C69CD" box="[536,627,226,251]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
and
<taxonomicName id="96F64D53FF9DFFD549CAFF3F520569CD" authorityName="Dunkle" authorityYear="1986" box="[684,794,226,251]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD549CAFF3F520569CD" box="[684,794,226,251]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
can be easily confounded in the field due to similar col- oration, as is the case for many members of the genus. However,
<taxonomicName id="96F64D53FF9DFFD54813FEDB52D06829" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[885,975,262,287]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54813FEDB52D06829" box="[885,975,262,287]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="96F64D53FF9DFFD54F02FEDB55CE6829" authorityName="Dunkle" authorityYear="1986" box="[1124,1233,262,287]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD54F02FEDB55CE6829" box="[1124,1233,262,287]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
by comparison of the paraproct in lateral and dorsal views (
<figureCitation id="C9CD2A55FF9DFFD54917FEF453A06872" box="[625,703,297,324]" captionStart="FIGURE 3" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[186,1400,352,1861]" captionTargetId="figure@3.[174,1413,337,1882]" captionTargetPageId="3" captionText="FIGURE 3. Philogenia gaiae sp. nov., paratype ♂ from Ecuador, Tiputini at RWG; (3a) appendages in lateral view; (3b) detail of right paraproct in lateral view; (3c) appendages in dorsal view. Philogenia minteri, ♂ from Ecuador, Limoncocha at RWG; (3d) appendages in lateral view; (3e) detail of right paraproct in lateral view; (3f) appendages in dorsal view. (Photos 3a,c, d, f by RWG)" figureDoi="http://doi.org/10.5281/zenodo.3479029" httpUri="https://zenodo.org/record/3479029/files/figure.png" pageId="6" pageNumber="418">Figs. 3</figureCitation>
af).
<bibRefCitation id="35674B21FF9DFFD54998FEF452FD6875" author="Bick, G. H. &amp; Bick, J. C." box="[766,994,297,323]" pageId="6" pageNumber="418" pagination="9 - 32" refId="ref3996" refString="Bick, G. H. &amp; Bick, J. C. (1988) A review of males of the genus Philogenia, with descriptions of five new species from South America (Zygoptera: Megapodagrionidae). Odonatologica, 17 (1), 9 - 32." type="journal article" year="1988">Bick &amp; Bick (1988)</bibRefCitation>
defined the paraprocts of
<taxonomicName id="96F64D53FF9DFFD54E68FEF754636875" authorityName="Dunkle" authorityYear="1986" box="[1294,1404,298,323]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD54E68FEF754636875" box="[1294,1404,298,323]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
as “broad and deformed looking”, a unique trait in the
<taxonomicName id="96F64D53FF9DFFD549B4FE9052026851" authorityName="Hagen" authorityYear="1869" box="[722,797,333,359]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="helena">
<emphasis id="6382EAC2FF9DFFD549B4FE9052026851" box="[722,797,333,359]" italics="true" pageId="6" pageNumber="418">helena</emphasis>
</taxonomicName>
group. Species of this group usually have paraprocts with an acute apex (except for
<taxonomicName id="96F64D53FF9DFFD54ADCFEAF535268BD" authorityName="Dunkle" authorityYear="1990" box="[442,589,370,395]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="compressa">
<emphasis id="6382EAC2FF9DFFD54ADCFEAF535268BD" box="[442,589,370,395]" italics="true" pageId="6" pageNumber="418">P. compressa</emphasis>
</taxonomicName>
,
<taxonomicName id="96F64D53FF9DFFD5493AFEAF53B268BD" authorityName="Dunkle" authorityYear="1986" box="[604,685,370,395]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD5493AFEAF53B268BD" box="[604,685,370,395]" italics="true" pageId="6" pageNumber="418">minteri</emphasis>
</taxonomicName>
and
<taxonomicName id="96F64D53FF9DFFD54982FEAF523D68BD" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[740,802,370,395]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54982FEAF523D68BD" box="[740,802,370,395]" italics="true" pageId="6" pageNumber="418">gaiae</emphasis>
</taxonomicName>
), which canbe straight, as in
<taxonomicName id="96F64D53FF9DFFD54F08FEAF55E868BD" authorityName="Selys" authorityYear="1886" box="[1134,1271,369,395]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="raphaella">
<emphasis id="6382EAC2FF9DFFD54F08FEAF55E868BD" box="[1134,1271,369,395]" italics="true" pageId="6" pageNumber="418">P. raphaella</emphasis>
</taxonomicName>
(
<figureCitation id="C9CD2A55FF9DFFD54E61FEAC545568BA" box="[1287,1354,369,396]" captionStart="FIGURE 4" captionStartId="4.[151,250,1420,1445]" captionTargetBox="[164,1423,344,1389]" captionTargetId="figure@4.[151,1435,337,1401]" captionTargetPageId="4" captionText="FIGURE 4. Philogenia raphaella, holotype ♂ from Colombia, Bogota at BMNH; (4a) appendages in lateral view; (4b) append- ages in dorsal view. Philogenia macuma, ♂ from Ecuador, Jatun Yacu at RWG; (4c) appendages in lateral view; (4d) appendages in dorsal view. (Photos 4a, b by Ben Price; 4c, d by RWG)" figureDoi="http://doi.org/10.5281/zenodo.3479031" httpUri="https://zenodo.org/record/3479031/files/figure.png" pageId="6" pageNumber="418">Fig. 4</figureCitation>
ab) or curved dorsally at apex, as in
<taxonomicName id="96F64D53FF9DFFD54ABBFE4B53496899" authorityName="Dunkle" authorityYear="1986" box="[477,598,406,431]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9DFFD54ABBFE4B53496899" box="[477,598,406,431]" italics="true" pageId="6" pageNumber="418">P. macuma</emphasis>
</taxonomicName>
(
<figureCitation id="C9CD2A55FF9DFFD54905FE4853AF6886" box="[611,688,405,432]" captionStart="FIGURE 4" captionStartId="4.[151,250,1420,1445]" captionTargetBox="[164,1423,344,1389]" captionTargetId="figure@4.[151,1435,337,1401]" captionTargetPageId="4" captionText="FIGURE 4. Philogenia raphaella, holotype ♂ from Colombia, Bogota at BMNH; (4a) appendages in lateral view; (4b) append- ages in dorsal view. Philogenia macuma, ♂ from Ecuador, Jatun Yacu at RWG; (4c) appendages in lateral view; (4d) appendages in dorsal view. (Photos 4a, b by Ben Price; 4c, d by RWG)" figureDoi="http://doi.org/10.5281/zenodo.3479031" httpUri="https://zenodo.org/record/3479031/files/figure.png" pageId="6" pageNumber="418">Figs. 4</figureCitation>
cd). Apex of paraprocts of
<taxonomicName id="96F64D53FF9DFFD54882FE4B55226899" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[996,1085,406,431]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54882FE4B55226899" box="[996,1085,406,431]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
is broad and is similar to that of
<taxonomicName id="96F64D53FF9DFFD54BF1FE67501C68E5" authorityName="Dunkle" authorityYear="1986" box="[151,259,442,467]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD54BF1FE67501C68E5" box="[151,259,442,467]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
, however, in
<taxonomicName id="96F64D53FF9DFFD54AFCFE6750EC68E5" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[410,499,442,467]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54AFCFE6750EC68E5" box="[410,499,442,467]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
the paraprocts are roundly curved mediodorsally ending in a club-like structure and in
<taxonomicName id="96F64D53FF9DFFD54BD2FE03503E68C1" authorityName="Dunkle" authorityYear="1986" box="[180,289,478,503]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD54BD2FE03503E68C1" box="[180,289,478,503]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
the paraprocts are truncated and “deformed-looking” (
<emphasis id="6382EAC2FF9DFFD548EAFE0352D568C1" box="[908,970,478,503]" italics="true" pageId="6" pageNumber="418">sensu</emphasis>
<bibRefCitation id="35674B21FF9DFFD548B7FE00557868CE" author="Dunkle, S. W." box="[977,1127,477,504]" pageId="6" pageNumber="418" pagination="43 - 50" refId="ref4178" refString="Dunkle, S. W. (1986) Four new species of Philogenia damselflies from Ecuador and Colombia (Zygoptera: Megapodagrioni- dae). Odonatologica, 15 (1), 43 - 50." type="journal article" year="1986">Dunkle 1986</bibRefCitation>
). In dorsal view (
<figureCitation id="C9CD2A55FF9DFFD54E4AFE00548A68CE" box="[1324,1429,477,504]" captionStart="FIGURE 3" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[186,1400,352,1861]" captionTargetId="figure@3.[174,1413,337,1882]" captionTargetPageId="3" captionText="FIGURE 3. Philogenia gaiae sp. nov., paratype ♂ from Ecuador, Tiputini at RWG; (3a) appendages in lateral view; (3b) detail of right paraproct in lateral view; (3c) appendages in dorsal view. Philogenia minteri, ♂ from Ecuador, Limoncocha at RWG; (3d) appendages in lateral view; (3e) detail of right paraproct in lateral view; (3f) appendages in dorsal view. (Photos 3a,c, d, f by RWG)" figureDoi="http://doi.org/10.5281/zenodo.3479029" httpUri="https://zenodo.org/record/3479029/files/figure.png" pageId="6" pageNumber="418">Fig. 3c, f</figureCitation>
) paraprocts of
<taxonomicName id="96F64D53FF9DFFD54A48FDDF50986B2D" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[302,391,514,539]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54A48FDDF50986B2D" box="[302,391,514,539]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
terminate in an acute arrow-like structure, whereas in
<taxonomicName id="96F64D53FF9DFFD548B8FDDF55566B2D" authorityName="Dunkle" authorityYear="1986" box="[990,1097,514,539]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD548B8FDDF55566B2D" box="[990,1097,514,539]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
the paraprocts are broader and rounded apically. The cerci, however, appear to be the same in both species, although slightly larger in
<taxonomicName id="96F64D53FF9DFFD54E7FFDFB546D6B09" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[1305,1394,550,575]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54E7FFDFB546D6B09" box="[1305,1394,550,575]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="514936D0FF9DFFD54BA1FD9455CE6D01" blockId="6.[151,1437,152,1079]" pageId="6" pageNumber="418">
The female of
<taxonomicName id="96F64D53FF9DFFD54A15FD9750D06B55" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[371,463,586,611]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54A15FD9750D06B55" box="[371,463,586,611]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
is similar to the female of
<taxonomicName id="96F64D53FF9DFFD5486BFD9752956B55" authorityName="Dunkle" authorityYear="1986" box="[781,906,586,611]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9DFFD5486BFD9752956B55" box="[781,906,586,611]" italics="true" pageId="6" pageNumber="418">P. macuma</emphasis>
</taxonomicName>
(
<figureCitation id="C9CD2A55FF9DFFD548FAFD9452F56B55" box="[924,1002,584,611]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="6" pageNumber="418">Figs. 5</figureCitation>
aj;
<figureCitation id="C9CD2A55FF9DFFD54F47FD9455706B52" box="[1057,1135,585,612]" captionStart="FIGURE 6" captionStartId="6.[151,250,1952,1977]" captionTargetBox="[162,1429,1126,1864]" captionTargetId="figure@6.[151,1436,1114,1878]" captionTargetPageId="6" captionText="FIGURE 6. Philogenia gaiae sp. nov., allotype ♀; (6a) lateral view of S8S10; (6b) wings. Philogenia macuma, ♀ from Ecua- dor, Jatún Sacha at ECOEVO; (6c) lateral view of S8S10; (6d) wings." figureDoi="http://doi.org/10.5281/zenodo.3479035" httpUri="https://zenodo.org/record/3479035/files/figure.png" pageId="6" pageNumber="418">Figs. 6</figureCitation>
ad). Differences between the two occur in the intersternites, ovipositor valve shape and dorsal view of the posterior lobe of prothorax. In
<taxonomicName id="96F64D53FF9DFFD54EEEFDB351E96B9D" authorityName="Dunkle" authorityYear="1986" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9DFFD54EEEFDB351E96B9D" italics="true" pageId="6" pageNumber="418">P. macuma</emphasis>
</taxonomicName>
, the anterior portion of intersternite is straight (
<figureCitation id="C9CD2A55FF9DFFD5486BFD4C52446B9D" box="[781,859,656,683]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="6" pageNumber="418">Fig. 5f</figureCitation>
), differing from the anterior irregular configuration of
<taxonomicName id="96F64D53FF9DFFD54BDEFD6B500C6BF9" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[184,275,694,719]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54BDEFD6B500C6BF9" box="[184,275,694,719]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
(
<figureCitation id="C9CD2A55FF9DFFD54A43FD6850666BF9" box="[293,377,692,719]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="6" pageNumber="418">Fig. 5e</figureCitation>
). Furthermore, the medial cleft is more pronounced and the setifer is broader in
<taxonomicName id="96F64D53FF9DFFD54E7FFD6B548A6BF9" authorityName="Dunkle" authorityYear="1986" box="[1305,1429,694,719]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9DFFD54E7FFD6B548A6BF9" box="[1305,1429,694,719]" italics="true" pageId="6" pageNumber="418">P. macuma</emphasis>
</taxonomicName>
. Use of the sclerotized intersternite as a diagnostic character should be used with caution since it can be invisible in young females, broken in mature females or vary individually. In dorsal view, the posterior lobe of the prothorax of
<taxonomicName id="96F64D53FF9DFFD54BF1FCFF51EE6A0D" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[151,241,802,827]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54BF1FCFF51EE6A0D" box="[151,241,802,827]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
is almost identical to the female of
<taxonomicName id="96F64D53FF9DFFD549E4FCFF53E36A0D" authorityName="Dunkle" authorityYear="1986" box="[642,764,802,827]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9DFFD549E4FCFF53E36A0D" box="[642,764,802,827]" italics="true" pageId="6" pageNumber="418">P. macuma</emphasis>
</taxonomicName>
, except in
<taxonomicName id="96F64D53FF9DFFD54811FCFF52CF6A0D" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[887,976,802,827]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54811FCFF52CF6A0D" box="[887,976,802,827]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
the lateral projections are slightly longer (
<figureCitation id="C9CD2A55FF9DFFD54BF9FC9850086A69" box="[159,279,836,863]" captionStart="FIGURE 5" captionStartId="5.[151,250,1864,1889]" captionTargetBox="[157,1396,180,1833]" captionTargetId="figure@5.[151,1405,166,1844]" captionTargetPageId="5" captionText="FIGURE 5. Philogenia gaiae sp. nov., allotype ♀; (5a) habitus; (5b) lateral view of head and prothorax; (5c) head; (5d) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5e) intersternite. Philogenia macuma, ♀ from Ecuador, Jatún Sacha at ECOEVO; (5f) intersternite; (5g) head; (5h) dorsal view of head and pterothorax showing a detail of the lateral projection of the posterior lobe of prothorax; (5i) lateral view of head and prothorax; (5j) habitus." figureDoi="http://doi.org/10.5281/zenodo.3479033" httpUri="https://zenodo.org/record/3479033/files/figure.png" pageId="6" pageNumber="418">Figs. 5d, h</figureCitation>
for a comparison). The posterior lobe of females of both species are also similar to
<taxonomicName id="96F64D53FF9DFFD54FA6FC9B54326A69" authorityName="Dunkle" authorityYear="1986" box="[1216,1325,838,863]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD54FA6FC9B54326A69" box="[1216,1325,838,863]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
. We were unable to examine any female from this species, however, by the illustration present in
<bibRefCitation id="35674B21FF9DFFD54F3DFCB455E86AB5" author="Dunkle, S. W." box="[1115,1271,873,899]" pageId="6" pageNumber="418" pagination="43 - 50" refId="ref4178" refString="Dunkle, S. W. (1986) Four new species of Philogenia damselflies from Ecuador and Colombia (Zygoptera: Megapodagrioni- dae). Odonatologica, 15 (1), 43 - 50." type="journal article" year="1986">Dunkle (1986</bibRefCitation>
,
<figureCitation id="C9CD2A55FF9DFFD54E64FCB454746AB5" box="[1282,1387,873,899]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="3.[151,250,1902,1927]" captionStartId-1="4.[151,250,1420,1445]" captionTargetBox-0="[186,1400,352,1861]" captionTargetBox-1="[164,1423,344,1389]" captionTargetId-0="figure@3.[174,1413,337,1882]" captionTargetId-1="figure@4.[151,1435,337,1401]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 3. Philogenia gaiae sp. nov., paratype ♂ from Ecuador, Tiputini at RWG; (3a) appendages in lateral view; (3b) detail of right paraproct in lateral view; (3c) appendages in dorsal view. Philogenia minteri, ♂ from Ecuador, Limoncocha at RWG; (3d) appendages in lateral view; (3e) detail of right paraproct in lateral view; (3f) appendages in dorsal view. (Photos 3a,c, d, f by RWG)" captionText-1="FIGURE 4. Philogenia raphaella, holotype ♂ from Colombia, Bogota at BMNH; (4a) appendages in lateral view; (4b) append- ages in dorsal view. Philogenia macuma, ♂ from Ecuador, Jatun Yacu at RWG; (4c) appendages in lateral view; (4d) appendages in dorsal view. (Photos 4a, b by Ben Price; 4c, d by RWG)" figureDoi-0="http://doi.org/10.5281/zenodo.3479029" figureDoi-1="http://doi.org/10.5281/zenodo.3479031" httpUri-0="https://zenodo.org/record/3479029/files/figure.png" httpUri-1="https://zenodo.org/record/3479031/files/figure.png" pageId="6" pageNumber="418">Figs. 34</figureCitation>
) the female of
<taxonomicName id="96F64D53FF9DFFD54A6EFC53506B6A91" authorityName="Dunkle" authorityYear="1986" box="[264,372,910,935]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="6382EAC2FF9DFFD54A6EFC53506B6A91" box="[264,372,910,935]" italics="true" pageId="6" pageNumber="418">P. minteri</emphasis>
</taxonomicName>
also possess lateral projections on the posterior lobe. Additionally, the tips of the genital valves in
<taxonomicName id="96F64D53FF9DFFD54BD5FC6F50136AFD" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[179,268,946,971]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9DFFD54BD5FC6F50136AFD" box="[179,268,946,971]" italics="true" pageId="6" pageNumber="418">P. gaiae</emphasis>
</taxonomicName>
are almost straight, surpassing S10 for a distance slightly longer than the segment, whereas in
<taxonomicName id="96F64D53FF9DFFD54E45FC6F54836AFD" authorityName="Dunkle" authorityYear="1986" box="[1315,1436,946,971]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9DFFD54E45FC6F54836AFD" box="[1315,1436,946,971]" italics="true" pageId="6" pageNumber="418">P. macuma</emphasis>
</taxonomicName>
the tips of the genital valves form a concave angulation and the ovipositor surpasses S10 for a distance subequal to the segment length (
<figureCitation id="C9CD2A55FF9DFFD54A1DFC2450EC6D25" box="[379,499,1017,1043]" captionStart="FIGURE 6" captionStartId="6.[151,250,1952,1977]" captionTargetBox="[162,1429,1126,1864]" captionTargetId="figure@6.[151,1436,1114,1878]" captionTargetPageId="6" captionText="FIGURE 6. Philogenia gaiae sp. nov., allotype ♀; (6a) lateral view of S8S10; (6b) wings. Philogenia macuma, ♀ from Ecua- dor, Jatún Sacha at ECOEVO; (6c) lateral view of S8S10; (6d) wings." figureDoi="http://doi.org/10.5281/zenodo.3479035" httpUri="https://zenodo.org/record/3479035/files/figure.png" pageId="6" pageNumber="418">Figs. 6a, c</figureCitation>
). However, it is worth mentioning that ovipositor length may vary individually and given the paucity of females available for this study, this trait should be considered with caution.
</paragraph>
<caption id="05896658FF9DFFD54BF1F87D52756EEB" ID-DOI="http://doi.org/10.5281/zenodo.3479035" ID-Zenodo-Dep="3479035" httpUri="https://zenodo.org/record/3479035/files/figure.png" pageId="6" pageNumber="418" startId="6.[151,250,1952,1977]" targetBox="[162,1429,1126,1864]" targetPageId="6">
<paragraph id="514936D0FF9DFFD54BF1F87D52756EEB" blockId="6.[151,1436,1952,2013]" pageId="6" pageNumber="418">
<emphasis id="6382EAC2FF9DFFD54BF1F87D500D6E8F" bold="true" box="[151,274,1952,1977]" pageId="6" pageNumber="418">FIGURE 6.</emphasis>
<taxonomicName id="96F64D53FF9DFFD54A7EF87C50DA6E8E" authority="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera, 2019" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[280,453,1953,1977]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="gaiae" status="sp. nov.">
<emphasis id="6382EAC2FF9DFFD54A7EF87C50DA6E8E" box="[280,453,1953,1977]" italics="true" pageId="6" pageNumber="418">Philogenia gaiae</emphasis>
</taxonomicName>
<emphasis id="6382EAC2FF9DFFD54AADF87C53046E8F" bold="true" box="[459,539,1953,1977]" pageId="6" pageNumber="418">
<taxonomicNameLabel id="78B157B9FF9DFFD54AADF87C53046E8F" box="[459,539,1953,1977]" pageId="6" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype ♀; (6a) lateral view of S8S10; (6b) wings.
<taxonomicName id="96F64D53FF9DFFD54F5CF87C541A6E8E" authorityName="Dunkle" authorityYear="1986" box="[1082,1285,1953,1977]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="418" phylum="Arthropoda" rank="species" species="macuma">
<emphasis id="6382EAC2FF9DFFD54F5CF87C541A6E8E" box="[1082,1285,1953,1977]" italics="true" pageId="6" pageNumber="418">Philogenia macuma</emphasis>
</taxonomicName>
, ♀ from Ecua- dor, Jatún Sacha at ECOEVO; (6c) lateral view of S8S10; (6d) wings.
</paragraph>
</caption>
<paragraph id="514936D0FF9CFFD44BA1FF45508668BD" blockId="7.[151,1437,152,395]" pageId="7" pageNumber="419">
<emphasis id="6382EAC2FF9CFFD44BA1FF4550A06985" bold="true" box="[199,447,152,179]" pageId="7" pageNumber="419">Habitat and ecology.</emphasis>
As is typical for the genus,
<taxonomicName id="96F64D53FF9CFFD44865FF4752416985" authorityName="Vilela &amp; Guillermo-Ferreira &amp; Encalada &amp; Cordero-Rivera" authorityYear="2019" box="[771,862,154,179]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="419" phylum="Arthropoda" rank="species" species="gaiae">
<emphasis id="6382EAC2FF9CFFD44865FF4752416985" box="[771,862,154,179]" italics="true" pageId="7" pageNumber="419">P. gaiae</emphasis>
</taxonomicName>
where found along mostly shaded trails in the un- derstory of the forest (
<figureCitation id="C9CD2A55FF9CFFD44AFFFF6050C069E1" box="[409,479,189,215]" captionStart="FIGURE 7" captionStartId="7.[151,250,1285,1310]" captionTargetBox="[151,1436,409,1265]" captionTargetId="figure@7.[151,1436,409,1265]" captionTargetPageId="7" captionText="FIGURE 7. Philogenia gaiae sp. nov., holotype ♂ and allotype ♀ in copula at the Tiputini Biodiversity Station. Photography taken by ACR at 16:15, using only natural light, no flashes." figureDoi="http://doi.org/10.5281/zenodo.3479037" httpUri="https://zenodo.org/record/3479037/files/figure.png" pageId="7" pageNumber="419">Fig. 7</figureCitation>
). We never found individuals near the streams, but we noted that some forest spots were preferred over others. Both sexes fly very low (
<quantity id="960E9B35FF9CFFD44997FF3C527B69CD" box="[753,868,224,251]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="7" pageNumber="419" unit="cm" value="35.0" valueMax="50.0" valueMin="20.0">2050 cm</quantity>
from the ground vegetation) and fast, and usually the only visible part is the whitish pruinescence at the end of the abdomen. Rosser Garrison and Natalia von El- lenrieder (pers. comm.) also found specimens deep within the forest understory, usually one at a time. They were unable to follow individuals that were disturbed and, despite their best efforts, were only able capture about half of all specimens they saw
</paragraph>
</subSubSection>
</treatment>
</document>