743 lines
93 KiB
XML
743 lines
93 KiB
XML
<document id="D587C6DFB5B3AE213B3934D9BAD38692" ID-CLB-Dataset="53840" ID-DOI="10.11646/phytotaxa.601.3.3" ID-GBIF-Dataset="891014bc-6b64-4705-9fec-bd35af7b209c" ID-ISSN="1179-3163" ID-Zenodo-Dep="8141764" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1689234199257" checkinUser="plazi" docAuthor="Toolmal, Nopparut, Suddee, Somran, Culham, Alastair, Utteridge, Timothy M. A. & Schuiteman, André" docDate="2023" docId="D62787BA2071FFA92C9510F0FAB542F7" docLanguage="en" docName="phytotaxa.601.3.3.pdf" docOrigin="Phytotaxa 601 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.601.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Thrixspermum alboluteum Toolmal, Schuit. & Culham 2023, spec. nov." docType="treatment" docVersion="4" lastPageNumber="230" masterDocId="2A1EFFC22073FFAE2C1D1144FFAF463A" masterDocTitle="Taxonomic notes on Thrixspermum (Orchidaceae: Aeridinae), with five new species from Thailand" masterLastPageNumber="262" masterPageNumber="223" pageNumber="225" updateTime="1689362489109" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="165ABE3AD2473F4F54065C3BCDE4173B">
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<mods:title id="19D07909EB7353F0E683033338B1B594">Taxonomic notes on Thrixspermum (Orchidaceae: Aeridinae), with five new species from Thailand</mods:title>
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<mods:namePart id="47006B12B7F84DFCDC3EF7C4CA1D6BE0">Toolmal, Nopparut</mods:namePart>
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<mods:nameIdentifier id="637F9C14AEAF094F06C0318EAA0C563A" type="ORCID">0009-0001-4886-7477</mods:nameIdentifier>
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<mods:affiliation id="B8E1650F651F4AFBE8C03C6F450D5903">Royal Botanic Gardens, Kew, Richmond, TW 9 3 AE, United Kingdom & Herbarium, School of Biological Sciences, University of Reading, Whiteknights, Reading, Berkshire, RG 6 6 EX, United Kingdom & Thai Traditional Medicine Herbarium (TTM), 693 Bamrung Muang Rd., Pom Prap Sattru Phai, Bangkok 10100, Thailand & n. toolmal @ kew. org, n. toolmal @ pgr. reading. ac. uk; https: // orcid. org / 0009 - 0001 - 4886 - 7477</mods:affiliation>
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<mods:nameIdentifier id="8A41AF9AB14DBFCD056FF97451ACDBC8" type="email">n.toolmal@pgr.reading.ac.uk</mods:nameIdentifier>
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<mods:namePart id="72B10D738529CEFF6D44E7044CD91745">Suddee, Somran</mods:namePart>
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<mods:affiliation id="4CF97D16418D3B0FF6E44A2B91B1EC2C">Forest Herbarium (BKF), Department of National Parks, Wildlife and Plant Conservation, 61 Phahonyothin Road, Chatuchak, Bangkok 10900, Thailand & somrans @ hotmail. com; https: // orcid. org / 0000 - 0003 - 4031 - 9721</mods:affiliation>
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<mods:nameIdentifier id="67F72B97A392DA7CFA52C7CDD43C0B6E" type="email">somrans@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="967D93E7562D73AAAFEDAC2469F9EB35">Culham, Alastair</mods:namePart>
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<mods:affiliation id="1AFDCA0066FBE59A1F01D9B7A5C48869">Herbarium, School of Biological Sciences, University of Reading, Whiteknights, Reading, Berkshire, RG 6 6 EX, United Kingdom & a. culham @ reading. ac. uk; https: // orcid. org / 0000 - 0002 - 7440 - 0133</mods:affiliation>
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<mods:nameIdentifier id="0732B51F47AECAF12035CF5D446B372E" type="email">a.culham@reading.ac.uk</mods:nameIdentifier>
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<mods:namePart id="44673ED14D2ECDEC8F667278637F8EC1">Utteridge, Timothy M. A.</mods:namePart>
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<mods:affiliation id="A0A789860067BAAAE3E3FD27C7746CF9">Royal Botanic Gardens, Kew, Richmond, TW 9 3 AE, United Kingdom & t. utteridge @ kew. org; https: // orcid. org / 0000 - 0003 - 2823 - 0337</mods:affiliation>
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<mods:namePart id="5AF3D44DE7E75CED234FD738DE459EF0">Schuiteman, André</mods:namePart>
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<mods:nameIdentifier id="92631389521057FFE70EB73CEB6388A3" type="email">a.schuiteman@kew.org</mods:nameIdentifier>
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<mods:title id="05F8D18448818E62980F783D09347DFD">Phytotaxa</mods:title>
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<mods:number id="E455E6014A927B4F23CC658D98A74374">601</mods:number>
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<emphasis id="6CFAEABE2071FFAC2C9510F0FF3147F4" bold="true" box="[136,158,436,462]" pageId="2" pageNumber="225">1.</emphasis>
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<taxonomicName id="998E4D2F2071FFAC2CB910F0FCB447F4" authority="Toolmal, Schuit. & Culham" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[164,795,436,463]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="225" phylum="Tracheophyta" rank="species" species="alboluteum" status="spec. nov.">
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<emphasis id="6CFAEABE2071FFAC2CB910F0FE7447F5" bold="true" box="[164,475,436,463]" italics="true" pageId="2" pageNumber="225">Thrixspermum alboluteum</emphasis>
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Toolmal, Schuit. & Culham
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,
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<taxonomicNameLabel id="77C957C52071FFAC2F3810F1FC3C47F4" box="[805,915,437,462]" pageId="2" pageNumber="225" rank="species">spec. nov.</taxonomicNameLabel>
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<paragraph id="5E3136AC2071FFAC2F8410F0FBC147F5" blockId="2.[136,1134,436,463]" box="[921,1134,436,463]" pageId="2" pageNumber="225">
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(
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<figureCitation id="C6B52A292071FFAC2FBC10F0FBA147F4" box="[929,1038,436,462]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[136,234,1912,1934]" captionStartId-1="4.[136,234,1375,1397]" captionStartId-2="5.[136,234,1980,2002]" captionTargetBox-0="[264,1323,498,1881]" captionTargetBox-1="[151,1436,190,1350]" captionTargetBox-2="[164,1423,569,1956]" captionTargetId-0="figure-169@3.[260,1327,494,1885]" captionTargetId-1="figure-22@4.[151,1436,187,1350]" captionTargetId-2="figure-219@5.[164,1423,569,1956]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Thrixspermum alboluteum. A. Habit. B. Leaf apex. C. Flower (front view).D. Rachis with flowers. E. Column & column-foot with petal & lateral sepal (lateral view). F. Sac with warts on the backwall (close-up). G. Lip (lateral view). H. Lip (longitudinal section). I. Lip (adaxial view, natural position). J. Lip (adaxial view, spread out). K. Dissected perianth. L. Column with pedicel (ventral view). M. Operculum (dorsal view). N.Anther with pollinia, stipe & viscidium (ventral view). O. Pollinia (two pairs). P. Stipe & viscidium. (Drawn from holotype, N. Toolmal et al. 20122001, TTM 0006394). Illustration by Phattaravee Prommanut." captionText-1="FIGURE 2. Thrixspermum alboluteum. A.Flower (front view).B. Flower (oblique view). C. Flower (ventral view).D. Dissected perianth. E. Tepals with column and column-foot.F. Column & column-foot with pedicel (ventral & dorsal views).G. Lip, adaxial view (left: natural position; right: spread out). H. Lip, abaxial view (left: natural position; right: spread out) with scattered orange warts. I. Lip (left: lateral view). J. Lip (showing long capitate hair on the mid-lobe). K. Anther with pollinia (left: ventral view; right: dorsal view). L. Operculum (left: ventral view; right: dorsal view). M. Stipe & viscidium. N. Pollinia (two pairs). O. Pollinia (individual). Photographed by Wattana Tanming (A–J) & Nopparut Toolmal (K–O). Illustration assembled by Yongyuth Rattanasootr. A–J from living collection (QSBG) and K–O from spirit collection of N. Toolmal et al. 20122001 TTM 0006394." captionText-2="FIGURE 3. Thrixspermum alboluteum. A. Plant habit. B. Leaf (adaxial view). C. Leaf apex. D. Flower, rachis and floral bracts (ventral view). E. Flower (ventral view). Photographed by Nopparut Toolmal (A–C & E. In situ) & Wattana Tanming (D & E. Cultivation, QSBG). Illustration assembled by Yongyuth Rattanasootr." figureDoi-0="http://doi.org/10.5281/zenodo.8141766" figureDoi-1="http://doi.org/10.5281/zenodo.8141770" figureDoi-2="http://doi.org/10.5281/zenodo.8141772" httpUri-0="https://zenodo.org/record/8141766/files/figure.png" httpUri-1="https://zenodo.org/record/8141770/files/figure.png" httpUri-2="https://zenodo.org/record/8141772/files/figure.png" pageId="2" pageNumber="225">Figs. 1–3</figureCitation>
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,
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<figureCitation id="C6B52A292071FFAC280610F0FBF147F4" box="[1051,1118,436,462]" captionStart="FIGURE 4" captionStartId="6.[136,234,1430,1452]" captionTargetBox="[169,1418,196,1400]" captionTargetId="figure-22@6.[164,1423,190,1406]" captionTargetPageId="6" captionText="FIGURE 4. A comparison of allied species. A–B. Thrixspermum alboluteum. C–D. Thrixspermum patkaiense. E–F. Thrixspermum formosanum. Photographed by Wattana Tanming (A–B. Cultivation, QSBG), Nopparut Toolmal (C–D. In situ) & Morris Ng (E–F. Cultivation, Taiwan). Illustration assembled by Yongyuth Rattanasootr." figureDoi="http://doi.org/10.5281/zenodo.8141774" httpUri="https://zenodo.org/record/8141774/files/figure.png" pageId="2" pageNumber="225">4A–B</figureCitation>
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)
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.
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<materialsCitation id="EEE63CF12071FFAC2C951344FB484403" collectingDate="2020-12-20" collectionCode="N" collectorName="Ban Krang Camp" country="Thailand" county="Type" elevation="312" location="Kaeng Krachan National Park" municipality="South-Western Thailand" pageId="2" pageNumber="225" specimenCount="1" stateProvince="Phetchaburi" typeStatus="holotype">
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:—
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<collectingCountry id="2699763C2071FFAC2CCB1344FEF9442F" box="[214,342,512,533]" name="Thailand" pageId="2" pageNumber="225">THAILAND</collectingCountry>
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<collectingMunicipality id="BE55ACD62071FFAC2D7E1344FDFC442C" box="[355,595,512,534]" pageId="2" pageNumber="225">South-Western Thailand</collectingMunicipality>
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,
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,
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,
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<collectorName id="F37B537A2071FFAC289C1344FA9B442F" box="[1153,1332,512,533]" pageId="2" pageNumber="225">Ban Krang Camp</collectorName>
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<collectingDate id="3A74E9842071FFAC298E1344FEF74403" pageId="2" pageNumber="225" value="2020-12-20">20 December 2020</collectingDate>
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|
||
<location id="5B5160772071FFAC2DED1360FD9A4403" LSID="urn:lsid:plazi:treatment:D62787BA2071FFA92C9510F0FAB542F7:5B5160772071FFAC2DED1360FD9A4403" box="[496,565,548,569]" country="Thailand" name="Raksue" pageId="2" pageNumber="225" stateProvince="Phetchaburi">Raksue</location>
|
||
&
|
||
<collectionCode id="389FAE692071FFAC2E491360FDC64403" box="[596,617,548,569]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="2" pageNumber="225" type="Herbarium">W</collectionCode>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="6CFAEABE2071FFAC2E6F1360FC844403" box="[626,811,548,569]" italics="true" pageId="2" pageNumber="225">
|
||
<location id="5B5160772071FFAC2E6F1360FD6A4403" LSID="urn:lsid:plazi:treatment:D62787BA2071FFA92C9510F0FAB542F7:5B5160772071FFAC2E6F1360FD6A4403" box="[626,709,548,569]" country="Thailand" name="Tanming" pageId="2" pageNumber="225" stateProvince="Phetchaburi">Tanming</location>
|
||
20122001
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="8135880E2071FFAC2F2A1360FC3F4400" box="[823,912,548,570]" pageId="2" pageNumber="225">holotype</typeStatus>
|
||
: TTM-Spirit; isotypes:
|
||
<collectionCode id="389FAE692071FFAC286C1360FB0F4403" box="[1137,1184,548,569]" country="Thailand" httpUri="http://grbio.org/cool/d4ij-e7pz" name="National Park, Wildlife and Plant Conservation Department" pageId="2" pageNumber="225" type="Herbarium">BKF</collectionCode>
|
||
,
|
||
<collectionCode id="389FAE692071FFAC28B71360FB4E4403" box="[1194,1249,548,569]" country="Thailand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13640" lsid="urn:lsid:biocol.org:col:13640" name="Queen Sirikit Botanic Garden" pageId="2" pageNumber="225">QBG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272071FFAC2C95130CFB264464" box="[136,1161,584,606]" pageId="2" pageNumber="225" type="reference_group">
|
||
<paragraph id="5E3136AC2071FFAC2C95130CFB264464" blockId="2.[136,1451,512,606]" box="[136,1161,584,606]" pageId="2" pageNumber="225">
|
||
<treatmentCitationGroup id="7E9E11822071FFAC2C95130CFB264464" box="[136,1161,584,606]" pageId="2" pageNumber="225">
|
||
<taxonomicName id="998E4D2F2071FFAC2C95130CFE3D4467" ID-CoL="7C9YB" authority="Seidenfaden" box="[136,402,584,605]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="225" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2071FFAC2C95130CFE3D4467" box="[136,402,584,605]" italics="true" pageId="2" pageNumber="225">Thrixspermum musciflorum</emphasis>
|
||
</taxonomicName>
|
||
auct. non A.S. Rao & J. Joseph:
|
||
<treatmentCitation id="DF2F10BD2071FFAC2ED3130CFC124464" author="Seidenfaden, G." box="[718,957,584,606]" page="166" pageId="2" pageNumber="225" year="1988">
|
||
<bibRefCitation id="3A1F4B5D2071FFAC2ED3130CFC214467" author="Seidenfaden, G." box="[718,910,584,606]" pageId="2" pageNumber="225" pagination="1 - 398" refId="ref25710" refString="Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine vandoid genera. Opera Botanica 95: 1 - 398." type="journal article" year="1988">Seidenfaden (1988)</bibRefCitation>
|
||
: 166
|
||
</treatmentCitation>
|
||
, Fig. 101, Pl. XVIIb.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272071FFAC2C9513C8FD9E4560" pageId="2" pageNumber="225" type="diagnosis">
|
||
<paragraph id="5E3136AC2071FFAC2C9513C8FD9E4560" blockId="2.[136,1452,652,2083]" pageId="2" pageNumber="225">
|
||
<taxonomicName id="998E4D2F2071FFAC2C9513C8FE1A449C" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[136,437,652,678]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="225" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2071FFAC2C9513C8FE1A449C" box="[136,437,652,678]" italics="true" pageId="2" pageNumber="225">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="998E4D2F2071FFAC2EDD13C9FCC3449C" authorityName="Schltr. (Schlechter" authorityYear="1919" baseAuthorityName="Hayata" baseAuthorityYear="1911" box="[704,876,652,678]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="225" phylum="Tracheophyta" rank="species" species="formosanum">
|
||
<emphasis id="6CFAEABE2071FFAC2EDD13C9FCC3449C" box="[704,876,652,678]" italics="true" pageId="2" pageNumber="225">T. formosanum</emphasis>
|
||
</taxonomicName>
|
||
by the distinct concave triangular mid-lobe with emarginate apex, covered with long clavate hairs adaxially and in the apical part (vs. indistinct mid-lobe with glandular hairs adaxially), the obliquely obovate lateral lobes 4.0–5.0 mm long with rounded apex (vs. triangular 1.5–3.0 mm long with obtuse apex), bright yellow tepals, lip white, with brownish orange spots on the sac, and with scattered small orange warts inside the sac (vs. white tepals, lip white, marked with pinkish lines, with two orange-yellow bars along the apical margin, and without warts).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272071FFAD2CA01220FE1147E2" lastPageId="3" lastPageNumber="226" pageId="2" pageNumber="225" type="description">
|
||
<paragraph id="5E3136AC2071FFAD2CA01220FE1147E2" blockId="2.[136,1452,652,2083]" lastBlockId="3.[136,1452,158,472]" lastPageId="3" lastPageNumber="226" pageId="2" pageNumber="225">
|
||
Epiphytic herb.
|
||
<emphasis id="6CFAEABE2071FFAC2D6D1220FE1E4544" box="[368,433,868,894]" italics="true" pageId="2" pageNumber="225">Stems</emphasis>
|
||
patent, 3.5–8.0 cm long, with 5–14 internodes, terete, slightly laterally compressed, curved upwards in upper half, covered with the leaf sheaths, 4–13-leaved; internodes 1.2–4.4 ×
|
||
<quantity id="99769B492071FFAC289912CCFABF4599" box="[1156,1296,904,931]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="3.6" metricValueMin="1.2" pageId="2" pageNumber="225" unit="mm" value="2.4" valueMax="3.6" valueMin="1.2">1.2–3.6 mm</quantity>
|
||
.
|
||
<emphasis id="6CFAEABE2071FFAC290012CDFAF24598" box="[1309,1373,905,930]" italics="true" pageId="2" pageNumber="225">Roots</emphasis>
|
||
terete, thick, elongate,
|
||
<quantity id="99769B492071FFAC2D2312E8FED945FD" box="[318,374,940,967]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="225" unit="cm" value="4.0">4 cm</quantity>
|
||
to more than
|
||
<quantity id="99769B492071FFAC2E0512E8FDF045FD" box="[536,607,940,967]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="225" unit="cm" value="30.0">30 cm</quantity>
|
||
long,
|
||
<quantity id="99769B492071FFAC2EB412E8FC9E45FD" box="[681,817,940,967]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="2.4" metricValueMin="0.5" pageId="2" pageNumber="225" unit="mm" value="1.45" valueMax="2.4" valueMin="0.5">0.5–2.4 mm</quantity>
|
||
diam., arising from the basal part of the stem and from nodes opposite the leaves, not or sparsely branching.
|
||
<emphasis id="6CFAEABE2071FFAC2ECB1294FCCD45D0" box="[726,866,976,1002]" italics="true" pageId="2" pageNumber="225">Leaf sheaths</emphasis>
|
||
green, spotted or suffused with purplish red, slightly bilaterally compressed, longer than the stem internodes.
|
||
<emphasis id="6CFAEABE2071FFAC2F1712B1FCF74234" box="[778,856,1013,1038]" italics="true" pageId="2" pageNumber="225">Leaves</emphasis>
|
||
green, spotted or suffused with purplish red, patent, narrowly oblong to linear, 12–60 ×
|
||
<quantity id="99769B492071FFAC2E0F155CFDDF4209" box="[530,624,1048,1075]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="2" pageNumber="225" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
|
||
, sessile, slightly narrowed towards the base, leaves on the same side of the stem 1.4–7.0 mm apart, dorsiventrally flattened, conduplicate, V-shaped in cross-section, apex unequally 2-lobed with sharply acute lobes, with a small mucro in the middle, thick, rigid, fleshy, coriaceous, the midrib distinct on the abaxial side.
|
||
<emphasis id="6CFAEABE2071FFAC2CD915C0FECA42A4" box="[196,357,1156,1182]" italics="true" pageId="2" pageNumber="225">Inflorescences</emphasis>
|
||
one to eight per plant, patent, racemose,
|
||
<quantity id="99769B492071FFAC2F2815C0FC2642A5" box="[821,905,1156,1183]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="2" pageNumber="225" unit="cm" value="4.0" valueMax="6.0" valueMin="2.0">2–6 cm</quantity>
|
||
long, long pedunculate, longer than leaf, arising from the nodes at the opposite side of the leaves, green, slightly spotted purplish red; peduncle slender, slightly upwards curved from the middle,
|
||
<quantity id="99769B492071FFAC2D871588FDAE42DD" box="[410,513,1228,1255]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="2" pageNumber="225" unit="cm" value="3.25" valueMax="5.0" valueMin="1.5">1.5–5 cm</quantity>
|
||
long, 0.4–2.0 mm diam., with 2–3 internodes, the basal internode somewhat shorter, 0.5–2.0 cm long,
|
||
<quantity id="99769B492071FFAC2D4C15B4FE754331" box="[337,474,1264,1291]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="225" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
|
||
diam., the middle internode longer, 1.5–4.0 cm long, 0.4–1.0 mm diam., the terminal internode swollen distally,
|
||
<quantity id="99769B492071FFAC2DAF1450FD814315" box="[434,558,1300,1327]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="2.3" metricValueMin="0.5" pageId="2" pageNumber="225" unit="cm" value="1.4" valueMax="2.3" valueMin="0.5">0.5–2.3 cm</quantity>
|
||
long, broadening to 2.0 mm diam. just below rachis; peduncle scales 2–3, green, broadly ovate to triangular, incurved, concave, thickened, slightly boat-shaped, clasping the peduncle, 1.2–2.0 ×
|
||
<quantity id="99769B492071FFAC2967147CFF70434D" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="2" pageNumber="225" unit="mm" value="1.15" valueMax="1.5" valueMin="0.8">0.8– 1.5 mm</quantity>
|
||
, apex acute, keeled; rachis relatively short,
|
||
<quantity id="99769B492071FFAC2EA31418FC95434D" box="[702,826,1372,1399]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="2" pageNumber="225" unit="cm" value="0.75" valueMax="1.2" valueMin="0.3">0.3–1.2 cm</quantity>
|
||
long, depending on the age of the inflorescence, densely 4–17-flowered, with one flower open at a time, swollen,
|
||
<quantity id="99769B492071FFAC2F3F14C4FC2D43A1" box="[802,898,1408,1435]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="2" pageNumber="225" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
|
||
diam., clearly thicker than the peduncle. Floral bracts quaquaversal, green, broadly ovate to triangular, incurved, concave, thickened, slightly boat-shaped, clasping the rachis, 1.5–2.0 × 1.0–
|
||
<quantity id="99769B492071FFAC2DBF148CFE5743D9" box="[418,504,1480,1507]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="225" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, apex acute, keeled.
|
||
<emphasis id="6CFAEABE2071FFAC2EC6148CFC1D43D8" box="[731,946,1480,1506]" italics="true" pageId="2" pageNumber="225">Pedicel-with-ovary</emphasis>
|
||
yellowish green, cylindrical, 6-grooved, 2.5– 7.0 mm long, 1.0 mm diam.
|
||
<emphasis id="6CFAEABE2071FFAC2DCD14A8FD83403C" box="[464,556,1516,1542]" italics="true" pageId="2" pageNumber="225">Flowers</emphasis>
|
||
resupinate, opening widely, lasting only one day, membranaceous,
|
||
<quantity id="99769B492071FFAC293014A8FA04403D" box="[1325,1451,1516,1543]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="2" pageNumber="225" unit="cm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 cm</quantity>
|
||
across, mildly sweet-scented; tepals bright yellow, lip white, spotted with brownish orange on both surfaces of the sac and at apex of the callus, the sac with orange warts inside on the back wall which shine through on the outside, column and anther white, pollinia bright yellow, stipe and viscidium creamy white.
|
||
<emphasis id="6CFAEABE2071FFAC2FCE171CFBCC4048" box="[979,1123,1624,1650]" italics="true" pageId="2" pageNumber="225">Dorsal sepal</emphasis>
|
||
obovate to elliptic, 3.5–8.0 ×
|
||
<quantity id="99769B492071FFAC2C951738FF4740AD" box="[136,232,1660,1687]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="2" pageNumber="225" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
|
||
, concave, slightly boat-shaped, apex obtuse, glabrous, 5-nerved, nerves reticulate branching.
|
||
<emphasis id="6CFAEABE2071FFAC29161738FA0440AC" box="[1291,1451,1660,1686]" italics="true" pageId="2" pageNumber="225">Lateral sepals</emphasis>
|
||
broadly ovate, subfalcate, 4–7 ×
|
||
<quantity id="99769B492071FFAC2DE717E4FDF64081" box="[506,601,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="225" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
|
||
, obliquely dilated at the lower margin, slightly concave, with a low abaxial keel, the base broadly connected with the column-foot, obtuse, glabrous, 5-nerved, nerves reticulate branching.
|
||
<emphasis id="6CFAEABE2071FFAC29781780FA0440E4" box="[1381,1451,1732,1758]" italics="true" pageId="2" pageNumber="225">Petals</emphasis>
|
||
oblanceolate, 4.4–4.5 × 1.8–2.0 mm, slightly oblique, curving downward from the middle, shorter and narrower than the sepals, slightly concave, obtuse, glabrous, 3-nerved, nerves reticulate branching.
|
||
<emphasis id="6CFAEABE2071FFAC28201649FBCD411C" box="[1085,1122,1805,1830]" italics="true" pageId="2" pageNumber="225">Lip</emphasis>
|
||
suborbicular in outline when spread, 3-lobed, 5–7 × 2.0–
|
||
<quantity id="99769B492071FFAC2DDD1674FDB94171" box="[448,534,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="225" unit="mm" value="3.5">3.5 mm</quantity>
|
||
in natural position,
|
||
<quantity id="99769B492071FFAC2EE41674FCF84171" box="[761,855,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="225" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
|
||
wide when spread, with a broad, sac-like spur at the base; spur oblong-conical, 2.0–2.2 ×
|
||
<quantity id="99769B492071FFAC2E351610FD1D4155" box="[552,690,1876,1903]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="2" pageNumber="225" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
|
||
, apex shallowly bilobulate with obtuse lobules (0.3 ×
|
||
<quantity id="99769B492071FFAC290D1610FAFA4154" box="[1296,1365,1876,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="225" unit="mm" value="1.0">1 mm</quantity>
|
||
), inside towards the base of the callus with scattered small orange warts, and densely covered with long clavate hairs (
|
||
<quantity id="99769B492071FFAC2948163CFA0341A9" box="[1365,1452,1912,1939]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="225" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long) towards the lip blade; lateral lobes obliquely obovate, distinctly surpassing the mid-lobe, 4–5 ×
|
||
<quantity id="99769B492071FFAC293816D8FAC4418D" box="[1317,1387,1948,1975]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="225" unit="mm" value="3.0">3 mm</quantity>
|
||
, with rounded apex, erect, curving forwards and bent inwards, clasping the column, adaxial surface sparsely covered with
|
||
<quantity id="99769B492071FFAC2C9516A0FF4F41C5" box="[136,224,2020,2047]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="225" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long clavate hairs in the basal part and along the apical margins, abaxial surface glabrous; mid-lobe short, broadly triangular to rounded, 0.5–1.0 ×
|
||
<quantity id="99769B492071FFAC2E53194CFD034E19" box="[590,684,2056,2083]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="225" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
, somewhat concave, densely covered with
|
||
<quantity id="99769B492071FFAC2896194CFB4E4E19" box="[1163,1249,2056,2083]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="225" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long clavate hairs on the adaxial surface and along the margins, rather thick (1.0–
|
||
<quantity id="99769B492070FFAD2F4E11DAFC024682" box="[851,941,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="226" unit="mm" value="1.5">1.5 mm</quantity>
|
||
), fleshy, entire, apex emarginate (each lobule 0.5 ×
|
||
<quantity id="99769B492070FFAD2CD01186FE8646E6" box="[205,297,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="226" unit="mm" value="1.5">1.5 mm</quantity>
|
||
); lip adaxially between the lateral lobes sparsely covered with long clavate hairs, glabrous abaxially, a little below the mid-lobe with a trapeziform callus 1.0–
|
||
<quantity id="99769B492070FFAD2F3C11A2FCD6473B" box="[801,889,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="226" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long,
|
||
<quantity id="99769B492070FFAD2FD811A2FB8A473B" box="[965,1061,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="226" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
|
||
diam., apex bilobulate, decurved.
|
||
<emphasis id="6CFAEABE2070FFAD2C95104EFF4E471E" box="[136,225,266,292]" italics="true" pageId="3" pageNumber="226">Column</emphasis>
|
||
cylindrical, short, stout, 2 × 1.5–2.0 mm, laterally winged; column-foot short, broad, 1.0–2.5 ×
|
||
<quantity id="99769B492070FFAD2900104EFA07471F" box="[1309,1448,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="3" pageNumber="226" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
|
||
.
|
||
<emphasis id="6CFAEABE2070FFAD2C95106AFF7B4772" box="[136,212,302,328]" italics="true" pageId="3" pageNumber="226">Anther</emphasis>
|
||
suborbicular in outline, glabrous, 1.0 ×
|
||
<quantity id="99769B492070FFAD2E8A106AFD5F4773" box="[663,752,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="226" unit="mm" value="1.2">1.2 mm</quantity>
|
||
.
|
||
<taxonomicName id="998E4D2F2070FFAD2EE6106AFCFA4772" box="[763,853,302,328]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="3" pageNumber="226" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6CFAEABE2070FFAD2EE6106AFCFA4772" box="[763,853,302,328]" italics="true" pageId="3" pageNumber="226">Pollinia</emphasis>
|
||
</taxonomicName>
|
||
in two pairs, those in a pair unequal, elliptic, curved; the larger 0.8 ×
|
||
<quantity id="99769B492070FFAD2D591016FE304757" box="[324,415,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="226" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; the smaller, 0.6 ×
|
||
<quantity id="99769B492070FFAD2E9D1016FD734757" box="[640,732,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="226" unit="mm" value="0.4">0.4 mm</quantity>
|
||
; stipe broadly ovate, 0.5×
|
||
<quantity id="99769B492070FFAD280C1016FBC44757" box="[1041,1131,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="226" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, concave along the median, slightly translucent; viscidium transverse-linear, 0.2 ×
|
||
<quantity id="99769B492070FFAD2EEF1032FCE447AB" box="[754,843,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="226" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, bent into a crescent shape, slightly translucent.
|
||
<emphasis id="6CFAEABE2070FFAD296F1033FA0347AA" box="[1394,1452,375,400]" italics="true" pageId="3" pageNumber="226">Fruit</emphasis>
|
||
green, narrowly cylindrical,
|
||
<quantity id="99769B492070FFAD2DD610DEFDB3478F" box="[459,540,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="226" unit="mm" value="70.0">70 mm</quantity>
|
||
long,
|
||
<quantity id="99769B492070FFAD2E7810DEFD07478F" box="[613,680,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="226" unit="mm" value="4.0">4 mm</quantity>
|
||
diam., with 6 longitudinal ridges. Described from living plants and flowers, and spirit material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0AF166242070FFAD2C95163CFBE64E24" ID-DOI="http://doi.org/10.5281/zenodo.8141766" ID-Zenodo-Dep="8141766" httpUri="https://zenodo.org/record/8141766/files/figure.png" pageId="3" pageNumber="226" startId="3.[136,234,1912,1934]" targetBox="[264,1323,498,1881]" targetPageId="3" targetType="figure">
|
||
<paragraph id="5E3136AC2070FFAD2C95163CFBE64E24" blockId="3.[136,1452,1912,2078]" pageId="3" pageNumber="226">
|
||
<emphasis id="6CFAEABE2070FFAD2C95163CFEAD41B7" bold="true" box="[136,258,1912,1934]" pageId="3" pageNumber="226">FIGURE 1.</emphasis>
|
||
<taxonomicName id="998E4D2F2070FFAD2D1B163CFDAC41B7" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[262,515,1912,1933]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="226" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2070FFAD2D1B163CFDAC41B7" box="[262,515,1912,1933]" italics="true" pageId="3" pageNumber="226">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
. A. Habit. B. Leaf apex. C. Flower (front view). D. Rachis with flowers. E. Column & column-foot with petal & lateral sepal (lateral view). F. Sac with warts on the backwall (close-up). G. Lip (lateral view). H. Lip (longitudinal section). I. Lip (adaxial view, natural position). J. Lip (adaxial view, spread out). K. Dissected perianth. L. Column with pedicel (ventral view). M. Operculum (dorsal view). N. Anther with pollinia, stipe & viscidium (ventral view). O.
|
||
<taxonomicName id="998E4D2F2070FFAD2FD016A0FBB741C0" box="[973,1048,2020,2042]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="3" pageNumber="257" phylum="Tracheophyta" rank="genus">Pollinia</taxonomicName>
|
||
(two pairs). P. Stipe & viscidium. (Drawn from holotype,
|
||
<emphasis id="6CFAEABE2070FFAD2D00194DFD8C4E24" box="[285,547,2056,2078]" italics="true" pageId="3" pageNumber="226">N. Toolmal et al. 20122001</emphasis>
|
||
, TTM 0006394). Illustration by Phattaravee Prommanut.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0AF166242077FFAA2C95141BFCE84077" ID-DOI="http://doi.org/10.5281/zenodo.8141770" ID-Zenodo-Dep="8141770" httpUri="https://zenodo.org/record/8141770/files/figure.png" pageId="4" pageNumber="227" startId="4.[136,234,1375,1397]" targetBox="[151,1436,190,1350]" targetPageId="4" targetType="figure">
|
||
<paragraph id="5E3136AC2077FFAA2C95141BFCE84077" blockId="4.[136,1453,1375,1613]" pageId="4" pageNumber="227">
|
||
<emphasis id="6CFAEABE2077FFAA2C95141BFEAE434E" bold="true" box="[136,257,1375,1397]" pageId="4" pageNumber="227">FIGURE 2.</emphasis>
|
||
<taxonomicName id="998E4D2F2077FFAA2D18141BFDAD434E" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[261,514,1375,1396]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2077FFAA2D18141BFDAD434E" box="[261,514,1375,1396]" italics="true" pageId="4" pageNumber="227">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
. A. Flower (front view). B. Flower (oblique view). C. Flower (ventral view). D. Dissected perianth. E. Tepals with column and column-foot. F. Column & column-foot with pedicel (ventral & dorsal views). G. Lip, adaxial view (left: natural position; right: spread out). H. Lip, abaxial view (left: natural position; right: spread out) with scattered orange warts. I. Lip (left: lateral view). J. Lip (showing long capitate hair on the mid-lobe). K. Anther with pollinia (left: ventral view; right: dorsal view). L. Operculum (left: ventral view; right: dorsal view). M. Stipe & viscidium. N.
|
||
<taxonomicName id="998E4D2F2077FFAA2EE214ABFCE5403F" box="[767,842,1519,1541]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="4" pageNumber="257" phylum="Tracheophyta" rank="genus">Pollinia</taxonomicName>
|
||
(two pairs). O.
|
||
<taxonomicName id="998E4D2F2077FFAA2FF914ABFB80403F" box="[996,1071,1519,1541]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="4" pageNumber="257" phylum="Tracheophyta" rank="genus">Pollinia</taxonomicName>
|
||
(individual). Photographed by Wattana Tanming (A–J) & Nopparut Toolmal (K–O). Illustration assembled by Yongyuth Rattanasootr. A–J from living collection (QSBG) and K–O from spirit collection of
|
||
<emphasis id="6CFAEABE2077FFAA2DBA177CFD024076" box="[423,685,1591,1613]" italics="true" pageId="4" pageNumber="227">N. Toolmal et al. 20122001</emphasis>
|
||
TTM 0006394.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="169465272077FFAA2CA0173FFCA640E4" pageId="4" pageNumber="227" type="distribution">
|
||
<paragraph id="5E3136AC2077FFAA2CA0173FFCA640E4" blockId="4.[136,1452,1659,2082]" pageId="4" pageNumber="227">
|
||
Distribution:—
|
||
<collectingCountry id="2699763C2077FFAA2D74173FFE5240AF" box="[361,509,1659,1685]" name="Thailand" pageId="4" pageNumber="227">THAILAND</collectingCountry>
|
||
. Eastern:
|
||
<collectingRegion id="9C4AF84E2077FFAA2E72173FFC6F40AF" box="[623,960,1659,1685]" country="Thailand" name="Nakhon Ratchasima" pageId="4" pageNumber="227">Nakhon Ratchasima Province</collectingRegion>
|
||
(Khao Yai National Park); South-Western:
|
||
<collectingRegion id="9C4AF84E2077FFAA2C9517DBFED54083" box="[136,378,1695,1721]" country="Thailand" name="Uthai Thani" pageId="4" pageNumber="227">Uthai Thani Province</collectingRegion>
|
||
(Huai Kha Khaeng Wildlife Sanctuary);
|
||
<collectingRegion id="9C4AF84E2077FFAA2F5A17DBFB974083" box="[839,1080,1695,1721]" country="Thailand" name="Phetchaburi" pageId="4" pageNumber="227">Phetchaburi Province</collectingRegion>
|
||
(Kaeng Krachan National Park);
|
||
<collectingRegion id="9C4AF84E2077FFAA2C951787FE4E40E7" box="[136,481,1731,1757]" country="Thailand" name="Prachuap Khiri Khan" pageId="4" pageNumber="227">Prachuap Khiri Khan Province</collectingRegion>
|
||
(Khan Kra Dai)
|
||
<emphasis id="6CFAEABE2077FFAA2E831787FCAD40E7" bold="true" box="[670,770,1731,1757]" pageId="4" pageNumber="227">
|
||
(
|
||
<figureCitation id="C6B52A292077FFAA2EBB1787FD5640E7" box="[678,761,1731,1757]" captionStart="FIGURE 25" captionStartId="36.[136,234,607,629]" captionTargetId="figure-22@36.[386,1202,175,583]" captionTargetPageId="36" captionText="FIGURE 25. Distribution map of Thrixspermum alboluteum in Thailand." figureDoi="http://doi.org/10.5281/zenodo.8148372" httpUri="https://zenodo.org/record/8148372/files/figure.png" pageId="4" pageNumber="227">Fig. 25</figureCitation>
|
||
)
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272077FFAA2CA017A3FD9A418F" pageId="4" pageNumber="227" type="biology_ecology">
|
||
<paragraph id="5E3136AC2077FFAA2CA017A3FAB0411C" blockId="4.[136,1452,1659,2082]" pageId="4" pageNumber="227">
|
||
Phenology:—Flowering in January, March, April and May (in natural habitat, four records); April (in cultivation, one record). Flowering is probably intermittent throughout the year, as in most species of
|
||
<taxonomicName id="998E4D2F2077FFAA2868164FFAB7411F" authorityName="Lour. (Loureiro" authorityYear="1790" box="[1141,1304,1803,1829]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6CFAEABE2077FFAA2868164FFAB7411F" box="[1141,1304,1803,1829]" italics="true" pageId="4" pageNumber="227">Thrixspermum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5E3136AC2077FFAA2CA0166BFD9A418F" blockId="4.[136,1452,1659,2082]" pageId="4" pageNumber="227">
|
||
Habitat and ecology:—Epiphyte in dry evergreen and mixed deciduous forests, at
|
||
<quantity id="99769B492077FFAA287C166BFB4D4170" box="[1121,1250,1839,1866]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="4" pageNumber="227" unit="m" value="450.0" valueMax="600.0" valueMin="300.0">300–600 m</quantity>
|
||
elevation, locally common along streams and surrounding zones, growing on twigs of small trees, such as
|
||
<taxonomicName id="998E4D2F2077FFAA28491610FEA541A8" authority="Jack (1820: 509)" authorityName="Jack" authorityPageNumber="509" authorityYear="1820" class="Magnoliopsida" family="Lythraceae" genus="Lagerstroemia" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="species" species="floribunda">
|
||
<emphasis id="6CFAEABE2077FFAA28491610FADB4157" box="[1108,1396,1875,1901]" italics="true" pageId="4" pageNumber="227">Lagerstroemia floribunda</emphasis>
|
||
Jack (1820: 509)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="998E4D2F2077FFAA2D0A1633FE3941AB" box="[279,406,1911,1937]" class="Magnoliopsida" family="Lythraceae" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="family">Lythraceae</taxonomicName>
|
||
) and (introduced)
|
||
<taxonomicName id="998E4D2F2077FFAA2E7C1633FCB341AB" box="[609,796,1911,1937]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="species" species="guajava">
|
||
<emphasis id="6CFAEABE2077FFAA2E7C1633FCB341AB" box="[609,796,1911,1937]" italics="true" pageId="4" pageNumber="227">Psidium guajava</emphasis>
|
||
</taxonomicName>
|
||
L. (
|
||
<bibRefCitation id="3A1F4B5D2077FFAA2F5A1633FB8841A8" author="Linnaeus, C." box="[839,1063,1911,1938]" pageId="4" pageNumber="227" refId="ref24938" refString="Linnaeus, C. (1753) Species Plantarum, vol. 1. Holmiae, Impensis Laurentii Salvii, 560 pp." type="book" year="1753">Linnaeus 1753: 470</bibRefCitation>
|
||
) (
|
||
<taxonomicName id="998E4D2F2077FFAA2821163CFB1741A8" box="[1084,1208,1912,1938]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
|
||
), in exposed positions at about 1–3 meters above the ground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272077FFAB2CA016FBFCC6441B" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227" type="conservation">
|
||
<paragraph id="5E3136AC2077FFAB2CA016FBFCC6441B" blockId="4.[136,1452,1659,2082]" lastBlockId="5.[136,1452,158,545]" lastPageId="5" lastPageNumber="228" pageId="4" pageNumber="227">
|
||
Conservation status:—
|
||
<taxonomicName id="998E4D2F2077FFAA2DA116FBFD4B41E3" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[444,740,1983,2009]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="227" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2077FFAA2DA116FBFD4B41E3" box="[444,740,1983,2009]" italics="true" pageId="4" pageNumber="227">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
has been recorded from four localities in Eastern
|
||
<collectingCountry id="2699763C2077FFAA291616FBFAC041E3" box="[1291,1391,1983,2009]" name="Thailand" pageId="4" pageNumber="227">Thailand</collectingCountry>
|
||
from Khao Yai National Park and South-Western
|
||
<collectingCountry id="2699763C2077FFAA2E6416A7FD7041C7" box="[633,735,2019,2045]" name="Thailand" pageId="4" pageNumber="227">Thailand</collectingCountry>
|
||
, namely from the Huai Kha Khaeng Wildlife Sanctuary, Kaeng Krachan National Park and Khan Kra Dai Cave, with an Extent of occurrence (EOO) of
|
||
<quantity id="99769B492077FFAA2871194CFABA4E1B" box="[1132,1301,2055,2082]" metricMagnitude="7" metricUnit="m" metricValue="4.9773942" pageId="4" pageNumber="227" unit="km" value="49773.942">49,773.942 km</quantity>
|
||
<superScript id="A9FB9BE42077FFAA29081943FAB24E2F" attach="left" box="[1301,1309,2055,2069]" fontSize="6" pageId="4" pageNumber="227">2</superScript>
|
||
and an Area of occupancy (AOO) of
|
||
<quantity id="99769B492076FFAB2D8A11DAFDA14682" box="[407,526,158,184]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="228" unit="km" value="16.0">16.000 km</quantity>
|
||
<superScript id="A9FB9BE42076FFAB2E1311DAFDB94696" attach="left" box="[526,534,158,172]" fontSize="6" pageId="5" pageNumber="228">2</superScript>
|
||
. Although the species occurs in a strictly protected area in Khao Yai National Park and Kaeng Krachan National Park, it has a very small known population there of less than 10 mature individuals. The natural habitat in Kaeng Krachan National Park was also destroyed by being developed into a campsite and visitor centre, and visitors also collected the plants. It is, however, possible that the new species can be found distributed in other similar habitats in both sites. Khan Kra Dai Cave is currently being developed for the local temple; it is surrounded by plantations and is becoming more popular among tourists, all of which poses a considerably threat to the population of this species. However, it seems to be fairly common in Huai Kha Khaeng Wildlife Sanctuary, with more than 1,000 mature individuals present in several places, located in a diverse forest of the wildlife sanctuary and with strict protection (Priwan Srisom and Salil Sittisatjathum, pers. comm.). It is, therefore, provisionally assigned the conservation status of Vulnerable (VU) according to IUCN criteria (Vulnerable: B1a, c; C2a; D1, 2). More insight into the distribution of this species is needed for a reliable assessment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0AF166242076FFAB2C9516F8FDCE4E20" ID-DOI="http://doi.org/10.5281/zenodo.8141772" ID-Zenodo-Dep="8141772" httpUri="https://zenodo.org/record/8141772/files/figure.png" pageId="5" pageNumber="228" startId="5.[136,234,1980,2002]" targetBox="[164,1423,569,1956]" targetPageId="5" targetType="figure">
|
||
<paragraph id="5E3136AC2076FFAB2C9516F8FDCE4E20" blockId="5.[136,1452,1980,2074]" pageId="5" pageNumber="228">
|
||
<emphasis id="6CFAEABE2076FFAB2C9516F8FEAC41EB" bold="true" box="[136,259,1980,2002]" pageId="5" pageNumber="228">FIGURE 3.</emphasis>
|
||
<taxonomicName id="998E4D2F2076FFAB2D1416F8FDA841EB" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[265,519,1980,2001]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="228" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2076FFAB2D1416F8FDA841EB" box="[265,519,1980,2001]" italics="true" pageId="5" pageNumber="228">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
. A. Plant habit. B. Leaf (adaxial view). C. Leaf apex. D. Flower, rachis and floral bracts (ventral view). E. Flower (ventral view). Photographed by Nopparut Toolmal (A–C & E. In situ) & Wattana Tanming (D & E. Cultivation, QSBG). Illustration assembled by Yongyuth Rattanasootr.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0AF166242075FFA82C9514D2FC9C43CE" ID-DOI="http://doi.org/10.5281/zenodo.8141774" ID-Zenodo-Dep="8141774" httpUri="https://zenodo.org/record/8141774/files/figure.png" pageId="6" pageNumber="229" startId="6.[136,234,1430,1452]" targetBox="[169,1418,196,1400]" targetPageId="6" targetType="figure">
|
||
<paragraph id="5E3136AC2075FFA82C9514D2FC9C43CE" blockId="6.[136,1452,1430,1524]" pageId="6" pageNumber="229">
|
||
<emphasis id="6CFAEABE2075FFA82C9514D2FEA94391" bold="true" box="[136,262,1430,1452]" pageId="6" pageNumber="229">FIGURE 4.</emphasis>
|
||
A comparison of allied species. A–B.
|
||
<taxonomicName id="998E4D2F2075FFA82E9214D2FC3E4391" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[655,913,1430,1451]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2075FFA82E9214D2FC3E4391" box="[655,913,1430,1451]" italics="true" pageId="6" pageNumber="229">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
. C–D.
|
||
<taxonomicName id="998E4D2F2075FFA82FC314D2FB754391" box="[990,1242,1430,1451]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="patkaiense">
|
||
<emphasis id="6CFAEABE2075FFA82FC314D2FB754391" box="[990,1242,1430,1451]" italics="true" pageId="6" pageNumber="229">Thrixspermum patkaiense</emphasis>
|
||
</taxonomicName>
|
||
. E–F.
|
||
<taxonomicName id="998E4D2F2075FFA8290214D2FEAF43F5" authorityName="Schltr. (Schlechter" authorityYear="1919" baseAuthorityName="Hayata" baseAuthorityYear="1911" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="formosanum">
|
||
<emphasis id="6CFAEABE2075FFA8290214D2FEAF43F5" italics="true" pageId="6" pageNumber="229">Thrixspermum formosanum</emphasis>
|
||
</taxonomicName>
|
||
. Photographed by Wattana Tanming (A–B. Cultivation, QSBG), Nopparut Toolmal (C–D. In situ) & Morris Ng (E–F. Cultivation, Taiwan). Illustration assembled by Yongyuth Rattanasootr.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="169465272075FFA82CA01767FF5D405B" pageId="6" pageNumber="229" type="etymology">
|
||
<paragraph id="5E3136AC2075FFA82CA01767FF5D405B" blockId="6.[136,1452,1571,2065]" pageId="6" pageNumber="229">
|
||
Etymology:—From the Latin
|
||
<taxonomicName id="998E4D2F2075FFA82E081767FDF44007" authorityName="Schltr. (Schlechter" authorityYear="1911" baseAuthorityName="Ridl." box="[533,603,1571,1597]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="album">
|
||
<emphasis id="6CFAEABE2075FFA82E081767FDF44007" box="[533,603,1571,1597]" italics="true" pageId="6" pageNumber="229">album</emphasis>
|
||
</taxonomicName>
|
||
, white, and
|
||
<emphasis id="6CFAEABE2075FFA82EF31767FC954007" box="[750,826,1571,1597]" italics="true" pageId="6" pageNumber="229">luteum</emphasis>
|
||
, yellow, referring to the yellow tepals and the mainly white lip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272075FFA82CA0172FFF444093" pageId="6" pageNumber="229" type="vernacular_names">
|
||
<paragraph id="5E3136AC2075FFA82CA0172FFF444093" blockId="6.[136,1452,1571,2065]" pageId="6" pageNumber="229">
|
||
Vernacular (
|
||
<collectingCountry id="2699763C2075FFA82D5A172FFE1F40BF" box="[327,432,1643,1669]" name="Thailand" pageId="6" pageNumber="229">Thailand</collectingCountry>
|
||
):—
|
||
<emphasis id="6CFAEABE2075FFA82DC4172FFD6E40BF" bold="true" box="[473,705,1643,1669]" pageId="6" pageNumber="229">ตะขาบเหลืองตะนาวศรี</emphasis>
|
||
(Takhap Lueang Tanao Si),
|
||
<emphasis id="6CFAEABE2075FFA8281A172FFB4440BF" bold="true" box="[1031,1259,1643,1669]" pageId="6" pageNumber="229">ตะขาบเหลืองปากหยัก</emphasis>
|
||
(Takhap Lueang Pakyak).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272075FFA82CA017F7FE7941F3" pageId="6" pageNumber="229" type="materials_examined">
|
||
<paragraph id="5E3136AC2075FFA82CA017F7FE7941F3" blockId="6.[136,1452,1571,2065]" pageId="6" pageNumber="229">
|
||
Additional specimens examined:—
|
||
<collectingCountry id="2699763C2075FFA82E5117F7FD4E40F7" box="[588,737,1715,1741]" name="Thailand" pageId="6" pageNumber="229">THAILAND</collectingCountry>
|
||
. Eastern:
|
||
<collectingRegion id="9C4AF84E2075FFA82F4F17F7FB0940F7" box="[850,1190,1715,1741]" country="Thailand" name="Nakhon Ratchasima" pageId="6" pageNumber="229">Nakhon Ratchasima Province</collectingRegion>
|
||
, Pak Chong, Khao Yai National Park,
|
||
<emphasis id="6CFAEABE2075FFA82D2F1793FE4240CB" box="[306,493,1751,1777]" italics="true" pageId="6" pageNumber="229">Cumberlege 526</emphasis>
|
||
(C 8433!, K 24716!,
|
||
<bibRefCitation id="3A1F4B5D2075FFA82EFF1793FC4940CB" author="Seidenfaden, G." box="[738,998,1751,1777]" pageId="6" pageNumber="229" pagination="1 - 398" refId="ref25710" refString="Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine vandoid genera. Opera Botanica 95: 1 - 398." type="journal article" year="1988">Seidenfaden 1988: 166</bibRefCitation>
|
||
as
|
||
<taxonomicName id="998E4D2F2075FFA82810179CFB1A40CB" box="[1037,1205,1751,1777]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2075FFA82810179CFB1A40CB" box="[1037,1205,1751,1777]" italics="true" pageId="6" pageNumber="229">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<emphasis id="6CFAEABE2075FFA828D01793FA2740CB" box="[1229,1416,1751,1777]" italics="true" pageId="6" pageNumber="229">Cumberlege 527</emphasis>
|
||
(C 8428!, K 27388!,
|
||
<bibRefCitation id="3A1F4B5D2075FFA82D4A17BFFDF1412F" author="Seidenfaden, G." box="[343,606,1787,1813]" pageId="6" pageNumber="229" pagination="1 - 398" refId="ref25710" refString="Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine vandoid genera. Opera Botanica 95: 1 - 398." type="journal article" year="1988">Seidenfaden 1988: 166</bibRefCitation>
|
||
as
|
||
<taxonomicName id="998E4D2F2075FFA82E9A17B8FC9E412F" box="[647,817,1787,1813]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2075FFA82E9A17B8FC9E412F" box="[647,817,1787,1813]" italics="true" pageId="6" pageNumber="229">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
), Haew Suwat Waterfall,
|
||
<emphasis id="6CFAEABE2075FFA8284117BFFF6F4103" italics="true" pageId="6" pageNumber="229">Seidenfaden & Smitinand GT 5790</emphasis>
|
||
(C 8353!, 8354!,
|
||
<bibRefCitation id="3A1F4B5D2075FFA82D8C165BFD384103" author="Seidenfaden, G." box="[401,663,1823,1849]" pageId="6" pageNumber="229" pagination="1 - 398" refId="ref25710" refString="Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine vandoid genera. Opera Botanica 95: 1 - 398." type="journal article" year="1988">Seidenfaden 1988: 162</bibRefCitation>
|
||
as
|
||
<taxonomicName id="998E4D2F2075FFA82EDC1664FCC74103" box="[705,872,1823,1849]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="trichoglottis">
|
||
<emphasis id="6CFAEABE2075FFA82EDC1664FCC74103" box="[705,872,1823,1849]" italics="true" pageId="6" pageNumber="229">T. trichoglottis</emphasis>
|
||
</taxonomicName>
|
||
); South-Western:
|
||
<collectingRegion id="9C4AF84E2075FFA82825165BFA9F4103" box="[1080,1328,1823,1849]" country="Thailand" name="Uthai Thani" pageId="6" pageNumber="229">Uthai Thani Province</collectingRegion>
|
||
, Huai Kha Khaeng Wildlife Sanctuary,
|
||
<emphasis id="6CFAEABE2075FFA82DDF1607FDB14167" box="[450,542,1859,1885]" italics="true" pageId="6" pageNumber="229">K.S. s.n.</emphasis>
|
||
,
|
||
<date id="2A30106C2075FFA82E361607FD6B4167" box="[555,708,1859,1885]" pageId="6" pageNumber="229" value="1991-04-03">03 April 1991</date>
|
||
(BCU s.n.!),
|
||
<collectingRegion id="9C4AF84E2075FFA82F471607FBE34167" box="[858,1100,1859,1885]" country="Thailand" name="Phetchaburi" pageId="6" pageNumber="229">Phetchaburi Province</collectingRegion>
|
||
, Kaeng Krachan National Park,
|
||
<emphasis id="6CFAEABE2075FFA82C951623FEB541BB" box="[136,282,1895,1921]" italics="true" pageId="6" pageNumber="229">S. Raksue 79</emphasis>
|
||
,
|
||
<date id="2A30106C2075FFA82D341623FE6841BB" box="[297,455,1895,1921]" pageId="6" pageNumber="229" value="2008-04-16">16 April 2008</date>
|
||
(BKF s.n.!),
|
||
<collectingRegion id="9C4AF84E2075FFA82E7D1623FC1141BB" box="[608,958,1895,1921]" country="Thailand" name="Prachuap Khiri Khan" pageId="6" pageNumber="229">Prachuap Khiri Khan Province</collectingRegion>
|
||
, Khan Kra Dai,
|
||
<emphasis id="6CFAEABE2075FFA82867162CFB4E41BB" box="[1146,1249,1895,1921]" italics="true" pageId="6" pageNumber="229">Put 2318</emphasis>
|
||
,
|
||
<date id="2A30106C2075FFA828ED1623FA0441BB" box="[1264,1451,1895,1921]" pageId="6" pageNumber="229" value="1929-01-16">16 January 1929</date>
|
||
(C 8379!; K s.n.!,
|
||
<bibRefCitation id="3A1F4B5D2075FFA82D4816CFFDF8419F" author="Seidenfaden, G." box="[341,599,1931,1957]" pageId="6" pageNumber="229" pagination="1 - 398" refId="ref25710" refString="Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine vandoid genera. Opera Botanica 95: 1 - 398." type="journal article" year="1988">Seidenfaden 1988: 166</bibRefCitation>
|
||
as
|
||
<taxonomicName id="998E4D2F2075FFA82E6116C8FC8B419F" box="[636,804,1931,1957]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2075FFA82E6116C8FC8B419F" box="[636,804,1931,1957]" italics="true" pageId="6" pageNumber="229">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<emphasis id="6CFAEABE2075FFA82F2616CFFC51419F" box="[827,1022,1931,1957]" italics="true" pageId="6" pageNumber="229">A.F.G. Kerr 0736</emphasis>
|
||
,
|
||
<date id="2A30106C2075FFA8281116CFFB18419F" box="[1036,1207,1931,1957]" pageId="6" pageNumber="229" value="1929-03-21">21 March 1929</date>
|
||
(K s.n.!,
|
||
<bibRefCitation id="3A1F4B5D2075FFA8293C16CFFF5641F3" author="Seidenfaden, G." pageId="6" pageNumber="229" pagination="1 - 398" refId="ref25710" refString="Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine vandoid genera. Opera Botanica 95: 1 - 398." type="journal article" year="1988">Seidenfaden 1988: 166</bibRefCitation>
|
||
as
|
||
<taxonomicName id="998E4D2F2075FFA82D0316F4FE6941F3" box="[286,454,1967,1993]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2075FFA82D0316F4FE6941F3" box="[286,454,1967,1993]" italics="true" pageId="6" pageNumber="229">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="169465272075FFA92CA01697FAB542F7" lastPageId="7" lastPageNumber="230" pageId="6" pageNumber="229" type="discussion">
|
||
<paragraph id="5E3136AC2075FFA92CA01697FBE34756" blockId="6.[136,1452,1571,2065]" lastBlockId="7.[136,1452,158,1229]" lastPageId="7" lastPageNumber="230" pageId="6" pageNumber="229">
|
||
Taxonomic notes:—
|
||
<taxonomicName id="998E4D2F2075FFA82DBF1697FD6341D7" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[418,716,2003,2029]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="229" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2075FFA82DBF1697FD6341D7" box="[418,716,2003,2029]" italics="true" pageId="6" pageNumber="229">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
may be recognised by the combination of the obliquely obovate lateral lobes of the lip surpassing the triangular, concave and emarginate mid-lobe, the long clavate hairs on the lip, yellow flowers with a white lip, and orange warts inside the spur. It is morphologically similar to
|
||
<taxonomicName id="998E4D2F2074FFA928DB11DBFAC24682" authorityName="Schltr. (Schlechter" authorityYear="1919" baseAuthorityName="Hayata" baseAuthorityYear="1911" box="[1222,1389,158,184]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="formosanum">
|
||
<emphasis id="6CFAEABE2074FFA928DB11DBFAC24682" box="[1222,1389,158,184]" italics="true" pageId="7" pageNumber="230">T. formosanum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CFAEABE2074FFA9296911DAFF7E46E6" bold="true" pageId="7" pageNumber="230">
|
||
(
|
||
<figureCitation id="C6B52A292074FFA9296111DAFF6646E6" captionStart="FIGURE 4" captionStartId="6.[136,234,1430,1452]" captionTargetBox="[169,1418,196,1400]" captionTargetId="figure-22@6.[164,1423,190,1406]" captionTargetPageId="6" captionText="FIGURE 4. A comparison of allied species. A–B. Thrixspermum alboluteum. C–D. Thrixspermum patkaiense. E–F. Thrixspermum formosanum. Photographed by Wattana Tanming (A–B. Cultivation, QSBG), Nopparut Toolmal (C–D. In situ) & Morris Ng (E–F. Cultivation, Taiwan). Illustration assembled by Yongyuth Rattanasootr." figureDoi="http://doi.org/10.5281/zenodo.8141774" httpUri="https://zenodo.org/record/8141774/files/figure.png" pageId="7" pageNumber="230">Fig. 4E–F</figureCitation>
|
||
)
|
||
</emphasis>
|
||
, a species from
|
||
<taxonomicName id="998E4D2F2074FFA92D631186FE1546E6" box="[382,442,194,220]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="262" phylum="Tracheophyta" rank="genus">India</taxonomicName>
|
||
, southern
|
||
<collectingCountry id="2699763C2074FFA92E351186FDC146E6" box="[552,622,194,220]" name="China" pageId="7" pageNumber="230">China</collectingCountry>
|
||
, North
|
||
<collectingCountry id="2699763C2074FFA92EA61186FCB546E6" box="[699,794,194,220]" name="Vietnam" pageId="7" pageNumber="230">Vietnam</collectingCountry>
|
||
and
|
||
<collectingCountry id="2699763C2074FFA92F561186FC3246E6" box="[843,925,194,220]" name="Taiwan" pageId="7" pageNumber="230">Taiwan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="3A1F4B5D2074FFA92FB41186FBF246E6" author="Lin, T. P. & Hsu, C. C." box="[937,1117,194,221]" pageId="7" pageNumber="230" pagination="59 - 72" refId="ref24868" refString="Lin, T. P. & Hsu, C. C. (1977) The genus Thrixspermum Lour. of Taiwan (Orchidaceae). Taiwania 22 (1): 59 - 72." type="journal article" year="1977">Lin & Hsu 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3A1F4B5D2074FFA9287B1186FAB546E6" author="Chen, S. C. & Liu, Z. J. & Zhu, G. H. & Lang, K. Y. & Tsi, Z. H. & Luo, Y. B. & Jin, X. H. & Cribb, P. J. & Wood, J. J. & Gale, S. W. & Ormerod, P. & Vermeulen, J. J. & Wood, H. P. & Clayton, D. & Bell, A." box="[1126,1306,194,221]" pageId="7" pageNumber="230" refId="ref23617" refString="Chen, S. C., Liu, Z. J., Zhu, G. H., Lang, K. Y., Tsi, Z. H., Luo, Y. B., Jin, X. H., Cribb, P. J., Wood, J. J., Gale, S. W., Ormerod, P., Vermeulen, J. J., Wood, H. P., Clayton, D. & Bell, A. (2009) Orchidaceae. In: Wu, Z. Y., Raven, P. H. & Hong, D. Y. (eds.) Flora of China, vol. 25. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis, 570 pp." type="book" year="2009">
|
||
Chen
|
||
<emphasis id="6CFAEABE2074FFA928B81187FB7B46E6" box="[1189,1236,194,220]" italics="true" pageId="7" pageNumber="230">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
, the Editorial Committee
|
||
<bibRefCitation id="3A1F4B5D2074FFA92D1611A2FDEE473A" author="Editorial Committee of the Flora of Taiwan" box="[267,577,230,256]" pageId="7" pageNumber="230" refId="ref23902" refString="Editorial Committee of the Flora of Taiwan. (2000) Flora of Taiwan, vol. 5 Angiosperms Monocotyledon s (2 nd ed.). Department of Botany, National Taiwan University, Taipei, Taiwan, 1143 pp." type="book" year="2000">
|
||
of the Flora of
|
||
<collectingCountry id="2699763C2074FFA92DB311A2FDAF473A" box="[430,512,230,256]" name="Taiwan" pageId="7" pageNumber="230">Taiwan</collectingCountry>
|
||
2000
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3A1F4B5D2074FFA92E5111A2FCB9473A" author="Lin, W. M. & Wang, Y. F." box="[588,790,230,257]" pageId="7" pageNumber="230" refId="ref24905" refString="Lin, W. M. & Wang, Y. F. (2014) The wild orchids of Taiwan an illustrated guide. KBCC press, Taiwan, 923 pp." type="book" year="2014">Lin & Wang 2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3A1F4B5D2074FFA92F3D11A2FB85473B" author="Gogoi, K. & Yonzone, R." box="[800,1066,230,257]" pageId="7" pageNumber="230" pagination="11 - 16" refId="ref24043" refString="Gogoi, K. & Yonzone, R. (2015) Thrixspermum formosanum (Hayata) Schltr. (Orchidaceae) from Karbi Anglong (Assam): A new record for India. The MIOS Journal 16 (1): 11 - 16." type="journal article" year="2015">Gogoi & Yonzone 2015</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3A1F4B5D2074FFA9282911A2FA08473A" author="Kaliamoorthy, S. & Saravanan, T. S." box="[1076,1447,230,256]" pageId="7" pageNumber="230" pagination="319 - 322" refId="ref24671" refString="Kaliamoorthy, S. & Saravanan, T. S. (2019) Additions to the orchid flora of Peninsular India. Rheedea 29 (4): 319 - 322. https: // dx. doi. org / 10.22244 / rheedea. 2019.29.4.05" type="journal article" year="2019">Kaliamoorthy & Saravanan 2019</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3A1F4B5D2074FFA92C95104EFEF3471E" author="Khanal, M. & Chettri, M. & Kumar, D. & Chhetri, P. & Rai, S. K." box="[136,348,266,292]" pageId="7" pageNumber="230" pagination="345 - 347" refId="ref24740" refString="Khanal, M., Chettri, M., Kumar, D., Chhetri, P. & Rai, S. K. (2022) Thrixspermum formosanum (Hayata) Schltr. (Orchidaceae) - A new record for the state flora of Sikkim, India. Pleione 16 (3): 345 - 347." type="journal article" year="2022">
|
||
Khanal
|
||
<emphasis id="6CFAEABE2074FFA92CFD104FFEB6471E" box="[224,281,266,292]" italics="true" pageId="7" pageNumber="230">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
), because of its small stature with oblong or lanceolate to linear leaves, inflorescences longer than the leaves, and sepals and petals similarly shaped; it differs in having yellow flowers, a lip with brownish orange spots and orange warts, a distinct mid-lobe with emarginate apex, and obovate lateral lobes.
|
||
</paragraph>
|
||
<paragraph id="5E3136AC2074FFA92CA01032FAD5455E" blockId="7.[136,1452,158,1229]" pageId="7" pageNumber="230">
|
||
Having critically compared the original descriptions and the
|
||
<typeStatus id="8135880E2074FFA92F601033FC0247AB" box="[893,941,375,401]" pageId="7" pageNumber="230">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="998E4D2F2074FFA9284A1033FAAD47AA" authorityName="Schltr. (Schlechter" authorityYear="1919" baseAuthorityName="Hayata" baseAuthorityYear="1911" box="[1111,1282,374,400]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="formosanum">
|
||
<emphasis id="6CFAEABE2074FFA9284A1033FAAD47AA" box="[1111,1282,374,400]" italics="true" pageId="7" pageNumber="230">T. formosanum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="6CFAEABE2074FFA929081033FA0347AA" box="[1301,1452,374,400]" italics="true" pageId="7" pageNumber="230">T. Kawakami</emphasis>
|
||
&
|
||
<emphasis id="6CFAEABE2074FFA92CBB10DFFE88478E" box="[166,295,411,436]" italics="true" pageId="7" pageNumber="230">N. Mori s.n</emphasis>
|
||
.,
|
||
<collectingCountry id="2699763C2074FFA92D2010DFFE0D478F" box="[317,418,411,437]" name="Taiwan" pageId="7" pageNumber="230">Formosa</collectingCountry>
|
||
, 1907, K 000891322!,
|
||
<typeStatus id="8135880E2074FFA92EBA10DFFD78478F" box="[679,727,411,437]" pageId="7" pageNumber="230">type</typeStatus>
|
||
of
|
||
<taxonomicName id="998E4D2F2074FFA92EE310DEFBCB478E" authority="Hayata" authorityName="Hayata" box="[766,1124,410,436]" class="Liliopsida" family="Orchidaceae" genus="Sarcochilus" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="formosanus">
|
||
<emphasis id="6CFAEABE2074FFA92EE310DEFBA3478E" box="[766,1036,410,436]" italics="true" pageId="7" pageNumber="230">Sarcochilus formosanus</emphasis>
|
||
Hayata
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="998E4D2F2074FFA928B910DFFAE3478E" box="[1188,1356,410,436]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2074FFA928B910DFFAE3478E" box="[1188,1356,410,436]" italics="true" pageId="7" pageNumber="230">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="6CFAEABE2074FFA9294310DEFF7047E2" italics="true" pageId="7" pageNumber="230">Joseph 45621A</emphasis>
|
||
, between Umran & Umsaw beside the Gauhati-Shillong highway, Nongpoh,
|
||
<collectingRegion id="9C4AF84E2074FFA9285F10FAFB6E47E2" box="[1090,1217,446,472]" country="India" name="Meghalaya" pageId="7" pageNumber="230">Meghalaya</collectingRegion>
|
||
,
|
||
<taxonomicName id="998E4D2F2074FFA928D110FAFAA747E2" box="[1228,1288,446,472]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="262" phylum="Tracheophyta" rank="genus">India</taxonomicName>
|
||
,
|
||
<date id="2A30106C2074FFA9290910FAFA0847E2" box="[1300,1447,446,472]" pageId="7" pageNumber="230" value="1966-07-23">23 July 1966</date>
|
||
,
|
||
<collectingRegion id="9C4AF84E2074FFA92C9510A6FF4747C6" box="[136,232,482,508]" country="India" name="Assam" pageId="7" pageNumber="230">ASSAM</collectingRegion>
|
||
0000000209!, image seen) and the relevant literature of both species, we conclude that the two taxa are conspecific; they agree in being small plants with stems
|
||
<quantity id="99769B492074FFA92F181342FCC1441B" box="[773,878,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="7" pageNumber="230" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
|
||
long, white flowers 1.0–
|
||
<quantity id="99769B492074FFA928941342FB7B441B" box="[1161,1236,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="230" unit="cm" value="1.2">1.2 cm</quantity>
|
||
across, an obovate to elliptic dorsal-sepal, ovate lateral-sepals, oblanceolate petals, a white lip, marked with pinkish to brownish stripes, triangular lateral lobes, and an indistinct mid-lobe with orangish markings and glandular hairs. In
|
||
<taxonomicName id="998E4D2F2074FFA928D4130AFAAC4452" box="[1225,1283,590,616]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="262" phylum="Tracheophyta" rank="genus">India</taxonomicName>
|
||
and
|
||
<collectingCountry id="2699763C2074FFA92925130AFA234452" box="[1336,1420,590,616]" name="Bhutan" pageId="7" pageNumber="230">Bhutan</collectingCountry>
|
||
,
|
||
<taxonomicName id="998E4D2F2074FFA9298A130BFEBB44B6" authorityName="Schltr. (Schlechter" authorityYear="1919" baseAuthorityName="Hayata" baseAuthorityYear="1911" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="formosanum">
|
||
<emphasis id="6CFAEABE2074FFA9298A130BFEBB44B6" italics="true" pageId="7" pageNumber="230">T. formosanum</emphasis>
|
||
</taxonomicName>
|
||
has often been identified as
|
||
<taxonomicName id="998E4D2F2074FFA92E4C1337FD5744B6" box="[593,760,626,652]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2074FFA92E4C1337FD5744B6" box="[593,760,626,652]" italics="true" pageId="7" pageNumber="230">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="3A1F4B5D2074FFA92F1B1336FC4A44B6" author="Rao, A. S. & Joseph, J." box="[774,997,626,652]" pageId="7" pageNumber="230" pagination="123 - 125" refId="ref25459" refString="Rao, A. S. & Joseph, J. (1969) A new orchid species from K. & J. Hills, Assam - Thrixspermum muscaeflorum A. S. Rao & Joseph. Bulletin of the Botanical Survey of India 11 (1 - 2): 123 - 125." type="journal article" year="1969">Rao & Joseph 1969</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3A1F4B5D2074FFA92FED1336FBD244B6" author="Kataki, S. K." box="[1008,1149,626,652]" pageId="7" pageNumber="230" refId="ref24718" refString="Kataki, S. K. (1986) Orchids of Meghalaya. Government of Meghalaya, Shillong, 258 pp." type="book" year="1986">Kataki 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3A1F4B5D2074FFA928951336FAE344B6" author="Chowdhery, H. J." box="[1160,1356,626,652]" pageId="7" pageNumber="230" refId="ref23771" refString="Chowdhery, H. J. (1998) Orchid Flora of Arunachal Pradesh. Bishen Singh Mahendra Pal Singh, India, 824 pp." type="book" year="1998">Chowdhery 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3A1F4B5D2074FFA9294A1337FECB448A" author="Bose, T. K. & Bhattacharjee, S. K. & Das, P. & Basak, U. C." pageId="7" pageNumber="230" refId="ref23479" refString="Bose, T. K., Bhattacharjee, S. K., Das, P. & Basak, U. C. (1999) Orchids of India, Second Edition. Naya Prokash, Calcutta, India, 570 pp." type="book" year="1999">Bose & Bhattacharjee 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3A1F4B5D2074FFA92D6D13D2FDF7448A" author="Dalstrom, S. & Gyeltshen, C. & Gyeltshen, N." box="[368,600,662,688]" pageId="7" pageNumber="230" refId="ref23851" refString="Dalstrom, S., Gyeltshen, C. & Gyeltshen, N. (2017) A Century of New Orchid Records in Bhutan. The National Biodiversity Centre, Ministry of Agriculture and Forests, Royal Government of Bhutan, Serbithang, Thimphu, Bhutan, 210 pp." type="book" year="2017">
|
||
Dalström
|
||
<emphasis id="6CFAEABE2074FFA92DC213D3FDB8448A" box="[479,535,662,688]" italics="true" pageId="7" pageNumber="230">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3A1F4B5D2074FFA92E7913D2FD48448A" author="Misra, S." box="[612,743,662,688]" pageId="7" pageNumber="230" refId="ref25016" refString="Misra, S. (2019) Orchids of India; A Handbook. Bishen Singh Mahendra Pal Singh, Dehra Dun, India, 652 pp." type="book" year="2019">Misra 2019</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3A1F4B5D2074FFA92EEE13D2FC46448A" author="Barbhuiya, H. A. & Verma, D. & Dey, S. & Salunkhe, C. K." box="[755,1001,662,688]" pageId="7" pageNumber="230" refId="ref23308" refString="Barbhuiya, H. A., Verma, D., Dey, S. & Salunkhe, C. K. (2021) An Illustrated Guide to the Orchids of Assam. Balipara foundation, Assam, India, 806 pp." type="book" year="2021">
|
||
Barbhuiya
|
||
<emphasis id="6CFAEABE2074FFA92F7213D3FC08448A" box="[879,935,662,688]" italics="true" pageId="7" pageNumber="230">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
). We also find no significant differences between
|
||
<taxonomicName id="998E4D2F2074FFA92CF313FFFE3844EE" authorityName="Schltr. (Schlechter" authorityYear="1919" baseAuthorityName="Hayata" baseAuthorityYear="1911" box="[238,407,698,724]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="formosanum">
|
||
<emphasis id="6CFAEABE2074FFA92CF313FFFE3844EE" box="[238,407,698,724]" italics="true" pageId="7" pageNumber="230">T. formosanum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="998E4D2F2074FFA92DCD13FFFBDC44EE" authority="Joseph & Vajravelu (1981)" authorityName="Joseph & Vajravelu" authorityYear="1981" box="[464,1139,698,725]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="variety" species="musciflorum" variety="nilagiricum">
|
||
<emphasis id="6CFAEABE2074FFA92DCD13FFFDD644EE" box="[464,633,698,724]" italics="true" pageId="7" pageNumber="230">T. musciflorum</emphasis>
|
||
var.
|
||
<emphasis id="6CFAEABE2074FFA92EAE13FEFC9944EE" box="[691,822,698,724]" italics="true" pageId="7" pageNumber="230">nilagiricum</emphasis>
|
||
<bibRefCitation id="3A1F4B5D2074FFA92F2313FEFBDC44EE" author="Joseph, J. & Vajravelu, E." box="[830,1139,698,724]" pageId="7" pageNumber="230" pagination="648 - 651" refId="ref24629" refString="Joseph, J. & Vajravelu, E. (1981) Thrixspermum muscaeflorum var. nilagiricum Joseph & Vajravelu, a new variety from Nilgiri Hills, South India. Indian Forester 107 (10): 648 - 651." type="journal article" year="1981">Joseph & Vajravelu (1981)</bibRefCitation>
|
||
</taxonomicName>
|
||
. In contrast, the species we here describe as
|
||
<taxonomicName id="998E4D2F2074FFA92D23139BFE7844C2" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[318,471,734,760]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2074FFA92D23139BFE7844C2" box="[318,471,734,760]" italics="true" pageId="7" pageNumber="230">T. alboluteum</emphasis>
|
||
</taxonomicName>
|
||
had been misidentified as
|
||
<taxonomicName id="998E4D2F2074FFA92EE0139BFC0C44C2" box="[765,931,734,760]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2074FFA92EE0139BFC0C44C2" box="[765,931,734,760]" italics="true" pageId="7" pageNumber="230">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="3A1F4B5D2074FFA92FD6139AFB3944C2" author="Seidenfaden, G." box="[971,1174,734,760]" pageId="7" pageNumber="230" pagination="1 - 398" refId="ref25710" refString="Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine vandoid genera. Opera Botanica 95: 1 - 398." type="journal article" year="1988">Seidenfaden 1988</bibRefCitation>
|
||
. His drawing, taxonomic notes and photographs (C-SEIDENFADEN no. 24-157!, 24-158!, 24-159!, 24-161!, 24-162!, 24-163!, 24-192!, 24- 193!, 24-194!) perfectly match our plant from Kaeng Krachan National Park. We also note that the species illustrated as
|
||
<taxonomicName id="998E4D2F2074FFA92CBB120FFEE1455E" box="[166,334,842,868]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="musciflorum">
|
||
<emphasis id="6CFAEABE2074FFA92CBB120FFEE1455E" box="[166,334,842,868]" italics="true" pageId="7" pageNumber="230">T. musciflorum</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="3A1F4B5D2074FFA92D6F120EFDD2455E" author="Singh, S. K. & Agrawala, D. K. & Jalal, J. S. & Dash, S. S. & Mao, A. A. & Singh, P." box="[370,637,842,869]" pageId="7" pageNumber="230" refId="ref25814" refString="Singh, S. K., Agrawala, D. K., Jalal, J. S., Dash, S. S., Mao, A. A. & Singh, P. (2019) Orchids of India - A Pictorial Guide. Botanical Survey of India, Kolkata, 547 pp." type="book" year="2019">
|
||
Singh
|
||
<emphasis id="6CFAEABE2074FFA92DA7120FFE44455E" box="[442,491,842,868]" italics="true" pageId="7" pageNumber="230">et al</emphasis>
|
||
. (2019: 492)
|
||
</bibRefCitation>
|
||
is
|
||
<taxonomicName id="998E4D2F2074FFA92E80120FFAD9455E" authority="Vik. Kumar, D. Verma & A. N. Rao (Kumar et al. 2017)" box="[669,1398,842,869]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="indicum">
|
||
<emphasis id="6CFAEABE2074FFA92E80120FFCBC455E" box="[669,787,842,868]" italics="true" pageId="7" pageNumber="230">T. indicum</emphasis>
|
||
Vik.Kumar, D.Verma & A.N.Rao (
|
||
<bibRefCitation id="3A1F4B5D2074FFA92881120EFAC1455E" author="Kumar, V. & Verma, D. & Rao, A. N." box="[1180,1390,842,869]" pageId="7" pageNumber="230" pagination="79 - 84" refId="ref24802" refString="Kumar, V., Verma, D. & Rao, A. N. (2017) Thrixspermum indicum (Orchidaceae), a new species from Northeast India. Phytotaxa 292: 79 - 84." type="journal article" year="2017">
|
||
Kumar
|
||
<emphasis id="6CFAEABE2074FFA928EF120FFA8B455E" box="[1266,1316,842,868]" italics="true" pageId="7" pageNumber="230">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5E3136AC2074FFA92CA0122AFAB542F7" blockId="7.[136,1452,158,1229]" pageId="7" pageNumber="230">
|
||
<taxonomicName id="998E4D2F2074FFA92CA0122AFE4945B2" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[189,486,878,904]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2074FFA92CA0122AFE4945B2" box="[189,486,878,904]" italics="true" pageId="7" pageNumber="230">Thrixspermum alboluteum</emphasis>
|
||
</taxonomicName>
|
||
is also similar to
|
||
<taxonomicName id="998E4D2F2074FFA92EB2122BFC1C45B2" authority="K. Gogoi" authorityName="K. Gogoi" box="[687,947,878,905]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="patkaiense">
|
||
<emphasis id="6CFAEABE2074FFA92EB2122BFCEC45B2" box="[687,835,878,904]" italics="true" pageId="7" pageNumber="230">T. patkaiense</emphasis>
|
||
K. Gogoi
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="3A1F4B5D2074FFA92FDE122AFBC645B2" author="Gogoi, K." box="[963,1129,878,904]" pageId="7" pageNumber="230" pagination="9 - 16" refId="ref24090" refString="Gogoi, K. (2022) Thrixspermum patkaiensis sp. nov. (Vandae: Aeridinae) from Assam, Northeast India. Lankesteriana 22 (1): 9 - 16. https: // doi. org / 10.15517 / lank. v 22 i 1.50589" type="journal article" year="2022">Gogoi 2022: 9</bibRefCitation>
|
||
, as ‘
|
||
<taxonomicName id="998E4D2F2074FFA92883122AFAB345B2" box="[1182,1308,878,904]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="patkaiensis">
|
||
<emphasis id="6CFAEABE2074FFA92883122AFAB345B2" box="[1182,1308,878,904]" italics="true" pageId="7" pageNumber="230">patkaiensis</emphasis>
|
||
</taxonomicName>
|
||
’), a recently described species from north-eastern
|
||
<taxonomicName id="998E4D2F2074FFA92E2812D6FDDE4596" box="[565,625,914,940]" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="262" phylum="Tracheophyta" rank="genus">India</taxonomicName>
|
||
, which also occurs in central
|
||
<collectingCountry id="2699763C2074FFA92FD612D7FBAE4597" box="[971,1025,915,941]" name="Laos" pageId="7" pageNumber="230">Laos</collectingCountry>
|
||
(
|
||
<bibRefCitation id="3A1F4B5D2074FFA9280E12D6FB7C4596" author="Averyanov, L. V. & Averyanova, A. L. & Sinh, N. K. & Orlov, N. L. & Maisak, T. V. & Tien, N. H." box="[1043,1235,914,940]" pageId="7" pageNumber="230" pagination="24 - 41" refId="ref23165" refString="Averyanov, L. V., Averyanova, A. L., Sinh, N. K., Orlov, N. L., Maisak, T. V. & Tien, N. H. (2018) New and rare orchid species (Orchidaceae) in the flora of Cambodia and Laos. Novitates Systematicae Plantarum Vascularium 49: 24 - 41. https: // doi. org / 10.31111 / novitates / 2018.49.24" type="journal article" year="2018">Averyanov 2018</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="998E4D2F2074FFA9291F12D7FAC34596" box="[1282,1388,914,940]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="hystrix">
|
||
<emphasis id="6CFAEABE2074FFA9291F12D7FAC34596" box="[1282,1388,914,940]" italics="true" pageId="7" pageNumber="230">T. hystrix</emphasis>
|
||
</taxonomicName>
|
||
auct. non (Blume) Rchb.f.),
|
||
<collectingCountry id="2699763C2074FFA92D9712F3FE5845EB" box="[394,503,951,977]" name="Myanmar" pageId="7" pageNumber="230">Myanmar</collectingCountry>
|
||
(photos by H. Kurzweil seen), and
|
||
<collectingCountry id="2699763C2074FFA92F9112F2FC5F45EA" box="[908,1008,950,976]" name="Thailand" pageId="7" pageNumber="230">Thailand</collectingCountry>
|
||
(pers. obs. Nopparut Toolmal). Like
|
||
<taxonomicName id="998E4D2F2074FFA9298A12F3FEA945CE" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2074FFA9298A12F3FEA945CE" italics="true" pageId="7" pageNumber="230">T. alboluteum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="998E4D2F2074FFA92D09129FFE0745CE" box="[276,424,986,1012]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="patkaiense">
|
||
<emphasis id="6CFAEABE2074FFA92D09129FFE0745CE" box="[276,424,986,1012]" italics="true" pageId="7" pageNumber="230">T. patkaiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CFAEABE2074FFA92DB3129EFD9545CE" bold="true" box="[430,570,986,1012]" pageId="7" pageNumber="230">
|
||
(
|
||
<figureCitation id="C6B52A292074FFA92DAB129EFD9C45CE" box="[438,563,986,1012]" captionStart="FIGURE 4" captionStartId="6.[136,234,1430,1452]" captionTargetBox="[169,1418,196,1400]" captionTargetId="figure-22@6.[164,1423,190,1406]" captionTargetPageId="6" captionText="FIGURE 4. A comparison of allied species. A–B. Thrixspermum alboluteum. C–D. Thrixspermum patkaiense. E–F. Thrixspermum formosanum. Photographed by Wattana Tanming (A–B. Cultivation, QSBG), Nopparut Toolmal (C–D. In situ) & Morris Ng (E–F. Cultivation, Taiwan). Illustration assembled by Yongyuth Rattanasootr." figureDoi="http://doi.org/10.5281/zenodo.8141774" httpUri="https://zenodo.org/record/8141774/files/figure.png" pageId="7" pageNumber="230">Fig. 4C–D</figureCitation>
|
||
)
|
||
</emphasis>
|
||
has yellow tepals but it has a yellow lip marked with longitudinal purplish red stripes on both surfaces of the lateral lobes towards the spur, the mid-lobe yellowish with an orange band along the apical margins, and a callus that is distinctly raised at the apex (vs. a white lip with different markings, see description, and a nearly flat callus). In
|
||
<taxonomicName id="998E4D2F2074FFA92DD91503FDF0425A" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[452,607,1094,1120]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2074FFA92DD91503FDF0425A" box="[452,607,1094,1120]" italics="true" pageId="7" pageNumber="230">T. alboluteum</emphasis>
|
||
</taxonomicName>
|
||
the lateral lobes are obovate and broadly rounded (vs. broadly ovate), the mid-lobe is densely covered with long capitate hairs on the adaxial surface and along the margin but not abaxially (vs. covered on both surfaces). The spur of
|
||
<taxonomicName id="998E4D2F2074FFA92E6315CBFCB54292" authorityName="Toolmal, Schuit. & Culham" authorityYear="2023" box="[638,794,1166,1192]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="alboluteum">
|
||
<emphasis id="6CFAEABE2074FFA92E6315CBFCB54292" box="[638,794,1166,1192]" italics="true" pageId="7" pageNumber="230">T. alboluteum</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="998E4D2F2074FFA9281A15CBFB334292" box="[1031,1180,1166,1192]" class="Liliopsida" family="Orchidaceae" genus="Thrixspermum" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="230" phylum="Tracheophyta" rank="species" species="patkaiense">
|
||
<emphasis id="6CFAEABE2074FFA9281A15CBFB334292" box="[1031,1180,1166,1192]" italics="true" pageId="7" pageNumber="230">T. patkaiense</emphasis>
|
||
</taxonomicName>
|
||
in being proportionally narrower, 2.0–2.2 ×
|
||
<quantity id="99769B492074FFA92D7115F6FE5C42F7" box="[364,499,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="7" pageNumber="230" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
|
||
(vs. 2.5 × 3.0–
|
||
<quantity id="99769B492074FFA92E8615F6FD5A42F6" box="[667,757,1202,1228]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="230" unit="mm" value="4.2">4.2 mm</quantity>
|
||
). Both species have orange warts inside the spur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |