treatments-xml/data/CC/2C/3C/CC2C3C1FF753FFBDFF47F949FC05FA8D.xml

<document id="AA4514C734BA1E918038249F61C61E63" ID-CLB-Dataset="29049" ID-DOI="10.11646/zootaxa.4457.4.10" ID-GBIF-Dataset="3d9c7c7e-1cb9-4035-bc5a-5e26e2108d02" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457993" ID-ZooBank="6E360D2D-C969-4593-B25A-56B7F6086479" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539267563033" checkinUser="plazi" docAuthor="Kalawate, Aparna &amp; Hillert, Oliver" docDate="2018" docId="CC2C3C1FF753FFBDFF47F949FC05FA8D" docLanguage="en" docName="zootaxa.4457.4.10.pdf" docOrigin="Zootaxa 4457 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolboceras sahyadriensis Kalawate &amp; Hillert 2018, new species" docType="treatment" docUuid="AACC6761-BF45-4606-AA24-7DDFD2EDDDDB" docUuidSource="ZooBank" docVersion="8" lastPageNumber="598" masterDocId="30154467F753FFBEFFD0FFF4FF99FFE1" masterDocTitle="A new species and new records of the genus Bolboceras Kirby, 1819 (Coleoptera: Geotrupidae: Bolboceratinae) from India" masterLastPageNumber="599" masterPageNumber="595" pageNumber="595" updateTime="1698577153616" updateUser="ExternalLinkService">
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<mods:title id="492E531E74329E5C669D0203EB94399E">A new species and new records of the genus Bolboceras Kirby, 1819 (Coleoptera: Geotrupidae: Bolboceratinae) from India</mods:title>
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<mods:namePart id="6351732AA6658BC64FDE7216EB2A9D97">Kalawate, Aparna</mods:namePart>
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<heading id="1F723A65F753FFBEFF47F949FB9EF939" bold="true" box="[151,1031,1725,1752]" fontSize="11" level="1" pageId="0" pageNumber="595" reason="1">
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<taxonomicName id="8385F68AF753FFBEFF47F949FD5DF939" ID-CoL="MBDM" authority="Kalawate &amp; Hillert" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[151,708,1725,1752]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="595" phylum="Arthropoda" rank="species" species="sahyadriensis" status="sp. nov.">
<emphasis id="76F1511BF753FFBEFF47F949FE50F936" bold="true" box="[151,457,1725,1751]" italics="true" pageId="0" pageNumber="595">Bolboceras sahyadriensis</emphasis>
Kalawate &amp; Hillert
</taxonomicName>
, 
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(
<figureCitation id="DCBE918CF753FFBEFCA0F94AFC66F939" box="[880,1023,1726,1752]" captionStart="FIGURE 2" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[151,1424,362,1892]" captionTargetId="figure@2.[151,1436,350,1910]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Bolboceras sahyadriensis Kalawate &amp; Hillert, new species. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H)." httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="0" pageNumber="595">Figs. 2A–H</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<paragraph id="443A8D09F753FFBEFF47F914FBF9F916" blockId="0.[151,1436,1725,2026]" box="[151,1120,1760,1783]" pageId="0" pageNumber="595">
<materialsCitation id="F4ED8754F753FFBEFF47F914FBF9F916" ID-GBIF-Occurrence="1917076320" box="[151,1120,1760,1783]" country="India" elevation="731" latitude="17.229" location="Bhosgaon" longLatPrecision="76" longitude="73.952" municipality="Satara District" pageId="0" pageNumber="595" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
<emphasis id="76F1511BF753FFBEFF47F914FEB5F916" bold="true" box="[151,300,1760,1783]" pageId="0" pageNumber="595">
<typeStatus id="9B3E33ABF753FFBEFF47F914FF56F916" box="[151,207,1760,1783]" pageId="0" pageNumber="595">Type</typeStatus>
locality.
</emphasis>
<location id="415ADBD2F753FFBEFEE4F914FE38F916" LSID="urn:lsid:plazi:treatment:CC2C3C1FF753FFBDFF47F949FC05FA8D:415ADBD2F753FFBEFEE4F914FE38F916" box="[308,417,1760,1783]" country="India" latitude="17.229" longLatPrecision="76" longitude="73.952" municipality="Satara District" name="Bhosgaon" pageId="0" pageNumber="595" stateProvince="Maharashtra">Bhosgaon</location>
, 
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, 
<collectingRegion id="864143EBF753FFBEFD82F914FD43F916" box="[594,730,1760,1783]" country="India" name="Maharashtra" pageId="0" pageNumber="595">Maharashtra</collectingRegion>
, 
<collectingCountry id="3C92CD99F753FFBEFD35F914FC85F916" box="[741,796,1760,1783]" name="India" pageId="0" pageNumber="595">India</collectingCountry>
[
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, 
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, 
<elevation id="CFA86A3AF753FFBEFBDCF914FBCCF916" box="[1036,1109,1760,1783]" metricMagnitude="2" metricUnit="m" metricValue="7.31" pageId="0" pageNumber="595" unit="m" value="731.0">
<quantity id="837D20ECF753FFBEFBDCF914FBCCF916" box="[1036,1109,1760,1783]" metricMagnitude="2" metricUnit="m" metricValue="7.31" pageId="0" pageNumber="595" unit="m" value="731.0">731 m</quantity>
</elevation>
].
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</paragraph>
<paragraph id="443A8D09F753FFBEFF17F8F7FF68F863" blockId="0.[151,1436,1725,2026]" pageId="0" pageNumber="595">
<materialsCitation id="F4ED8754F753FFBEFF17F8F7FE75F8BE" ID-GBIF-Occurrence="1917076316" collectingDate="2017-07-15" country="India" county="Forest Rest House" elevation="731" latitude="17.229" location="Patan" longLatPrecision="76" longitude="73.952" municipality="Bhosgaon" pageId="0" pageNumber="595" specimenCount="2" stateProvince="Maharashtra" typeStatus="holotype">
<emphasis id="76F1511BF753FFBEFF17F8F7FEE9F8FB" bold="true" box="[199,368,1795,1818]" pageId="0" pageNumber="595">
<typeStatus id="9B3E33ABF753FFBEFF17F8F7FF66F8FB" box="[199,255,1795,1818]" pageId="0" pageNumber="595">Type</typeStatus>
material.
</emphasis>
<typeStatus id="9B3E33ABF753FFBEFEAAF8F7FE79F8FB" box="[378,480,1795,1818]" pageId="0" pageNumber="595" type="holotype">Holotype</typeStatus>
, male, labeled: “ 
<collectingCountry id="3C92CD99F753FFBEFD4BF8F7FD4FF8FB" box="[667,726,1795,1818]" name="India" pageId="0" pageNumber="595">India</collectingCountry>
, 
<collectingCounty id="AD5BF585F753FFBEFD34F8F7FC28F8FB" box="[740,945,1795,1818]" pageId="0" pageNumber="595">Forest Rest House</collectingCounty>
, near 
<collectingMunicipality id="A45E1773F753FFBEFC27F8F7FBFDF8FB" box="[1015,1124,1795,1818]" pageId="0" pageNumber="595">Bhosgaon</collectingMunicipality>
, 
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taluk, 
<location id="415ADBD2F753FFBEFB2EF8F7FA01F8FB" LSID="urn:lsid:plazi:treatment:CC2C3C1FF753FFBDFF47F949FC05FA8D:415ADBD2F753FFBEFB2EF8F7FA01F8FB" box="[1278,1432,1795,1818]" country="India" county="Forest Rest House" latitude="17.229" longLatPrecision="76" longitude="73.952" municipality="Bhosgaon" name="Satara district" pageId="0" pageNumber="595" stateProvince="Maharashtra">Satara district</location>
, 
<collectingRegion id="864143EBF753FFBEFF47F8D2FE85F8DC" box="[151,284,1830,1853]" country="India" name="Maharashtra" pageId="0" pageNumber="595">Maharashtra</collectingRegion>
(
<geoCoordinate id="21B1EBCEF753FFBEFEFFF8D2FE01F8DC" box="[303,408,1830,1853]" direction="north" orientation="latitude" pageId="0" pageNumber="595" precision="55" value="17.229">17.229°N</geoCoordinate>
, 
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; altitude 
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<quantity id="837D20ECF753FFBEFDAAF8D2FD5FF8DC" box="[634,710,1830,1853]" metricMagnitude="2" metricUnit="m" metricValue="7.31" pageId="0" pageNumber="595" unit="m" value="731.0">731 m</quantity>
</elevation>
), 
<collectingDate id="207F5221F753FFBEFD0EF8D2FCECF8DC" box="[734,885,1830,1853]" pageId="0" pageNumber="595" value="2017-07-15">
<date id="303BABC9F753FFBEFD0EF8D2FCECF8DC" box="[734,885,1830,1853]" pageId="0" pageNumber="595" value="2017-07-15">15 July 2017</date>
</collectingDate>
, Aparna Kalawate lgt.” (deposited in ZSI-WRC, catalogue number ENT-1/2916)
</materialsCitation>
; 
<materialsCitation id="F4ED8754F753FFBEFE28F8BCFF68F863" ID-GBIF-Occurrence="1917076328" collectingDate="2017-07-15" country="India" county="Forest Rest House" elevation="731" latitude="17.229" location="Patan" longLatPrecision="76" longitude="73.952" municipality="Bhosgaon" pageId="0" pageNumber="595" specimenCount="1" stateProvince="Maharashtra" typeStatus="paratype">
<typeStatus id="9B3E33ABF753FFBEFE28F8BCFDCDF8BE" box="[504,596,1864,1887]" pageId="0" pageNumber="595" type="paratype">paratype</typeStatus>
male, same data as holotype (deposited in ZSI-WRC, catalogue number ENT- 1/2952).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0C9FDE82F753FFBDFF17F87AFD2DFE9E" lastPageId="3" lastPageNumber="598" pageId="0" pageNumber="595" type="description">
<paragraph id="443A8D09F753FFBEFF17F87AFABFF826" blockId="0.[151,1436,1725,2026]" pageId="0" pageNumber="595">
<emphasis id="76F1511BF753FFBEFF17F87AFD86F844" bold="true" box="[199,543,1934,1957]" pageId="0" pageNumber="595">
Description of 
<typeStatus id="9B3E33ABF753FFBEFEA6F87AFE41F844" box="[374,472,1934,1957]" pageId="0" pageNumber="595" type="holotype">holotype</typeStatus>
male.
</emphasis>
Body distinctly convex, surface brown, elytra light brown and shiny; pronotum microsculptured basally, punctation distinct but sparse in anterior half; elytron microsculptured (
<figureCitation id="DCBE918CF753FFBEFB4AF844FA82F826" box="[1178,1307,1968,1991]" captionStart="FIGURE 2" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[151,1424,362,1892]" captionTargetId="figure@2.[151,1436,350,1910]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Bolboceras sahyadriensis Kalawate &amp; Hillert, new species. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H)." httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="0" pageNumber="595">Figs. 2A–B</figureCitation>
).
</paragraph>
<paragraph id="443A8D09F753FFBEFF17F827FA02F80B" blockId="0.[151,1436,1725,2026]" box="[199,1435,2003,2026]" pageId="0" pageNumber="595">
<emphasis id="76F1511BF753FFBEFF17F827FE9DF80B" bold="true" box="[199,260,2003,2026]" pageId="0" pageNumber="595">Head</emphasis>
(
<figureCitation id="DCBE918CF753FFBEFEC1F827FEFFF80B" box="[273,358,2003,2026]" captionStart="FIGURE 2" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[151,1424,362,1892]" captionTargetId="figure@2.[151,1436,350,1910]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Bolboceras sahyadriensis Kalawate &amp; Hillert, new species. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H)." httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="0" pageNumber="595">Fig. 2C</figureCitation>
). Labrum with slightly concave anterior margin; transverse ridge indistinct; lateral perimarginal ridge
</paragraph>
<paragraph id="443A8D09F752FFBFFF47FF6CFE1FFE9E" blockId="1.[151,1435,152,625]" pageId="1" pageNumber="596">of clypeus convergent to a single, anteromedian, weakly transverse tubercle. Perimarginal clypeal ridge distinct, kshaped; frons densely punctate; vertex distinctly but sparsely punctate; transversal interocular ridge straight, long (approximately 0.75 times of interocular distance), non-tuberculate, not reaching paraocular ridges, occupying approximately basal third of eyes, lateral slope distinct, crest equally developed. Surface behind transverse interocular ridge smooth with few irregular punctures; anterior edge of eye canthus regularly convex, surface rugopunctate, with distinctly pointed tubercle; paraocular ridge fine, almost straight, reaching beyond level of interocular elevation, but not reaching base of head.</paragraph>
<paragraph id="443A8D09F752FFBFFF17FE7EFF74FD90" blockId="1.[151,1435,152,625]" pageId="1" pageNumber="596">
<emphasis id="76F1511BF752FFBFFF17FE7EFEA0FE40" bold="true" box="[199,313,394,417]" pageId="1" pageNumber="596">Pronotum</emphasis>
(
<figureCitation id="DCBE918CF752FFBFFE9DFE7EFE31FE40" box="[333,424,394,417]" captionStart="FIGURE 2" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[151,1424,362,1892]" captionTargetId="figure@2.[151,1436,350,1910]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Bolboceras sahyadriensis Kalawate &amp; Hillert, new species. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H)." httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="1" pageNumber="596">Fig. 2D</figureCitation>
). Slightly depressed anteromedially in lateral view; discoparamedian protrusions conically pointed; discolateral protrusions distinctly elevated; discomedian impressions distinct, deeply impressed, posteriorly evenly rounded; discolateral impressions distinct, shallow, rounded; finely punctate midline impression at basomedian surface. Base of pronotum finely micropunctate; pronotal punctures are of two 
<typeStatus id="9B3E33ABF752FFBFFBDDFE06FBD0FDE8" box="[1037,1097,498,521]" pageId="1" pageNumber="596">types</typeStatus>
; first 
<typeStatus id="9B3E33ABF752FFBFFB5DFE06FB22FDE8" box="[1165,1211,498,521]" pageId="1" pageNumber="596">type</typeStatus>
of punctures larger, abundant laterally, anteriorly, and along margins, a clusters of 20–25 distinct punctures formed from edge of lateral (punctate) fovea on basomedian surface; second 
<typeStatus id="9B3E33ABF752FFBFFD7FFDCCFD44FDAE" box="[687,733,568,591]" pageId="1" pageNumber="596">type</typeStatus>
of punctures are sparse and small; margin of base of pronotum distinct.
</paragraph>
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<paragraph id="443A8D09F752FFBFFF47F847FC5EF80B" blockId="1.[151,1436,1971,2026]" pageId="1" pageNumber="596">
<emphasis id="76F1511BF752FFBFFF47F847FE9FF828" bold="true" box="[151,262,1971,1993]" pageId="1" pageNumber="596">FIGURE 1</emphasis>
. Map of collection sites of 
<taxonomicName id="8385F68AF752FFBFFDDEF847FC5CF829" authority="Kalawate &amp; Hillert" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[526,965,1971,1992]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="sahyadriensis" status="sp. nov.">
<emphasis id="76F1511BF752FFBFFDDEF847FC9CF829" box="[526,773,1971,1992]" italics="true" pageId="1" pageNumber="596">Bolboceras sahyadriensis</emphasis>
Kalawate &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="6DC2EC60F752FFBFFC1FF847FBDCF828" box="[975,1093,1971,1993]" pageId="1" pageNumber="596" rank="species">
<emphasis id="76F1511BF752FFBFFC1FF847FBDCF828" bold="true" box="[975,1093,1971,1993]" pageId="1" pageNumber="596">new species</emphasis>
</taxonomicNameLabel>
(square), 
<taxonomicName id="8385F68AF752FFBFFB76F847FEE4F80B" authority="Gupta &amp; Chandra, 2016" authorityName="Gupta &amp; Chandra" authorityYear="2016" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="76F1511BF752FFBFFB76F847FA05F829" box="[1190,1436,1971,1992]" italics="true" pageId="1" pageNumber="596">Bolboceras bilaspuricans</emphasis>
Gupta &amp; Chandra, 2016
</taxonomicName>
(triangles), and 
<taxonomicName id="8385F68AF752FFBFFDCAF821FCF2F80B" authority="Krikken, 2013" authorityName="Krikken" authorityYear="2013" box="[538,875,2005,2026]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="insulare">
<emphasis id="76F1511BF752FFBFFDCAF821FD43F80B" box="[538,730,2005,2026]" italics="true" pageId="1" pageNumber="596">Bolboceras insulare</emphasis>
Krikken, 2013
</taxonomicName>
(circles).
</paragraph>
</caption>
<paragraph id="443A8D09F751FFBCFF17FF6CFD6DFF4E" blockId="2.[151,1435,152,279]" box="[199,756,152,175]" pageId="2" pageNumber="597">
<emphasis id="76F1511BF751FFBCFF17FF6CFEA0FF4E" bold="true" box="[199,313,152,175]" pageId="2" pageNumber="597">Scutellum</emphasis>
(
<figureCitation id="DCBE918CF751FFBCFE97FF6CFE02FF4E" box="[327,411,152,175]" captionStart="FIGURE 2" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[151,1424,362,1892]" captionTargetId="figure@2.[151,1436,350,1910]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Bolboceras sahyadriensis Kalawate &amp; Hillert, new species. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H)." httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="2" pageNumber="597">Fig. 2E</figureCitation>
). Triangulate, sparsely punctate.
</paragraph>
<paragraph id="443A8D09F751FFBCFF17FF4EFBA9FF30" blockId="2.[151,1435,152,279]" box="[199,1072,186,209]" pageId="2" pageNumber="597">
<emphasis id="76F1511BF751FFBCFF17FF4EFE89FF30" bold="true" box="[199,272,186,209]" pageId="2" pageNumber="597">Elytra</emphasis>
(
<figureCitation id="DCBE918CF751FFBCFECEFF4EFEF6FF30" box="[286,367,186,209]" captionStart="FIGURE 2" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[151,1424,362,1892]" captionTargetId="figure@2.[151,1436,350,1910]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Bolboceras sahyadriensis Kalawate &amp; Hillert, new species. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H)." httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="2" pageNumber="597">Fig. 2F</figureCitation>
). With weakly impressed, finely punctate striae; intervals smooth.
</paragraph>
<paragraph id="443A8D09F751FFBCFF17FF29FB3AFEF6" blockId="2.[151,1435,152,279]" pageId="2" pageNumber="597">
<emphasis id="76F1511BF751FFBCFF17FF29FF65FF15" bold="true" box="[199,252,221,244]" pageId="2" pageNumber="597">Legs</emphasis>
. Protibia with 6 distinct external denticles, diminishing in size from apex to base; apex of terminal spur robust, tapering and sharp apically. Mesotibiae and metatibiae with distinctly elevated, transverse carina.
</paragraph>
<caption id="10FADD81F751FFBCFF47F86CFF2EF80E" httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="2" pageNumber="597" startId="2.[151,244,1944,1966]" targetBox="[151,1424,362,1892]" targetPageId="2">
<paragraph id="443A8D09F751FFBCFF47F86CFF2EF80E" blockId="2.[151,1436,1944,2031]" pageId="2" pageNumber="597">
<emphasis id="76F1511BF751FFBCFF47F86CFE91F84F" bold="true" box="[151,264,1944,1966]" pageId="2" pageNumber="597">FIGURE 2</emphasis>
. Holotype of 
<taxonomicName id="8385F68AF751FFBCFE42F86CFCC9F84C" authority="Kalawate &amp; Hillert" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[402,848,1944,1965]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="sahyadriensis" status="sp. nov.">
<emphasis id="76F1511BF751FFBCFE42F86CFD12F84C" box="[402,651,1944,1965]" italics="true" pageId="2" pageNumber="597">Bolboceras sahyadriensis</emphasis>
Kalawate &amp; Hillert
</taxonomicName>
, 
<taxonomicNameLabel id="6DC2EC60F751FFBCFC8CF86CFC4AF84F" box="[860,979,1944,1966]" pageId="2" pageNumber="597" rank="species">
<emphasis id="76F1511BF751FFBCFC8CF86CFC4AF84F" bold="true" box="[860,979,1944,1966]" pageId="2" pageNumber="597">new species</emphasis>
</taxonomicNameLabel>
. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H).
</paragraph>
</caption>
<paragraph id="443A8D09F750FFBDFF17FF6CFBF7FF15" blockId="3.[151,1436,152,1388]" pageId="3" pageNumber="598">
<emphasis id="76F1511BF750FFBDFF17FF6CFEACFF4E" bold="true" box="[199,309,152,175]" pageId="3" pageNumber="598">Aedeagus</emphasis>
(
<figureCitation id="DCBE918CF750FFBDFE9AFF6CFE4BFF4E" box="[330,466,152,175]" captionStart="FIGURE 2" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[151,1424,362,1892]" captionTargetId="figure@2.[151,1436,350,1910]" captionTargetPageId="2" captionText="FIGURE 2. Holotype of Bolboceras sahyadriensis Kalawate &amp; Hillert, new species. A, dorsal habitus; B, lateral habitus; C, head, dorsal view; D, pronotum, dorsal view; E, scutellum; F, elytron; G, H, aedeagus. Scale bars = 2 mm (A–D, F); 1 mm (E); 0.5 mm (G, H)." httpUri="https://zenodo.org/record/1457997/files/figure.png" pageId="3" pageNumber="598">Figs. 2G–H</figureCitation>
). Parameres thin, strongly acuminate, divergent laterally, overlapped in basal part of aedeagus, reaching beyond paramedian aedeagal struts. Lateral aedeagal stalks robust, strongly sclerotized, finely hooked, recurved at apex; median aedeagal apparatus (upper lobe) plain, rounded anteriorly.
</paragraph>
<paragraph id="443A8D09F750FFBDFF17FEF4FD2DFE9E" blockId="3.[151,1436,152,1388]" pageId="3" pageNumber="598">
<emphasis id="76F1511BF750FFBDFF17FEF4FEECFEF6" bold="true" box="[199,373,256,279]" pageId="3" pageNumber="598">Measurements.</emphasis>
Body length: 
<quantity id="837D20ECF750FFBDFDF0FEF4FD11FEF6" box="[544,648,256,279]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="598" unit="mm" value="12.3">12.3 mm</quantity>
, width: 
<quantity id="837D20ECF750FFBDFD3BFEF4FCDEFEF6" box="[747,839,256,279]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="598" unit="mm" value="7.6">7.6 mm</quantity>
. Head, length in dorsal view excluding labrum and mandibles: 
<quantity id="837D20ECF750FFBDFEC6FED6FEF7FED8" box="[278,366,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="598" unit="mm" value="2.2">2.2 mm</quantity>
, width: 
<quantity id="837D20ECF750FFBDFE1AFED6FDBBFED8" box="[458,546,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="598" unit="mm" value="3.7">3.7 mm</quantity>
; interocular ridge: 
<quantity id="837D20ECF750FFBDFD26FED6FCD6FED8" box="[758,847,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="598" unit="mm" value="1.8">1.8 mm</quantity>
; ocular distance: 
<quantity id="837D20ECF750FFBDFBC2FED6FBF2FED8" box="[1042,1131,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="598" unit="mm" value="2.6">2.6 mm</quantity>
. Pronotum length: 
<quantity id="837D20ECF750FFBDFA90FED6FA0EFED8" box="[1344,1431,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="598" unit="mm" value="4.5">4.5 mm</quantity>
, width: 
<quantity id="837D20ECF750FFBDFF37FEB1FED8FEBD" box="[231,321,325,348]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="598" unit="mm" value="7.4">7.4 mm</quantity>
. Scutellar shield length: 
<quantity id="837D20ECF750FFBDFD8CFEB1FD2FFEBD" box="[604,694,325,348]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="598" unit="mm" value="1.4">1.4 mm</quantity>
, width: 
<quantity id="837D20ECF750FFBDFCC6FEB1FCE9FEBD" box="[790,880,325,348]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="598" unit="mm" value="1.5">1.5 mm</quantity>
. Elytra, sutural length: 4.0 mm, maximum width combined: 
<quantity id="837D20ECF750FFBDFEDEFE9CFEFAFE9E" box="[270,355,360,383]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="598" unit="mm" value="7.8">7.8 mm</quantity>
. Genital capsule width 
<quantity id="837D20ECF750FFBDFD8CFE9CFD28FE9E" box="[604,689,360,383]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="598" unit="mm" value="0.9">0.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0C9FDE82F750FFBDFF17FE7EFF7DFE25" pageId="3" pageNumber="598" type="etymology">
<paragraph id="443A8D09F750FFBDFF17FE7EFF7DFE25" blockId="3.[151,1436,152,1388]" pageId="3" pageNumber="598">
<emphasis id="76F1511BF750FFBDFF17FE7EFEDEFE40" bold="true" box="[199,327,394,417]" pageId="3" pageNumber="598">Etymology.</emphasis>
This species is named after “Sahyadri”, the Sanskrit and Marathi name for the 
<collectingRegion id="864143EBF750FFBDFB70FE7EFB6EFE40" box="[1184,1271,394,417]" country="Sri Lanka" name="Western" pageId="3" pageNumber="598">Western</collectingRegion>
Ghats where it occurs.
</paragraph>
</subSubSection>
<subSubSection id="0C9FDE82F750FFBDFF17FE24FA1BFE06" box="[199,1410,464,487]" pageId="3" pageNumber="598" type="distribution">
<paragraph id="443A8D09F750FFBDFF17FE24FA1BFE06" blockId="3.[151,1436,152,1388]" box="[199,1410,464,487]" pageId="3" pageNumber="598">
<emphasis id="76F1511BF750FFBDFF17FE24FEC0FE06" bold="true" box="[199,345,464,487]" pageId="3" pageNumber="598">Distribution.</emphasis>
Known only from the 
<typeStatus id="9B3E33ABF750FFBDFD9CFE24FDE3FE06" box="[588,634,464,487]" pageId="3" pageNumber="598">type</typeStatus>
locality: Bhosgaon, Satara District (
<collectingRegion id="864143EBF750FFBDFC2DFE24FBCCFE06" box="[1021,1109,464,487]" country="Sri Lanka" name="Western" pageId="3" pageNumber="598">Western</collectingRegion>
Ghats), 
<collectingRegion id="864143EBF750FFBDFB61FE24FAA0FE06" box="[1201,1337,464,487]" country="India" name="Maharashtra" pageId="3" pageNumber="598">Maharashtra</collectingRegion>
, 
<collectingCountry id="3C92CD99F750FFBDFA93FE24FAE4FE06" box="[1347,1405,464,487]" name="India" pageId="3" pageNumber="598">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0C9FDE82F750FFBDFF17FE06FC05FA8D" pageId="3" pageNumber="598" type="diagnosis">
<paragraph id="443A8D09F750FFBDFF17FE06FE4BFCFE" blockId="3.[151,1436,152,1388]" pageId="3" pageNumber="598">
<emphasis id="76F1511BF750FFBDFF17FE06FDF9FDE8" bold="true" box="[199,608,498,521]" pageId="3" pageNumber="598">Differential diagnosis and remarks.</emphasis>
The species of the 
<taxonomicName id="8385F68AF750FFBDFCE9FE06FC24FDE8" authorityName="Westwood" authorityYear="1848" box="[825,957,498,521]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="76F1511BF750FFBDFCE9FE06FC24FDE8" box="[825,957,498,521]" italics="true" pageId="3" pageNumber="598">B. nigricans</emphasis>
</taxonomicName>
species group are separated on the basis of characteristic perimarginal clypeal ridges. The clypeal ridges in males are convergent to a single anteromedian tubercle. The clypeal ridges are k-shaped or x-shaped in dorsal view. According to 
<bibRefCitation id="2014F0F8F750FFBDFC6BFDCCFBE9FDAE" box="[955,1136,568,591]" pageId="3" pageNumber="598" refId="ref2340" refString="Krikken, J. (2013 a) Oriental Bolboceras: introduction to the genus and taxonomy of the nigricans group of species (Coleoptera: Geotrupidae: Bolboceratinae). Tijdschrift voor Entomologie, 156, 35 - 56. https: // doi. org / 10.1163 / 22119434 - 00002018" type="journal article">Krikken (2013a)</bibRefCitation>
, the species in 
<taxonomicName id="8385F68AF750FFBDFAC8FDCCFA05FDAE" authorityName="Westwood" authorityYear="1848" box="[1304,1436,568,591]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="76F1511BF750FFBDFAC8FDCCFA05FDAE" box="[1304,1436,568,591]" italics="true" pageId="3" pageNumber="598">B. nigricans</emphasis>
</taxonomicName>
species group can be separated by characteristic x-shaped clypeal ridge (
<taxonomicName id="8385F68AF750FFBDFC66FDAEFB75FD90" authority="Krikken, 2013" authorityName="Krikken" authorityYear="2013" box="[950,1260,602,625]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="extraneum">
<emphasis id="76F1511BF750FFBDFC66FDAEFBDFFD90" box="[950,1094,602,625]" italics="true" pageId="3" pageNumber="598">B. extraneum</emphasis>
Krikken, 2013
</taxonomicName>
and 
<taxonomicName id="8385F68AF750FFBDFAF5FDAEFEAAFD75" authority="Krikken, 2013" authorityName="Krikken" authorityYear="2013" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="insulare">
<emphasis id="76F1511BF750FFBDFAF5FDAEFA02FD90" box="[1317,1435,601,625]" italics="true" pageId="3" pageNumber="598">B. insulare</emphasis>
Krikken, 2013
</taxonomicName>
) and k-shaped clypeal ridge (
<taxonomicName id="8385F68AF750FFBDFDA1FD89FC28FD75" authority="Westwood, 1848" authorityName="Westwood" authorityYear="1848" box="[625,945,637,660]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="76F1511BF750FFBDFDA1FD89FD6AFD75" box="[625,755,637,660]" italics="true" pageId="3" pageNumber="598">B. nigricans</emphasis>
Westwood, 1848
</taxonomicName>
, 
<taxonomicName id="8385F68AF750FFBDFC6CFD89FA95FD75" authority="Krikken, 2013" authorityName="Krikken" authorityYear="2013" box="[956,1292,636,660]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="76F1511BF750FFBDFC6CFD89FBF1FD75" box="[956,1128,636,660]" italics="true" pageId="3" pageNumber="598">B. malabaricum</emphasis>
Krikken, 2013
</taxonomicName>
, 
<taxonomicName id="8385F68AF750FFBDFAC8FD89FEA1FD56" authority="Krikken, 2013" authorityName="Krikken" authorityYear="2013" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="mimicans">
<emphasis id="76F1511BF750FFBDFAC8FD89FA05FD75" box="[1304,1436,637,660]" italics="true" pageId="3" pageNumber="598">B. mimicans</emphasis>
Krikken, 2013
</taxonomicName>
, 
<taxonomicName id="8385F68AF750FFBDFE98FD54FDF4FD56" authority="Krikken, 2013" authorityName="Krikken" authorityYear="2013" box="[328,621,671,695]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="schulzei">
<emphasis id="76F1511BF750FFBDFE98FD54FE59FD56" box="[328,448,671,695]" italics="true" pageId="3" pageNumber="598">B. schulzei</emphasis>
Krikken, 2013
</taxonomicName>
, 
<taxonomicName id="8385F68AF750FFBDFDADFD54FBD2FD56" authority="Gupta &amp; Chandra, 2016" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[637,1099,671,695]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="76F1511BF750FFBDFDADFD54FCA9FD56" box="[637,816,671,695]" italics="true" pageId="3" pageNumber="598">B. bilaspuricans</emphasis>
Gupta &amp; Chandra, 2016
</taxonomicName>
and 
<taxonomicName id="8385F68AF750FFBDFB59FD54FEA5FD38" authority="Gupta &amp; Chandra, 2016" authorityName="Gupta &amp; Chandra" authorityYear="2016" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="darjeelicans">
<emphasis id="76F1511BF750FFBDFB59FD54FAB7FD56" box="[1161,1326,671,695]" italics="true" pageId="3" pageNumber="598">B. darjeelicans</emphasis>
Gupta &amp; Chandra, 2016
</taxonomicName>
). The discomedian and discolateral impressions of pronotum are more or less deeply impressed (see 
<bibRefCitation id="2014F0F8F750FFBDFF47FD11FEA4FD1D" box="[151,317,741,764]" pageId="3" pageNumber="598" refId="ref2340" refString="Krikken, J. (2013 a) Oriental Bolboceras: introduction to the genus and taxonomy of the nigricans group of species (Coleoptera: Geotrupidae: Bolboceratinae). Tijdschrift voor Entomologie, 156, 35 - 56. https: // doi. org / 10.1163 / 22119434 - 00002018" type="journal article">Krikken 2013a</bibRefCitation>
). The pronotal impressions probably depend on body size. The allometric specimens show often only weakly developed characters.
</paragraph>
<paragraph id="443A8D09F750FFBDFF17FCDEFC05FA8D" blockId="3.[151,1436,152,1388]" pageId="3" pageNumber="598">
Among the species of 
<taxonomicName id="8385F68AF750FFBDFE64FCDEFDAFFCA0" authorityName="Westwood" authorityYear="1848" box="[436,566,810,833]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="76F1511BF750FFBDFE64FCDEFDAFFCA0" box="[436,566,810,833]" italics="true" pageId="3" pageNumber="598">B. nigricans</emphasis>
</taxonomicName>
species group, 
<taxonomicName id="8385F68AF750FFBDFD0CFCDEFC15FCA0" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[732,908,809,833]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="sahyadriensis">
<emphasis id="76F1511BF750FFBDFD0CFCDEFC15FCA0" box="[732,908,809,833]" italics="true" pageId="3" pageNumber="598">B. sahyadriensis</emphasis>
</taxonomicName>
is unique mainly due to its structure of aedeagus. The new species has a thin, strongly acuminate, and laterally divergent parameres that overlap the basal part of the aedeagus. The parameres extend beyond the paramedian aedeagal struts. The lateral aedeagal stalks are robust, strongly sclerotized, finely hooked, and recurved at the apex. The median aedeagal apparatus (upper lobe) is plain anteriorly. The basal genitalic capsule is broad. The closely related species, 
<taxonomicName id="8385F68AF750FFBDFC9CFC41FB99FC2D" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[844,1024,948,972]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="76F1511BF750FFBDFC9CFC41FB99FC2D" box="[844,1024,948,972]" italics="true" pageId="3" pageNumber="598">B. bilaspuricans</emphasis>
</taxonomicName>
and 
<taxonomicName id="8385F68AF750FFBDFBEEFC41FB69FC2D" authorityName="Krikken" authorityYear="2013" box="[1086,1264,948,972]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="76F1511BF750FFBDFBEEFC41FB69FC2D" box="[1086,1264,948,972]" italics="true" pageId="3" pageNumber="598">B. malabaricum</emphasis>
</taxonomicName>
have a similar structure of the aedeagus. The new species is, however, differentiated from them by the presence of strongly acuminate paramere tip that extends beyond the paramedian aedeagal struts; basal genitalic capsule broad. In 
<taxonomicName id="8385F68AF750FFBDFB14FC0EFAEAFBF0" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[1220,1395,1017,1041]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="76F1511BF750FFBDFB14FC0EFAEAFBF0" box="[1220,1395,1017,1041]" italics="true" pageId="3" pageNumber="598">B. bilaspuricans</emphasis>
</taxonomicName>
the tip of parameres are subangular, reaching just beyond the lateral aedeagal stalks, basal genitalic capsule not broad; whereas, in 
<emphasis id="76F1511BF750FFBDFEC7FBB4FE53FBB6" box="[279,458,1087,1111]" italics="true" pageId="3" pageNumber="598">
<taxonomicName id="8385F68AF750FFBDFEC7FBB4FE5FFBB6" authorityName="Krikken" authorityYear="2013" box="[279,454,1087,1111]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="malabaricum">B. malabaricum</taxonomicName>
,
</emphasis>
the parameres are acuminate and extended to the lateral aedeagal stalks; the basal genitalic capsule smaller. 
<taxonomicName id="8385F68AF750FFBDFE97FB95FDC0FB98" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[327,601,1121,1145]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="sahyadriensis">
<emphasis id="76F1511BF750FFBDFE97FB95FDC0FB98" box="[327,601,1121,1145]" italics="true" pageId="3" pageNumber="598">Bolboceras sahyadriensis</emphasis>
</taxonomicName>
is morphologically distinct from its close relatives by the presence of the long and non-tuberculate transverse interocular ridge, the evenly developed crest, the greater height of the interocular ridge; the slightly concave anterior margin of the labrum, and the indistinct transverse ridge. Conversely, the transverse interocular ridge of 
<taxonomicName id="8385F68AF750FFBDFEA7FB3EFDBEFB00" authorityName="Krikken" authorityYear="2013" box="[375,551,1225,1249]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="76F1511BF750FFBDFEA7FB3EFDBEFB00" box="[375,551,1225,1249]" italics="true" pageId="3" pageNumber="598">B. malabaricum</emphasis>
</taxonomicName>
is very short and tuberculate (medially raised), unevenly crested, height of the interocular ridge high; the labrum has a slightly concave anterior margin and the transverse ridge is distinct. The transverse interocular ridge of 
<taxonomicName id="8385F68AF750FFBDFE3BFAE4FD04FAC6" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[491,669,1295,1319]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="76F1511BF750FFBDFE3BFAE4FD04FAC6" box="[491,669,1295,1319]" italics="true" pageId="3" pageNumber="598">B. bilaspuricans</emphasis>
</taxonomicName>
is long and non-tuberculate, evenly crested, height of the interocular ridge is low; the labrum with slightly concave anterior margin and the transverse ridge is distinct. For differentiation of 
<taxonomicName id="8385F68AF750FFBDFF47FAA1FEDFFA8D" authorityName="Kalawate &amp; Hillert" authorityYear="2018" box="[151,326,1364,1388]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="sahyadriensis">
<emphasis id="76F1511BF750FFBDFF47FAA1FEDFFA8D" box="[151,326,1364,1388]" italics="true" pageId="3" pageNumber="598">B. sahyadriensis</emphasis>
</taxonomicName>
from 
<taxonomicName id="8385F68AF750FFBDFE58FAA1FDAAFA8D" authorityName="Krikken" authorityYear="2013" box="[392,563,1364,1388]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="malabaricum">
<emphasis id="76F1511BF750FFBDFE58FAA1FDAAFA8D" box="[392,563,1364,1388]" italics="true" pageId="3" pageNumber="598">B. malabaricum</emphasis>
</taxonomicName>
and 
<taxonomicName id="8385F68AF750FFBDFDB8FAA1FC8FFA8D" authorityName="Gupta &amp; Chandra" authorityYear="2016" box="[616,790,1364,1388]" class="Insecta" family="Geotrupidae" genus="Bolboceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="bilaspuricans">
<emphasis id="76F1511BF750FFBDFDB8FAA1FC8FFA8D" box="[616,790,1364,1388]" italics="true" pageId="3" pageNumber="598">B. bilaspuricans</emphasis>
</taxonomicName>
see 
<tableCitation id="0907B8B2F750FFBDFC96FAA1FC0CFA8D" box="[838,917,1365,1388]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Character matrix for the separation of males of B. sahyadriensis Kalawate &amp; Hillert, new species and similar species in the B. nigricans species group" pageId="3" pageNumber="598">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>