428 lines
51 KiB
XML
428 lines
51 KiB
XML
<document id="53683F10B6A56BD4314A3A16FAC99C3E" ID-DOI="10.1080/00222933.2020.1759721" ID-ISSN="1464-5262" ID-Zenodo-Dep="4609053" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615923843413" checkinUser="carolina" docAuthor="Bevilaqua, Marcus & Fonseca, Claudio Ruy Vasconcelos da" docDate="2020" docId="C4008788690E9E18FE8CFCBC81526AE0" docLanguage="en" docName="JNATHIST.54.5-6.321-350.pdf" docOrigin="Journal of Natural History 54 (5 - 6)" docSource="http://dx.doi.org/10.1080/00222933.2020.1759721" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Passalus (Pertinax) beneshi Hincks 1950" docType="treatment" docVersion="3" lastPageNumber="327" masterDocId="3839FFF0690D9E1FFFCAFF9A83526814" masterDocTitle="Redescription of the species of Passalus Fabricius, 1792 (Coleoptera: Passalidae) described by Walter Douglas Hincks (1906 - 1961) deposited in the Museum of the University of Manchester" masterLastPageNumber="350" masterPageNumber="321" pageNumber="323" updateTime="1698887187464" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B98035747183C3488922C10E91BCDA4F">Redescription of the species of Passalus Fabricius, 1792 (Coleoptera: Passalidae) described by Walter Douglas Hincks (1906 - 1961) deposited in the Museum of the University of Manchester</mods:title>
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<mods:namePart id="3840A826C08D1A889F9DE78851086A5C">Bevilaqua, Marcus</mods:namePart>
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<mods:namePart id="7F957C57FC6454AA59D7433976ADEA51">Fonseca, Claudio Ruy Vasconcelos da</mods:namePart>
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<mods:typeOfResource id="691843C9219A6240D8CFC55EC7689270">text</mods:typeOfResource>
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<mods:title id="38677B6381AB3CF5CB6235B6ABFAEB54">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="1D5DB023E1E462605463652844837634">
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<mods:date id="C2C250F444231A126F88753BD6845628">2020</mods:date>
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<mods:number id="0AF1CF570EC7A5B9F6BC01DE532CCB5C">2020-07-27</mods:number>
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<mods:detail id="4D50F5F13569616AB78488BC0CB32259" type="volume">
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<mods:number id="2EAB6D63C301018F7D1726F3ED26EC7D">54</mods:number>
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<mods:number id="7E452CB1D76F1CE8DB7A8FBAFAA2E0E5">5 - 6</mods:number>
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<mods:start id="45640167A026401EE718A5E2B468F9AE">321</mods:start>
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<mods:url id="448E8BB56095276DE09BC7A7963E7444">http://dx.doi.org/10.1080/00222933.2020.1759721</mods:url>
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<mods:classification id="72DE8622BCFBCDDF1DBE82CC6CDB8C17">journal article</mods:classification>
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<mods:identifier id="B892797DC6C73B005C12A27CAF503834" type="DOI">10.1080/00222933.2020.1759721</mods:identifier>
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<mods:identifier id="5626435BFD8EAACD20DC10E296DCAE1A" type="ISSN">1464-5262</mods:identifier>
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<treatment id="C4008788690E9E18FE8CFCBC81526AE0" LSID="urn:lsid:plazi:treatment:C4008788690E9E18FE8CFCBC81526AE0" httpUri="http://treatment.plazi.org/id/C4008788690E9E18FE8CFCBC81526AE0" lastPageId="7" lastPageNumber="327" pageId="3" pageNumber="323">
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<subSubSection id="04B36515690E9E1CFE8CFCBC801C6BAC" pageId="3" pageNumber="323" type="nomenclature">
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<paragraph id="4C16369E690E9E1CFE8CFCBC808F6B2B" blockId="3.[326,989,806,831]" box="[326,989,806,831]" pageId="3" pageNumber="323">
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<heading id="175E81F2690E9E1CFE8CFCBC808F6B2B" box="[326,989,806,831]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="323" reason="2">
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<taxonomicName id="8BA94D1D690E9E1CFE8CFCBC806F6B2B" authority="Hincks, 1950" authorityName="Hincks" authorityYear="1950" box="[326,829,806,831]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="323" phylum="Arthropoda" rank="species" species="beneshi" subGenus="Pertinax">
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<emphasis id="7EDDEA8C690E9E1CFE8CFCBC806F6B2B" bold="true" box="[326,829,806,831]" pageId="3" pageNumber="323">
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<emphasis id="7EDDEA8C690E9E1CFE8CFCBC82FD6B2B" bold="true" box="[326,431,806,831]" italics="true" pageId="3" pageNumber="323">Passalus</emphasis>
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(
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<emphasis id="7EDDEA8C690E9E1CFE75FCBC81776B2B" bold="true" box="[447,549,806,831]" italics="true" pageId="3" pageNumber="323">Pertinax</emphasis>
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)
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<emphasis id="7EDDEA8C690E9E1CFDFCFCBC81C66B2B" bold="true" box="[566,660,806,831]" italics="true" pageId="3" pageNumber="323">beneshi</emphasis>
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<bibRefCitation id="28384B6F690E9E1CFD57FCBC806F6B2B" author="Hincks WD" box="[669,829,806,831]" pageId="3" pageNumber="323" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks, 1950</bibRefCitation>
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</emphasis>
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</taxonomicName>
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(
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<figureCitation id="D4922A1B690E9E1CFC87FCBC80876B2B" box="[845,981,806,831]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="4.[160,225,1482,1505]" captionStartId-1="5.[160,225,1329,1352]" captionTargetBox-0="[163,1153,142,1451]" captionTargetBox-1="[163,1153,142,1297]" captionTargetId-0="figure-56@4.[163,1153,142,1451]" captionTargetId-1="figure-115@5.[163,1153,142,1298]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." captionText-1="Figure 2. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4609057" figureDoi-1="http://doi.org/10.5281/zenodo.4609061" httpUri-0="https://zenodo.org/record/4609057/files/figure.png" httpUri-1="https://zenodo.org/record/4609061/files/figure.png" pageId="3" pageNumber="323">
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Figures 1
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<emphasis id="7EDDEA8C690E9E1CFC73FCBC80956B2B" box="[953,967,806,831]" italics="true" pageId="3" pageNumber="323">–</emphasis>
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2
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</figureCitation>
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)
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</heading>
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</paragraph>
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<paragraph id="4C16369E690E9E1CFF0AFCC1801C6BAC" blockId="3.[160,1156,859,952]" pageId="3" pageNumber="323">
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<bibRefCitation id="28384B6F690E9E1CFF0AFCC182DC6B67" author="Hincks WD" box="[192,398,859,883]" pageId="3" pageNumber="323" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks 1950: 1041</bibRefCitation>
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, fig. 4;
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<bibRefCitation id="28384B6F690E9E1CFE14FCC180526B67" author="Hincks WD & Dibb JR" box="[478,768,859,883]" pageId="3" pageNumber="323" pagination="1 - 33" refId="ref13308" refString="Hincks WD, Dibb JR. 1958. Supplementa. Passalidae. Coleopterorum Catalogus. Berlin: W. Junk, J. Schenkling, es-Gravenhage; p. 1 - 33." type="book chapter" year="1958">Hincks and Dibb 1958: 15</bibRefCitation>
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;
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<bibRefCitation id="28384B6F690E9E1CFCC1FCC187496B67" author="Reyes-Castillo P" box="[779,1051,859,883]" pageId="3" pageNumber="323" pagination="1 - 240" refId="ref13746" refString="Reyes-Castillo P. 1970. Coleoptera, Passalidae: morfologia y division en grandes grupos; generos americanos. Folia Entomol Mex. 20 (22): 1 - 240." type="journal article" year="1970">Reyes-Castillo 1970: 204</bibRefCitation>
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;
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<bibRefCitation id="28384B6F690E9E1CFBEDFCC182476B82" author="Boucher S" pageId="3" pageNumber="323" pagination="113 - 120" refId="ref12922" refString="Boucher S. 2015. Splitting of the polyphyletic genus Passalus Fabricius, s. auct. I. (Coleoptera, Passalidae). Bull Soc Entomol France. 120: 113 - 120." type="journal article" year="2015">Boucher 2015: 118</bibRefCitation>
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;
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<bibRefCitation id="28384B6F690E9E1CFEECFCE4805B6B82" author="Amat-Garcia G & Reyes-Castillo P" box="[294,777,894,918]" pageId="3" pageNumber="323" pagination="139 - 151" refId="ref12711" refString="Amat-Garcia G, Reyes-Castillo P. 2002. Los Coleoptera Passalidae de Colombia. In: Costa M, Vanin SA, Lobo JM, Melic A, editors. Proyecto de Red Iberoamericana de Biogeografia y Entomologia Sistematica PrIBES 2002. Monografias Tercer Milenio. Vol. II. Zaragoza: Sociedad Entomologica Aragones (SEA); p. 139 - 151." type="book chapter" year="2002">Amat-Garcia and Reyes-Castillo, 2002:141</bibRefCitation>
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;
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<bibRefCitation id="28384B6F690E9E1CFCD0FCE482506BAC" author="Reyes-Castillo P & Amat-Garcia G" pageId="3" pageNumber="323" pagination="35 - 50" refId="ref13821" refString="Reyes-Castillo P, Amat-Garcia G. 2003. Passalidae (Coleoptera) de Colombia. In: Onore G, Reyes- Castillo P, Zunino M, editors. Escarabajos de Latinoamerica: estado del conocimiento. Monografias tercer milenio. Vol. III. Zaragoza: Sociedad Entomologica Aragones (SEA); p. 35 - 50." type="book chapter" year="2003">Reyes-Castillo and Amat-García 2003: 37</bibRefCitation>
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;
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<bibRefCitation id="28384B6F690E9E1CFEC6FC3A81116BAC" author="Amat-Garcia G & Vargas EB & Reyes-Castillo P" box="[268,579,928,952]" pageId="3" pageNumber="323" pagination="173 - 181" refId="ref12776" refString="Amat-Garcia G, Vargas EB, Reyes-Castillo P. 2004. Lista de especies de los escarabajos pasalidos (Coleoptera: Passalidae) de Colombia. Biota Colombiana. 5 (2): 173 - 181." type="journal article" year="2004">Amat-García et al. 2004:176</bibRefCitation>
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||
;
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||
<bibRefCitation id="28384B6F690E9E1CFD84FC3A81AD6BAC" author="Miles C" box="[590,767,928,952]" pageId="3" pageNumber="323" pagination="109 - 126" refId="ref13705" refString="Miles C. 2017. The Passalidae collection of the Manchester Museum, UK, with a complete type catalogue. Russ Entomol J. 26 (2): 109 - 126. doi: 10.15298 / rusentj. 26.2.03" type="journal article" year="2017">Miles 2017: 117</bibRefCitation>
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, fig 11.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="04B36515690E9E1CFF6AFC7080B86CE2" pageId="3" pageNumber="323" type="diagnosis">
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<paragraph id="4C16369E690E9E1CFF6AFC7080B86CE2" blockId="3.[160,1156,1002,1270]" pageId="3" pageNumber="323">
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<emphasis id="7EDDEA8C690E9E1CFF6AFC7082486C10" bold="true" box="[160,282,1002,1028]" italics="true" pageId="3" pageNumber="323">Diagnosis.</emphasis>
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Small size species; convex body; anterior edge of the front slightly convex; coarsely punctuated mediofrontal area; laterofrontal + mediofrontal tubercles connected to the inner tubercles by the anterofrontal ridges; small and flattened central tubercle; acute and protruded pronotum anterior angles; coarse and abundant pronotum punctuation, extending beyond three-fourths of pronotal width; glabrous prepisternum; large, wide, wellmarked and deep mesosternal scars; not defined metasternal disk, without pubescence and with reduced punctuation to a small group in the posterior region; median lobe almost entirely sclerotised and basal piece fused with parameres on ventral surface.
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</paragraph>
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</subSubSection>
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<subSubSection id="04B36515690E9E18FF6AFAB2829968DD" lastPageId="7" lastPageNumber="327" pageId="3" pageNumber="323" type="description">
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<paragraph id="4C16369E690E9E19FF6AFAB280C66CCD" blockId="3.[160,1156,1320,1727]" lastBlockId="6.[160,1156,141,1241]" lastPageId="6" lastPageNumber="326" pageId="3" pageNumber="323">
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<emphasis id="7EDDEA8C690E9E1CFF6AFAB282146D56" bold="true" box="[160,326,1320,1346]" italics="true" pageId="3" pageNumber="323">Redescription.</emphasis>
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<emphasis id="7EDDEA8C690E9E1CFE9CFAB082C66D56" bold="true" box="[342,404,1322,1346]" pageId="3" pageNumber="323">Body</emphasis>
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(
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<figureCitation id="D4922A1B690E9E1CFE6FFAB081566D56" box="[421,516,1322,1346]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="3" pageNumber="323">Figure 1</figureCitation>
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):
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<emphasis id="7EDDEA8C690E9E1CFDD0FAB381396D56" box="[538,619,1321,1346]" italics="true" pageId="3" pageNumber="323">habitus</emphasis>
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: convex;
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<emphasis id="7EDDEA8C690E9E1CFD13FAB380536D56" box="[729,769,1321,1346]" italics="true" pageId="3" pageNumber="323">size</emphasis>
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: small (>
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<quantity id="8B519B7B690E9E1CFCA2FAB080EE6D56" box="[872,956,1322,1346]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="323" unit="mm" value="20.0">20 mm</quantity>
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in length).
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<emphasis id="7EDDEA8C690E9E1CFB8EFAB087D16D56" bold="true" box="[1092,1155,1322,1346]" pageId="3" pageNumber="323">Head</emphasis>
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(
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<figureCitation id="D4922A1B690E9E1CFF62FAD782556D71" box="[168,263,1357,1381]" captionStart="Figure 2" captionStartId="5.[160,225,1329,1352]" captionTargetBox="[163,1153,142,1297]" captionTargetId="figure-115@5.[163,1153,142,1298]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4609061" httpUri="https://zenodo.org/record/4609061/files/figure.png" pageId="3" pageNumber="323">Figure 2</figureCitation>
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(a,c)):
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<emphasis id="7EDDEA8C690E9E1CFE86FAD682F26D71" box="[332,416,1356,1381]" italics="true" pageId="3" pageNumber="323">Labrum</emphasis>
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: anterior border straight.
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<emphasis id="7EDDEA8C690E9E1CFD05FAD680706D71" box="[719,802,1356,1381]" italics="true" pageId="3" pageNumber="323">Clypeus</emphasis>
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: hidden under the frons; with anterior angles under the lateral + mediofrontal tubercles.
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<emphasis id="7EDDEA8C690E9E1CFCF5FAF487766D93" box="[831,1060,1390,1415]" italics="true" pageId="3" pageNumber="323">Anterior frontal edge</emphasis>
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: slightly convex, with a notch in the median region.
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<emphasis id="7EDDEA8C690E9E1CFD66FA0B87736DBE" box="[684,1057,1425,1450]" italics="true" pageId="3" pageNumber="323">Secondary mediofrontal tubercles</emphasis>
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: absent.
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<emphasis id="7EDDEA8C690E9E1CFF6AFA2E827C6DD9" box="[160,302,1460,1485]" italics="true" pageId="3" pageNumber="323">Laterofrontal</emphasis>
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+
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<emphasis id="7EDDEA8C690E9E1CFEAAFA2E810D6DD9" box="[352,607,1460,1485]" italics="true" pageId="3" pageNumber="323">mediofrontal tubercles</emphasis>
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: conspicuous, acute, and projected forward.
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<emphasis id="7EDDEA8C690E9E1CFF6AFA4C82346DFB" box="[160,358,1494,1519]" italics="true" pageId="3" pageNumber="323">Mediofrontal area</emphasis>
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: caliciform 2.1x wider than long, flat, and deep; with coarse punctuations scattered throughout the anterior region.
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<emphasis id="7EDDEA8C690E9E1CFD15FA6380C76E06" box="[735,917,1529,1554]" italics="true" pageId="3" pageNumber="323">Cephalic nodule</emphasis>
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: large but not well marked and rounded shape.
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<emphasis id="7EDDEA8C690E9E1CFE36F98681F36E21" box="[508,673,1564,1589]" italics="true" pageId="3" pageNumber="323">Inner tubercles</emphasis>
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: large, conspicuous with obtuse apexes, smaller than the laterofrontal + mediofrontal tubercles, of which they are distinctly separated but located closer to these than the apex of the central tubercle.
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<emphasis id="7EDDEA8C690E9E1CFC3BF9FB83B36E89" italics="true" pageId="3" pageNumber="323">Anterofrontal ridges</emphasis>
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: straight, smaller, and weaker than the posterofrontal ridges.
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<emphasis id="7EDDEA8C690E9E1CFC51F91E872F6E89" box="[923,1149,1668,1693]" italics="true" pageId="3" pageNumber="323">Posterofrontal ridges</emphasis>
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: high, strong, and slightly sinuous, beginning at the apex of the central tubercle.
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<emphasis id="7EDDEA8C69099E1BFF6AF9A282236E45" box="[160,369,1592,1617]" italics="true" pageId="4" pageNumber="324">Laterofrontal areas</emphasis>
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: flat and smooth surface.
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<emphasis id="7EDDEA8C69099E1BFD50F9A2801A6E45" box="[666,840,1592,1617]" italics="true" pageId="4" pageNumber="324">Central tubercle</emphasis>
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: conical, low, with non-free apex.
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<emphasis id="7EDDEA8C69099E1BFF25F9C1815B6E60" box="[239,521,1627,1652]" italics="true" pageId="4" pageNumber="324">Lateroposterior tubercles</emphasis>
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: conspicuous, close to the central tubercle, with a rounded shape.
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<emphasis id="7EDDEA8C69099E1BFEBCF9E4817B6E83" box="[374,553,1662,1687]" italics="true" pageId="4" pageNumber="324">Postfrontal area</emphasis>
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: flat, with a surface containing many fine punctuations.
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<emphasis id="7EDDEA8C69099E1BFF24F93A82936EAD" box="[238,449,1696,1721]" italics="true" pageId="4" pageNumber="324">Postfrontal groove</emphasis>
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: well marked and with a slight notch located medially.
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<emphasis id="7EDDEA8C69089E1AFF6AFA2E82F66DD9" box="[160,420,1460,1485]" italics="true" pageId="5" pageNumber="325">Lateropostfrontal areas</emphasis>
|
||
: shallow, with smooth and glabrous surface; containing some punctuations.
|
||
<emphasis id="7EDDEA8C69089E1AFE83FA4C81406DFB" box="[329,530,1494,1519]" italics="true" pageId="5" pageNumber="325">Epicranial sutures</emphasis>
|
||
: poorly marked.
|
||
<emphasis id="7EDDEA8C69089E1AFD2CFA4C80D56DFB" box="[742,903,1494,1519]" italics="true" pageId="5" pageNumber="325">Epicranial pits</emphasis>
|
||
: deep in the region closest to the anterior angles of the head and shallow when close to the laterofrontal areas, but very visible.
|
||
<emphasis id="7EDDEA8C69089E1AFE67F98681B16E21" box="[429,739,1564,1589]" italics="true" pageId="5" pageNumber="325">Anterior angles of the head</emphasis>
|
||
: well developed, with acute apices, slightly smaller than the laterofrontal + mediofrontal tubercles.
|
||
<emphasis id="7EDDEA8C69089E1AFCB5F9A487746E43" box="[895,1062,1598,1623]" italics="true" pageId="5" pageNumber="325">Canthus ocular</emphasis>
|
||
: apexes rounded not reaching half eye.
|
||
<emphasis id="7EDDEA8C69089E1AFDDEF9FB81296E6E" box="[532,635,1633,1658]" italics="true" pageId="5" pageNumber="325">Antennas</emphasis>
|
||
: trilamellate, with robust lamellae, the distal being wider than the other two.
|
||
<emphasis id="7EDDEA8C69089E1AFDDDF91F81F66E89" bold="true" box="[535,676,1669,1693]" pageId="5" pageNumber="325">Mouthparts</emphasis>
|
||
(
|
||
<figureCitation id="D4922A1B69089E1AFD7EF91F804E6E89" box="[692,796,1669,1693]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="5" pageNumber="325">Figures 1</figureCitation>
|
||
(b), 2(b)):
|
||
<taxonomicName id="8BA94D1D69089E1AFC43F91E80996E89" box="[905,971,1668,1693]" class="Cestoda" family="Diphyllobothriidae" genus="Ligula" kingdom="Animalia" order="Diphyllobothriidea" pageId="5" pageNumber="325" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="7EDDEA8C69089E1AFC43F91E80996E89" box="[905,971,1668,1693]" italics="true" pageId="5" pageNumber="325">Ligula</emphasis>
|
||
</taxonomicName>
|
||
: tridentate with a larger middle tooth and narrower than the lateral teeth.
|
||
<emphasis id="7EDDEA8C69089E1AFC92F93C87646EAB" box="[856,1078,1702,1727]" italics="true" pageId="5" pageNumber="325">Hypostomal process</emphasis>
|
||
: wide, glabrous and separated from mentum.
|
||
<emphasis id="7EDDEA8C690B9E19FDA4FF1780DC68B2" box="[622,910,141,166]" italics="true" pageId="6" pageNumber="326">Mentum mediobasal area</emphasis>
|
||
: not dilated, smooth and glabrous, with protruding anterior region without notching in the middle.
|
||
<emphasis id="7EDDEA8C690B9E19FBE2FF2A827A68F8" italics="true" pageId="6" pageNumber="326">Mentum lateral lobes</emphasis>
|
||
: rounded apexes.
|
||
<emphasis id="7EDDEA8C690B9E19FDCBFF4981BE68F8" box="[513,748,211,236]" italics="true" pageId="6" pageNumber="326">Mentum lateral scars</emphasis>
|
||
: shallow, glabrous, inpunctate and rounded shape.
|
||
<emphasis id="7EDDEA8C690B9E19FEAAFF6F8282691A" box="[352,464,245,270]" italics="true" pageId="6" pageNumber="326">Mandibles</emphasis>
|
||
: incisor lobe with three well-formed teeth at apex; robust suprainternal teeth; inconspicuous infrabasal pits.
|
||
<emphasis id="7EDDEA8C690B9E19FD11FE8280786925" box="[731,810,280,305]" italics="true" pageId="6" pageNumber="326">Maxilla</emphasis>
|
||
: lacinia bidentate at the apex.
|
||
<emphasis id="7EDDEA8C690B9E19FF6AFEA6824B6940" bold="true" box="[160,281,316,340]" pageId="6" pageNumber="326">Prothorax</emphasis>
|
||
. Pronotum (
|
||
<figureCitation id="D4922A1B690B9E19FE78FEA6814B6940" box="[434,537,316,340]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="6" pageNumber="326">Figures 1</figureCitation>
|
||
(a,c), 2(a
|
||
<emphasis id="7EDDEA8C690B9E19FD48FEA181DD6947" box="[642,655,315,339]" italics="true" pageId="6" pageNumber="326">–</emphasis>
|
||
c)):
|
||
<emphasis id="7EDDEA8C690B9E19FD77FEA180046940" box="[701,854,315,340]" italics="true" pageId="6" pageNumber="326">Anterior edge</emphasis>
|
||
: straight.
|
||
<emphasis id="7EDDEA8C690B9E19FC18FEA1872C6940" box="[978,1150,315,340]" italics="true" pageId="6" pageNumber="326">Anterior angles</emphasis>
|
||
: acute and fairly protracted.
|
||
<emphasis id="7EDDEA8C690B9E19FE2AFEC781C46962" box="[480,662,349,374]" italics="true" pageId="6" pageNumber="326">Marginal groove</emphasis>
|
||
: well marked, deep and with coarse punctures throughout its length, without dilation at the apex; reaching four-fifth of pronotal width.
|
||
<emphasis id="7EDDEA8C690B9E19FF3FFE3982C169A8" box="[245,403,419,444]" italics="true" pageId="6" pageNumber="326">Lateral fossae</emphasis>
|
||
: small, well marked and softly deep; slightly rounded shape.
|
||
<emphasis id="7EDDEA8C690B9E19FF6AFE5F82C469CA" box="[160,406,453,478]" italics="true" pageId="6" pageNumber="326">Pronotal punctuations</emphasis>
|
||
: coarse and fairly dense, diffused throughout pronotum, concentrated mainly above the lateral fossae. Prosternum (
|
||
<figureCitation id="D4922A1B690B9E19FD26FE73801D6A15" box="[748,847,489,513]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="6" pageNumber="326">Figures 1</figureCitation>
|
||
(b), 2(d)):
|
||
<emphasis id="7EDDEA8C690B9E19FC0AFE7287026A15" box="[960,1104,488,513]" italics="true" pageId="6" pageNumber="326">Prepisternum</emphasis>
|
||
: not dilated in the anterior region; glabrous.
|
||
<emphasis id="7EDDEA8C690B9E19FDA8FD9181B56A30" box="[610,743,523,548]" italics="true" pageId="6" pageNumber="326">Prepimerum</emphasis>
|
||
: glabrous.
|
||
<emphasis id="7EDDEA8C690B9E19FCA9FD9180A16A30" box="[867,1011,523,548]" italics="true" pageId="6" pageNumber="326">Prosternelum</emphasis>
|
||
: rhomboidal shape with narrow apex, slightly obtuse.
|
||
<emphasis id="7EDDEA8C690B9E19FD47FDB480636A52" bold="true" box="[653,817,558,582]" pageId="6" pageNumber="326">Mesosternum</emphasis>
|
||
(
|
||
<figureCitation id="D4922A1B690B9E19FC8FFDB480FC6A52" box="[837,942,558,582]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="6" pageNumber="326">Figures 1</figureCitation>
|
||
(b), 2(d)): smooth, shiny, and glabrous.
|
||
<emphasis id="7EDDEA8C690B9E19FE50FDCA810D6A7D" box="[410,607,592,617]" italics="true" pageId="6" pageNumber="326">Mesosternal scars</emphasis>
|
||
: long, broad, well marked and deep; without punctuations or pubescence.
|
||
<emphasis id="7EDDEA8C690B9E19FE39FDE981C66A9F" bold="true" box="[499,660,627,651]" pageId="6" pageNumber="326">Metasternum</emphasis>
|
||
(
|
||
<figureCitation id="D4922A1B690B9E19FD6EFDE9805B6A9F" box="[676,777,627,651]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="6" pageNumber="326">Figures 1</figureCitation>
|
||
(b), 2(e)):
|
||
<emphasis id="7EDDEA8C690B9E19FCB3FDE8877F6A9F" box="[889,1069,626,651]" italics="true" pageId="6" pageNumber="326">Metasternal disc</emphasis>
|
||
: poorly delimited, but more dilated than the lateral region of the metasternum; punctuations in the posteromedian region.
|
||
<emphasis id="7EDDEA8C690B9E19FE29FD2280526AC5" box="[483,768,696,721]" italics="true" pageId="6" pageNumber="326">Metasternal punctuations</emphasis>
|
||
: group of punctures only in the lateroposterior region.
|
||
<emphasis id="7EDDEA8C690B9E19FE60FD4081E66AE7" box="[426,692,730,755]" italics="true" pageId="6" pageNumber="326">Metasternal pubescence</emphasis>
|
||
: absent.
|
||
<emphasis id="7EDDEA8C690B9E19FCEAFD4087146AE7" box="[800,1094,730,755]" italics="true" pageId="6" pageNumber="326">Metasternal lateral groove</emphasis>
|
||
: narrow, thinner than mesotibiae, deep, without punctures or bristles.
|
||
<emphasis id="7EDDEA8C690B9E19FC6EFD6480B86B02" bold="true" box="[932,1002,766,790]" pageId="6" pageNumber="326">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="D4922A1B690B9E19FC31FD6487076B02" box="[1019,1109,766,790]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="6" pageNumber="326">Figure 1</figureCitation>
|
||
(a)): Approximately 2.5x longer and 1.1x wider than pronotum.
|
||
<emphasis id="7EDDEA8C690B9E19FC94FCBA80CE6B2D" box="[862,924,800,825]" italics="true" pageId="6" pageNumber="326">Striae</emphasis>
|
||
: narrower than the interstriae, marked with round and fine punctures, well defined in both the dorsal and lateral striae.
|
||
<taxonomicName id="8BA94D1D690B9E19FEF3FCFF82CD6B6A" authorityName="Walker" authorityYear="1861" box="[313,415,869,894]" class="Insecta" family="Uraniidae" genus="Epipleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="326" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7EDDEA8C690B9E19FEF3FCFF82CD6B6A" box="[313,415,869,894]" italics="true" pageId="6" pageNumber="326">Epipleura</emphasis>
|
||
</taxonomicName>
|
||
: glabrous.
|
||
<emphasis id="7EDDEA8C690B9E19FDD7FCFF813E6B6A" box="[541,620,869,894]" italics="true" pageId="6" pageNumber="326">Humeri</emphasis>
|
||
: glabrous.
|
||
<emphasis id="7EDDEA8C690B9E19FD21FCFC80706B6A" bold="true" box="[747,802,870,894]" pageId="6" pageNumber="326">Legs</emphasis>
|
||
(
|
||
<figureCitation id="D4922A1B690B9E19FCF8FCFC80C76B6A" box="[818,917,870,894]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="6" pageNumber="326">Figures 1</figureCitation>
|
||
(a,b), 2(e)):
|
||
<emphasis id="7EDDEA8C690B9E19FBD3FCFF872F6B6A" box="[1049,1149,869,894]" italics="true" pageId="6" pageNumber="326">Profemur</emphasis>
|
||
: ventral anterior border with well-marked groove, not reaching the apex of the profemur; ventral posterior border with few setae only near the apex.
|
||
<emphasis id="7EDDEA8C690B9E19FC5AFC3080A36BD7" box="[912,1009,938,963]" italics="true" pageId="6" pageNumber="326">Protibiae</emphasis>
|
||
: not dilated.
|
||
<emphasis id="7EDDEA8C690B9E19FF6AFC5782446BF2" box="[160,278,973,998]" italics="true" pageId="6" pageNumber="326">Mesotibiae</emphasis>
|
||
: three small spines on the outer face.
|
||
<emphasis id="7EDDEA8C690B9E19FD20FC57800D6BF2" box="[746,863,973,998]" italics="true" pageId="6" pageNumber="326">Metatibiae</emphasis>
|
||
: one small spine on the outer face.
|
||
<emphasis id="7EDDEA8C690B9E19FEEAFC6B82C46C1D" bold="true" box="[288,406,1009,1033]" pageId="6" pageNumber="326">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="D4922A1B690B9E19FE6CFC6B82AD6C1D" box="[422,511,1009,1033]" captionStart="Figure 1" captionStartId="4.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-56@4.[163,1153,142,1451]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="6" pageNumber="326">Figure 1</figureCitation>
|
||
(b)): sternite VII with full groove and well marked; rough sides.
|
||
<emphasis id="7EDDEA8C690B9E19FF20FB8982336C3F" bold="true" box="[234,353,1043,1067]" pageId="6" pageNumber="326">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="D4922A1B690B9E19FEBEFB8982846C3F" box="[372,470,1043,1067]" captionStart="Figure 2" captionStartId="5.[160,225,1329,1352]" captionTargetBox="[163,1153,142,1297]" captionTargetId="figure-115@5.[163,1153,142,1298]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) beneshi Hincks, 1950 male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4609061" httpUri="https://zenodo.org/record/4609061/files/figure.png" pageId="6" pageNumber="326">Figure 2</figureCitation>
|
||
(f)): Median lobe wider than the parameres and the basal piece together and almost as long as these. In ventral view completely sclerotised with a recess in the basal region; basal piece and parameres fused forming a tegmen; parameres projections reaching less than one-third of the length of the median lobe; arched parameres anterior edge. In lateral view, the apex of the parameres is acute and slightly rounded. In dorsal view with widely separated parameres.
|
||
</paragraph>
|
||
<caption id="18D6661669099E1BFF6AFA5082016DEB" ID-DOI="http://doi.org/10.5281/zenodo.4609057" ID-Zenodo-Dep="4609057" httpUri="https://zenodo.org/record/4609057/files/figure.png" pageId="4" pageNumber="324" startId="4.[160,225,1482,1505]" targetBox="[163,1153,142,1451]" targetPageId="4">
|
||
<paragraph id="4C16369E69099E1BFF6AFA5082016DEB" blockId="4.[160,1156,1482,1535]" pageId="4" pageNumber="324">
|
||
<emphasis id="7EDDEA8C69099E1BFF6AFA5083AB6DF5" bold="true" box="[160,249,1482,1505]" pageId="4" pageNumber="324">Figure 1.</emphasis>
|
||
<taxonomicName id="8BA94D1D69099E1BFF34FA50812F6DF5" authority="Hincks, 1950" authorityName="Hincks" authorityYear="1950" box="[254,637,1482,1505]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="324" phylum="Arthropoda" rank="species" species="beneshi" subGenus="Pertinax">
|
||
<emphasis id="7EDDEA8C69099E1BFF34FA50821C6DF5" box="[254,334,1482,1505]" italics="true" pageId="4" pageNumber="324">Passalus</emphasis>
|
||
(
|
||
<emphasis id="7EDDEA8C69099E1BFE91FA5082FB6DF5" box="[347,425,1482,1505]" italics="true" pageId="4" pageNumber="324">Pertinax</emphasis>
|
||
)
|
||
<emphasis id="7EDDEA8C69099E1BFE7EFA5082AF6DF5" box="[436,509,1482,1505]" italics="true" pageId="4" pageNumber="324">beneshi</emphasis>
|
||
<bibRefCitation id="28384B6F69099E1BFDC8FA50812F6DF5" author="Hincks WD" box="[514,637,1482,1505]" pageId="4" pageNumber="324" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks, 1950</bibRefCitation>
|
||
</taxonomicName>
|
||
male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="18D6661669089E1AFF6AFAAB80ED6D72" ID-DOI="http://doi.org/10.5281/zenodo.4609061" ID-Zenodo-Dep="4609061" httpUri="https://zenodo.org/record/4609061/files/figure.png" pageId="5" pageNumber="325" startId="5.[160,225,1329,1352]" targetBox="[163,1153,142,1297]" targetPageId="5">
|
||
<paragraph id="4C16369E69089E1AFF6AFAAB80ED6D72" blockId="5.[160,1155,1328,1382]" pageId="5" pageNumber="325">
|
||
<emphasis id="7EDDEA8C69089E1AFF6AFAAB83A86D5C" bold="true" box="[160,250,1329,1352]" pageId="5" pageNumber="325">Figure 2.</emphasis>
|
||
<taxonomicName id="8BA94D1D69089E1AFECBFAAA81D56D5C" authority="Hincks, 1950" authorityName="Hincks" authorityYear="1950" box="[257,647,1328,1352]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="325" phylum="Arthropoda" rank="species" species="beneshi" subGenus="Pertinax">
|
||
<emphasis id="7EDDEA8C69089E1AFECBFAAA82036D53" box="[257,337,1328,1351]" italics="true" pageId="5" pageNumber="325">Passalus</emphasis>
|
||
(
|
||
<emphasis id="7EDDEA8C69089E1AFE95FAAA82FF6D53" box="[351,429,1328,1351]" italics="true" pageId="5" pageNumber="325">Pertinax</emphasis>
|
||
)
|
||
<emphasis id="7EDDEA8C69089E1AFE71FAAA81566D53" box="[443,516,1328,1351]" italics="true" pageId="5" pageNumber="325">beneshi</emphasis>
|
||
<bibRefCitation id="28384B6F69089E1AFDC0FAAB81D56D5C" author="Hincks WD" box="[522,647,1329,1352]" pageId="5" pageNumber="325" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks, 1950</bibRefCitation>
|
||
</taxonomicName>
|
||
male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4C16369E690B9E18FF6AFA8E829968DD" blockId="6.[160,1156,1300,1707]" lastBlockId="7.[160,1155,142,201]" lastPageId="7" lastPageNumber="327" pageId="6" pageNumber="326">
|
||
<emphasis id="7EDDEA8C690B9E19FF6AFA8E827D6D3A" bold="true" box="[160,303,1300,1326]" italics="true" pageId="6" pageNumber="326">Dimensions.</emphasis>
|
||
Total length:
|
||
<quantity id="8B519B7B690B9E19FE17FA8C81606D3A" box="[477,562,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="326" unit="mm" value="17.0">17 mm</quantity>
|
||
; cephalic length:
|
||
<quantity id="8B519B7B690B9E19FCC8FA8C800C6D3A" box="[770,862,1302,1326]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="326" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; cephalic width:
|
||
<quantity id="8B519B7B690B9E19FBEFFA8C87D26D3A" box="[1061,1152,1302,1326]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="326" unit="mm" value="3.6">3.6 mm</quantity>
|
||
; mediofrontal area length:
|
||
<quantity id="8B519B7B690B9E19FE1FFAA381636D45" box="[469,561,1337,1361]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="326" unit="mm" value="0.7">0.7 mm</quantity>
|
||
; mediofrontal area width:
|
||
<quantity id="8B519B7B690B9E19FCA7FAA380986D45" box="[877,970,1337,1361]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="326" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; canthus ocular length:
|
||
<quantity id="8B519B7B690B9E19FF3DFAC682026D60" box="[247,336,1372,1396]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="326" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; canthus ocular width:
|
||
<quantity id="8B519B7B690B9E19FD90FAC681E16D60" box="[602,691,1372,1396]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="326" unit="mm" value="0.3">0.3 mm</quantity>
|
||
; area of the mediofrontal area:
|
||
<quantity id="8B519B7B690B9E19FBD7FAC687216D60" box="[1053,1139,1372,1396]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="326" unit="mm" value="0.7">0.7 mm</quantity>
|
||
<superScript id="BBDC9BD6690B9E19FBB9FACC872F6D72" attach="left" box="[1139,1149,1366,1382]" fontSize="7" pageId="6" pageNumber="326">2</superScript>
|
||
; mandibles external angle: 141º; antennal club length:
|
||
<quantity id="8B519B7B690B9E19FCEFFAE480D36D82" box="[805,897,1406,1430]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="326" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; antennal club width:
|
||
<quantity id="8B519B7B690B9E19FF6AFA3B83AE6DAD" box="[160,252,1441,1465]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="326" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; distal lamella length:
|
||
<quantity id="8B519B7B690B9E19FDC0FA3B81346DAD" box="[522,614,1441,1465]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="326" unit="mm" value="0.4">0.4 mm</quantity>
|
||
; medial lamella length:
|
||
<quantity id="8B519B7B690B9E19FC4EFA3B80B26DAD" box="[900,992,1441,1465]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="326" unit="mm" value="0.2">0.2 mm</quantity>
|
||
; width of the mentum at the lateral scars:
|
||
<quantity id="8B519B7B690B9E19FE29FA5E81696DC8" box="[483,571,1476,1500]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="326" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; mentum mediobasal area width:
|
||
<quantity id="8B519B7B690B9E19FC76FA5E87466DC8" box="[956,1044,1476,1500]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="326" unit="mm" value="0.8">0.8 mm</quantity>
|
||
; diameter of the mentum lateral scars:
|
||
<quantity id="8B519B7B690B9E19FE22FA7C81136DEA" box="[488,577,1510,1534]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="326" unit="mm" value="0.4">0.4 mm</quantity>
|
||
; pronotal length:
|
||
<quantity id="8B519B7B690B9E19FCC6FA7C80376DEA" box="[780,869,1510,1534]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="326" unit="mm" value="3.9">3.9 mm</quantity>
|
||
; pronotal width:
|
||
<quantity id="8B519B7B690B9E19FBE2FA7C87D26DEA" box="[1064,1152,1510,1534]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="326" unit="mm" value="5.2">5.2 mm</quantity>
|
||
; length of the pronotal anterior groove:
|
||
<quantity id="8B519B7B690B9E19FDB9F993819F6E35" box="[627,717,1545,1569]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="326" unit="mm" value="1.8">1.8 mm</quantity>
|
||
; elytral length:
|
||
<quantity id="8B519B7B690B9E19FC4FF993808D6E35" box="[901,991,1545,1569]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="6" pageNumber="326" unit="mm" value="9.9">9.9 mm</quantity>
|
||
; elytral width:
|
||
<quantity id="8B519B7B690B9E19FF6AF9B683A56E50" box="[160,247,1580,1604]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="326" unit="mm" value="6.1">6.1 mm</quantity>
|
||
; humeral width:
|
||
<quantity id="8B519B7B690B9E19FE79F9B681586E50" box="[435,522,1580,1604]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="326" unit="mm" value="5.5">5.5 mm</quantity>
|
||
; profemur length:
|
||
<quantity id="8B519B7B690B9E19FD11F9B680606E50" box="[731,818,1580,1604]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="326" unit="mm" value="2.2">2.2 mm</quantity>
|
||
; length of the anterior ventral groove of the profemur:
|
||
<quantity id="8B519B7B690B9E19FE65F9D481566E72" box="[431,516,1614,1638]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="326" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; protibiae width:
|
||
<quantity id="8B519B7B690B9E19FD09F9D4804A6E72" box="[707,792,1614,1638]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="326" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; protarsal length:
|
||
<quantity id="8B519B7B690B9E19FC16F9D487606E72" box="[988,1074,1614,1638]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="326" unit="mm" value="1.7">1.7 mm</quantity>
|
||
; length of the last protarsomer:
|
||
<quantity id="8B519B7B690B9E19FE72F9EB81436E9D" box="[440,529,1649,1673]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="326" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; mesotibiae width:
|
||
<quantity id="8B519B7B690B9E19FD38F9EB80196E9D" box="[754,843,1649,1673]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="326" unit="mm" value="0.3">0.3 mm</quantity>
|
||
; metasternal lateral groove width:
|
||
<quantity id="8B519B7B690B9E19FF3BF90982196EBF" box="[241,331,1683,1707]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="326" unit="mm" value="0.2">0.2 mm</quantity>
|
||
; metasternal disc length:
|
||
<quantity id="8B519B7B690B9E19FDBDF90981806EBF" box="[631,722,1683,1707]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="326" unit="mm" value="3.9">3.9 mm</quantity>
|
||
; abdomen length:
|
||
<quantity id="8B519B7B690B9E19FC65F90987586EBF" box="[943,1034,1683,1707]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="326" unit="mm" value="4.5">4.5 mm</quantity>
|
||
; aedeagus length:
|
||
<quantity id="8B519B7B690A9E18FF36FF14820B68B2" box="[252,345,142,166]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="327" unit="mm" value="1.7">1.7 mm</quantity>
|
||
; aedeagus width:
|
||
<quantity id="8B519B7B690A9E18FDFFFF1481C168B2" box="[565,659,142,166]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="327" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; median lobe length:
|
||
<quantity id="8B519B7B690A9E18FC55FF1480AE68B2" box="[927,1020,142,166]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="327" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; paremeres projection lenght:
|
||
<quantity id="8B519B7B690A9E18FEBAFF2B829A68DD" box="[368,456,177,201]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="327" unit="mm" value="0.1">0.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B36515690A9E18FF6AFF6E81E36962" pageId="7" pageNumber="327" type="materials_examined">
|
||
<paragraph id="4C16369E690A9E18FF6AFF6E81E36962" blockId="7.[160,1155,244,374]" pageId="7" pageNumber="327">
|
||
<emphasis id="7EDDEA8C690A9E18FF6AFF6E817D691A" bold="true" box="[160,559,244,270]" italics="true" pageId="7" pageNumber="327">Material examined: type material.</emphasis>
|
||
<typeStatus id="9312883C690A9E18FDF5FF6C81F4691A" box="[575,678,246,270]" pageId="7" pageNumber="327" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<specimenCount id="5AAFFD17690A9E18FD7EFF6C819A691A" box="[692,712,246,270]" pageId="7" pageNumber="327" type="male">♂</specimenCount>
|
||
) F2439.48.
|
||
<collectingCountry id="34BE760E690A9E18FC8CFF6C80FD691A" box="[838,943,246,270]" name="Panama" pageId="7" pageNumber="327">PANAMA</collectingCountry>
|
||
: XX Plantation/Rep of
|
||
<collectingRegion id="8E6DF87C690A9E18FF00FE83827B6925" box="[202,297,281,305]" country="Panama" name="Panama" pageId="7" pageNumber="327">Panama</collectingRegion>
|
||
/2/17/30//Blackwelder/Collection//via B. Benesh/North Chicago//
|
||
<taxonomicName id="8BA94D1D690A9E18FBD3FE83872E6925" authorityName="Fabricius" authorityYear="1792" box="[1049,1148,281,305]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="genus">Passalus</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="8BA94D1D690A9E18FF6CFEA682576940" authorityName="Kaup" authorityYear="1869" box="[166,261,316,340]" class="Insecta" family="Passalidae" genus="Pertinax" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="genus">Pertinax</taxonomicName>
|
||
) Type/
|
||
<taxonomicName id="8BA94D1D690A9E18FE92FEA681596940" authority="Hincks" authorityName="Hincks" authorityYear="1950" box="[344,523,316,340]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="beneshi">beneshi Hincks</taxonomicName>
|
||
/det. W.D. Hincks.//Manchester Museum/
|
||
<typeStatus id="9312883C690A9E18FC11FEA687336940" box="[987,1121,316,340]" pageId="7" pageNumber="327" type="holotype">HOLOTYPE</typeStatus>
|
||
//
|
||
<specimenCount id="5AAFFD17690A9E18FBA5FEA187D16947" box="[1135,1155,315,339]" pageId="7" pageNumber="327" type="male">♂</specimenCount>
|
||
dissec/S. Boucher det. 2012//F2439.48 (
|
||
<collectionCode id="2AB8AE5B690A9E18FD92FEC481F56962" box="[600,679,350,374]" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="7" pageNumber="327">MMUE</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B36515690A9E18FF6AFE3881566A98" pageId="7" pageNumber="327" type="discussion">
|
||
<paragraph id="4C16369E690A9E18FF6AFE3881566A98" blockId="7.[160,1155,418,652]" pageId="7" pageNumber="327">
|
||
<emphasis id="7EDDEA8C690A9E18FF6AFE38825E69A8" bold="true" box="[160,268,418,444]" italics="true" pageId="7" pageNumber="327">Remarks.</emphasis>
|
||
<taxonomicName id="8BA94D1D690A9E18FED7FE39828169A8" authorityName="Hincks" authorityYear="1950" box="[285,467,419,444]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="beneshi">
|
||
<emphasis id="7EDDEA8C690A9E18FED7FE39828169A8" box="[285,467,419,444]" italics="true" pageId="7" pageNumber="327">Passalus beneshi</emphasis>
|
||
</taxonomicName>
|
||
as well as
|
||
<taxonomicName id="8BA94D1D690A9E18FD85FE39806E69A8" authority="(Kuwert)" baseAuthorityName="Kuwert" box="[591,828,419,444]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="irregularis">
|
||
<emphasis id="7EDDEA8C690A9E18FD85FE39818469A8" box="[591,726,419,444]" italics="true" pageId="7" pageNumber="327">P. irregularis</emphasis>
|
||
(Kuwert)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8BA94D1D690A9E18FCBDFE39870A69A8" authority="Hincks" authorityName="Hincks, 1950:" authorityYear="1044" box="[887,1112,419,444]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="umbriensis">
|
||
<emphasis id="7EDDEA8C690A9E18FCBDFE39875469A8" box="[887,1030,419,444]" italics="true" pageId="7" pageNumber="327">P. umbriensis</emphasis>
|
||
Hincks
|
||
</taxonomicName>
|
||
are part of a group of species that present small body size and large eyes with abundant pronotal punctuations and reduced mesotibial spines.
|
||
<taxonomicName id="8BA94D1D690A9E18FD34FE7280386A15" box="[766,874,488,513]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="benehsi">
|
||
<emphasis id="7EDDEA8C690A9E18FD34FE7280386A15" box="[766,874,488,513]" italics="true" pageId="7" pageNumber="327">P. benehsi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="8BA94D1D690A9E18FC3CFE7287D16A15" authorityName="Hincks, 1950:" authorityYear="1044" box="[1014,1155,488,513]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="umbriensis">
|
||
<emphasis id="7EDDEA8C690A9E18FC3CFE7287D16A15" box="[1014,1155,488,513]" italics="true" pageId="7" pageNumber="327">P. umbriensis</emphasis>
|
||
</taxonomicName>
|
||
by presenting conspicuous inner tubercles and anterofrontal ridges, notch in the middle of the frons with not raised edges, more abundant pronotal punctuations reaching more than three-fourths of pronotal length; of
|
||
<taxonomicName id="8BA94D1D690A9E18FD43FDCA80466A7D" box="[649,788,592,617]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="irregularis">
|
||
<emphasis id="7EDDEA8C690A9E18FD43FDCA80466A7D" box="[649,788,592,617]" italics="true" pageId="7" pageNumber="327">P. irregularis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished mainly by the
|
||
<taxonomicName id="8BA94D1D690A9E18FF6AFDEE82586A98" authorityName="Walker" authorityYear="1861" box="[160,266,628,652]" class="Insecta" family="Uraniidae" genus="Epipleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="347" phylum="Arthropoda" rank="genus">epipleura</taxonomicName>
|
||
and humeri glabrous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B36515690A9E18FF6AFD2281526AE0" pageId="7" pageNumber="327" type="distribution">
|
||
<paragraph id="4C16369E690A9E18FF6AFD2281526AE0" blockId="7.[160,1155,696,756]" pageId="7" pageNumber="327">
|
||
<emphasis id="7EDDEA8C690A9E18FF6AFD2282616AC6" bold="true" box="[160,307,696,722]" italics="true" pageId="7" pageNumber="327">Distribution.</emphasis>
|
||
<collectingCountry id="34BE760E690A9E18FE89FD2382CF6AC5" box="[323,413,697,721]" name="Panama" pageId="7" pageNumber="327">Panama</collectingCountry>
|
||
(locality
|
||
<typeStatus id="9312883C690A9E18FDCFFD23816A6AC5" box="[517,568,697,721]" pageId="7" pageNumber="327">type</typeStatus>
|
||
) and
|
||
<collectingCountry id="34BE760E690A9E18FDB0FD2381B46AC5" box="[634,742,697,721]" name="Colombia" pageId="7" pageNumber="327">Colombia</collectingCountry>
|
||
(
|
||
<collectingRegion id="8E6DF87C690A9E18FD3EFD2380176AC5" box="[756,837,697,721]" country="Colombia" name="Boyaca" pageId="7" pageNumber="327">Boyacá</collectingRegion>
|
||
and
|
||
<collectingRegion id="8E6DF87C690A9E18FC4BFD2387776AC5" box="[897,1061,697,721]" country="Colombia" name="Cundinamarca" pageId="7" pageNumber="327">Cundinamarca</collectingRegion>
|
||
) (Reyes- Castillo and Amat-García 2003).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |