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<mods:title id="552C2323CDFD93A8E5EB566B222DA9BB">First description of the male of Cryptothele verrucosa L. Koch, 1872 (Araneae), the type species of the genus</mods:title>
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<mods:namePart id="9D60DA0662DA1B57E4F57B147E7717FC">Marusik, Yuri M.</mods:namePart>
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<mods:namePart id="30D2418728D900D68013F0A3AF8D1B6F">Omelko, Mikhail M.</mods:namePart>
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<treatment id="B7AC82CAFDCBE3795C4B7528886DEDD2" ID-GBIF-Taxon="152049585" LSID="urn:lsid:plazi:treatment:B7AC82CAFDCBE3795C4B7528886DEDD2" httpUri="http://treatment.plazi.org/id/B7AC82CAFDCBE3795C4B7528886DEDD2" lastPageId="4" lastPageNumber="35" pageId="1" pageNumber="32" scope_order="Araneae">
<subSubSection id="9F290D9B3534FC697BAF487919B205D5" pageId="1" pageNumber="32" type="nomenclature">
<paragraph id="B5768C7109B098C7031A49A52AEB1B3F" pageId="1" pageNumber="32">
<taxonomicName id="3C4356D1F671A351D6BEA2371C0DBB0C" ID-CoL="3235Q" LSID="http://species-id.net/wiki/Cryptothele_verrucosa" authority="L. Koch, 1872" authorityName="L. Koch" authorityYear="1872" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa L. Koch, 1872</taxonomicName>
Figs 1-3, 6-11
</paragraph>
</subSubSection>
<subSubSection id="17446DA657D1C6BFA8CAB9F38462B532" pageId="1" pageNumber="32" type="reference_group">
<paragraph id="0387F18C218E3394967CCB87C11868E6" pageId="1" pageNumber="32">
<taxonomicName id="F51CC700B68C7C8B610728E80BAA03E6" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
L. Koch, 1872: 240, pl. 20, f. 2 (D♀).
</paragraph>
</subSubSection>
<subSubSection id="37B33937C3F10D4E713536C7E7311B53" pageId="1" pageNumber="32" type="material examined">
<paragraph id="FCC44A751E4F089C0EBA7B7028A6D9C2" pageId="1" pageNumber="32">Material examined.</paragraph>
<paragraph id="631A47B913ABF7F30EF20D43E9C22C53" pageId="1" pageNumber="32">FIJI: 1♂ (ZMUT AA 5.828), Viti Levu, Suva rain forest, 26.05.1973 (J.M. Ackerman).</paragraph>
</subSubSection>
<subSubSection id="BA10E140806EF94C01E4CC7C29459C5A" pageId="1" pageNumber="32" type="diagnosis">
<paragraph id="558DA4D94C4F17F1D269E5525DBF2299" pageId="1" pageNumber="32">Diagnosis.</paragraph>
<paragraph id="87A3F70D3431BD5C0E92576B7FC98111" pageId="1" pageNumber="32">
<taxonomicName id="94F3E81D05F567384D407FA450A8F065" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
differs from
<taxonomicName id="B8BA81D2B33F0F6B65E8E127E2782DD1" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
, the only properly described species in the genus, by lack of carapace pattern, much more heavy camouflage of dirt that covers whole body, straight row of posterior eyes, anterior lateral eyes spaced by one diameter (1/2 of diameter in
<taxonomicName id="8D956B22E02DF3376F99AAE1A6B336B8" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
), shape of tibial apophysis (cf. Figs 11, 12), short and broad embolus with two processes (long and filamentous in
<taxonomicName id="6FFBFEAAD308A673BC29B63D88423EC2" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
, Fig. 12).
</paragraph>
<caption id="AF6630F8E9E29FD1449CFC40D5F4AD91" pageId="1" pageNumber="32">
<paragraph id="A338E3AFFB2EAB413BC1BDF4026E3108" pageId="1" pageNumber="32">
Figures 1-5. General appearance of males of
<taxonomicName id="F84B82794648EF5CBBFEDD247E8D3127" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
(1-3) and
<taxonomicName id="CC3E06D3D8F172F1C551226F333F327B" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
(4-5). 1, 4 dorsal 2 ventral 3, 5 frontal 4-5 after
<bibRefCitation id="6B6CBFA68EC34F98008C6DB25FD0F422" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="5" pageNumber="36" pagination="183 - 186" title="Redescription of Cryptothele alluaudi Simon, 1893 (Araneae)." volume="21, 2" year="2012">Marusik and Omelko (2012)</bibRefCitation>
. Abbreviations: AL anterior lateral eye; AM anterior median eye; At anal tubercle, Sn spinneret.
</paragraph>
</caption>
<caption id="C69F79BB0F96FE66FCB90DBEE1A55662" pageId="1" pageNumber="32">
<paragraph id="F575E5FE71D66BA9DBBABC6D2A7256D5" pageId="1" pageNumber="32">
Figures 6-12. Male palp of
<taxonomicName id="5A785A9865D3BEC4A31FFBFE69E0AF2A" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
(6-11) and
<taxonomicName id="3DE74BDB0F050314A626D2D2AFABE377" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
(12). 6 bulbus, ventral 7 bulbus, ventro-prolateral 8, 10 bulbus, retrolateral 9 bulbus, prolateral 11 palp with removed bulbus, retrolateral 12 palp, ventral (after
<bibRefCitation id="F43D1A03F03B1EEEBDFB22710B2FCD24" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="5" pageNumber="36" pagination="183 - 186" title="Redescription of Cryptothele alluaudi Simon, 1893 (Araneae)." volume="21, 2" year="2012">Marusik and Omelko (2012)</bibRefCitation>
). Abbreviations: Co conductor; Ea terminal process of embolus; Eb embolus base; Em embolus; Ep posterior process of embolus; Sd seminal duct; Sp subtegular process; St subtegulum; Te triangle extesion of tegulum; Ts threads of subtegulum.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E55F10CA8FC9C5B4452416EEED6AEF62" lastPageId="4" lastPageNumber="35" pageId="2" pageNumber="33" type="description">
<paragraph id="0987B5D2B8A351F80B236B975277223E" pageId="2" pageNumber="33">
<pageBreakToken id="B0E0C2E47D70B4B9B54C81FB77776145" pageId="2" pageNumber="33" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph id="AAD868FB628FE6AB41D1884921EFCABB" pageId="2" pageNumber="33">Measurements. Total length 5.5, carapace 3.0 long, 2.28 wide. AME 0.14, AME-AME 0.15. Position of metatarsal trichobothria IV 0.9.</paragraph>
<paragraph id="221FB92D540F889FA1FF2BFABDA11C66" pageId="2" pageNumber="33">
Whole body, including sternum and venter of abdomen covered by comouflaging dirt (Figs 1-2). Carapace brown, without pattern, with two rows of hairs aside of median line, these rows are visible after removing the comouflaging dirt. AME spaced by one diameter (Fig. 3), anterior eyes form inverted trapezium (ALE row wider than AME row), posterior eye row almost straight, cephalic area with pit behind posterior median eyes. Leg subeaqual in length, formula 1423. Legs heavily built, with thick
<normalizedToken id="9C4DF5EADE67C855A49F3A317503171A" originalValue="femurtibia">femur-tibia</normalizedToken>
and twice as thin
<normalizedToken id="C7759EFF66B2A136102E173331F26A1E" originalValue="metatasrustarsus">metatasrus-tarsus</normalizedToken>
, border between tarsus and metatarsus poorly visible, metatarsi with terminal trichobothria.
</paragraph>
<paragraph id="C26650644C9D9FF6A0B095D6D5CA3AEF" pageId="2" pageNumber="33">Length of legs and leg joints.</paragraph>
<paragraph id="FCC8A2E8D2679C2A89F3AC4BAEE890D6" pageId="3" pageNumber="34">
<pageBreakToken id="7FE984C3AEEE126D48E5D91705DF51F8" pageId="3" pageNumber="34" start="start">Abdomen</pageBreakToken>
oval, with two spinnerets (Sn) about the size of anal tubercle (At).
</paragraph>
<paragraph id="38CECBF486D1DC5CF0BECF4739A5F0AE" lastPageId="4" lastPageNumber="35" pageId="3" pageNumber="34">
Palp as shown in Figs 6-11. The single specimen examined has both palps expanded. Tibia (Fig. 11) with long retrolateral apophysis tapering in terminal 1/3, dorsal side of apophysis with shallow blunt outgrowth, retrolateral side of tibia with trichobothrium in proximal part; cymbium oval, with trichobothrium on retrolateral side. Subtegulum (
<pageBreakToken id="71622978368C86C2E6170E8C64545179" pageId="4" pageNumber="35" start="start">St</pageBreakToken>
) (Figs 8-10) large (as long as tegulum in lateral view), cone-shaped with three threads (Ts); prolaterally with process (Sp) directed to notch of embolic base. Tegulum nearly oval with triangle extesion (Te) in terminal part (Figs 6, 8, 10). Embolus (Em) broad, longer than tegulum, heavily built in the base (Eb), terminal part lamellated with two processes: digitiform posterior process (Ep) and triangle shape terminal process (Ea), seminal duct (Sd) broad and heavily sclerotised in the base of embolus, and very fine in lamellar part of embolus.
</paragraph>
</subSubSection>
<subSubSection id="E1B4962719283D751D5E20DE28B78A48" pageId="4" pageNumber="35" type="comments">
<paragraph id="C8E6F0649DDEA140237ECF7839493A52" pageId="4" pageNumber="35">Comments.</paragraph>
<paragraph id="62D2A7A4D1B334E8E5C3CC2BDDA9C77B" pageId="4" pageNumber="35">
Thanks to the discovery of the male of
<taxonomicName id="4F6492C4C2054E416DE88BF9FB800E5C" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
(the easternmost species of the genus) it has became possible to compare it with the westernmost species,
<taxonomicName id="C0A44E3ADD0DB857075D501367DF51D2" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
. General appearance of the two species is rather similar (Figs 1-5). They differ in amount of camouflage cover which is almost absent in
<taxonomicName id="8477BF0EB55273AD82C59A20860B7724" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
,but
<taxonomicName id="3F1397B9CD13418E8CA188DD78C1DBC6" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
is covered heavily on dorsal and ventral sides. Both male and female of
<taxonomicName id="6B29FBC64EA7E7724C32DA0F6AEF51A1" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
have a distinct pattern on the carapace. Such a pattern is absent in the studied male of
<taxonomicName id="F4BE9F53D6CA577ABF938ECAB4E00EFB" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
(we have removed the camouflage cover); it is also absent in the conspecific female, judging from L.
<normalizedToken id="720871E907E1200478194BEE0211C3C4" originalValue="Kochs">Koch's</normalizedToken>
description. Male of
<taxonomicName id="83BFFF5DB5E72AFC8A016F0FE2D69EF0" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
has relatively longer and thinner legs (cf. Figs 1 and 4) and less spaced anterior lateral eyes (Figs 3 and 5). The posterior eye row is straight in
<taxonomicName id="CB438FEC4D4D1C262957E9B5E2251F79" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
(Fig. 1) and recurved in
<taxonomicName id="7285FD066D5D6BB3C7C40DB0D9B45A11" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
(Fig. 4). Male palps in two species are strikingly different.
<taxonomicName id="6C8EDB7ED9B15FB41EB099A672FA83E4" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
has a long filamentous spiraled embolus and conductor (Co) (Fig. 12). In
<taxonomicName id="D3966694582748711FD75EDB9871B29C" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
the embolus is flat and broad. Extension of tegulum in
<taxonomicName id="9D1B276CEFF5B8565995746FEE7252B8" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
is weakly sclerotized and has subparallel margins, while in
<taxonomicName id="BBA6A3533D9FDBFE4B6BF7F18B952C7E" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
it is
<normalizedToken id="B1B4C2CE78D33E7DAD727CC5E678C111" originalValue="triangleshaped">triangle-shaped</normalizedToken>
and strongly sclerotized.
</paragraph>
<paragraph id="83486DDF00EB1F4A1FB45847B574C481" pageId="4" pageNumber="35">
In the collection of the Zoological Museum, University of Turku we had the opportunity to examine males of two unidentified species of
<taxonomicName id="65CD3BE049C68C8C7F8E67E6A6F224B3" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="genus">Cryptothele</taxonomicName>
, one from Thailand (which seems new to science) and another from Indonesia (probably
<taxonomicName id="5B9BD422E91E92D175FCB364A4E65B18" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele sundaica" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="sundaica">Cryptothele sundaica</taxonomicName>
Thorell, 1890). Males of both species have conformation of the palp similar to that in
<taxonomicName id="8E4FFB070AAE202D9735D958E64C3AFF" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
(a thin and long embolus, a well developed conductor, etc.). This may indicate that
<taxonomicName id="1A0AB5F61CC03D2CE12CE09314D62DDB" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
,
<taxonomicName id="FA6A83996AE3964E0A4AC9A96B0520A7" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele sundaica" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="sundaica">Cryptothele sundaica</taxonomicName>
and the mentioned undescribed species most likely are not congeneric with
<taxonomicName id="784965F21093A0963359D8D2B4A4DAE6" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele verrucosa" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosa">Cryptothele verrucosa</taxonomicName>
and the genus could be split in the future into two separate genera.
</paragraph>
</subSubSection>
<subSubSection id="AD1CDB4728D7463331AF551BB6C64C2E" pageId="4" pageNumber="35" type="comparative material examined">
<paragraph id="F11422DA5FD2483AE679B9EDC42C9D50" pageId="4" pageNumber="35">Comparative material examined.</paragraph>
<paragraph id="48ED6A7D4B49C271584E97E98328120D" pageId="4" pageNumber="35">
<taxonomicName id="2874C1F82F1B7E3283EBA21629F00863" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="genus">Cryptothele</taxonomicName>
sp. (presumably new) 1♂ (ZMUT: AA 5.812), Thailand, Chanthaburi Pr., Kho Yai N.P., Wang Chum Pee, rain forest, 27.10-22.11.1976 (P.Lehtinen) covered with dirt, even cymbium, but RTA like in
<taxonomicName id="1EBEF13D12B201641923A462B663A26D" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele alluaudi" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Cryptothele alluaudi</taxonomicName>
, embolus and conductor long.
</paragraph>
<paragraph id="41118E7A4C5D867F9906607DE644E7D6" pageId="4" pageNumber="35">
<taxonomicName id="698BF0F6E7AD4AAF8A3F8E00F4EE6BE1" class="Arachnida" family="Zodariidae" genus="Cryptothele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptothele sundaica" order="Araneae" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="sundaica">Cryptothele sundaica</taxonomicName>
?: 2♂ 1♀ (ZMUT: AA 5.806), Indonesia, Sumatera Barat, Paykumbuh d., Lubu Bangku, low jungle, 7.12.1980 (P.T.Lehtinen).
</paragraph>
</subSubSection>
</treatment>
</document>