598 lines
69 KiB
XML
598 lines
69 KiB
XML
<document id="23FC9C54AD1EF88AEC4FF8ADF79B702B" ID-DOI="10.11646/zootaxa.4061.1.3" ID-GBIF-Dataset="91860246-ab68-4534-9afe-35b432f66fdc" ID-ISSN="1175-5326" ID-Zenodo-Dep="270357" ID-ZooBank="F92B9552-F0FD-436F-8531-8C692AE9411B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458391222201" checkinUser="plazi" docAuthor="Rivera-Correa, Mauricio, García-Burneo, Karla & Grant, Taran" docDate="2016" docId="A54A87BB8976FFA9FF28FCE78342FE57" docLanguage="en" docName="zootaxa.4061.1.3.pdf.imf" docOrigin="Zootaxa 4061 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hyloscirtus diabolus Rivera-Correa, García-Burneo & Grant, 2016, sp. nov." docType="treatment" docVersion="5" lastPageNumber="35" masterDocId="5973FFC38977FFAFFFBFFFD58610FFA2" masterDocTitle="A new red-eyed of stream treefrog of Hyloscirtus (Anura: Hylidae) from Peru, with comments on the taxonomy of the genus" masterLastPageNumber="41" masterPageNumber="29" pageNumber="30" updateTime="1698185462904" updateUser="plazi">
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<mods:titleInfo id="B027A41C4E7160C9CB80E160A6BA8FAF">
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<mods:title id="A09113461DA808CBB615502E19E51764">A new red-eyed of stream treefrog of Hyloscirtus (Anura: Hylidae) from Peru, with comments on the taxonomy of the genus</mods:title>
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<mods:name id="BBD1E405CCAB526F53FDF42340DE39BD" type="personal">
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<mods:namePart id="E4DADCB6E84C587A48AF1C3D66451694">Rivera-Correa, Mauricio</mods:namePart>
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<mods:namePart id="E5DD0852CD5748BA14EFA24FEC43E7CA">García-Burneo, Karla</mods:namePart>
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<mods:name id="3F384B981AD57760629FEF25E638A341" type="personal">
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<mods:roleTerm id="C68DE4783FDC78B47A3C191F97F8FAC4">Author</mods:roleTerm>
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<mods:namePart id="3D3F44CEF164CF911F13C2D1D786D890">Grant, Taran</mods:namePart>
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<mods:typeOfResource id="5D6BF9CEC563CAB274345CB996C9EF77">text</mods:typeOfResource>
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<mods:title id="8618FF8EF0CF7E928B3538E68AC37A3E">Zootaxa</mods:title>
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<mods:date id="94C7FCD7CA8CEAD8F22470E9BA3BA5E9">2016</mods:date>
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<mods:number id="5A89BB38E7DCB8C09A888817C08713FA">4061</mods:number>
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<mods:identifier id="83C8585F9420D6A7D2357DA8B8012060" type="DOI">10.11646/zootaxa.4061.1.3</mods:identifier>
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<mods:identifier id="4E89089EE9876431AB5A031CD4DF243F" type="GBIF-Dataset">91860246-ab68-4534-9afe-35b432f66fdc</mods:identifier>
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<treatment id="A54A87BB8976FFA9FF28FCE78342FE57" ID-DOI="http://doi.org/10.5281/zenodo.5664005" ID-GBIF-Taxon="127638489" ID-Zenodo-Dep="5664005" LSID="urn:lsid:plazi:treatment:A54A87BB8976FFA9FF28FCE78342FE57" httpUri="http://treatment.plazi.org/id/A54A87BB8976FFA9FF28FCE78342FE57" lastPageId="6" lastPageNumber="35" pageId="1" pageNumber="30">
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<subSubSection id="65F965268976FFAEFF28FCE78701FCCC" pageId="1" pageNumber="30" type="nomenclature">
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<paragraph id="2D5C36AD8976FFAEFF28FCE787E4FCEE" blockId="1.[151,500,818,878]" box="[151,500,818,844]" pageId="1" pageNumber="30">
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<heading id="761481C18976FFAEFF28FCE787E4FCEE" bold="true" box="[151,500,818,844]" fontSize="11" level="1" pageId="1" pageNumber="30" reason="1">
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<emphasis id="1F97EABF8976FFAEFF28FCE787E4FCEE" bold="true" box="[151,500,818,844]" pageId="1" pageNumber="30">
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<taxonomicName id="EAE34D2E8976FFAEFF28FCE7879FFCEE" ID-CoL="3NGPY" box="[151,399,818,844]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="diabolus" status="sp. nov.">
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<emphasis id="1F97EABF8976FFAEFF28FCE7879FFCEE" bold="true" box="[151,399,818,844]" italics="true" pageId="1" pageNumber="30">Hyloscirtus diabolus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="04A457C48976FFAEFE29FCE787E4FCEE" box="[406,500,818,844]" pageId="1" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2D5C36AD8976FFAEFF28FC808701FCCC" blockId="1.[151,500,818,878]" box="[151,273,853,878]" pageId="1" pageNumber="30">
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(
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<figureCitation id="B5D82A288976FFAEFF20FC808719FCCC" box="[159,265,853,878]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[151,250,1381,1403]" captionStartId-1="5.[151,250,1308,1330]" captionStartId-2="6.[151,250,1975,1997]" captionTargetBox-0="[262,1322,201,1346]" captionTargetBox-1="[155,1428,680,1276]" captionTargetBox-2="[221,1370,547,1936]" captionTargetId-0="figure@4.[254,1333,193,1359]" captionTargetId-1="figure@5.[150,1436,669,1286]" captionTargetId-2="figure@6.[208,1379,533,1953]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Hyloscirtus diabolus sp. nov. in life (CORBIDI 12885, holotype, adult male, 82.3 mm SVL). Photos: K. García- Burneo." captionText-1="FIGURE 2. Dorsal and ventral views (in preservative) of the holotype of Hyloscirtus diabolus sp. nov. (CORBIDI 12885; adult male, 82.3 mm SVL)." captionText-2="FIGURE 3. Hyloscirtus diabolus sp. nov. (CORBIDI 12885, holotype). (A) Head in lateral view (B) head in dorsal view; (C) left hand in ventral view; (D) left foot in ventral view. Scale bar = 10 mm." httpUri-0="https://zenodo.org/record/270358/files/figure.png" httpUri-1="https://zenodo.org/record/270359/files/figure.png" httpUri-2="https://zenodo.org/record/270360/files/figure.png" pageId="1" pageNumber="30">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="65F965268976FFAEFF28FC488735FBC8" pageId="1" pageNumber="30" type="materials_examined">
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<paragraph id="2D5C36AD8976FFAEFF28FC48847AFC5C" blockId="1.[151,1437,925,1964]" pageId="1" pageNumber="30">
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<emphasis id="1F97EABF8976FFAEFF28FC48871BFC14" bold="true" box="[151,267,925,950]" pageId="1" pageNumber="30">
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<typeStatus id="F258880F8976FFAEFF28FC488716FC14" box="[151,262,925,950]" pageId="1" pageNumber="30" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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CORBIDI 12885, adult male,
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<collectingCountry id="55F4763D8976FFAEFDDEFC4B8487FC17" box="[609,663,926,949]" name="Peru" pageId="1" pageNumber="30">Peru</collectingCountry>
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, Departamento de Amazonas, Provincia de Chachapoyas, Distrito La Jalca, km 10 from La Colpa–Rodríguez de Mendoza (-6.398944, -77.226278,
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<quantity id="EA1B9B488976FFAEFBF0FC1782BBFC7B" box="[1103,1195,962,986]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="30" unit="m" value="2300.0">2300 m</quantity>
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a.s.l.) Collected
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<date id="595D106D8976FFAEFA3FFC17875CFC5C" pageId="1" pageNumber="30" value="2012-11-14">14 November 2012</date>
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by Karla García-Burneo.
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</paragraph>
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<paragraph id="2D5C36AD8976FFAEFF78FBDC8735FBC8" blockId="1.[151,1437,925,1964]" pageId="1" pageNumber="30">
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<emphasis id="1F97EABF8976FFAEFF78FBDC872BFB80" bold="true" box="[199,315,1033,1058]" pageId="1" pageNumber="30">
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<typeStatus id="F258880F8976FFAEFF78FBDC8726FB80" box="[199,310,1033,1058]" pageId="1" pageNumber="30" type="paratype">Paratype</typeStatus>
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.
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</emphasis>
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MUSA 3864, adult male,
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<collectingCountry id="55F4763D8976FFAEFD3EFBDF84A7FB83" box="[641,695,1034,1057]" name="Peru" pageId="1" pageNumber="30">Peru</collectingCountry>
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, Departamento de Amazonas, Provincia de Pedro Ruiz Gallo, Comunidad Shipasbama (-5.845833, -78.071944,
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<quantity id="EA1B9B488976FFAEFD6FFBFB8538FBE7" box="[720,808,1070,1094]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="1" pageNumber="30" unit="m" value="2180.0">2180 m</quantity>
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a.s.l.). Collected on
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<date id="595D106D8976FFAEFBA5FBFB82E7FBE4" box="[1050,1271,1069,1094]" pageId="1" pageNumber="30" value="2011-09-18">18 September 2011</date>
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by Roy Santa Cruz Farfán.
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</paragraph>
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</subSubSection>
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<subSubSection id="65F965268976FFAEFF78FBA0829FF809" pageId="1" pageNumber="30" type="diagnosis">
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<paragraph id="2D5C36AD8976FFAEFF78FBA08256FACE" blockId="1.[151,1437,925,1964]" pageId="1" pageNumber="30">
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<emphasis id="1F97EABF8976FFAEFF78FBA08751FB2C" bold="true" box="[199,321,1141,1166]" pageId="1" pageNumber="30">Diagnosis.</emphasis>
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<taxonomicName id="EAE34D2E8976FFAEFEF6FBA08423FB2F" box="[329,563,1141,1165]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="diabolus">
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<emphasis id="1F97EABF8976FFAEFEF6FBA08423FB2F" box="[329,563,1141,1165]" italics="true" pageId="1" pageNumber="30">Hyloscirtus diabolus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="B5D82A288976FFAEFDFDFBA08493FB2C" box="[578,643,1141,1166]" captionStart="FIGURE 1" captionStartId="4.[151,250,1381,1403]" captionTargetBox="[262,1322,201,1346]" captionTargetId="figure@4.[254,1333,193,1359]" captionTargetPageId="4" captionText="FIGURE 1. Hyloscirtus diabolus sp. nov. in life (CORBIDI 12885, holotype, adult male, 82.3 mm SVL). Photos: K. García- Burneo." httpUri="https://zenodo.org/record/270358/files/figure.png" pageId="1" pageNumber="30">Fig. 1</figureCitation>
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) can be diagnosed by the following characters: large size (SVL
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<quantity id="EA1B9B488976FFAEFAE2FBA38713FB13" metricMagnitude="-2" metricUnit="m" metricValue="8.575" metricValueMax="8.92" metricValueMin="8.23" pageId="1" pageNumber="30" unit="mm" value="85.75" valueMax="89.2" valueMin="82.3">82.3– 89.2 mm</quantity>
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, n = 2); vertical keel in the snout and upper lip cleft in the region of the premaxilla; vomerine odontophores with a conical-shape and a gap medially, each process with two or three prominent teeth; large, thick, supracloacal flap and supratympanic fold present; finger webbing formula:
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<date id="595D106D8976FFAEFBB1FB308221FB5E" box="[1038,1073,1252,1276]" pageId="1" pageNumber="30">I 2</date>
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½ –2½
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<date id="595D106D8976FFAEFBCEFB30828DFB5E" box="[1137,1181,1252,1276]" pageId="1" pageNumber="30">II 1</date>
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2/3–2 2/3
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<date id="595D106D8976FFAEFB53FB308332FB5E" box="[1260,1314,1252,1276]" pageId="1" pageNumber="30">III 2</date>
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+–
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<date id="595D106D8976FFAEFA86FB31837AFB5E" box="[1337,1386,1252,1276]" pageId="1" pageNumber="30">2 IV</date>
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, toe webbing formula:
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<date id="595D106D8976FFAEFEDBFAD98791FA81" box="[356,385,1291,1315]" pageId="1" pageNumber="30">I 1</date>
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1/2 –2-
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<date id="595D106D8976FFAEFE7CFAD987FAFA81" box="[451,490,1291,1315]" pageId="1" pageNumber="30">II 1</date>
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-–2
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<date id="595D106D8976FFAEFDAEFAD98451FA81" box="[529,577,1291,1315]" pageId="1" pageNumber="30">III 1</date>
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1/2 –2 1/2
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<date id="595D106D8976FFAEFD31FAD984D0FA81" box="[654,704,1291,1315]" pageId="1" pageNumber="30">IV 2</date>
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1/2–
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<date id="595D106D8976FFAEFD5AFADE851EFA81" box="[741,782,1287,1315]" bridgedPair="-" pageId="1" pageNumber="30">1- V</date>
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; nuptial pad absent; hypertrophied forelimbs and enlarged, curved and protruding spine prepollex present; fleshy calcar tubercle present; dorsum, thighs, shanks, finger, toes and discs brown with abundant tiny yellow marks; iris dark red without reticulation.
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</paragraph>
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<paragraph id="2D5C36AD8976FFAEFF78FAA3829FF809" blockId="1.[151,1437,925,1964]" pageId="1" pageNumber="30">
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<emphasis id="1F97EABF8976FFAEFF78FAA38771FA2D" bold="true" box="[199,353,1398,1423]" pageId="1" pageNumber="30">Comparison.</emphasis>
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The only species of the
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<taxonomicName id="EAE34D2E8976FFAEFD3AFAAD852CFA2D" box="[645,828,1399,1423]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="larinopygion">
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<emphasis id="1F97EABF8976FFAEFD3AFAAD852CFA2D" box="[645,828,1399,1423]" italics="true" pageId="1" pageNumber="30">H. larinopygion</emphasis>
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</taxonomicName>
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species group with hypertrophied forelimbs in adult males and broad elliptical prepollex, modified as a projecting spine, are
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<taxonomicName id="EAE34D2E8976FFAEFC6FFA498364FA11" authority="Kizirian et al. 2003" authorityName="Kizirian et al." authorityYear="2003" box="[976,1396,1435,1460]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
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<emphasis id="1F97EABF8976FFAEFC6FFA498268FA11" box="[976,1144,1435,1459]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
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(Kizirian
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<emphasis id="1F97EABF8976FFAEFB51FA498338FA11" box="[1262,1320,1435,1459]" italics="true" pageId="1" pageNumber="30">et al.</emphasis>
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2003)
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</taxonomicName>
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,
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<taxonomicName id="EAE34D2E8976FFAEFA3EFA498416FA75" authority="Almendariz et al. 2014" authorityName="Almendariz et al." authorityYear="2014" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="condor">
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<emphasis id="1F97EABF8976FFAEFA3EFA4986F6FA75" italics="true" pageId="1" pageNumber="30">H. condor</emphasis>
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Almendáriz
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<emphasis id="1F97EABF8976FFAEFE3DFA1587D0FA75" box="[386,448,1471,1495]" italics="true" pageId="1" pageNumber="30">et al.</emphasis>
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2014
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</taxonomicName>
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, and
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<emphasis id="1F97EABF8976FFAEFDF4FA1584CAFA75" box="[587,730,1471,1495]" italics="true" pageId="1" pageNumber="30">
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<taxonomicName id="EAE34D2E8976FFAEFDF4FA1584C6FA75" box="[587,726,1471,1495]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="diabolus">H. diabolus</taxonomicName>
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.
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</emphasis>
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The remaining species of the group lack the hypertrophied forelimbs and their prepollex is trapezoidal (Kizirian
|
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<emphasis id="1F97EABF8976FFAEFCBBFA31852FFA59" box="[772,831,1507,1531]" italics="true" pageId="1" pageNumber="30">et al.</emphasis>
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2003; Rivera-Correa and Faivovich 2013;
|
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<tableCitation id="606103168976FFAEFA89FA36839BFA59" box="[1334,1419,1507,1532]" captionStart="TABLE 1" captionStartId="2.[119,201,519,540]" captionTargetBox="[119,1872,616,1607]" captionTargetPageId="2" captionText="TABLE 1. Cοmparisοn οf sοme diagnοstic characters in species οf the Hyloscirtus larinopygion grοup. Sοurces ∶ (1) Almendariz et al. 2014; (2) Ardila-Rοbayο et al. (1993); (3) Cοlοma et al. (2012); (4) Duellman (1973); (5) Duellman and Altig (1978); (6) Duellman and Berger (1982); (7) Duellman and Hillis (1990); (8) Duellman and Cοlοma (1993); (9) Kizirian et al. (2003); (10) Mueses-Cisnerοs and Anganοy-Criοllο (2008); (11) Mueses-Cisnerοs and Perdοmο-Castillο (2011); (12) Ruiz-Carranza and Lynch (1982); (*) this study." httpUri="http://table.plazi.org/id/799C66258975FFADFFC8FDD28140FDF0" pageId="1" pageNumber="30" tableUuid="799C66258975FFADFFC8FDD28140FDF0">Table 1</tableCitation>
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).
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<taxonomicName id="EAE34D2E8976FFAEFF28F9D2876FF9BD" box="[151,383,1543,1567]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="diabolus">
|
||
<emphasis id="1F97EABF8976FFAEFF28F9D2876FF9BD" box="[151,383,1543,1567]" italics="true" pageId="1" pageNumber="30">Hyloscirtus diabolus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="EAE34D2E8976FFAEFDAFF9DD8491F9BD" box="[528,641,1543,1567]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="condor">
|
||
<emphasis id="1F97EABF8976FFAEFDAFF9DD8491F9BD" box="[528,641,1543,1567]" italics="true" pageId="1" pageNumber="30">H. condor</emphasis>
|
||
</taxonomicName>
|
||
by having a strongly well developed, fleshy, heel tubercle (absent in
|
||
<taxonomicName id="EAE34D2E8976FFAEFA3EF9DD86F6F9E1" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="condor">
|
||
<emphasis id="1F97EABF8976FFAEFA3EF9DD86F6F9E1" italics="true" pageId="1" pageNumber="30">H. condor</emphasis>
|
||
</taxonomicName>
|
||
), dark red iris (iris golden with fine tan reticulations in
|
||
<taxonomicName id="EAE34D2E8976FFAEFCECF9F985D4F9E1" box="[851,964,1579,1603]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="condor">
|
||
<emphasis id="1F97EABF8976FFAEFCECF9F985D4F9E1" box="[851,964,1579,1603]" italics="true" pageId="1" pageNumber="30">H. condor</emphasis>
|
||
</taxonomicName>
|
||
), gular region and belly dark reddish gray with pink marks in preservation (belly olive without marks and chest is light gray in
|
||
<taxonomicName id="EAE34D2E8976FFAEFBFAF98582A6F9C5" box="[1093,1206,1615,1639]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="condor">
|
||
<emphasis id="1F97EABF8976FFAEFBFAF98582A6F9C5" box="[1093,1206,1615,1639]" italics="true" pageId="1" pageNumber="30">H. condor</emphasis>
|
||
</taxonomicName>
|
||
), vertical keel in the snout and upper lip cleft in the region of the premaxilla (absent in
|
||
<taxonomicName id="EAE34D2E8976FFAEFC1BF9A18208F929" box="[932,1048,1651,1675]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="condor">
|
||
<emphasis id="1F97EABF8976FFAEFC1BF9A18208F929" box="[932,1048,1651,1675]" italics="true" pageId="1" pageNumber="30">H. condor</emphasis>
|
||
</taxonomicName>
|
||
) and greater adult male SVL (
|
||
<taxonomicName id="EAE34D2E8976FFAEFA3EF9A186F6F90D" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="condor">
|
||
<emphasis id="1F97EABF8976FFAEFA3EF9A186F6F90D" italics="true" pageId="1" pageNumber="30">H. condor</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="EA1B9B488976FFAEFF49F943878CF90D" box="[246,412,1686,1711]" metricMagnitude="-2" metricUnit="m" metricValue="6.93" metricValueMax="7.38" metricValueMin="6.4799999999999995" pageId="1" pageNumber="30" unit="mm" value="69.3" valueMax="73.8" valueMin="64.8">64.8–73.8 mm</quantity>
|
||
, n =
|
||
<quantity id="EA1B9B488976FFAEFE69F94D8416F90D" box="[470,518,1686,1711]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="1" pageNumber="30" unit="in" value="7.0">7 in</quantity>
|
||
). The new species differs from
|
||
<taxonomicName id="EAE34D2E8976FFAEFCD3F94D8204F90D" box="[876,1044,1687,1711]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
|
||
<emphasis id="1F97EABF8976FFAEFCD3F94D8204F90D" box="[876,1044,1687,1711]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
|
||
</taxonomicName>
|
||
in dorsal and lateral color pattern: dorsum gray or brown, flanks gray brown, bearing pink-orange marks more conspicuous in axillary and inguinal regions, lateral and lower margin of cloaca, knee, elbow, and outer margin of forearm white in
|
||
<taxonomicName id="EAE34D2E8976FFAEFB01F9358374F955" box="[1214,1380,1759,1783]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
|
||
<emphasis id="1F97EABF8976FFAEFB01F9358374F955" box="[1214,1380,1759,1783]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
|
||
</taxonomicName>
|
||
; and ventral color pattern: gular region and belly dark reddish gray with pink marks in preservation to
|
||
<taxonomicName id="EAE34D2E8976FFAEFB6DF8D18344F8B9" box="[1234,1364,1795,1819]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="diabolus">
|
||
<emphasis id="1F97EABF8976FFAEFB6DF8D18344F8B9" box="[1234,1364,1795,1819]" italics="true" pageId="1" pageNumber="30">H. diabolus</emphasis>
|
||
</taxonomicName>
|
||
(belly gray-blue and chest is light gray in
|
||
<taxonomicName id="EAE34D2E8976FFAEFD9BF8FD84DBF89D" box="[548,715,1831,1855]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
|
||
<emphasis id="1F97EABF8976FFAEFD9BF8FD84DBF89D" box="[548,715,1831,1855]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
|
||
</taxonomicName>
|
||
), digital discs brown (digital discs white in
|
||
<taxonomicName id="EAE34D2E8976FFAEFB05F8FD8370F89D" box="[1210,1376,1831,1855]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
|
||
<emphasis id="1F97EABF8976FFAEFB05F8FD8370F89D" box="[1210,1376,1831,1855]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
|
||
</taxonomicName>
|
||
), iris dark red (iris yellow-gold with fine black reticulations in
|
||
<taxonomicName id="EAE34D2E8976FFAEFC80F89985FAF8C1" box="[831,1002,1867,1891]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
|
||
<emphasis id="1F97EABF8976FFAEFC80F89985FAF8C1" box="[831,1002,1867,1891]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
|
||
</taxonomicName>
|
||
), upper lip cleft in the region of the premaxilla (absent in
|
||
<taxonomicName id="EAE34D2E8976FFAEFE2FF8A58428F825" box="[400,568,1903,1927]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
|
||
<emphasis id="1F97EABF8976FFAEFE2FF8A58428F825" box="[400,568,1903,1927]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
|
||
</taxonomicName>
|
||
) and greater adult male SVL (
|
||
<taxonomicName id="EAE34D2E8976FFAEFC24F8A58253F825" box="[923,1091,1903,1927]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="tapichalaca">
|
||
<emphasis id="1F97EABF8976FFAEFC24F8A58253F825" box="[923,1091,1903,1927]" italics="true" pageId="1" pageNumber="30">H. tapichalaca</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="EA1B9B488976FFAEFBEBF8BB82EBF825" box="[1108,1275,1902,1927]" metricMagnitude="-2" metricUnit="m" metricValue="6.145" metricValueMax="6.38" metricValueMin="5.91" pageId="1" pageNumber="30" unit="mm" value="61.45" valueMax="63.8" valueMin="59.1">59.1–63.8 mm</quantity>
|
||
, n = 8). For a summary of others diagnostic characters of the
|
||
<taxonomicName id="EAE34D2E8976FFAEFD15F841854DF809" box="[682,861,1939,1963]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="larinopygion">
|
||
<emphasis id="1F97EABF8976FFAEFD15F841854DF809" box="[682,861,1939,1963]" italics="true" pageId="1" pageNumber="30">H. larinopygion</emphasis>
|
||
</taxonomicName>
|
||
species group, see
|
||
<tableCitation id="606103168976FFAEFB8AF8468299F809" box="[1077,1161,1939,1964]" captionStart="TABLE 1" captionStartId="2.[119,201,519,540]" captionTargetBox="[119,1872,616,1607]" captionTargetPageId="2" captionText="TABLE 1. Cοmparisοn οf sοme diagnοstic characters in species οf the Hyloscirtus larinopygion grοup. Sοurces ∶ (1) Almendariz et al. 2014; (2) Ardila-Rοbayο et al. (1993); (3) Cοlοma et al. (2012); (4) Duellman (1973); (5) Duellman and Altig (1978); (6) Duellman and Berger (1982); (7) Duellman and Hillis (1990); (8) Duellman and Cοlοma (1993); (9) Kizirian et al. (2003); (10) Mueses-Cisnerοs and Anganοy-Criοllο (2008); (11) Mueses-Cisnerοs and Perdοmο-Castillο (2011); (12) Ruiz-Carranza and Lynch (1982); (*) this study." httpUri="http://table.plazi.org/id/799C66258975FFADFFC8FDD28140FDF0" pageId="1" pageNumber="30" tableUuid="799C66258975FFADFFC8FDD28140FDF0">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="799C66258975FFADFFC8FDD28140FDF0" ID-Table-UUID="799C66258975FFADFFC8FDD28140FDF0" httpUri="http://table.plazi.org/id/799C66258975FFADFFC8FDD28140FDF0" pageId="2" pageNumber="31" targetBox="[119,1872,616,1607]" targetIsTable="true" targetPageId="2">
|
||
<paragraph id="2D5C36AD8975FFADFFC8FDD28140FDF0" blockId="2.[119,1872,518,594]" pageId="2" pageNumber="31">
|
||
<emphasis id="1F97EABF8975FFADFFC8FDD286F3FDBE" bold="true" box="[119,227,519,540]" pageId="2" pageNumber="31">TABLE 1.</emphasis>
|
||
Cοmparisοn οf sοme diagnοstic characters in species οf the
|
||
<taxonomicName id="EAE34D2E8975FFADFC86FDD3823DFDB9" box="[825,1069,518,539]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="larinopygion">
|
||
<emphasis id="1F97EABF8975FFADFC86FDD3823DFDB9" box="[825,1069,518,539]" italics="true" pageId="2" pageNumber="31">Hyloscirtus larinopygion</emphasis>
|
||
</taxonomicName>
|
||
grοup. Sοurces∶ (1) Almendariz
|
||
<emphasis id="1F97EABF8975FFADFACDFDD283B4FDB9" box="[1394,1444,518,540]" italics="true" pageId="2" pageNumber="31">et al.</emphasis>
|
||
2014; (2) Ardila-Rοbayο
|
||
<emphasis id="1F97EABF8975FFADF91AFDD280C8FDB9" box="[1701,1752,518,540]" italics="true" pageId="2" pageNumber="31">et al.</emphasis>
|
||
(1993); (3) Cοlοma
|
||
<emphasis id="1F97EABF8975FFADFF76FDF486EBFD94" box="[201,251,545,566]" italics="true" pageId="2" pageNumber="31">et al.</emphasis>
|
||
(2012); (4) Duellman (1973); (5) Duellman and Altig (1978); (6) Duellman and Berger (1982); (7) Duellman and Hillis (1990); (8) Duellman and Cοlοma (1993); (9) Kizirian et al. (2003); (10) Mueses-Cisnerοs and Anganοy-Criοllο (2008); (11) Mueses-Cisnerοs and Perdοmο-Castillο (2011); (12) Ruiz-Carranza and Lynch (1982); (*) this study.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D5C36AD8975FFADFFC8FDBD810CF9B0" pageId="2" pageNumber="31">
|
||
<table id="5FE3C40D89750050FFC8FDBD8140F9E5" box="[119,1872,616,1607]" gridcols="8" gridrows="6" pageId="2" pageNumber="31">
|
||
<tr id="93D334EF89750050FFC8FDBD8140FCAF" box="[119,1872,616,781]" gridrow="0" pageId="2" pageNumber="31">
|
||
<th id="D0025D9389750050FFC8FDBD8702FCAF" box="[119,274,616,781]" gridcol="0" gridrow="0" pageId="2" pageNumber="31">
|
||
Species
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FD7D86E2FD1E" box="[119,242,679,701]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="antioquia">
|
||
<emphasis id="1F97EABF8975FFADFFC8FD7D86E2FD1E" box="[119,242,679,701]" italics="true" pageId="2" pageNumber="31">H. antioquia</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="D0025D9389750050FE80FDBD847DFCAF" box="[319,621,616,781]" gridcol="1" gridrow="0" pageId="2" pageNumber="31">Dοrsal pattern Reddish brοwn; with οr withοut scattered οrange blοtches</th>
|
||
<th id="D0025D9389750050FD30FDBD850CFCAF" box="[655,796,616,781]" gridcol="2" gridrow="0" pageId="2" pageNumber="31">Fοrelimbs Nο hypertrοphied</th>
|
||
<th id="D0025D9389750050FCFEFDBD85ECFCAF" box="[833,1020,616,781]" gridcol="3" gridrow="0" pageId="2" pageNumber="31">Iris cοlοr Gray with burgundy reticulatiοns</th>
|
||
<th id="D0025D9389750050FBA5FDBD82D0FCAF" box="[1050,1216,616,781]" gridcol="4" gridrow="0" pageId="2" pageNumber="31">vοmerine teeth In cοntact οr nοt, 12‾ 20 teeth⁄each</th>
|
||
<th id="D0025D9389750050FB5FFDBD83D3FCAF" box="[1248,1475,616,781]" gridcol="5" gridrow="0" pageId="2" pageNumber="31">Nuptial pad Medial and distal cοmpοnent; dark brοwn epidermal prοϳectiοns</th>
|
||
<th id="D0025D9389750050FA5EFDBD8045FCAF" box="[1505,1621,616,781]" gridcol="6" gridrow="0" pageId="2" pageNumber="31">Prepοllex Τrapezοidal</th>
|
||
<th id="D0025D9389750050F9CEFDBD8140FCAF" box="[1649,1872,616,781]" gridcol="7" gridrow="0" pageId="2" pageNumber="31">Calcar Sοurce tubercle Present * (2)</th>
|
||
</tr>
|
||
<tr id="93D334EF89750050FFC8FCFB8140FC36" box="[119,1872,814,916]" gridrow="1" pageId="2" pageNumber="31">
|
||
<th id="D0025D9389750050FFC8FCFB8702FC36" box="[119,274,814,916]" gridcol="0" gridrow="1" pageId="2" pageNumber="31">
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FCFB86E1FCE1" box="[119,241,814,835]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="caucanus">
|
||
<emphasis id="1F97EABF8975FFADFFC8FCFB86E1FCE1" box="[119,241,814,835]" italics="true" pageId="2" pageNumber="31">H. caucanus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D0025D9389750050FE80FCFB847DFC36" box="[319,621,814,916]" gridcol="1" gridrow="1" pageId="2" pageNumber="31">Brοwn οr reddish brοwn with dark brοwn scattered blοtches; with οr withοut middοrsal stripe</td>
|
||
<td id="D0025D9389750050FD30FCFB850CFC36" box="[655,796,814,916]" gridcol="2" gridrow="1" pageId="2" pageNumber="31">Nο hypertrοphied</td>
|
||
<td id="D0025D9389750050FCFEFCFB85ECFC36" box="[833,1020,814,916]" gridcol="3" gridrow="1" pageId="2" pageNumber="31">Pale yellοw with brοwn reticulatiοns</td>
|
||
<td id="D0025D9389750050FBA5FCFB82D0FC36" box="[1050,1216,814,916]" gridcol="4" gridrow="1" pageId="2" pageNumber="31">In cοntact, 10‾14 teeth⁄each</td>
|
||
<td id="D0025D9389750050FB5FFCFB83D3FC36" box="[1248,1475,814,916]" gridcol="5" gridrow="1" pageId="2" pageNumber="31">Distal cοmpοnent οnly; creamy white</td>
|
||
<td id="D0025D9389750050FA5EFCFB8045FC36" box="[1505,1621,814,916]" gridcol="6" gridrow="1" pageId="2" pageNumber="31">Τrapezοidal</td>
|
||
<td id="D0025D9389750050F9CEFCFB8140FC36" box="[1649,1872,814,916]" gridcol="7" gridrow="1" pageId="2" pageNumber="31">Present οr * Absent (2)</td>
|
||
</tr>
|
||
<tr id="93D334EF89750050FFC8FC618140FB02" box="[119,1872,948,1184]" gridrow="2" pageId="2" pageNumber="31">
|
||
<th id="D0025D9389750050FFC8FC618702FB02" box="[119,274,948,1184]" gridcol="0" gridrow="2" pageId="2" pageNumber="31">
|
||
<emphasis id="1F97EABF8975FFADFFC8FC6186F9FB27" italics="true" pageId="2" pageNumber="31">
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FC6186CAFC6B" box="[119,218,948,969]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="condor">H. condor</taxonomicName>
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FBD086F0FBBB" box="[119,224,1028,1050]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="criptico">H. criptico</taxonomicName>
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FBA486F9FB27" box="[119,233,1136,1158]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="diabolus">H. diabolus</taxonomicName>
|
||
</emphasis>
|
||
</th>
|
||
<td id="D0025D9389750050FE80FC61847DFB02" box="[319,621,948,1184]" gridcol="1" gridrow="2" pageId="2" pageNumber="31">Light brοwn with abundant yellοw marks Grayish brοwn, densely stippled with minute cream flecks and bright οrange blοtches Brοwn with abundant yellοw marks</td>
|
||
<td id="D0025D9389750050FD30FC61850CFB02" box="[655,796,948,1184]" gridcol="2" gridrow="2" pageId="2" pageNumber="31">Hypertrοphied Nο hypertrοphied Hypertrοphied</td>
|
||
<td id="D0025D9389750050FCFEFC6185ECFB02" box="[833,1020,948,1184]" gridcol="3" gridrow="2" pageId="2" pageNumber="31">Gοlden with minute reticulatiοns Dark gray Dark red, withοut reticulatiοns</td>
|
||
<td id="D0025D9389750050FBA5FC6182D0FB02" box="[1050,1216,948,1184]" gridcol="4" gridrow="2" pageId="2" pageNumber="31">Nο cοntact, 3‾4 teeth⁄each In cοntact οr nοt, 8‾15 teeth⁄each Nο cοntact, 2‾3 teeth⁄each</td>
|
||
<td id="D0025D9389750050FB5FFC6183D3FB02" box="[1248,1475,948,1184]" gridcol="5" gridrow="2" pageId="2" pageNumber="31">Absent Absent Absent</td>
|
||
<td id="D0025D9389750050FA5EFC618045FB02" box="[1505,1621,948,1184]" gridcol="6" gridrow="2" pageId="2" pageNumber="31">Spine Τrapezοidal Spine</td>
|
||
<td id="D0025D9389750050F9CEFC618140FB02" box="[1649,1872,948,1184]" gridcol="7" gridrow="2" pageId="2" pageNumber="31">Absent (1) Present * (3) Present *</td>
|
||
</tr>
|
||
<tr id="93D334EF89750050FFC8FB158140FAD2" box="[119,1872,1216,1392]" gridrow="3" pageId="2" pageNumber="31">
|
||
<th id="D0025D9389750050FFC8FB158702FAD2" box="[119,274,1216,1392]" gridcol="0" gridrow="3" pageId="2" pageNumber="31">
|
||
<emphasis id="1F97EABF8975FFADFFC8FB1486C2FA98" italics="true" pageId="2" pageNumber="31">
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FB148702FB77" box="[119,274,1216,1238]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="larinopygion">H. larinopygion</taxonomicName>
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FAF086C2FA98" box="[119,210,1317,1338]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="lindae">H. lindae</taxonomicName>
|
||
</emphasis>
|
||
</th>
|
||
<td id="D0025D9389750050FE80FB15847DFAD2" box="[319,621,1216,1392]" gridcol="1" gridrow="3" pageId="2" pageNumber="31">Light brοwn tο dark brοwn; with οr withοut dark brοwn reticulatiοn Dark metallic brοwn</td>
|
||
<td id="D0025D9389750050FD30FB15850CFAD2" box="[655,796,1216,1392]" gridcol="2" gridrow="3" pageId="2" pageNumber="31">Nο hypertrοphied Nο hypertrοphied</td>
|
||
<td id="D0025D9389750050FCFEFB1585ECFAD2" box="[833,1020,1216,1392]" gridcol="3" gridrow="3" pageId="2" pageNumber="31">Gοlden οr silver with black reticulatiοns Dull bluish gray with minute black flecks</td>
|
||
<td id="D0025D9389750050FBA5FB1582D0FAD2" box="[1050,1216,1216,1392]" gridcol="4" gridrow="3" pageId="2" pageNumber="31">In cοntact οr nοt, 8‾15 teeth⁄each In cοntact, 11‾13 teeth⁄each</td>
|
||
<td id="D0025D9389750050FB5FFB1583D3FAD2" box="[1248,1475,1216,1392]" gridcol="5" gridrow="3" pageId="2" pageNumber="31">Medial and distal cοmpοnent; creamy white Distal cοmpοnent; creamy white</td>
|
||
<td id="D0025D9389750050FA5EFB158045FAD2" box="[1505,1621,1216,1392]" gridcol="6" gridrow="3" pageId="2" pageNumber="31">Τrapezοidal Τrapezοidal</td>
|
||
<td id="D0025D9389750050F9CEFB158140FAD2" box="[1649,1872,1216,1392]" gridcol="7" gridrow="3" pageId="2" pageNumber="31">Present οr * Absent (4) Present οr * Absent (3, 5)</td>
|
||
</tr>
|
||
<tr id="93D334EF89750050FFC8FA458140FA63" box="[119,1872,1424,1473]" gridrow="4" pageId="2" pageNumber="31">
|
||
<th id="D0025D9389750050FFC8FA458702FA63" box="[119,274,1424,1473]" gridcol="0" gridrow="4" pageId="2" pageNumber="31">
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FA4486C1FA07" box="[119,209,1424,1446]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="pacha">
|
||
<emphasis id="1F97EABF8975FFADFFC8FA4486C1FA07" box="[119,209,1424,1446]" italics="true" pageId="2" pageNumber="31">H. pacha</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D0025D9389750050FE80FA45847DFA63" box="[319,621,1424,1473]" gridcol="1" gridrow="4" pageId="2" pageNumber="31">Dark brοwn with metallic οrange flecks</td>
|
||
<td id="D0025D9389750050FD30FA45850CFA63" box="[655,796,1424,1473]" gridcol="2" gridrow="4" pageId="2" pageNumber="31">Nο hypertrοphied</td>
|
||
<td id="D0025D9389750050FCFEFA4585ECFA63" box="[833,1020,1424,1473]" gridcol="3" gridrow="4" pageId="2" pageNumber="31">Οlive brοwn</td>
|
||
<td id="D0025D9389750050FBA5FA4582D0FA63" box="[1050,1216,1424,1473]" gridcol="4" gridrow="4" pageId="2" pageNumber="31">In cοntact, 11‾15 teeth⁄each</td>
|
||
<td id="D0025D9389750050FB5FFA4583D3FA63" box="[1248,1475,1424,1473]" gridcol="5" gridrow="4" pageId="2" pageNumber="31">Absent</td>
|
||
<td id="D0025D9389750050FA5EFA458045FA63" box="[1505,1621,1424,1473]" gridcol="6" gridrow="4" pageId="2" pageNumber="31">Τrapezοidal</td>
|
||
<td id="D0025D9389750050F9CEFA458140FA63" box="[1649,1872,1424,1473]" gridcol="7" gridrow="4" pageId="2" pageNumber="31">Present * (3, 7)</td>
|
||
</tr>
|
||
<tr id="93D334EF89750050FFC8FA348140F9E5" box="[119,1872,1505,1607]" gridrow="5" pageId="2" pageNumber="31">
|
||
<th id="D0025D9389750050FFC8FA348702F9E5" box="[119,274,1505,1607]" gridcol="0" gridrow="5" pageId="2" pageNumber="31">
|
||
<taxonomicName id="EAE34D2E8975FFADFFC8FA34871AFA54" box="[119,266,1505,1526]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="31" phylum="Chordata" rank="species" species="pantostictus">
|
||
<emphasis id="1F97EABF8975FFADFFC8FA34871AFA54" box="[119,266,1505,1526]" italics="true" pageId="2" pageNumber="31">H. pantostictus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D0025D9389750050FE80FA34847DF9E5" box="[319,621,1505,1607]" gridcol="1" gridrow="5" pageId="2" pageNumber="31">Οlive-brοwn with small οrange spοts</td>
|
||
<td id="D0025D9389750050FD30FA34850CF9E5" box="[655,796,1505,1607]" gridcol="2" gridrow="5" pageId="2" pageNumber="31">Nο hypertrοphied</td>
|
||
<td id="D0025D9389750050FCFEFA3485ECF9E5" box="[833,1020,1505,1607]" gridcol="3" gridrow="5" pageId="2" pageNumber="31">Dark gray</td>
|
||
<td id="D0025D9389750050FBA5FA3482D0F9E5" box="[1050,1216,1505,1607]" gridcol="4" gridrow="5" pageId="2" pageNumber="31">In cοntact οr nοt, 10‾14 teeth⁄each</td>
|
||
<td id="D0025D9389750050FB5FFA3483D3F9E5" box="[1248,1475,1505,1607]" gridcol="5" gridrow="5" pageId="2" pageNumber="31">Medial and distal cοmpοnent, light brοwn epidermal prοϳectiοns</td>
|
||
<td id="D0025D9389750050FA5EFA348045F9E5" box="[1505,1621,1505,1607]" gridcol="6" gridrow="5" pageId="2" pageNumber="31">Τrapezοidal</td>
|
||
<td id="D0025D9389750050F9CEFA348140F9E5" box="[1649,1872,1505,1607]" gridcol="7" gridrow="5" pageId="2" pageNumber="31">Present οr * Absent (3, 5)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="65F965268975FFAAF9B4F9BD8355F82C" lastPageId="5" lastPageNumber="34" pageId="2" pageNumber="31" type="description">
|
||
<paragraph id="2D5C36AD8975FFABF9B4F9BD85ABF80F" blockId="2.[1547,1851,1640,1662]" lastBlockId="4.[151,1437,1472,2037]" lastPageId="4" lastPageNumber="33" pageId="2" pageNumber="31">
|
||
<emphasis id="1F97EABF8975FFADF9B4F9BD812BF9DC" box="[1547,1851,1640,1662]" italics="true" pageId="2" pageNumber="31">……continued on the next page</emphasis>
|
||
<emphasis id="1F97EABF8973FFABFF78FA1587F3FA7B" bold="true" box="[199,483,1472,1497]" pageId="4" pageNumber="33">
|
||
Description of
|
||
<typeStatus id="F258880F8973FFABFECAFA1587CEFA7B" box="[373,478,1472,1497]" pageId="4" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male,
|
||
<quantity id="EA1B9B488973FFABFDCBFA1484C8FA7A" box="[628,728,1473,1497]" metricMagnitude="-2" metricUnit="m" metricValue="8.23" pageId="4" pageNumber="33" unit="mm" value="82.3">82.3 mm</quantity>
|
||
SVL (
|
||
<figureCitation id="B5D82A288973FFABFC9EFA158576FA7B" box="[801,870,1472,1497]" captionStart="FIGURE 2" captionStartId="5.[151,250,1308,1330]" captionTargetBox="[155,1428,680,1276]" captionTargetId="figure@5.[150,1436,669,1286]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal and ventral views (in preservative) of the holotype of Hyloscirtus diabolus sp. nov. (CORBIDI 12885; adult male, 82.3 mm SVL)." httpUri="https://zenodo.org/record/270359/files/figure.png" pageId="4" pageNumber="33">Fig. 2</figureCitation>
|
||
). Body robust. Head 13% wider than long; head as wide as body; head width 35% of SVL; head length 27% of SVL. Snout rounded in dorsal view, truncate in profile (
|
||
<figureCitation id="B5D82A288973FFABFF20F9DD86F0F983" box="[159,224,1544,1569]" captionStart="FIGURE 3" captionStartId="6.[151,250,1975,1997]" captionTargetBox="[221,1370,547,1936]" captionTargetId="figure@6.[208,1379,533,1953]" captionTargetPageId="6" captionText="FIGURE 3. Hyloscirtus diabolus sp. nov. (CORBIDI 12885, holotype). (A) Head in lateral view (B) head in dorsal view; (C) left hand in ventral view; (D) left foot in ventral view. Scale bar = 10 mm." httpUri="https://zenodo.org/record/270360/files/figure.png" pageId="4" pageNumber="33">Fig. 3</figureCitation>
|
||
A–B); canthus rostralis rounded, distinct; loreal region weakly concave, nearly vertical; lips rounded, faintly flared; nostrils slightly protuberant, directed anterolaterally, located at level of anterior margin of lower jaw. Vertical keel on snout and upper lip cleft in region of premaxilla. Dorsal surface of internarial region concave. Interorbital distance slightly greater than upper eyelid width. Eye prominent, diameter greater than eye–nostril distance. Tympanum barely visible, round, positioned vertically such that is not visible from above; diameter 51% of eye diameter; tympanic annulus inconspicuous. Supratympanic fold prominent, extending from posterior to eye to posterior margin of arm insertion, slightly covering dorsal edge of tympanic membrane. Region between head and suprascapula depressed. Vomerine odontophores conical, separated by gap medially, positioned posterior to choanae; each process with 2–3 prominent teeth. Choanae small, ovoid, separated by a minimum distance greater than 3.5 times their maximum diameter. Tongue cordiform, barely free around lateral and posterior margin. Vocal slits longitudinal, originating on lateral to tongue and extending extending to the corner of the mouth. Vocal sac evident externally, large, single, median, subgular. Mental gland absent.
|
||
</paragraph>
|
||
<paragraph id="2D5C36AD8974FFACFFC2FD58810DFA65" pageId="3" pageNumber="32">
|
||
<table id="5FE3C40D89740050FFC2FD58810DF9BA" box="[125,1821,653,1560]" gridcols="7" gridrows="6" pageId="3" pageNumber="32">
|
||
<tr id="93D334EF89740050FFC2FD58810DFC93" box="[125,1821,653,817]" gridrow="0" pageId="3" pageNumber="32">
|
||
<th id="D0025D9389740050FFC2FD588471FC93" box="[125,609,653,817]" gridcol="0" gridrow="0" pageId="3" pageNumber="32">
|
||
<emphasis id="1F97EABF8974FFACFFC2FD5886CFFD00" bold="true" box="[125,223,653,674]" pageId="3" pageNumber="32">TABLE 1</emphasis>
|
||
. (Cοntinued) Species Dοrsal pattern
|
||
<taxonomicName id="EAE34D2E8974FFACFFC2FD338768FD59" authority="Black" authorityName="Black" box="[125,376,741,763]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="princecharlesi">
|
||
<emphasis id="1F97EABF8974FFACFFC2FD338733FD58" box="[125,291,741,763]" italics="true" pageId="3" pageNumber="32">H. princecharlesi</emphasis>
|
||
Black
|
||
</taxonomicName>
|
||
with many rοund-οval οrange blοtches
|
||
</th>
|
||
<th id="D0025D9389740050FD34FD588505FC93" box="[651,789,653,817]" gridcol="1" gridrow="0" pageId="3" pageNumber="32">Fοrelimbs Nο hypertrοphied</th>
|
||
<th id="D0025D9389740050FC85FD5885F5FC93" box="[826,997,653,817]" gridcol="2" gridrow="0" pageId="3" pageNumber="32">Iris cοlοr Dark gray</th>
|
||
<th id="D0025D9389740050FBAFFD5882A3FC93" box="[1040,1203,653,817]" gridcol="3" gridrow="0" pageId="3" pageNumber="32">vοmerine teeth In cοntact οr nοt, 11‾15 teeth⁄each</th>
|
||
<th id="D0025D9389740050FB6CFD58839FFC93" box="[1235,1423,653,817]" gridcol="4" gridrow="0" pageId="3" pageNumber="32">Nuptial pad Medial and distal cοmpοnent; creamy white</th>
|
||
<th id="D0025D9389740050FA70FD588051FC93" box="[1487,1601,653,817]" gridcol="5" gridrow="0" pageId="3" pageNumber="32">Prepοllex</th>
|
||
<th id="D0025D9389740050F9E2FD58810DFC93" box="[1629,1821,653,817]" gridcol="6" gridrow="0" pageId="3" pageNumber="32">Calcar Sοurce tubercle Absent * (3)</th>
|
||
</tr>
|
||
<tr id="93D334EF89740050FFC2FC84810DFC15" box="[125,1821,849,951]" gridrow="1" pageId="3" pageNumber="32">
|
||
<th id="D0025D9389740050FFC2FC848471FC15" box="[125,609,849,951]" gridcol="0" gridrow="1" pageId="3" pageNumber="32">
|
||
H.
|
||
<taxonomicName id="EAE34D2E8974FFACFF25FC848700FCC4" box="[154,272,849,870]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="psarolaimus">
|
||
<emphasis id="1F97EABF8974FFACFF25FC848700FCC4" box="[154,272,849,870]" italics="true" pageId="3" pageNumber="32">psarolaimus</emphasis>
|
||
</taxonomicName>
|
||
Grayish brοwn with dark brοwn and minute cream flecks; with οr withοut middοrsal stripe
|
||
</th>
|
||
<td id="D0025D9389740050FD34FC848505FC15" box="[651,789,849,951]" gridcol="1" gridrow="1" pageId="3" pageNumber="32">Nο hypertrοphied</td>
|
||
<td id="D0025D9389740050FC85FC8485F5FC15" box="[826,997,849,951]" gridcol="2" gridrow="1" pageId="3" pageNumber="32">Dull brοnze with black reticulatiοn</td>
|
||
<td id="D0025D9389740050FBAFFC8482A3FC15" box="[1040,1203,849,951]" gridcol="3" gridrow="1" pageId="3" pageNumber="32">In cοntact, 10‾18 teeth⁄each</td>
|
||
<td id="D0025D9389740050FB6CFC84839FFC15" box="[1235,1423,849,951]" gridcol="4" gridrow="1" pageId="3" pageNumber="32">Absent</td>
|
||
<td id="D0025D9389740050FA70FC848051FC15" box="[1487,1601,849,951]" gridcol="5" gridrow="1" pageId="3" pageNumber="32">Τrapezοidal</td>
|
||
<td id="D0025D9389740050F9E2FC84810DFC15" box="[1629,1821,849,951]" gridcol="6" gridrow="1" pageId="3" pageNumber="32">Present * (3, 7)</td>
|
||
</tr>
|
||
<tr id="93D334EF89740050FFC2FC0D810DFBFA" box="[125,1821,984,1112]" gridrow="2" pageId="3" pageNumber="32">
|
||
<th id="D0025D9389740050FFC2FC0D8471FBFA" box="[125,609,984,1112]" gridcol="0" gridrow="2" pageId="3" pageNumber="32">
|
||
<taxonomicName id="EAE34D2E8974FFACFFC2FC0D871CFBA5" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="ptychodactylus">
|
||
<emphasis id="1F97EABF8974FFACFFC2FC0D871CFBA5" italics="true" pageId="3" pageNumber="32">H. Οrange-tan tο reddish brοwn ptychodactylus</emphasis>
|
||
</taxonomicName>
|
||
with minute οrange-tan flecks enclοsed in black markings; with οr withοut middοrsal stripe
|
||
</th>
|
||
<td id="D0025D9389740050FD34FC0D8505FBFA" box="[651,789,984,1112]" gridcol="1" gridrow="2" pageId="3" pageNumber="32">Nο hypertrοphied</td>
|
||
<td id="D0025D9389740050FC85FC0D85F5FBFA" box="[826,997,984,1112]" gridcol="2" gridrow="2" pageId="3" pageNumber="32">Pale blue</td>
|
||
<td id="D0025D9389740050FBAFFC0D82A3FBFA" box="[1040,1203,984,1112]" gridcol="3" gridrow="2" pageId="3" pageNumber="32">In cοntact, 11‾15 teeth⁄each</td>
|
||
<td id="D0025D9389740050FB6CFC0D839FFBFA" box="[1235,1423,984,1112]" gridcol="4" gridrow="2" pageId="3" pageNumber="32">Absent</td>
|
||
<td id="D0025D9389740050FA70FC0D8051FBFA" box="[1487,1601,984,1112]" gridcol="5" gridrow="2" pageId="3" pageNumber="32">Τrapezοidal</td>
|
||
<td id="D0025D9389740050F9E2FC0D810DFBFA" box="[1629,1821,984,1112]" gridcol="6" gridrow="2" pageId="3" pageNumber="32">Present * (3, 7)</td>
|
||
</tr>
|
||
<tr id="93D334EF89740050FFC2FBAD810DFB7C" box="[125,1821,1144,1246]" gridrow="3" pageId="3" pageNumber="32">
|
||
<th id="D0025D9389740050FFC2FBAD8471FB7C" box="[125,609,1144,1246]" gridcol="0" gridrow="3" pageId="3" pageNumber="32">
|
||
<taxonomicName id="EAE34D2E8974FFACFFC2FBAC871CFB2F" box="[125,268,1144,1166]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="sarampiona">
|
||
<emphasis id="1F97EABF8974FFACFFC2FBAC871CFB2F" box="[125,268,1144,1166]" italics="true" pageId="3" pageNumber="32">H. sarampiona</emphasis>
|
||
</taxonomicName>
|
||
Pale οlive with οrange spοts
|
||
</th>
|
||
<td id="D0025D9389740050FD34FBAD8505FB7C" box="[651,789,1144,1246]" gridcol="1" gridrow="3" pageId="3" pageNumber="32">Nο hypertrοphied</td>
|
||
<td id="D0025D9389740050FC85FBAD85F5FB7C" box="[826,997,1144,1246]" gridcol="2" gridrow="3" pageId="3" pageNumber="32">Gοld with thin black reticulatiοn</td>
|
||
<td id="D0025D9389740050FBAFFBAD82A3FB7C" box="[1040,1203,1144,1246]" gridcol="3" gridrow="3" pageId="3" pageNumber="32">In cοntact οr nοt, 8‾16 teeth⁄each</td>
|
||
<td id="D0025D9389740050FB6CFBAD839FFB7C" box="[1235,1423,1144,1246]" gridcol="4" gridrow="3" pageId="3" pageNumber="32">Medial and distal cοmpοnent; dark brοwn epidermal prοϳectiοns</td>
|
||
<td id="D0025D9389740050FA70FBAD8051FB7C" box="[1487,1601,1144,1246]" gridcol="5" gridrow="3" pageId="3" pageNumber="32">Τrapezοidal</td>
|
||
<td id="D0025D9389740050F9E2FBAD810DFB7C" box="[1629,1821,1144,1246]" gridcol="6" gridrow="3" pageId="3" pageNumber="32">Present οr * Absent (12)</td>
|
||
</tr>
|
||
<tr id="93D334EF89740050FFC2FB2A810DFAD5" box="[125,1821,1279,1399]" gridrow="4" pageId="3" pageNumber="32">
|
||
<th id="D0025D9389740050FFC2FB2A8471FAD5" box="[125,609,1279,1399]" gridcol="0" gridrow="4" pageId="3" pageNumber="32">
|
||
<emphasis id="1F97EABF8974FFACFFC2FB2A871BFAF9" italics="true" pageId="3" pageNumber="32">
|
||
<taxonomicName id="EAE34D2E8974FFACFFC2FB2A8705FAB6" box="[125,277,1279,1300]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="staufferorum">H. staufferorum</taxonomicName>
|
||
Unifοrm dark brοwn
|
||
<taxonomicName id="EAE34D2E8974FFACFFC2FA92871BFAF9" box="[125,267,1350,1372]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="tapichalaca">H. tapichalaca</taxonomicName>
|
||
</emphasis>
|
||
Gray οr dark brοwn
|
||
</th>
|
||
<td id="D0025D9389740050FD34FB2A8505FAD5" box="[651,789,1279,1399]" gridcol="1" gridrow="4" pageId="3" pageNumber="32">Nο hypertrοphied Hypertrοphied</td>
|
||
<td id="D0025D9389740050FC85FB2A85F5FAD5" box="[826,997,1279,1399]" gridcol="2" gridrow="4" pageId="3" pageNumber="32">Metallic brοwn Υellοw-gοld with black reticulatiοn</td>
|
||
<td id="D0025D9389740050FBAFFB2A82A3FAD5" box="[1040,1203,1279,1399]" gridcol="3" gridrow="4" pageId="3" pageNumber="32">In cοntact, 16‾25 teeth⁄each Nο cοntact, 4‾6 teeth⁄each</td>
|
||
<td id="D0025D9389740050FB6CFB2A839FFAD5" box="[1235,1423,1279,1399]" gridcol="4" gridrow="4" pageId="3" pageNumber="32">Creamy white Absent</td>
|
||
<td id="D0025D9389740050FA70FB2A8051FAD5" box="[1487,1601,1279,1399]" gridcol="5" gridrow="4" pageId="3" pageNumber="32">Spine</td>
|
||
<td id="D0025D9389740050F9E2FB2A810DFAD5" box="[1629,1821,1279,1399]" gridcol="6" gridrow="4" pageId="3" pageNumber="32">Present * (3, 8) Present * (9)</td>
|
||
</tr>
|
||
<tr id="93D334EF89740050FFC2FA42810DF9BA" box="[125,1821,1431,1560]" gridrow="5" pageId="3" pageNumber="32">
|
||
<th id="D0025D9389740050FFC2FA428471F9BA" box="[125,609,1431,1560]" gridcol="0" gridrow="5" pageId="3" pageNumber="32">
|
||
<taxonomicName id="EAE34D2E8974FFACFFC2FA4D86F4FA0E" box="[125,228,1431,1453]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="32" phylum="Chordata" rank="species" species="tigrinus">
|
||
<emphasis id="1F97EABF8974FFACFFC2FA4D86F4FA0E" box="[125,228,1431,1453]" italics="true" pageId="3" pageNumber="32">H. tigrinus</emphasis>
|
||
</taxonomicName>
|
||
Υellοw-green οr yellοwish brοwn with transverse black stripes, sοmetimes reticulated; with οr withοut middοrsal stripe
|
||
</th>
|
||
<td id="D0025D9389740050FD34FA428505F9BA" box="[651,789,1431,1560]" gridcol="1" gridrow="5" pageId="3" pageNumber="32">Nο hypertrοphied</td>
|
||
<td id="D0025D9389740050FC85FA4285F5F9BA" box="[826,997,1431,1560]" gridcol="2" gridrow="5" pageId="3" pageNumber="32">Light gray οr yellοw with black reticulatiοn</td>
|
||
<td id="D0025D9389740050FBAFFA4282A3F9BA" box="[1040,1203,1431,1560]" gridcol="3" gridrow="5" pageId="3" pageNumber="32">In cοntact, 10‾15 teeth⁄each</td>
|
||
<td id="D0025D9389740050FB6CFA42839FF9BA" box="[1235,1423,1431,1560]" gridcol="4" gridrow="5" pageId="3" pageNumber="32">Absent</td>
|
||
<td id="D0025D9389740050FA70FA428051F9BA" box="[1487,1601,1431,1560]" gridcol="5" gridrow="5" pageId="3" pageNumber="32">Τrapezοidal</td>
|
||
<td id="D0025D9389740050F9E2FA42810DF9BA" box="[1629,1821,1431,1560]" gridcol="6" gridrow="5" pageId="3" pageNumber="32">Present * (10, 11)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="799C66258973FFABFF28FAB086F9FA3B" httpUri="https://zenodo.org/record/270358/files/figure.png" pageId="4" pageNumber="33" targetBox="[262,1322,201,1346]" targetPageId="4">
|
||
<paragraph id="2D5C36AD8973FFABFF28FAB086F9FA3B" blockId="4.[151,1436,1380,1433]" pageId="4" pageNumber="33">
|
||
<emphasis id="1F97EABF8973FFABFF28FAB0871DFAD8" bold="true" box="[151,269,1381,1403]" pageId="4" pageNumber="33">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="EAE34D2E8973FFABFEA4FAB187FEFAD8" box="[283,494,1380,1402]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="33" phylum="Chordata" rank="species" species="diabolus" status="sp. nov.">
|
||
<emphasis id="1F97EABF8973FFABFEA4FAB187FEFAD8" box="[283,494,1380,1402]" italics="true" pageId="4" pageNumber="33">Hyloscirtus diabolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="04A457C48973FFABFE4AFAB08456FAD8" box="[501,582,1381,1402]" pageId="4" pageNumber="33" rank="species">
|
||
<emphasis id="1F97EABF8973FFABFE4AFAB08456FAD8" bold="true" box="[501,582,1381,1402]" pageId="4" pageNumber="33">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in life (CORBIDI 12885, holotype, adult male, 82.3 mm SVL). Photos: K. García- Burneo.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D5C36AD8973FFAAFF78F86C87A3FDCB" blockId="4.[151,1437,1472,2037]" lastBlockId="5.[151,1437,151,617]" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
|
||
Upper arms and forearms hypertrophied; axillary membrane absent; ulnar fold and tubercles absent. Fingers short, thick, bearing large, ovoid discs, circumferential groove of each disc clearly defined; width of disc on Finger
|
||
<date id="595D106D8972FFAAFF28FF4E86C7FF11" box="[151,215,154,179]" pageId="5" pageNumber="34" value="1946-03">III 46</date>
|
||
% greater than tympanum diameter. Relative lengths of fingers I <II <IV <III.; webbing formula:
|
||
<date id="595D106D8972FFAAFA91FF4E836AFF11" box="[1326,1402,155,179]" pageId="5" pageNumber="34">I – II 1</date>
|
||
2/3– 2 2/3
|
||
<date id="595D106D8972FFAAFF7FFF1786E1FF78" box="[192,241,194,218]" pageId="5" pageNumber="34">III 2</date>
|
||
+–
|
||
<date id="595D106D8972FFAAFEB7FF178728FF7B" box="[264,312,194,218]" pageId="5" pageNumber="34">2 IV</date>
|
||
(
|
||
<figureCitation id="B5D82A288972FFAAFEF8FF14879AFF78" box="[327,394,193,218]" captionStart="FIGURE 3" captionStartId="6.[151,250,1975,1997]" captionTargetBox="[221,1370,547,1936]" captionTargetId="figure@6.[208,1379,533,1953]" captionTargetPageId="6" captionText="FIGURE 3. Hyloscirtus diabolus sp. nov. (CORBIDI 12885, holotype). (A) Head in lateral view (B) head in dorsal view; (C) left hand in ventral view; (D) left foot in ventral view. Scale bar = 10 mm." httpUri="https://zenodo.org/record/270360/files/figure.png" pageId="5" pageNumber="34">Fig. 3</figureCitation>
|
||
C). Distal and proximal subarticular tubercles large, single, rounded, conical; distal tubercles on Finger IV globular. Palmar surfaces with deep folds. Large supernumerary tubercles, high and round, distributed over base of fingers. Outer metacarpal tubercle diffuse. Inner metacarpal tubercle large, its shape contouring underlying distal prepollex that is modified into an enlarged, protruding spine. Nuptial pad absent.
|
||
<collectingCountry id="55F4763D8972FFAAFAA4FEF88343FEE4" box="[1307,1363,301,326]" name="India" pageId="5" pageNumber="34">Hind</collectingCountry>
|
||
limbs robust; tibia length 50% of SVL; foot length 47% of SVL. Calcar tubercle large, fleshy; tarsal fold or tubercles absent; metatarsal fold thick. Inner metatarsal tubercle large, ovoid; outer metatarsal tubercle small, round. Toes short, lateral fringes absent, discs smaller than those of fingers, slightly wider than digit. Relative length of toes: I <II <V <III <IV; toe webbing formula:
|
||
<date id="595D106D8972FFAAFDD9FE148495FE7B" box="[614,645,449,473]" pageId="5" pageNumber="34">I 1</date>
|
||
1/2 –2-
|
||
<date id="595D106D8972FFAAFD74FE1484E5FE7B" box="[715,757,449,473]" pageId="5" pageNumber="34">II 1</date>
|
||
-–2
|
||
<date id="595D106D8972FFAAFCA1FE148540FE7B" box="[798,848,449,473]" pageId="5" pageNumber="34">III 1</date>
|
||
1/2 –2 1/2
|
||
<date id="595D106D8972FFAAFC1EFE1485C5FE7B" box="[929,981,449,473]" pageId="5" pageNumber="34">IV 2</date>
|
||
1/2–
|
||
<date id="595D106D8972FFAAFC44FE148233FE7A" box="[1019,1059,445,473]" bridgedPair="-" pageId="5" pageNumber="34">1- V</date>
|
||
(
|
||
<figureCitation id="B5D82A288972FFAAFB8BFE158268FE7B" box="[1076,1144,448,473]" captionStart="FIGURE 3" captionStartId="6.[151,250,1975,1997]" captionTargetBox="[221,1370,547,1936]" captionTargetId="figure@6.[208,1379,533,1953]" captionTargetPageId="6" captionText="FIGURE 3. Hyloscirtus diabolus sp. nov. (CORBIDI 12885, holotype). (A) Head in lateral view (B) head in dorsal view; (C) left hand in ventral view; (D) left foot in ventral view. Scale bar = 10 mm." httpUri="https://zenodo.org/record/270360/files/figure.png" pageId="5" pageNumber="34">Fig. 3</figureCitation>
|
||
D). Subarticular tubercles large, round, conical; single row of smaller supernumerary tubercles along axis of each toe. Cloacal opening directed posteriorly at upper level of thighs; supracloacal flap large, thick; margins of vent with numerous small folds; cloacal sheath short. Dorsal skin, gular region, pectoral region and flanks smooth; belly and proximal ventral region of thighs granular.
|
||
</paragraph>
|
||
<caption id="799C66258972FFAAFF28FAC987BDFAF2" httpUri="https://zenodo.org/record/270359/files/figure.png" pageId="5" pageNumber="34" targetBox="[155,1428,680,1276]" targetPageId="5">
|
||
<paragraph id="2D5C36AD8972FFAAFF28FAC987BDFAF2" blockId="5.[151,1436,1308,1360]" pageId="5" pageNumber="34">
|
||
<emphasis id="1F97EABF8972FFAAFF28FAC9871FFA93" bold="true" box="[151,271,1308,1330]" pageId="5" pageNumber="34">FIGURE 2</emphasis>
|
||
. Dorsal and ventral views (in preservative) of the holotype of
|
||
<taxonomicName id="EAE34D2E8972FFAAFC1DFAC98267FA90" box="[930,1143,1308,1330]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="34" phylum="Chordata" rank="species" species="diabolus" status="sp. nov.">
|
||
<emphasis id="1F97EABF8972FFAAFC1DFAC98267FA90" box="[930,1143,1308,1330]" italics="true" pageId="5" pageNumber="34">Hyloscirtus diabolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="04A457C48972FFAAFB3FFAC982C2FA93" box="[1152,1234,1308,1329]" pageId="5" pageNumber="34" rank="species">
|
||
<emphasis id="1F97EABF8972FFAAFB3FFAC982C2FA93" bold="true" box="[1152,1234,1308,1329]" pageId="5" pageNumber="34">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(CORBIDI 12885; adult male, 82.3 mm SVL).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D5C36AD8972FFAAFF78FAA2832CFA5E" blockId="5.[151,1437,1399,2006]" pageId="5" pageNumber="34">
|
||
<emphasis id="1F97EABF8972FFAAFF78FAA287F7FA32" bold="true" box="[199,487,1399,1424]" pageId="5" pageNumber="34">
|
||
Color of
|
||
<typeStatus id="F258880F8972FFAAFE8FFAA28786FA32" box="[304,406,1399,1424]" pageId="5" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
Dorsum, flanks, anterior and posterior surfaces of thighs and shanks, calcar tubercle, dorsal surfaces of fingers and toes brown with an abundant tiny yellow marks; discs dark bluish gray ventrally. Gular region and ventral surfaces dirty gray with an abundant small yellow flecks and spots; ventral surfaces of hands, feet, and webbing dirty gray. Pericloacal region light gray. Iris solid dark red, without reticulations.
|
||
</paragraph>
|
||
<paragraph id="2D5C36AD8972FFAAFF78F9D28749F92E" blockId="5.[151,1437,1399,2006]" pageId="5" pageNumber="34">
|
||
<emphasis id="1F97EABF8972FFAAFF78F9D28478F982" bold="true" box="[199,616,1543,1568]" pageId="5" pageNumber="34">
|
||
Color of
|
||
<typeStatus id="F258880F8972FFAAFE86F9D2878FF982" box="[313,415,1543,1568]" pageId="5" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
Dorsum, flanks, anterior and posterior surfaces of thighs and shanks, dorsal surfaces of fingers and toes dark gray with an abundant light gray marks. Gular region and belly, ventral surfaces of thighs and shanks dark reddish gray with pink flecks and spots; hands, feet, webbing, and pericloacal region dirty gray.
|
||
</paragraph>
|
||
<paragraph id="2D5C36AD8972FFAAFF78F942839FF976" blockId="5.[151,1437,1399,2006]" pageId="5" pageNumber="34">
|
||
<emphasis id="1F97EABF8972FFAAFF78F9428467F912" bold="true" box="[199,631,1687,1712]" pageId="5" pageNumber="34">
|
||
Measurements of
|
||
<typeStatus id="F258880F8972FFAAFE20F9428415F912" box="[415,517,1687,1712]" pageId="5" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
(in mm).
|
||
</emphasis>
|
||
SVL 82.3; HL 22.4; HW 28.5; ED 9.0;
|
||
<collectingCountry id="55F4763D8972FFAAFBF3F94D8296F90D" box="[1100,1158,1688,1711]" name="India" pageId="5" pageNumber="34">END</collectingCountry>
|
||
5.7; NSD 3.7;
|
||
<collectingCountry id="55F4763D8972FFAAFA88F94D8379F90D" box="[1335,1385,1688,1711]" name="India" pageId="5" pageNumber="34">IND</collectingCountry>
|
||
5.8; AMD 12.1; TD 2.7; FAL 14.3; FAB 12.0; HAL 29.7; THL 39.8; TL 41.0; TAL 22.0; FL 38.7; TFD 5.0; FTD 3.8.
|
||
</paragraph>
|
||
<paragraph id="2D5C36AD8972FFAAFF78F90A8355F82C" blockId="5.[151,1437,1399,2006]" pageId="5" pageNumber="34">
|
||
<emphasis id="1F97EABF8972FFAAFF78F90A872BF95A" bold="true" box="[199,315,1759,1784]" pageId="5" pageNumber="34">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="F258880F8972FFAAFEC1F93587CFF95A" box="[382,479,1760,1784]" pageId="5" pageNumber="34" type="paratype">paratype</typeStatus>
|
||
MUSA 3864 (SVL =
|
||
<quantity id="EA1B9B488972FFAAFD55F9358543F95A" box="[746,851,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="8.92" pageId="5" pageNumber="34" unit="mm" value="89.2">89.2 mm</quantity>
|
||
) is remarkably larger than the
|
||
<typeStatus id="F258880F8972FFAAFB02F90A8332F95A" box="[1213,1314,1759,1784]" pageId="5" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
, possesses fewer yellow flecks and spots on the dorsum but more conspicuous on the loreal region, digits, flanks, thighs (
|
||
<figureCitation id="B5D82A288972FFAAFAD0F8D686B7F8E1" captionStart="FIGURE 4" captionStartId="7.[151,250,1077,1099]" captionTargetBox="[176,1405,193,1051]" captionTargetId="figure@7.[176,1406,193,1052]" captionTargetPageId="7" captionText="FIGURE 4. Hyloscirtus diabolus sp. nov. in life (MUSA 3864, paratype, adult male, 89.2 mm SVL). Note the dark marks (scars) in the anterior region of dorsum, which are consistent with the occurrence of male combat (see text). Photo: R. Santa Cruz." httpUri="https://zenodo.org/record/270361/files/figure.png" pageId="5" pageNumber="34">Fig. 4</figureCitation>
|
||
), and belly in life. Foot webbing is more extensive:
|
||
<date id="595D106D8972FFAAFD4CF8FE853DF8E1" box="[755,813,1835,1859]" pageId="5" pageNumber="34">I 2–2</date>
|
||
2/3
|
||
<date id="595D106D8972FFAAFCF7F8FE8560F8E1" box="[840,880,1835,1859]" pageId="5" pageNumber="34">II 1</date>
|
||
2/3–2 2/
|
||
<date id="595D106D8972FFAAFC13F8F285C6F8E0" box="[940,982,1831,1858]" pageId="5" pageNumber="34">3 III</date>
|
||
2–
|
||
<date id="595D106D8972FFAAFC45F8FE8239F8E0" box="[1018,1065,1835,1859]" pageId="5" pageNumber="34">2 IV</date>
|
||
and
|
||
<date id="595D106D8972FFAAFBDFF8FE826FF8E1" box="[1120,1151,1835,1859]" pageId="5" pageNumber="34">I 1</date>
|
||
- –1 1/
|
||
<date id="595D106D8972FFAAFB0DF8F282C5F8E0" box="[1202,1237,1831,1858]" pageId="5" pageNumber="34">2 II</date>
|
||
1–2
|
||
<date id="595D106D8972FFAAFAB2F8FE832EF8E1" box="[1293,1342,1835,1859]" pageId="5" pageNumber="34">III 1</date>
|
||
1/3 –2-
|
||
<date id="595D106D8972FFAAFAC1F8FE86B5F8C8" pageId="5" pageNumber="34">IV 2</date>
|
||
-–
|
||
<date id="595D106D8972FFAAFF03F88786F8F8CB" box="[188,232,1870,1898]" bridgedPair="-" pageId="5" pageNumber="34">1- V.</date>
|
||
Other measurements of the
|
||
<typeStatus id="F258880F8972FFAAFD97F8878498F8C8" box="[552,648,1874,1898]" pageId="5" pageNumber="34" type="paratype">paratype</typeStatus>
|
||
are: HL 23.9; HW 28.7; ED 8.8;
|
||
<collectingCountry id="55F4763D8972FFAAFBB7F8878252F8CB" box="[1032,1090,1874,1897]" name="India" pageId="5" pageNumber="34">END</collectingCountry>
|
||
6.0; NSD 3.9;
|
||
<collectingCountry id="55F4763D8972FFAAFB51F8878330F8CB" box="[1262,1312,1874,1897]" name="India" pageId="5" pageNumber="34">IND</collectingCountry>
|
||
5.8; AMD 12.8; TD 2.7; FAL 14.3; FAB 11.8; HAL 30.5; THL 40.7; TL 44.0; TAL 25.6; FL 42.4; TFD 5.9; FTD 4.8.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65F965268972FFA9FF78F84C8316FEC6" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34" type="distribution">
|
||
<paragraph id="2D5C36AD8972FFA9FF78F84C8316FEC6" blockId="5.[151,1437,1399,2006]" lastBlockId="6.[151,1436,151,501]" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
|
||
<emphasis id="1F97EABF8972FFAAFF78F84C8473F810" bold="true" box="[199,611,1945,1970]" pageId="5" pageNumber="34">Distribution and natural history</emphasis>
|
||
.
|
||
<taxonomicName id="EAE34D2E8972FFAAFDC5F84C857CF813" box="[634,876,1945,1969]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="34" phylum="Chordata" rank="species" species="diabolus">
|
||
<emphasis id="1F97EABF8972FFAAFDC5F84C857CF813" box="[634,876,1945,1969]" italics="true" pageId="5" pageNumber="34">Hyloscirtus diabolus</emphasis>
|
||
</taxonomicName>
|
||
is currently known only from two localities approximately
|
||
<quantity id="EA1B9B488972FFAAFEFEF86B8788F874" box="[321,408,1981,2006]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="34" unit="km" value="100.0">100 km</quantity>
|
||
straight-line distance apart in the highlands on the eastern flank of the Cordillera Central, northern Peruvian Andes (
|
||
<figureCitation id="B5D82A288971FFA9FE7BFF42841AFF12" box="[452,522,151,176]" captionStart="FIGURE 5" captionStartId="7.[151,250,1962,1984]" captionTargetBox="[159,1430,1216,1934]" captionTargetId="figure@7.[151,1435,1210,1941]" captionTargetPageId="7" captionText="FIGURE 5. Geographic distribution of Hyloscirtus diabolus sp. nov. (holotype locality, star; paratype locality, dot), H. condor (triangle), and H. tapichalaca (square)." httpUri="https://zenodo.org/record/270362/files/figure.png" pageId="6" pageNumber="35">Fig. 5</figureCitation>
|
||
). The two known individuals were found in dense cloud forest adjacent to very rocky streams with strong flow and 30–35º slope. The specimens were collected while calling at night, perched on tree branches ca.
|
||
<quantity id="EA1B9B488971FFA9FEDEFF3587A1FF55" box="[353,433,224,248]" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="35" unit="m" value="1.8">1.80 m</quantity>
|
||
above the ground and ca.
|
||
<quantity id="EA1B9B488971FFA9FD58FF358504FF55" box="[743,788,224,248]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="35" unit="m" value="2.0">2 m</quantity>
|
||
from the edge of the stream. When handled, they clung tightly to the substrate, acquired a position with the head directed toward the belly and limbs pressed to the abdomen, and secreted a white, viscous, sticky, liquid over the entire body. Advertisement calls, reproductive biology, and tadpoles are unknown. No other anuran species was found syntopically with
|
||
<taxonomicName id="EAE34D2E8971FFA9FB3BFE988316FEC6" box="[1156,1286,332,356]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="diabolus">
|
||
<emphasis id="1F97EABF8971FFA9FB3BFE988316FEC6" box="[1156,1286,332,356]" italics="true" pageId="6" pageNumber="35">H. diabolus</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65F965268971FFA9FF6EFEBA8342FE57" pageId="6" pageNumber="35" type="etymology">
|
||
<paragraph id="2D5C36AD8971FFA9FF6EFEBA8342FE57" blockId="6.[151,1436,151,501]" pageId="6" pageNumber="35">
|
||
<emphasis id="1F97EABF8971FFA9FF6EFEBA8747FE2A" bold="true" box="[209,343,367,392]" pageId="6" pageNumber="35">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="EAE34D2E8971FFA9FDF2FEA584BDFE2A" box="[589,685,368,392]" class="Amphibia" family="Hylidae" genus="Hyloscirtus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="diabolus">
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<emphasis id="1F97EABF8971FFA9FDF2FEA584BDFE2A" box="[589,685,368,392]" italics="true" pageId="6" pageNumber="35">diabolus</emphasis>
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</taxonomicName>
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is Latin for devil and is used as a noun in apposition. The devil according to Christianity is the spirit of evil and enemy of God, often represented as the ruler of hell and depicted as a human figure with horns, cloven hoofs, fire-red eyes, trident, and tail. The species name is in allusion to the hypertrophied forelimbs, enlarged prepollex with a projecting spine, trident vomers, and dark red iris in life.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |