treatments-xml/data/5C/37/A7/5C37A735FFEEBA74FF28B7EBF666D077.xml
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<document id="8E27BB7635E30D297824CCBB9D2B1C9C" ID-CLB-Dataset="46210" ID-DOI="10.5281/zenodo.278895" ID-GBIF-Dataset="3498da36-33bd-4135-a475-9c978c7f7abc" ID-ISSN="1175-5326" ID-Zenodo-Dep="278895" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460397671527" checkinUser="plazi" docAuthor="Borrero, Francisco J. &amp; Breure, Abraham S. H." docDate="2011" docId="5C37A735FFEEBA74FF28B7EBF666D077" docLanguage="en" docName="zt03054p059.pdf" docOrigin="Zootaxa 3054" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Plekocheilus (Plekocheilus) glandiformis Lea 1838, comb. nov." docType="treatment" docVersion="9" lastPageNumber="22" masterDocId="A00EDF4DFFFABA61FFBFB77CF716D113" masterDocTitle="The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas" masterLastPageNumber="59" masterPageNumber="1" pageNumber="21" updateTime="1698298882325" updateUser="ExternalLinkService">
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<mods:title id="5E95D898029760BF2A3B39F3BF7BC058">The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas</mods:title>
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<mods:namePart id="71D46BBFB6A8BEF1914A4ABB80B7694E">Borrero, Francisco J.</mods:namePart>
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<mods:namePart id="28658E2443F817EC8D94FEDC3B0547C8">Breure, Abraham S. H.</mods:namePart>
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<mods:date id="3E1A034EE37632A8110ACC027EE07782">2011</mods:date>
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<treatment id="5C37A735FFEEBA74FF28B7EBF666D077" ID-DOI="http://doi.org/10.5281/zenodo.6195060" ID-GBIF-Taxon="119418577" ID-Zenodo-Dep="6195060" LSID="urn:lsid:plazi:treatment:5C37A735FFEEBA74FF28B7EBF666D077" httpUri="http://treatment.plazi.org/id/5C37A735FFEEBA74FF28B7EBF666D077" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<subSubSection id="9C8445A8FFEEBA75FF28B7EBF658D1C1" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="D4211623FFEEBA75FF28B7EBF4BFD1A2" blockId="20.[151,937,151,210]" box="[151,937,151,177]" pageId="20" pageNumber="21">
<heading id="8F69A14FFFEEBA75FF28B7EBF4BFD1A2" bold="true" box="[151,937,151,177]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="E6EACA31FFEEBA75FF28B7EBF4BFD1A2" bold="true" box="[151,937,151,177]" pageId="20" pageNumber="21">
<taxonomicName id="139E6DA0FFEEBA75FF28B7EBF40AD1A2" authority="Lea, 1838" authorityName="Lea" authorityYear="1838" box="[151,796,151,177]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis" status="comb. nov." subGenus="Plekocheilus">
<emphasis id="E6EACA31FFEEBA75FF28B7EBF624D1A2" bold="true" box="[151,306,151,177]" italics="true" pageId="20" pageNumber="21">Plekocheilus</emphasis>
(
<emphasis id="E6EACA31FFEEBA75FEFCB7EBF6CBD1A2" bold="true" box="[323,477,151,177]" italics="true" pageId="20" pageNumber="21">Plekocheilus</emphasis>
)
<emphasis id="E6EACA31FFEEBA75FE51B7EBF59FD1A2" bold="true" box="[494,649,151,177]" italics="true" pageId="20" pageNumber="21">glandiformis</emphasis>
(
<bibRefCitation id="B00F6BD2FFEEBA75FD27B7EBF405D1A2" author="Lea" box="[664,787,151,177]" pageId="20" pageNumber="21" refString="Lea, I. (1838) Descriptions of new freshwater and land shells. Transactions of the American Philosophical Society, (n. s.) 6, 1 - 154, pls. 1 - 24." type="journal article" year="1838">Lea, 1838</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="FDD9774AFFEEBA75FC9CB7EBF4BFD1A2" box="[803,937,151,177]" pageId="20" pageNumber="21">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D4211623FFEEBA75FF28B7C6F658D1C1" blockId="20.[151,937,151,210]" box="[151,334,186,210]" pageId="20" pageNumber="21">
(
<figureCitation id="4CA50AA6FFEEBA75FF20B7C6F7F5D1C1" box="[159,227,186,210]" captionStart="FIGURE 8" captionStartId="21.[151,250,1630,1653]" captionTargetBox="[151,1436,431,1580]" captionTargetId="figure@21.[151,1436,431,1580]" captionTargetPageId="21" captionText="FIGURE 8. Plekocheilus s. str. A F. P. (P.) glandiformis (Lea, 1838). A C. Holotype, USNM 105045 (H 33 mm). D F. Holotype of couturesi Ancey, NMW 1955.158. 24074 (H 35.7 mm). G I. P. (P.) taquinensis (Pfeiffer, 1855), lectotype BMNH 1957.6. 3.1 (H 40.2 mm), photo P. Crabb. J O. P. (P.) bigener spec. nov. J L. Holotype RMNH 125626 (H 38.5 mm). M O. Paratype RMNH 125627." httpUri="https://zenodo.org/record/278903/files/figure.png" pageId="20" pageNumber="21">Figs 8</figureCitation>
AF, 9D)
</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFEEBA75FF28B67EF27FD067" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="D4211623FFEEBA75FF28B67EF3F6D024" blockId="20.[151,1436,258,373]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="F48E310DFFEEBA75FF28B67EF3F6D024" pageId="20" pageNumber="21">
<taxonomicName id="139E6DA0FFEEBA75FF28B67EF505D00B" ID-CoL="NTDC" authority="Lea, 1838: 83" authorityName="Lea" authorityPageNumber="83" authorityYear="1838" box="[151,531,258,280]" class="Gastropoda" family="Amphibulimidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEEBA75FF28B67EF663D00B" box="[151,373,258,280]" italics="true" pageId="20" pageNumber="21">Bulimus glandiformis</emphasis>
<treatmentCitation id="553F3032FFEEBA75FEC0B67FF505D00B" author="Lea" box="[383,531,259,280]" page="83" pageId="20" pageNumber="21" year="1838">
<bibRefCitation id="B00F6BD2FFEEBA75FEC0B67FF6FAD00B" author="Lea" box="[383,492,259,280]" pageId="20" pageNumber="21" refString="Lea, I. (1838) Descriptions of new freshwater and land shells. Transactions of the American Philosophical Society, (n. s.) 6, 1 - 154, pls. 1 - 24." type="journal article" year="1838">Lea, 1838</bibRefCitation>
: 83
</treatmentCitation>
</taxonomicName>
, pl. 23 fig. 92.
<typeStatus id="0B25A881FFEEBA75FD01B67FF5E6D00B" box="[702,752,259,280]" pageId="20" pageNumber="21">Type</typeStatus>
locality:
<collectingCountry id="AC8956B3FFEEBA75FCE9B67EF4AAD00B" box="[854,956,258,280]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
, between La Plata and Tocaima (see remarks below).
<typeStatus id="0B25A881FFEEBA75FEA5B65DF66ED024" box="[282,376,289,311]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
USNM 105045.
<taxonomicName id="139E6DA0FFEEBA75FD99B65DF479D02B" authority="Lea" authorityName="Lea" box="[550,879,289,312]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEEBA75FD99B65DF439D024" box="[550,815,289,311]" italics="true" pageId="20" pageNumber="21">Plekocheilus glandiformis</emphasis>
(Lea)
</taxonomicName>
;
<treatmentCitation id="553F3032FFEEBA75FCC5B65DF34FD024" author="Richardson" box="[890,1113,289,311]" page="314" pageId="20" pageNumber="21" year="1995">
<bibRefCitation id="B00F6BD2FFEEBA75FCC5B65DF33FD024" author="Richardson" box="[890,1065,289,311]" pageId="20" pageNumber="21" refString="Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458." type="journal article" year="1995">Richardson 1995</bibRefCitation>
: 314
</treatmentCitation>
(references).
</treatmentCitationGroup>
</paragraph>
<paragraph id="D4211623FFEEBA75FF28B63DF27FD067" blockId="20.[151,1436,258,373]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="F48E310DFFEEBA75FF28B63DF27FD067" pageId="20" pageNumber="21">
<taxonomicName id="139E6DA0FFEEBA75FF28B63DF6ECD046" ID-CoL="6HG25" authority="Ancey 1900: 42" authorityName="Ancey" authorityPageNumber="42" authorityYear="1900" box="[151,506,320,342]" class="Gastropoda" family="Amphibulimidae" genus="Eurytus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="couturesi">
<emphasis id="E6EACA31FFEEBA75FF28B63DF65CD045" box="[151,330,321,342]" italics="true" pageId="20" pageNumber="21">Eurytus couturesi</emphasis>
<treatmentCitation id="553F3032FFEEBA75FEECB63CF6ECD046" author="Ancey" box="[339,506,320,341]" page="42" pageId="20" pageNumber="21" year="1900">
<bibRefCitation id="B00F6BD2FFEEBA75FEECB63CF6C2D046" author="Ancey" box="[339,468,320,341]" pageId="20" pageNumber="21" refString="Ancey, C. F. (1900) Description of a land shell from South America. The Nautilus, 14, 42." type="journal article" year="1900">Ancey 1900</bibRefCitation>
: 42
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="0B25A881FFEEBA75FDB9B63DF52ED045" box="[518,568,321,342]" pageId="20" pageNumber="21">Type</typeStatus>
locality: “
<collectingCountry id="AC8956B3FFEEBA75FD1AB63CF5E4D045" box="[677,754,320,342]" name="Bolivia" pageId="20" pageNumber="21">Bolivie</collectingCountry>
” [see remarks].
<typeStatus id="0B25A881FFEEBA75FC22B63CF4EDD045" box="[925,1019,320,342]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
NMW 1955.158.24074;
<treatmentCitation id="553F3032FFEEBA75FB40B63CF671D067" author="Wood" page="39" pageId="20" pageNumber="21" year="2008">
<bibRefCitation id="B00F6BD2FFEEBA75FB40B63CF657D067" author="Wood" pageId="20" pageNumber="21" refString="Wood, H. &amp; Gallichan, J. (2008) The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organisms from the National Museum of Wales, Biotir Reports, 3, i - vi, 1 - 162." type="journal article" year="2008">Wood and Gallichan 2008</bibRefCitation>
: 39
</treatmentCitation>
, pl. 9 fig. 10, x.
<taxonomicName id="139E6DA0FFEEBA75FDB3B623F458D066" authority="Ancey" authorityName="Ancey" box="[524,846,351,373]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="couturesi">
<emphasis id="E6EACA31FFEEBA75FDB3B623F5E4D066" box="[524,754,351,373]" italics="true" pageId="20" pageNumber="21">Plekocheilus couturesi</emphasis>
(Ancey)
</taxonomicName>
;
<treatmentCitation id="553F3032FFEEBA75FCE7B622F32ED067" author="Richardson" box="[856,1080,350,372]" page="308" pageId="20" pageNumber="21" year="1995">
<bibRefCitation id="B00F6BD2FFEEBA75FCE7B622F31ED067" author="Richardson" box="[856,1032,350,372]" pageId="20" pageNumber="21" refString="Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458." type="journal article" year="1995">Richardson 1995</bibRefCitation>
: 308
</treatmentCitation>
(references). New synonymy.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFEEBA75FF28B6DCF256D2C6" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="D4211623FFEEBA75FF28B6DCF649D2A2" blockId="20.[151,1436,416,2026]" pageId="20" pageNumber="21">
<emphasis id="E6EACA31FFEEBA75FF28B6DCF610D0AA" bold="true" box="[151,262,416,441]" pageId="20" pageNumber="21">Material.</emphasis>
Cauca, Volcán de Puracé, Paletará,
<quantity id="1366BBC6FFEEBA75FD20B6DDF42BD0AA" box="[671,829,417,441]" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="20" pageNumber="21" unit="m" value="3100.0" valueMax="3200.0" valueMin="3000.0">30003200 m</quantity>
, leg./1951:
<collectionCode id="B28F8EE6FFEEBA75FC7DB6DEF305D0AA" LSID="urn:lsid:biocol.org:col:34795" box="[962,1043,418,441]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21">FMNH</collectionCode>
109715); “La Cumbre or Popayán (?),” F.W. Pennell &amp; E.P. Killip leg./1922:
<collectionCode id="B28F8EE6FFEEBA75FDC4B6B9F5D5D0CE" LSID="urn:lsid:biocol.org:col:34784" box="[635,707,453,477]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="20" pageNumber="21">ANSP</collectionCode>
132450,
<collectionCode id="B28F8EE6FFEEBA75FC92B6BAF468D0CE" LSID="urn:lsid:biocol.org:col:34795" box="[813,894,454,477]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21">FMNH</collectionCode>
78741; Chocó, Rio Nuquí, Haught leg.:
<collectionCode id="B28F8EE6FFEEBA75FAF4B6B9F28AD0CE" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,453,477]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
488861 (as
<taxonomicName id="139E6DA0FFEEBA75FEA1B695F50AD312" box="[286,540,489,513]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="jucundus">
<emphasis id="E6EACA31FFEEBA75FEA1B695F50AD312" box="[286,540,489,513]" italics="true" pageId="20" pageNumber="21">Plekocheilus jucundus</emphasis>
</taxonomicName>
); Quindío, Hacienda Alaska, Finca Bengala,
<quantity id="1366BBC6FFEEBA75FB96B695F372D312" box="[1065,1124,489,513]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="21" unit="km" value="3.0">3 km</quantity>
N of end of road Salento Cocora,
<quantity id="1366BBC6FFEEBA75FF47B571F644D336" box="[248,338,525,549]" metricMagnitude="3" metricUnit="m" metricValue="3.25" pageId="20" pageNumber="21" unit="m" value="3250.0">3250 m</quantity>
, C. Restrepo leg./1982:
<collectionCode id="B28F8EE6FFEEBA75FDD5B572F5B2D336" box="[618,676,526,549]" pageId="20" pageNumber="21">UVZ</collectionCode>
82252; “Quindío subpáramo,” F. Ortega leg./2004:
<collectionCode id="B28F8EE6FFEEBA75FB44B572F22ED336" box="[1275,1336,526,549]" pageId="20" pageNumber="21">UVZ</collectionCode>
; Tolima,
<collectingCountry id="AC8956B3FFEEBA75FF28B54DF7FFD35A" box="[151,233,561,585]" name="Bolivia" pageId="20" pageNumber="21">Bolivia</collectingCountry>
[see remarks]:
<collectionCode id="B28F8EE6FFEEBA75FE21B54EF6F3D35A" box="[414,485,562,585]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21">NMW</collectionCode>
1955.158.24074 (
<typeStatus id="0B25A881FFEEBA75FD09B54DF40CD35A" box="[694,794,561,585]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
of E.
<taxonomicName id="139E6DA0FFEEBA75FCD9B54EF333D35A" authority="Ancey" authorityName="Ancey" box="[870,1061,561,585]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="couturesi">
<emphasis id="E6EACA31FFEEBA75FCD9B54EF4DBD35A" box="[870,973,562,585]" italics="true" pageId="20" pageNumber="21">couturesi</emphasis>
Ancey
</taxonomicName>
); Roncesvalles, Vereda La Yerbabuena, Páramo El Oasis,
<quantity id="1366BBC6FFEEBA75FE7AB529F508D37E" box="[453,542,597,621]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="20" pageNumber="21" unit="m" value="3350.0">3350 m</quantity>
, Linares leg./2005:
<collectionCode id="B28F8EE6FFEEBA75FCBDB529F453D37E" box="[770,837,597,621]" pageId="20" pageNumber="21">ICNB</collectionCode>
10571;? Between La Plata and Tocaima [Tolima?], J.H. Gibbon leg:
<collectionCode id="B28F8EE6FFEEBA75FEE7B505F6BAD382" LSID="urn:lsid:biocol.org:col:34871" box="[344,428,633,657]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
; Valle del Cauca, Hacienda Las Vegas,
<collectingCountry id="AC8956B3FFEEBA75FCD7B505F4E6D382" box="[872,1008,633,657]" name="Saint Lucia" pageId="20" pageNumber="21">Santa Lucía</collectingCountry>
, Tuluá,
<quantity id="1366BBC6FFEEBA75FBF5B505F3B7D382" box="[1098,1185,633,657]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="21" unit="m" value="3000.0">3000 m</quantity>
, E. Velasco leg./1984:
<collectionCode id="B28F8EE6FFEEBA75FF28B5E2F7C7D3A6" box="[151,209,670,693]" pageId="20" pageNumber="21">UVZ</collectionCode>
84020; Páramo de Riofrio,
<quantity id="1366BBC6FFEEBA75FDADB5E1F5A6D3A6" box="[530,688,669,693]" metricMagnitude="3" metricUnit="m" metricValue="3.425" metricValueMax="3.5" metricValueMin="3.35" pageId="20" pageNumber="21" unit="m" value="3425.0" valueMax="3500.0" valueMin="3350.0">33503500 m</quantity>
, V. Rojas leg./1997:
<collectionCode id="B28F8EE6FFEEBA75FC1FB5E2F4CCD3A6" box="[928,986,670,693]" pageId="20" pageNumber="21">UVZ</collectionCode>
97023,
<collectionCode id="B28F8EE6FFEEBA75FB87B5E2F364D3A6" box="[1080,1138,670,693]" pageId="20" pageNumber="21">UVZ</collectionCode>
97024 (in
<taxonomicName id="139E6DA0FFEEBA75FB4EB5E1F286D3A6" box="[1265,1424,669,693]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
); Páramo El Duende, Riofrio,
<quantity id="1366BBC6FFEEBA75FE69B5BDF53AD3CA" box="[470,556,705,729]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="20" pageNumber="21" unit="m" value="3400.0">3400 m</quantity>
, L.A. Neira leg./2004:
<collectionCode id="B28F8EE6FFEEBA75FC92B5BEF47CD3CA" box="[813,874,706,729]" pageId="20" pageNumber="21">UVZ</collectionCode>
; Parque Nacional Farallones [= PNF], trail to Pico de Loro,
<quantity id="1366BBC6FFEEBA75FEBFB599F689D3EE" box="[256,415,741,765]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="20" pageNumber="21" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">24002500 m</quantity>
, F. Zapata leg./1981:
<collectionCode id="B28F8EE6FFEEBA75FD26B59AF5C5D3EE" box="[665,723,742,765]" pageId="20" pageNumber="21">UVZ</collectionCode>
81246; PNF, Cerro Teta de la Viuda,
<quantity id="1366BBC6FFEEBA75FB38B599F3F6D3EE" box="[1159,1248,741,765]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="21" unit="m" value="2800.0">2800 m</quantity>
, G. Bolívar leg./ 1981:
<collectionCode id="B28F8EE6FFEEBA75FF60B476F60ED232" box="[223,280,778,801]" pageId="20" pageNumber="21">UVZ</collectionCode>
81242; PNF, Campamento
<collectingCountry id="AC8956B3FFEEBA75FDEAB475F58AD232" box="[597,668,777,801]" name="South Korea" pageId="20" pageNumber="21">Corea</collectingCountry>
:
<collectionCode id="B28F8EE6FFEEBA75FD17B476F5F7D232" box="[680,737,778,801]" pageId="20" pageNumber="21">UVZ</collectionCode>
82022 (
<quantity id="1366BBC6FFEEBA75FCFFB475F48CD232" box="[832,922,777,801]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="21" unit="m" value="2300.0">2300 m</quantity>
, C. Restrepo leg./1982),
<collectionCode id="B28F8EE6FFEEBA75FB09B476F3E6D232" box="[1206,1264,778,801]" pageId="20" pageNumber="21">UVZ</collectionCode>
82197 (Zapata leg./1982),
<collectionCode id="B28F8EE6FFEEBA75FEA8B452F647D256" box="[279,337,814,837]" pageId="20" pageNumber="21">UVZ</collectionCode>
81133 (~
<quantity id="1366BBC6FFEEBA75FE06B451F502D256" box="[441,532,813,837]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="20" pageNumber="21" unit="m" value="2700.0">2700 m</quantity>
); Reserva Hato Viejo,
<quantity id="1366BBC6FFEEBA75FCAEB451F47ED256" box="[785,872,813,837]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="20" pageNumber="21" unit="m" value="2900.0">2900 m</quantity>
, R. García &amp; A.F. Giraldo leg./1996:
<collectionCode id="B28F8EE6FFEEBA75FAB1B452F251D256" box="[1294,1351,814,837]" pageId="20" pageNumber="21">UVZ</collectionCode>
97018; road to La Herrera, Florida [
<quantity id="1366BBC6FFEEBA75FE6BB42DF561D27A" box="[468,631,849,873]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.4" metricValueMin="2.5" pageId="20" pageNumber="21" unit="m" value="2950.0" valueMax="3400.0" valueMin="2500.0">25003400 m</quantity>
], L.A. Neira leg./1993:
<collectionCode id="B28F8EE6FFEEBA75FC3CB42EF4D6D27A" box="[899,960,850,873]" pageId="20" pageNumber="21">UVZ</collectionCode>
.?,
<collectingCountry id="AC8956B3FFEEBA75FC59B42DF34FD27A" box="[998,1113,849,873]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
:
<collectionCode id="B28F8EE6FFEEBA75FBDCB42DF3A2D27A" LSID="urn:lsid:biocol.org:col:34871" box="[1123,1204,849,873]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
532642 (ex Fulton);
<collectingCountry id="AC8956B3FFEEBA75FF28B40AF620D29E" box="[151,310,885,909]" name="Colombia" pageId="20" pageNumber="21">New Granada</collectingCountry>
:
<collectionCode id="B28F8EE6FFEEBA75FEFEB40AF686D29E" box="[321,400,886,909]" httpUri="http://grbio.org/cool/hx01-4v4f" name="Zoologisches Museum der Universitaet Zuerich" pageId="20" pageNumber="21">ZMUZ</collectionCode>
511629,
<collectionCode id="B28F8EE6FFEEBA75FE47B40AF551D29E" box="[504,583,886,909]" httpUri="http://grbio.org/cool/hx01-4v4f" name="Zoologisches Museum der Universitaet Zuerich" pageId="20" pageNumber="21">ZMUZ</collectionCode>
511630 (both not seen); “
<collectingCountry id="AC8956B3FFEEBA75FCD3B409F488D29E" box="[876,926,885,909]" name="United States of America" pageId="20" pageNumber="21">U.S.</collectingCountry>
<collectingCountry id="AC8956B3FFEEBA75FC1AB409F30CD29E" box="[933,1050,885,909]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
:”
<collectionCode id="B28F8EE6FFEEBA75FB90B40AF390D29E" box="[1071,1158,886,909]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="20" pageNumber="21">UMMZ</collectionCode>
147711 (ex Fulton, as
<taxonomicName id="139E6DA0FFEEBA75FA38B40AF644D2A2" authority="Ancey" authorityName="Ancey" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="couturesi">
<emphasis id="E6EACA31FFEEBA75FA38B40AF7E8D2A2" italics="true" pageId="20" pageNumber="21">P. couturesi</emphasis>
Ancey
</taxonomicName>
).
</paragraph>
<paragraph id="D4211623FFEEBA75FF7AB4C1F256D2C6" blockId="20.[151,1436,416,2026]" box="[197,1344,957,982]" pageId="20" pageNumber="21">
<emphasis id="E6EACA31FFEEBA75FF7AB4C1F68CD2C6" bold="true" box="[197,410,957,982]" pageId="20" pageNumber="21">Altitudinal range.</emphasis>
The localities presented indicate a range of
<quantity id="1366BBC6FFEEBA75FC37B4C1F335D2C6" box="[904,1059,957,981]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="3.5" metricValueMin="1.7" pageId="20" pageNumber="21" unit="m" value="2600.0" valueMax="3500.0" valueMin="1700.0">17003500 m</quantity>
(but see remarks below).
</paragraph>
</subSubSection>
<subSubSection id="9C8445A8FFEEBA74FF7AB49EF666D077" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="D4211623FFEEBA75FF7AB49EF517D42E" blockId="20.[151,1436,416,2026]" pageId="20" pageNumber="21">
<emphasis id="E6EACA31FFEEBA75FF7AB49EF62FD2E9" bold="true" box="[197,313,994,1018]" pageId="20" pageNumber="21">Remarks.</emphasis>
After having studied the
<typeStatus id="0B25A881FFEEBA75FDE7B49DF5ABD2EA" box="[600,701,993,1017]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
, we conclude that this species belongs to the nominate subgenus, contrary to the classification in
<emphasis id="E6EACA31FFEEBA75FE41B37AF507D50E" box="[510,529,1030,1053]" italics="true" pageId="20" pageNumber="21">P.</emphasis>
(
<taxonomicName id="139E6DA0FFEEBA75FD9BB37AF56DD50E" box="[548,635,1030,1053]" class="Gastropoda" family="Amphibulimidae" genus="Eurytus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="E6EACA31FFEEBA75FD9BB37AF56DD50E" box="[548,635,1030,1053]" italics="true" pageId="20" pageNumber="21">Eurytus</emphasis>
</taxonomicName>
) by
<bibRefCitation id="B00F6BD2FFEEBA75FD0EB37AF496D50E" author="Breure" box="[689,896,1029,1053]" pageId="20" pageNumber="21" refString="Breure, A. S. H. (1979) Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen, 168, 1 - 215." type="journal article" year="1979">Breure (1979: 30)</bibRefCitation>
. The sculpture is clearly malleated as in typical
<taxonomicName id="139E6DA0FFEEBA75FF28B355F630D552" box="[151,294,1065,1089]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="E6EACA31FFEEBA75FF28B355F630D552" box="[151,294,1065,1089]" italics="true" pageId="20" pageNumber="21">Plekocheilus</emphasis>
</taxonomicName>
(
<emphasis id="E6EACA31FFEEBA75FE87B356F65CD552" box="[312,330,1066,1089]" italics="true" pageId="20" pageNumber="21">P.</emphasis>
), but also shows granulation that is typically found in a large number of species in
<emphasis id="E6EACA31FFEEBA75FAB1B356F236D552" box="[1294,1312,1066,1089]" italics="true" pageId="20" pageNumber="21">P.</emphasis>
(
<taxonomicName id="139E6DA0FFEEBA75FA8BB356F29DD552" box="[1332,1419,1066,1089]" class="Gastropoda" family="Amphibulimidae" genus="Eurytus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="E6EACA31FFEEBA75FA8BB356F29DD552" box="[1332,1419,1066,1089]" italics="true" pageId="20" pageNumber="21">Eurytus</emphasis>
</taxonomicName>
). The figure of the sculpture in
<bibRefCitation id="B00F6BD2FFEEBA75FE57B331F40BD576" author="Pilsbry" box="[488,797,1101,1125]" pageId="20" pageNumber="21" refString="Pilsbry, H. A. (1895 - 1896) Manual of Conchology, (2) 10. American Bulimi and Bululimuli. Strophocheilus, Plekocheilus, Auris, Bulimulus. Academy of Natural Sciences of Philadelphia, Philadelphia, iv + 213 pp, 51 pls." type="book" year="1895">Pilsbry (1895: pl. 38 fig. 4)</bibRefCitation>
is somewhat misleading, as the granulation in
<taxonomicName id="139E6DA0FFEEBA75FA90B332F7C9D59A" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEEBA75FA90B332F7C9D59A" italics="true" pageId="20" pageNumber="21">P. glandiformis</emphasis>
</taxonomicName>
is less strong than suggested. This species is thus somewhat intermediate between the two subgenera, although we keep the distinction between the two and still classify
<taxonomicName id="139E6DA0FFEEBA75FC26B3EAF353D5BE" box="[921,1093,1173,1197]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEEBA75FC26B3EAF353D5BE" box="[921,1093,1173,1197]" italics="true" pageId="20" pageNumber="21">P. glandiformis</emphasis>
</taxonomicName>
witin the nominate subgenus. Some variation exists in the height of the spire, and in the general coloration (some specimens are a very dark olive brown, frequently with pink peristome). Owing to the fact that no anatomical or molecular data are yet available, we treat them here as part of a single taxon. Further study is clearly needed. Has been found in
<taxonomicName id="139E6DA0FFEEBA75FB7CB27DF276D40A" box="[1219,1376,1281,1305]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
, suggesting a partly arboreal habitat.
</paragraph>
<paragraph id="D4211623FFEEBA75FF7AB236F54FD792" blockId="20.[151,1436,416,2026]" pageId="20" pageNumber="21">
We consider the
<taxonomicName id="139E6DA0FFEEBA75FE38B236F522D472" box="[391,564,1353,1377]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEEBA75FE38B236F522D472" box="[391,564,1353,1377]" italics="true" pageId="20" pageNumber="21">P. glandiformis</emphasis>
</taxonomicName>
group a high altitude group (all precise localities are at
<quantity id="1366BBC6FFEEBA75FB7DB235F20CD472" box="[1218,1306,1353,1377]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="m" value="2000.0">2000 m</quantity>
or higher). Thus the
<typeStatus id="0B25A881FFEEBA75FEBDB212F624D495" box="[258,306,1390,1414]" pageId="20" pageNumber="21">type</typeStatus>
locality of
<taxonomicName id="139E6DA0FFEEBA75FE09B212F581D496" authority="Lea" authorityName="Lea" box="[438,663,1389,1413]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEEBA75FE09B212F574D496" box="[438,610,1389,1413]" italics="true" pageId="20" pageNumber="21">P. glandiformis</emphasis>
Lea
</taxonomicName>
, based on material collected by Gibbon, is likely in southern Antioquia; a finca Tocaima [
<geoCoordinate id="B1AA70E4FFEEBA75FE2AB2EDF50AD4BA" box="[405,540,1425,1449]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="5.538889">05°32'20&quot;N</geoCoordinate>
<geoCoordinate id="B1AA70E4FFEEBA75FD9DB2EDF5ABD4BA" box="[546,701,1425,1449]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-75.93222">075°55'56&quot;W</geoCoordinate>
] is located at
<quantity id="1366BBC6FFEEBA75FCE9B2EDF4B8D4BA" box="[854,942,1425,1449]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="20" pageNumber="21" unit="m" value="1950.0">1950 m</quantity>
. “La Plata” could be assumed to be La Plata [
<geoCoordinate id="B1AA70E4FFEEBA75FF21B2C9F632D4DE" box="[158,292,1461,1485]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="5.038611">05°02'19&quot;N</geoCoordinate>
<geoCoordinate id="B1AA70E4FFEEBA75FE95B2C9F6D3D4DE" box="[298,453,1461,1485]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-75.6775">075°40'39&quot;W</geoCoordinate>
] in Caldas (which is the nearest location of that name, ca.
<quantity id="1366BBC6FFEEBA75FBF4B2C9F384D4DE" box="[1099,1170,1461,1485]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="km" value="60.0">60 km</quantity>
SSW of Tocaima as the crow flies), crossing high altitudes in the border region of Antioquia/Risaralda/Caldas. However, doubt remains, as we do not know anything about the itinerary of Dr. Gibbon. There are two localities with the name La Cumbre in Cauca, both (S)SW of Popayán, respectively
<geoCoordinate id="B1AA70E4FFEEBA75FD31B15DF5FDD72A" box="[654,747,1569,1593]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="1.9166667">01°55'N</geoCoordinate>
<geoCoordinate id="B1AA70E4FFEEBA75FD4EB15DF474D72A" box="[753,866,1569,1593]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-76.95">076°57'W</geoCoordinate>
and
<geoCoordinate id="B1AA70E4FFEEBA75FC29B15DF30DD72A" box="[918,1051,1569,1593]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="1.7658334">01°45'57&quot;N</geoCoordinate>
<geoCoordinate id="B1AA70E4FFEEBA75FB9EB15DF3ACD72A" box="[1057,1210,1569,1593]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-76.784996">076°47'06&quot;W</geoCoordinate>
. Lot USNM 488861 (Chocó, Rio Nuquí) is problematic since it is the westernmost locality for this species, and is at a comparatively low elevation (probably at most
<quantity id="1366BBC6FFEEBA75FDBDB115F55BD792" box="[514,589,1641,1665]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="500.0">500 m</quantity>
).
</paragraph>
<paragraph id="D4211623FFEEBA74FF7AB1F1F63CD1E4" blockId="20.[151,1436,416,2026]" lastBlockId="21.[151,1437,151,356]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
The material of
<taxonomicName id="139E6DA0FFEEBA75FEC8B1F2F6EFD7B6" box="[375,505,1678,1701]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="couturesi">
<emphasis id="E6EACA31FFEEBA75FEC8B1F2F6EFD7B6" box="[375,505,1678,1701]" italics="true" pageId="20" pageNumber="21">P. couturesi</emphasis>
</taxonomicName>
was collected by G. Coutures, but we have been unable to find any details about his travels in South
<collectingCountry id="AC8956B3FFEEBA75FEF0B1CDF6A5D7DA" box="[335,435,1713,1737]" name="United States of America" pageId="20" pageNumber="21">America</collectingCountry>
. The sculpture of the
<typeStatus id="0B25A881FFEEBA75FD16B1CDF41DD7DA" box="[681,779,1713,1737]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
consists of weak malleation and very fine granulation (cf.
<figureCitation id="4CA50AA6FFEEBA75FF28B1A9F7CCD7FE" box="[151,218,1749,1773]" captionStart="FIGURE 4" captionStartId="9.[151,250,1088,1111]" captionTargetBox="[188,1423,269,1050]" captionTargetId="figure@9.[151,1436,235,1067]" captionTargetPageId="9" captionText="FIGURE 4. Dryptus distribution. A. D. pardalis. B. D. moritzianus and D. stuebeli." httpUri="https://zenodo.org/record/278899/files/figure.png" pageId="20" pageNumber="21">Fig. 4</figureCitation>
D). This is unlike the sculpture found in Bolivian
<taxonomicName id="139E6DA0FFEEBA75FCBFB1A9F499D7FE" box="[768,911,1749,1773]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="E6EACA31FFEEBA75FCBFB1A9F499D7FE" box="[768,911,1749,1773]" italics="true" pageId="20" pageNumber="21">Plekocheilus</emphasis>
</taxonomicName>
, but completely in accordance with material of two other species we now place in the group of
<taxonomicName id="139E6DA0FFEEBA75FD0BB186F476D602" box="[692,864,1785,1809]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEEBA75FD0BB186F476D602" box="[692,864,1785,1809]" italics="true" pageId="20" pageNumber="21">P. glandiformis</emphasis>
</taxonomicName>
. Thus we conclude that Anceys location has been misinterpreted and now consider this taxon as Colombian. When we realized that there could be a locality named Bolivia in
<collectingCountry id="AC8956B3FFEEBA75FE9EB03DF699D64A" box="[289,399,1857,1881]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
(Bolivie is the French version of this name), we found ten localities in the GNS Gazetteer (http://earth-info.nga.mil/gns/html/index.html). Considering this a high altitude group, we could eliminate nine localities. We now assume that the
<typeStatus id="0B25A881FFEEBA75FDA2B0F6F55BD6B1" box="[541,589,1930,1954]" pageId="20" pageNumber="21">type</typeStatus>
locality of
<taxonomicName id="139E6DA0FFEEBA75FD73B0F6F458D6B2" box="[716,846,1930,1953]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="couturesi">
<emphasis id="E6EACA31FFEEBA75FD73B0F6F458D6B2" box="[716,846,1930,1953]" italics="true" pageId="20" pageNumber="21">P. couturesi</emphasis>
</taxonomicName>
is likely to be in the border area between Tolima and Valle del Cauca, where a place
<collectingCountry id="AC8956B3FFEEBA75FE4BB0D1F550D6D6" box="[500,582,1965,1989]" name="Bolivia" pageId="20" pageNumber="21">Bolivia</collectingCountry>
exists at
<quantity id="1366BBC6FFEEBA75FD10B0D1F415D6D6" box="[687,771,1965,1989]" metricMagnitude="3" metricUnit="m" metricValue="3.29" pageId="20" pageNumber="21" unit="m" value="3290.0">3290 m</quantity>
(
<geoCoordinate id="B1AA70E4FFEEBA75FCADB0D1F481D6D6" box="[786,919,1965,1989]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="3.8933332">03°53'36&quot;N</geoCoordinate>
<geoCoordinate id="B1AA70E4FFEEBA75FC21B0D1F32CD6D6" box="[926,1082,1965,1989]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-75.78833">075°47'18&quot;W</geoCoordinate>
). A second specimen labeled as
<taxonomicName id="139E6DA0FFEEBA75FF28B0AEF67AD6F9" authority="Ancey" authorityName="Ancey" box="[151,364,2002,2026]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="couturesi">
<emphasis id="E6EACA31FFEEBA75FF28B0AEF60CD6FA" box="[151,282,2002,2025]" italics="true" pageId="20" pageNumber="21">P. couturesi</emphasis>
Ancey
</taxonomicName>
, and certainly corresponding to this taxon, is held at UMMZ (lot 147711). The label accompanying the single shell states “original lot,” suggesting that it may be a
<typeStatus id="0B25A881FFEFBA74FC01B7E4F334D1A3" box="[958,1058,152,176]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
. However, this could not be confirmed as the label may not be the original from C.F. Ancey, but a label written later by H.C. Fulton (J. Gallichan, pers.comm.).
</paragraph>
<paragraph id="D4211623FFEFBA74FF7AB678F666D077" blockId="21.[151,1437,151,356]" pageId="21" pageNumber="22">
The specimens from Valle del Cauca, Florida, road to La Herrera (which is in southwestern Tolima) probably are from the area around the Parque Nacional Natural Las Hermosas. These specimens are more slender than typical
<taxonomicName id="139E6DA0FFEFBA74FF01B631F67FD077" box="[190,361,332,356]" class="Gastropoda" family="Orthalicidae" genus="Plekocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="glandiformis">
<emphasis id="E6EACA31FFEFBA74FF01B631F67FD077" box="[190,361,332,356]" italics="true" pageId="21" pageNumber="22">P. glandiformis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>