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<document id="EA00E7C24C44B9F7BA908B03FE174BC7" ID-CLB-Dataset="30435" ID-DOI="10.11646/zootaxa.4399.4.1" ID-GBIF-Dataset="082fa0e5-df2b-45d9-b882-110560dc1d54" ID-ISSN="1175-5326" ID-Zenodo-Dep="1206764" ID-ZooBank="93DD4D04-12CF-4B57-B97B-FC3FB0648FE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521900032092" checkinUser="plazi" docAuthor="Carnimeo, Fernando Henrique &amp; Noll, Fernando Barbosa" docDate="2018" docId="35683C06FF82FF8214948C39FE31FC8E" docLanguage="en" docName="zootaxa.4399.4.1.pdf" docOrigin="Zootaxa 4399 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scotaena reversa Carnimeo &amp; Noll 2018, sp. nov." docType="treatment" docVersion="9" lastPageNumber="461" masterDocId="C951447EFF8BFF8814038A5FFF77FFDF" masterDocTitle="On the dumping ground genus Scotaena Klug, 1810 (Hymenoptera: Tiphiidae: Thynninae): Phylogeny, taxonomy and geographic distribution" masterLastPageNumber="490" masterPageNumber="451" pageNumber="460" updateTime="1698533209561" updateUser="ExternalLinkService">
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<mods:title id="6D44ABF16E22FBC02BC10F85E86F6131">On the dumping ground genus Scotaena Klug, 1810 (Hymenoptera: Tiphiidae: Thynninae): Phylogeny, taxonomy and geographic distribution</mods:title>
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<mods:namePart id="DA11DEFA8E1609A037F706418DD83153">Carnimeo, Fernando Henrique</mods:namePart>
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<mods:namePart id="DCEAA13196D4778D644F8A8975B7BBAD">Noll, Fernando Barbosa</mods:namePart>
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<mods:typeOfResource id="D05DB8D18293273BE01C2DCF644BAA05">text</mods:typeOfResource>
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<mods:title id="DF461E5CB9718E030940F5DCE7C782EB">Zootaxa</mods:title>
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<mods:date id="9F853D992936794875E738041552AAB4">2018</mods:date>
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<treatment id="35683C06FF82FF8214948C39FE31FC8E" ID-DOI="http://doi.org/10.5281/zenodo.4484718" ID-GBIF-Taxon="142524454" ID-Zenodo-Dep="4484718" LSID="urn:lsid:plazi:treatment:35683C06FF82FF8214948C39FE31FC8E" httpUri="http://treatment.plazi.org/id/35683C06FF82FF8214948C39FE31FC8E" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460">
<subSubSection id="F5DBDE9BFF82FF8114948C39FE9DF97D" pageId="9" pageNumber="460" type="nomenclature">
<paragraph id="BD7E8D10FF82FF8114948C39FD39F95E" blockId="9.[151,590,1638,1698]" box="[151,590,1638,1665]" pageId="9" pageNumber="460">
<heading id="E6363A7CFF82FF8114948C39FD39F95E" bold="true" box="[151,590,1638,1665]" fontSize="11" level="1" pageId="9" pageNumber="460" reason="1">
<emphasis id="8FB55102FF82FF8114948C39FD39F95E" bold="true" box="[151,590,1638,1665]" pageId="9" pageNumber="460">
<taxonomicName id="7AC1F693FF82FF8114948C39FE9EF95E" authority="Carnimeo" authorityName="Carnimeo &amp; Noll" authorityYear="2018" box="[151,489,1638,1665]" class="Insecta" family="Tiphiidae" genus="Scotaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="reversa" status="sp. nov.">
<emphasis id="8FB55102FF82FF8114948C39FE12F95F" bold="true" box="[151,357,1638,1664]" italics="true" pageId="9" pageNumber="460">Scotaena reversa</emphasis>
Carnimeo
</taxonomicName>
<taxonomicNameLabel id="9486EC79FF82FF8115F38C38FD39F95E" box="[496,590,1639,1665]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="BD7E8D10FF82FF8114948CD6FE9DF97D" blockId="9.[151,590,1638,1698]" box="[151,490,1673,1698]" pageId="9" pageNumber="460">
(
<figureCitation id="25FA9195FF82FF81149C8CD6FE57F97D" box="[159,288,1673,1698]" captionStart="FIGURES 2732" captionStartId="12.[151,264,1179,1202]" captionTargetBox="[151,1436,209,1146]" captionTargetId="figure@12.[151,1436,209,1146]" captionTargetPageId="12" captionText="FIGURES 2732. Scotaena reversa, holotype male. 27, frontal head; 28, dorsal habitus; 29, lateral habitus. Genital capsule: 30, dorsal view; 31, ventral view; 32, lateral view.Scale bar = 1mm." httpUri="https://zenodo.org/record/1206782/files/figure.png" pageId="9" pageNumber="460">Figs 2732</figureCitation>
,
<figureCitation id="25FA9195FF82FF8115288CD5FE95F97D" box="[299,482,1673,1698]" captionStart="FIGURES 90105" captionStartId="29.[151,264,1899,1922]" captionTargetBox="[173,1417,301,1880]" captionTargetId="figure@29.[173,1417,301,1881]" captionText="FIGURES 90105. Scotaena characters. 90, 96, 99: S. trifasciata, 91, 98, 100, 105 S. reversa; 92, 94, 97, 101, 103, 104: S. vetusta; 93, 95, 102: S. horni. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4484930" httpUri="https://zenodo.org/record/4484930/files/figure.png" pageId="9" pageNumber="460">91, 98, 100, 105</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F5DBDE9BFF82FF8114948C8DFBE3F8FA" pageId="9" pageNumber="460" type="materials_examined">
<paragraph id="BD7E8D10FF82FF8114948C8DFBE3F8FA" blockId="9.[151,1435,1745,1829]" pageId="9" pageNumber="460">
<materialsCitation id="0DA9874DFF82FF8114948C8DFA5DF938" ID-GBIF-Occurrence="1836204522" box="[151,1322,1745,1767]" collectingDate="1964-11-11" collectionCode="AMNH" collectorName="Kovacs" country="Brazil" location="Sao Jose do Barreiro" municipality="Serra da Bocaina" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="8FB55102FF82FF8114948C8DFF88F938" bold="true" box="[151,255,1746,1767]" pageId="9" pageNumber="460">
<typeStatus id="627A33B2FF82FF8114948C8DFF8CF938" box="[151,251,1746,1767]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="C5D6CD80FF82FF81150F8C8DFE1CF938" box="[268,363,1746,1767]" name="Brazil" pageId="9" pageNumber="460">BRAZIL</collectingCountry>
:
<collectingMunicipality id="5D1A176AFF82FF8115798C8EFD42F938" box="[378,565,1745,1767]" pageId="9" pageNumber="460">Serra da Bocaina</collectingMunicipality>
,
<location id="B81EDBCBFF82FF8116478C8EFC5EF938" LSID="urn:lsid:plazi:treatment:35683C06FF82FF8214948C39FE31FC8E:B81EDBCBFF82FF8116478C8EFC5EF938" box="[580,809,1745,1767]" country="Brazil" municipality="Serra da Bocaina" name="Sao Jose do Barreiro" pageId="9" pageNumber="460" stateProvince="Sao Paulo">São José do Barreiro</location>
,
<collectingRegion id="7F0543F2FF82FF81173A8C8EFCD0F938" box="[825,935,1745,1767]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="460">São Paulo</collectingRegion>
,
<collectingDate id="D93B5238FF82FF8117B58C8EFB52F938" box="[950,1061,1745,1767]" pageId="9" pageNumber="460" value="1964-11-11">
<date id="C97FABD0FF82FF8117B58C8EFB52F938" box="[950,1061,1745,1767]" pageId="9" pageNumber="460" value="1964-11-11">11/xi/1964</date>
</collectingDate>
,
<specimenCount id="ABC74699FF82FF8110358C8EFB22F938" box="[1078,1109,1745,1767]" pageId="9" pageNumber="460" type="male">1♂</specimenCount>
(
<collectorName id="1034E8C6FF82FF8110698C8DFBC2F938" box="[1130,1205,1746,1767]" pageId="9" pageNumber="460">
<emphasis id="8FB55102FF82FF8110698C8DFBC2F938" box="[1130,1205,1746,1767]" italics="true" pageId="9" pageNumber="460">Kovacs</emphasis>
</collectorName>
) (
<collectionCode id="DBD015D5FF82FF8110CC8C8EFA55F938" box="[1231,1314,1745,1767]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="460">AMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0DA9874DFF82FF81113A8C8DFCBCF8D9" ID-GBIF-Occurrence="1836204523" collectingDate="2014-01-04" collectorName="Parque Nacional da Serra dos Orgaos" country="Brazil" location="Parque Nacional da Serra" pageId="9" pageNumber="460" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<typeStatus id="627A33B2FF82FF81113A8C8DFAE1F938" box="[1337,1430,1746,1767]" pageId="9" pageNumber="460" type="paratype">
<emphasis id="8FB55102FF82FF81113A8C8DFAE1F938" box="[1337,1430,1746,1767]" italics="true" pageId="9" pageNumber="460">Paratype</emphasis>
</typeStatus>
,
<collectingCountry id="C5D6CD80FF82FF8114C48CAEFE50F8D9" box="[199,295,1777,1798]" name="Brazil" pageId="9" pageNumber="460">BRAZIL</collectingCountry>
:
<collectorName id="1034E8C6FF82FF8115328CAEFDC0F8D9" box="[305,695,1776,1798]" pageId="9" pageNumber="460">
<location id="B81EDBCBFF82FF8115328CAEFD4FF8D9" LSID="urn:lsid:plazi:treatment:35683C06FF82FF8214948C39FE31FC8E:B81EDBCBFF82FF8115328CAEFD4FF8D9" box="[305,568,1776,1798]" country="Brazil" name="Parque Nacional da Serra" pageId="9" pageNumber="460" stateProvince="Rio de Janeiro">Parque Nacional da Serra</location>
dos Órgãos
</collectorName>
,
<collectingRegion id="7F0543F2FF82FF8116C08CAFFC2BF8D9" box="[707,860,1776,1798]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="460">Rio de Janeiro</collectingRegion>
,
<collectingDate id="D93B5238FF82FF81176B8CAFFCBCF8D9" box="[872,971,1776,1798]" pageId="9" pageNumber="460" value="2014-01-04">
<date id="C97FABD0FF82FF81176B8CAFFCBCF8D9" box="[872,971,1776,1798]" pageId="9" pageNumber="460" value="2014-01-04">04/i/2014</date>
</collectingDate>
</materialsCitation>
<materialsCitation id="0DA9874DFF82FF8117DB8CAFFBE7F8FA" ID-GBIF-Occurrence="1836204524" collectingDate="2014-02-01" collectingDateMax="2014-02-27" collectingDateMin="2014-02-01" collectionCode="UFRJ" collectorName="R. F. Monteiro" country="Brazil" location="Parque Nacional do Itatiaia" pageId="9" pageNumber="460" specimenCount="4" specimenCount-male="4" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingDate id="D93B5238FF82FF8117DB8CAFFB36F8D9" box="[984,1089,1776,1798]" pageId="9" pageNumber="460" value="2014-02-01">
<date id="C97FABD0FF82FF8117DB8CAFFB36F8D9" box="[984,1089,1776,1798]" pageId="9" pageNumber="460" value="2014-02-01">01/ii/2014</date>
</collectingDate>
,
<specimenCount id="ABC74699FF82FF81104F8CAFFB1AF8D9" box="[1100,1133,1776,1798]" pageId="9" pageNumber="460" type="male">3♂</specimenCount>
(
<emphasis id="8FB55102FF82FF81107F8CAEFAFAF8D9" box="[1148,1421,1776,1798]" italics="true" pageId="9" pageNumber="460">
<collectorName id="1034E8C6FF82FF81107F8CAEFA7DF8D9" box="[1148,1290,1777,1798]" pageId="9" pageNumber="460">R.F. Monteiro</collectorName>
and eq. col.
</emphasis>
);
<location id="B81EDBCBFF82FF8114C48D4FFEA9F8FA" LSID="urn:lsid:plazi:treatment:35683C06FF82FF8214948C39FE31FC8E:B81EDBCBFF82FF8114C48D4FFEA9F8FA" box="[199,478,1807,1829]" country="Brazil" name="Parque Nacional do Itatiaia" pageId="9" pageNumber="460" stateProvince="Rio de Janeiro">Parque Nacional do Itatiaia</location>
,
<collectingRegion id="7F0543F2FF82FF8115EB8D50FD0AF8FA" box="[488,637,1807,1829]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="460">Rio de Janeiro</collectingRegion>
,
<collectingDate id="D93B5238FF82FF8116858D50FD86F8FA" box="[646,753,1807,1829]" pageId="9" pageNumber="460" value="2014-02-27">
<date id="C97FABD0FF82FF8116858D50FD86F8FA" box="[646,753,1807,1829]" pageId="9" pageNumber="460" value="2014-02-27">27/ii/2014</date>
</collectingDate>
,
<specimenCount id="ABC74699FF82FF8116F88D50FC6CF8FA" box="[763,795,1807,1829]" pageId="9" pageNumber="460" type="male">1♂</specimenCount>
(
<emphasis id="8FB55102FF82FF8117298D4FFB44F8FA" box="[810,1075,1807,1829]" italics="true" pageId="9" pageNumber="460">
<collectorName id="1034E8C6FF82FF8117298D4FFCC2F8FA" box="[810,949,1808,1829]" pageId="9" pageNumber="460">R.F. Monteiro</collectorName>
and eq. col.
</emphasis>
) (
<collectionCode id="DBD015D5FF82FF81104B8D50FBFEF8FA" box="[1096,1161,1807,1829]" country="Brazil" httpUri="http://grbio.org/cool/qstz-adte" name="Universidade Federal Rural do Rio de Janeiro" pageId="9" pageNumber="460">UFRJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F5DBDE9BFF82FF8214948D0EFEEFFD41" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460" type="description">
<paragraph id="BD7E8D10FF82FF8214948D0EFE87FECA" blockId="9.[151,1436,1873,2032]" lastBlockId="10.[151,1436,151,850]" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460">
<emphasis id="8FB55102FF82FF8114948D0EFE1EF8B5" bold="true" box="[151,361,1873,1898]" pageId="9" pageNumber="460">Male description.</emphasis>
Structure: Supra-antennal plate with slightly carinate margins. A slight elevation between the plates; clypeal apical emargination as deep as its sublateral margins. Base of clypeal emargination subtriangular, what origins a slight longitudinal bevel raised on the medial region of the clypeus; wings hyaline, ferruginous on the anterior margins, stigma and marginal cell. The anterior half of the forewing slightly castaneus; scutellum raised and pointed at the top; Dorsal surface of the propodeum with dense and coalescent punctures, laterals striated at the base; mesepisternal suture strongly raised apically, the lamellae narrow and acute; first metasomal tergite longer than wide. Smooth transverse fasciae on apical margins of tergites almost absent, very short and covered by a dense row of setae; epipygium sides rugose by elongate punctures, smooth centrally; hypopygium subquadrate, rounded laterally.
</paragraph>
<paragraph id="BD7E8D10FF81FF8214C48B42FD30FDD1" blockId="10.[151,1436,151,850]" pageId="10" pageNumber="461">
Genital capsule: Dorsal surface of the basal ring more than
<date id="C97FABD0FF81FF8217A78B41FCB7FEE9" box="[932,960,286,310]" pageId="10" pageNumber="461">2x</date>
longer than its ventral surface. With a longitudinal line medially in dorsal view; gonocoxite tapering towards the apex, with more than half the gonostylus extension. The apex rounded and almost contiguous, slightly separated medially, with a longitudinal line dorsally; aedeagus longer than the gonocoxite and shortest than the gonostylus. Expanded laterally in two longitudinal lamellae, curved ventrally and with a small apical lobe; gonostylus slightly wider at the base, narrowing medially and broadening again before the apex. With golden pubescence on the half apical region. The apex slightly rounded, triangularly curved ventrally.
</paragraph>
<paragraph id="BD7E8D10FF81FF8214C48846FDF5FDA6" blockId="10.[151,1436,151,850]" pageId="10" pageNumber="461">Coloration. body predominantly black, except for the following castaneus markings: flagellomeres VVII. A large spot on the scutellum dorsal surface. The metanotum centrally. The legs except for the outer surface of the femora. Diffuse spots on the sternites IIIV.</paragraph>
<paragraph id="BD7E8D10FF81FF8214C488DAFEEFFD41" blockId="10.[151,1436,151,850]" box="[199,408,645,670]" pageId="10" pageNumber="461">Female: unknown.</paragraph>
</subSubSection>
<subSubSection id="F5DBDE9BFF81FF8214C488F6FEB0FCD5" pageId="10" pageNumber="461" type="etymology">
<paragraph id="BD7E8D10FF81FF8214C488F6FEB0FCD5" blockId="10.[151,1436,151,850]" pageId="10" pageNumber="461">
<emphasis id="8FB55102FF81FF8214C488F6FE39FD1D" bold="true" box="[199,334,681,706]" pageId="10" pageNumber="461">Etymology.</emphasis>
In Latin
<emphasis id="8FB55102FF81FF8215B788F5FD65FD1E" box="[436,530,682,705]" italics="true" pageId="10" pageNumber="461">reversus</emphasis>
means “returned”, referring to the lack of the strong carina on the margins of the supra-antennal plate, which is a diagnostic character of the
<taxonomicName id="7AC1F693FF81FF8217578891FCCCFD3A" authorityName="Klug" authorityYear="1810" box="[852,955,718,741]" class="Insecta" family="Tiphiidae" genus="Scotaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="8FB55102FF81FF8217578891FCCCFD3A" box="[852,955,718,741]" italics="true" pageId="10" pageNumber="461">Scotaena</emphasis>
</taxonomicName>
but seems to be reverted or at least less pronounced in this species.
</paragraph>
</subSubSection>
<subSubSection id="F5DBDE9BFF81FF8214C4894AFE31FC8E" pageId="10" pageNumber="461" type="distribution">
<paragraph id="BD7E8D10FF81FF8214C4894AFE31FC8E" blockId="10.[151,1436,151,850]" pageId="10" pageNumber="461">
<emphasis id="8FB55102FF81FF8214C4894AFE17FCF1" bold="true" box="[199,352,789,814]" pageId="10" pageNumber="461">Distribution.</emphasis>
The known records of
<taxonomicName id="7AC1F693FF81FF8216798949FC48FCF2" authorityName="Carnimeo &amp; Noll" authorityYear="2018" box="[634,831,790,813]" class="Insecta" family="Tiphiidae" genus="Scotaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="reversa">
<emphasis id="8FB55102FF81FF8216798949FC48FCF2" box="[634,831,790,813]" italics="true" pageId="10" pageNumber="461">Scotaena reversa</emphasis>
</taxonomicName>
are restricted to the coastal region of the Brazilian Atlantic Forest.
</paragraph>
</subSubSection>
</treatment>
</document>